190 lines
22 KiB
XML
190 lines
22 KiB
XML
<document id="D048F588DCECF96B074ADDF8A770ECD4" ID-DOI="10.11646/zootaxa.3955.3.9" ID-GBIF-Dataset="a09f0c7f-f65f-47af-bebc-9371363b9653" ID-ISSN="1175-5326" ID-Zenodo-Dep="233117" ID-ZooBank="D018AFA7-2EA9-44BF-AB81-ECBAF132E7AE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461051944299" checkinUser="plazi" docAuthor="Häckel, Martin & Anichtchenko, Alexandr" docDate="2015" docId="B85E894010049909FF5C11E8FB68A8D8" docLanguage="en" docName="zt03955p434.pdf" docOrigin="Zootaxa 3955 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Craspedophorus kirschenhoferi Häckel & Anichtchenko, 2015, sp. nov." docType="treatment" docVersion="9" lastPageNumber="431" masterDocId="4467F1381000990DFFCB117FFFEEAE78" masterDocTitle="Two new Craspedophorus species (Coleoptera: Carabidae: Panagaeini) from the Philippines" masterLastPageNumber="434" masterPageNumber="427" pageNumber="431" updateTime="1698579237837" updateUser="plazi">
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<mods:titleInfo id="82F4B9FB912A728B6085FC4BA570491B">
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<mods:title id="4E20ADB46CF491FEE838BEC40E765E12">Two new Craspedophorus species (Coleoptera: Carabidae: Panagaeini) from the Philippines</mods:title>
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<mods:namePart id="110E7342BF4B97998B78C4A47C419F88">Häckel, Martin</mods:namePart>
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<mods:namePart id="98C4E58DABC8818478327DB26654FC01">Anichtchenko, Alexandr</mods:namePart>
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<mods:title id="070058D7995FEC06A8EE5C74F43A483E">Zootaxa</mods:title>
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<mods:part id="4CC9D2816EB2089C16276A7FC3771516">
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<mods:date id="9E42C37B528966ADBD39FE120E366785">2015</mods:date>
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<mods:detail id="A47AAA7F9F1A6A30FAAFC3C7F30A9E1F" type="volume">
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<mods:number id="B3F4678834B10799D790278BC28145B4">3955</mods:number>
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<treatment id="B85E894010049909FF5C11E8FB68A8D8" ID-DOI="http://doi.org/10.5281/zenodo.6111078" ID-GBIF-Taxon="119612162" ID-Zenodo-Dep="6111078" LSID="urn:lsid:plazi:treatment:B85E894010049909FF5C11E8FB68A8D8" httpUri="http://treatment.plazi.org/id/B85E894010049909FF5C11E8FB68A8D8" lastPageNumber="431" pageId="4" pageNumber="431">
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<subSubSection id="78ED6BDD10049909FF5C11E8FF05AEAB" pageId="4" pageNumber="431" type="nomenclature">
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<paragraph id="3048385610049909FF5C11E8FD95AECA" blockId="4.[151,635,151,211]" box="[151,635,151,178]" pageId="4" pageNumber="431">
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<heading id="6B008F3A10049909FF5C11E8FD95AECA" bold="true" box="[151,635,151,178]" fontSize="11" level="1" pageId="4" pageNumber="431" reason="1">
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<emphasis id="0283E44410049909FF5C11E8FD95AECA" bold="true" box="[151,635,151,178]" pageId="4" pageNumber="431">
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<taxonomicName id="F7F743D510049909FF5C11E8FDF8AEC9" ID-CoL="Z5WD" box="[151,534,151,177]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="431" phylum="Arthropoda" rank="species" species="kirschenhoferi" status="sp. nov.">
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<emphasis id="0283E44410049909FF5C11E8FDF8AEC9" bold="true" box="[151,534,151,177]" italics="true" pageId="4" pageNumber="431">Craspedophorus kirschenhoferi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="19B0593F10049909FDD611E7FD95AECA" box="[541,635,152,178]" pageId="4" pageNumber="431" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="3048385610049909FF5C11C5FF05AEAB" blockId="4.[151,635,151,211]" box="[151,235,186,211]" pageId="4" pageNumber="431">
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(
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<figureCitation id="A8CC24D310049909FF5411C5FF0CAEAB" box="[159,226,186,211]" captionStart="FIGURE 3. C" captionStartId="5.[151,250,1954,1976]" captionText="FIGURE 3. C. kirschenhoferi sp. nov.: A. Holotype, B. Lateral view of aedeagus." pageId="4" pageNumber="431">Fig. 3</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="78ED6BDD10049909FF5C107DFAB8AF47" pageId="4" pageNumber="431" type="materials_examined">
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<paragraph id="3048385610049909FF5C107DFAB8AF47" blockId="4.[151,1437,258,1696]" pageId="4" pageNumber="431">
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<emphasis id="0283E44410049909FF5C107DFEAEAF63" bold="true" box="[151,320,258,283]" pageId="4" pageNumber="431">
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<typeStatus id="EF4C86F410049909FF5C107DFF3CAF63" box="[151,210,258,283]" pageId="4" pageNumber="431">Type</typeStatus>
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material
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</emphasis>
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.
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<typeStatus id="EF4C86F410049909FE9B107DFE56AF63" box="[336,440,258,283]" pageId="4" pageNumber="431" type="holotype">Holotype</typeStatus>
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♂: “
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<collectingCountry id="48E078C610049909FE39107DFD9BAF63" box="[498,629,258,283]" name="Philippines" pageId="4" pageNumber="431">Philippines</collectingCountry>
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, Mindanao, Surigao del Norte.
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<date id="44491E9610049909FC29107CFBA7AF63" box="[994,1097,259,283]" pageId="4" pageNumber="431" value="2014-03">III. 2014</date>
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, local collector leg. (
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<collectionCode id="56E6A09310049909FAF7107CFA7EAF63" box="[1340,1424,259,283]" pageId="4" pageNumber="431">DUBC</collectionCode>
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).
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<typeStatus id="EF4C86F410049909FF5C1058FF12AF47" box="[151,252,295,319]" pageId="4" pageNumber="431" type="paratype">Paratype</typeStatus>
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, 1 ♀: “
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<collectingCountry id="48E078C610049909FE821059FE22AF47" box="[329,460,294,319]" name="Philippines" pageId="4" pageNumber="431">Philippines</collectingCountry>
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, Mindanao, Bukidnon. Mt. Kalatungan.
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<date id="44491E9610049909FC591058FBE5AF47" box="[914,1035,295,319]" pageId="4" pageNumber="431" value="2014-08">VIII. 2014</date>
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, local collector leg. (
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<collectionCode id="56E6A09310049909FB3E1058FAA5AF47" box="[1269,1355,295,319]" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="4" pageNumber="431">NMPC</collectionCode>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="78ED6BDD10049909FF0C1035FE8CA84C" pageId="4" pageNumber="431" type="description">
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<paragraph id="3048385610049909FF0C1035FD7EAFFE" blockId="4.[151,1437,258,1696]" pageId="4" pageNumber="431">
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<emphasis id="0283E44410049909FF0C1035FEB7AF1B" bold="true" box="[199,345,330,355]" pageId="4" pageNumber="431">Description.</emphasis>
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<typeStatus id="EF4C86F410049909FEAA1035FE27AF1B" box="[353,457,330,355]" pageId="4" pageNumber="431" type="holotype">Holotype</typeStatus>
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length 11.5-12.0 mm, width
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<quantity id="F70F95B310049909FCD21035FC4EAF1A" box="[793,928,330,355]" metricMagnitude="-3" metricUnit="m" metricValue="4.65" metricValueMax="4.8" metricValueMin="4.5" pageId="4" pageNumber="431" unit="mm" value="4.65" valueMax="4.8" valueMin="4.5">4.5-4.8 mm</quantity>
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. Pronotum 1.37 wider than long, 1.62 wider than head, elytra 1.22x wider than pronotum.
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</paragraph>
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<paragraph id="3048385610049909FF0C10EDFBBCAC38" blockId="4.[151,1437,258,1696]" pageId="4" pageNumber="431">Colour: Head, pronotum and elytra black. Elytra near margins densely covered by yellow setae, each elytron with two yellow spots. Humeral spot rounded, extending from 4th interval to margin and epipleura. Preapical spot smaller, rounded, reaching from 4th stria to 8th interval, separated from margin. Antennae, epistoma and legs black proximally; antennomeres IV–XI, tarsi and terminal parts of mandibles distally ferrugineous. Venter black, covered sparsely by long, yellow setae. Epipleura light yellowish, laterally near humeral spot.</paragraph>
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<paragraph id="3048385610049909FF0C1333FB4CACA9" blockId="4.[151,1437,258,1696]" pageId="4" pageNumber="431">
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<emphasis id="0283E44410049909FF0C1333FEEAAC1C" box="[199,260,588,612]" italics="true" pageId="4" pageNumber="431">Head</emphasis>
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large and wide, only 1.62x narrower than pronotum; frons medially smooth and weakly convex, laterally coarsely punctured, frontal margins rimmed by narrowly sulcate sharp furrows continuing posteriorly and medially between eyes; vertex coarsely punctured; clypeus smooth, shiny, medially upturned; anterior side of labrum concave; eyes strongly convex; temples barely indicated; neck convex, glossy, finely rugate.
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</paragraph>
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<paragraph id="3048385610049909FF0C13A2FEEBADB5" blockId="4.[151,1437,258,1696]" pageId="4" pageNumber="431">
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<emphasis id="0283E44410049909FF0C13A2FED8AC8C" box="[199,310,733,756]" italics="true" pageId="4" pageNumber="431">Pronotum</emphasis>
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transverse, 1.37x wider than long, almost hexagonal, widest immediately before middle, lateral angles round; anterior margin flat, parallel with base, as long as base; lateral margins narrowing weakly forward; anterior angles indistinct, widely rounded and not extending forward; posterior margins narrow obliquely, in front of posterior angles become weakly sinuate without any indentation; middle of base weakly sinuate, margin weakly oblique; anterior side of lateral rims indistinct, near base wider, more excavated, coarsely punctured; disc weakly convex and punctured more coarsely than rims; sagittal line indistinct, interrupted; basal pits very shallow, weakly indicated.
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</paragraph>
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<paragraph id="3048385610049909FF0C12A7FB71AA27" blockId="4.[151,1437,258,1696]" pageId="4" pageNumber="431">
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<emphasis id="0283E44410049909FF0C12A7FEE3AD88" box="[199,269,984,1008]" italics="true" pageId="4" pageNumber="431">Elytra</emphasis>
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moderately convex, almost parallel-sided, 1.22x wider than pronotum, weakly widening toward maximum width at middle, distinctly tuberose before apex; basal rim discernible only medial of 4th interval; humeri weakly developed, moderately oblique, widely rounded; striae deeply impressed, coarsely punctured; intervals convex, irregularly punctured; dorsum with black setae, on elytral spots setae yellowish red.
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</paragraph>
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<paragraph id="3048385610049909FF0C1513FAC7AAFB" blockId="4.[151,1437,258,1696]" box="[199,1321,1131,1156]" pageId="4" pageNumber="431">
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<emphasis id="0283E44410049909FF0C1513FEE2AAFB" box="[199,268,1132,1155]" italics="true" pageId="4" pageNumber="431">Venter</emphasis>
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black, glossy, metepisterna almost square, narrowed posteriorly, punctured, covered by setae.
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</paragraph>
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<paragraph id="3048385610049909FF0C15F1FBBBAADF" blockId="4.[151,1437,258,1696]" box="[199,1109,1166,1191]" pageId="4" pageNumber="431">
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Aedeagus (damaged in mounting), median lobe strongly curved laterally (
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<figureCitation id="A8CC24D310049909FBC815F1FBA9AADF" box="[1027,1095,1166,1191]" captionStart="FIGURE 3. C" captionStartId="5.[151,250,1954,1976]" captionText="FIGURE 3. C. kirschenhoferi sp. nov.: A. Holotype, B. Lateral view of aedeagus." pageId="4" pageNumber="431">Fig. 3</figureCitation>
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).
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</paragraph>
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<paragraph id="3048385610049909FF0C15CCFE8CA84C" blockId="4.[151,1437,258,1696]" pageId="4" pageNumber="431">
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<emphasis id="0283E44410049909FF0C15CCFE3AAAB4" bold="true" box="[199,468,1203,1228]" pageId="4" pageNumber="431">Differential diagnosis.</emphasis>
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<taxonomicName id="F7F743D510049909FE2A15CCFCA5AAB3" box="[481,843,1203,1227]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="431" phylum="Arthropoda" rank="species" species="kirschenhoferi" status="sp. nov.">
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<emphasis id="0283E44410049909FE2A15CCFCA5AAB3" box="[481,843,1203,1227]" italics="true" pageId="4" pageNumber="431">Craspedophorus kirschenhoferi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="19B0593F10049909FC9315CCFC56AAB4" box="[856,952,1203,1228]" pageId="4" pageNumber="431" rank="species">
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<emphasis id="0283E44410049909FC9315CCFC56AAB4" bold="true" box="[856,952,1203,1228]" pageId="4" pageNumber="431">sp. nov.</emphasis>
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</taxonomicNameLabel>
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resembles species of the
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<emphasis id="0283E44410049909FB3F15CCFA72AAB3" box="[1268,1436,1203,1227]" italics="true" pageId="4" pageNumber="431">
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“
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<taxonomicName id="F7F743D510049909FB3715CCFA72AAB3" box="[1276,1436,1203,1227]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="431" phylum="Arthropoda" rank="species" species="microspilotus">microspilotus</taxonomicName>
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</emphasis>
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species group sensu
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<bibRefCitation id="546645A710049909FEB515A9FD9FAA97" author="Kirschenhofer" box="[382,625,1238,1263]" pageId="4" pageNumber="431" refString="Kirschenhofer, E. (2000) Neue und wenig bekannte Panagaeini der ostlichen Palaarktis sowie der Orientalis. (Coleoptera, Carabidae). Entomofauna, 21 (29), 321 - 372." type="journal article" year="2000">Kirschenhofer (2000)</bibRefCitation>
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by its body size and shape, but differs clearly from most species of that group by markedly less transverse, hexagonal pronotum. The two species of that group with most similar pronotum,
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<taxonomicName id="F7F743D510049909FED8145FFCA4AB4F" authority="Hackel & Kirschenhofer, 2014" authorityName="Hackel & Kirschenhofer" authorityYear="2014" box="[275,842,1311,1336]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="431" phylum="Arthropoda" rank="species" species="chiangdaoensis">
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<emphasis id="0283E44410049909FED8145FFE0DAB4F" box="[275,483,1311,1335]" italics="true" pageId="4" pageNumber="431">C. chiangdaoensis</emphasis>
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<bibRefCitation id="546645A710049909FE271460FCA4AB4F" author="Hackel" box="[492,842,1311,1336]" pageId="4" pageNumber="431" refString="Hackel, M. & Kirchenhofer, E. (2014) A contribution to the knowledge of the subfamily Panagaeinae Hope, 1838 from Asia. Part 2. Genus Craspedophorus Hope, 1838, east Palearctic and Oriental species, and genus Tinoderus Chaudoir, 1879 (Coleoptera: Carabidae). Studies and Reports, Taxonomical Series, 10, 275 - 392." type="journal article" year="2014">Häckel & Kirschenhofer, 2014</bibRefCitation>
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</taxonomicName>
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and
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<taxonomicName id="F7F743D510049909FC4F145FFA76AB4F" authority="Hackel & Kirschenhofer, 2014" authorityName="Hackel & Kirschenhofer" authorityYear="2014" box="[900,1432,1311,1336]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="431" phylum="Arthropoda" rank="species" species="kiwlomensis">
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<emphasis id="0283E44410049909FC4F145FFBC1AB4F" box="[900,1071,1311,1335]" italics="true" pageId="4" pageNumber="431">C. kiwlomensis</emphasis>
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<bibRefCitation id="546645A710049909FBF31460FA76AB4F" author="Hackel" box="[1080,1432,1311,1336]" pageId="4" pageNumber="431" refString="Hackel, M. & Kirchenhofer, E. (2014) A contribution to the knowledge of the subfamily Panagaeinae Hope, 1838 from Asia. Part 2. Genus Craspedophorus Hope, 1838, east Palearctic and Oriental species, and genus Tinoderus Chaudoir, 1879 (Coleoptera: Carabidae). Studies and Reports, Taxonomical Series, 10, 275 - 392." type="journal article" year="2014">Häckel & Kirschenhofer, 2014</bibRefCitation>
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</taxonomicName>
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, differ inter alia by elytral pattern with distinctly larger spots.
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<taxonomicName id="F7F743D510049909FCA8143CFBC7AB23" box="[867,1065,1347,1371]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="431" phylum="Arthropoda" rank="species" species="kirschenhoferi" status="sp. nov.">
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<emphasis id="0283E44410049909FCA8143CFBC7AB23" box="[867,1065,1347,1371]" italics="true" pageId="4" pageNumber="431">C. kirschenhoferi</emphasis>
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</taxonomicName>
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<emphasis id="0283E44410049909FBF8143CFB61AB24" bold="true" box="[1075,1167,1347,1372]" pageId="4" pageNumber="431">
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<taxonomicNameLabel id="19B0593F10049909FBF8143CFB61AB24" box="[1075,1167,1347,1372]" pageId="4" pageNumber="431" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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also weakly resembles species of
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<taxonomicName id="F7F743D510049909FED81417FE27AB07" box="[275,457,1383,1407]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="431" phylum="Arthropoda" rank="species" species="basifasciatus">
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<emphasis id="0283E44410049909FED81417FE27AB07" box="[275,457,1383,1407]" italics="true" pageId="4" pageNumber="431">C. basifasciatus</emphasis>
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</taxonomicName>
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group sensu
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<bibRefCitation id="546645A710049909FDAD1418FC3AAB07" author="Hackel" box="[614,980,1383,1408]" pageId="4" pageNumber="431" refString="Hackel, M. & Kirchenhofer, E. (2014) A contribution to the knowledge of the subfamily Panagaeinae Hope, 1838 from Asia. Part 2. Genus Craspedophorus Hope, 1838, east Palearctic and Oriental species, and genus Tinoderus Chaudoir, 1879 (Coleoptera: Carabidae). Studies and Reports, Taxonomical Series, 10, 275 - 392." type="journal article" year="2014">Häckel & Kirschenhofer (2014)</bibRefCitation>
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by similar pronotum, but differs in the elytral pattern with spots without serrate margins. Another very similar but larger (
|
||
<quantity id="F70F95B310049909FB5D14F4FAEBABDB" box="[1174,1285,1419,1444]" metricMagnitude="-2" metricUnit="m" metricValue="1.35" pageId="4" pageNumber="431" unit="mm" value="13.5">13.5 mm</quantity>
|
||
) species,
|
||
<taxonomicName id="F7F743D510049909FA4914F4FDCCABBF" authority="Kirschenhofer, 2011" authorityName="Kirschenhofer" authorityYear="2011" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="431" phylum="Arthropoda" rank="species" species="soppongensis">
|
||
<emphasis id="0283E44410049909FA4914F4FEC0ABBF" italics="true" pageId="4" pageNumber="431">C. soppongensis</emphasis>
|
||
Kirschenhofer, 2011
|
||
</taxonomicName>
|
||
(
|
||
<emphasis id="0283E44410049909FDFC14D0FD7CABBF" box="[567,658,1455,1479]" italics="true" pageId="4" pageNumber="431">“kubani</emphasis>
|
||
species group) from northern
|
||
<collectingCountry id="48E078C610049909FC3614D0FB8FABB0" box="[1021,1121,1455,1480]" name="Thailand" pageId="4" pageNumber="431">Thailand</collectingCountry>
|
||
(
|
||
<figureCitation id="A8CC24D310049909FBBF14D0FB52ABBF" box="[1140,1212,1455,1480]" captionStart="FIGURE 4. C" captionStartId="6.[151,250,1952,1974]" captionTargetBox="[315,1272,215,1916]" captionTargetId="figure@6.[303,1283,193,1930]" captionTargetPageId="6" captionText="FIGURE 4. C. soppongenis Kirschenhofer, 2011 male, holotype. Scale bar: 10 mm." httpUri="https://zenodo.org/record/233120/files/figure.png" pageId="4" pageNumber="431">Fig. 4</figureCitation>
|
||
) has the pronotum similarly shaped but differently sculptured; the elytra shows a similar pattern but is differently sculptured; the sagittal line of pronotum is narrow and deeply impressed, and the pronotal basal pits are narrow, elongated and deeply impressed.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="78ED6BDD10049909FF0C1740FB41A803" pageId="4" pageNumber="431" type="etymology">
|
||
<paragraph id="3048385610049909FF0C1740FB41A803" blockId="4.[151,1437,258,1696]" pageId="4" pageNumber="431">
|
||
<emphasis id="0283E44410049909FF0C1740FEA0A820" bold="true" box="[199,334,1599,1624]" pageId="4" pageNumber="431">Etymology.</emphasis>
|
||
Named after our colleague and friend Erich Kirschenhofer (Perchtoldsdorf,
|
||
<collectingCountry id="48E078C610049909FB7F1740FAE5A820" box="[1204,1291,1599,1624]" name="Austria" pageId="4" pageNumber="431">Austria</collectingCountry>
|
||
), a specialist in the Panagaeini and
|
||
<taxonomicName id="F7F743D510049909FE45171CFD98A804" box="[398,630,1635,1660]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="431" phylum="Arthropoda" rank="tribe" tribe="Licinini">Licinini (Carabidae)</taxonomicName>
|
||
. The specific epithet is a noun in the genitive case.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="78ED6BDD10049909FF0C17F8FB68A8D8" box="[199,1158,1671,1696]" pageId="4" pageNumber="431" type="distribution">
|
||
<paragraph id="3048385610049909FF0C17F8FB68A8D8" blockId="4.[151,1437,258,1696]" box="[199,1158,1671,1696]" pageId="4" pageNumber="431">
|
||
<emphasis id="0283E44410049909FF0C17F8FE8EA8D8" bold="true" box="[199,352,1671,1696]" pageId="4" pageNumber="431">Distribution.</emphasis>
|
||
So far known only from two localities in Mindanao Island,
|
||
<collectingCountry id="48E078C610049909FBCB17F8FB6CA8D8" box="[1024,1154,1671,1696]" name="Philippines" pageId="4" pageNumber="431">Philippines</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |