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<document id="9CD60C8E000F1951EADC5D41A61E53B3" ID-DOI="10.5281/zenodo.174029" ID-GBIF-Dataset="d9d8d147-e8fc-4014-8af2-aa2fd28c20df" ID-ISSN="1175­5326" ID-Zenodo-Dep="174029" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459790304226" checkinUser="plazi" docAuthor="Schawaller, Wolfgang" docDate="2006" docId="7F54080DFFECFF86D56ADBE7FC3DFA9B" docLanguage="en" docName="zt01325p384.pdf" docOrigin="Zootaxa 1325" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Laena kenyirica Schawaller, 2006, n. sp." docType="treatment" docVersion="9" lastPageNumber="371" masterDocId="836D7075FFEDFF84D462DD29FF82FFF5" masterDocTitle="New species of the genus Laena Latreille (Coleoptera: Tenebrionidae) from Southeastern Asia *" masterLastPageNumber="384" masterPageNumber="369" pageNumber="370" updateTime="1698219154779" updateUser="plazi">
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<mods:title id="B779905CBA71BAF6A0D58A787181A057">New species of the genus Laena Latreille (Coleoptera: Tenebrionidae) from Southeastern Asia *</mods:title>
</mods:titleInfo>
<mods:name id="741E9C33DE07A0634BCA77D977E88731" type="personal">
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<mods:namePart id="971BD6EF3A8E91719E1FF5AC792FBBA0">Schawaller, Wolfgang</mods:namePart>
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<mods:title id="86957B3942299DB31F86466643109451">Zootaxa</mods:title>
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<mods:date id="62A66B5A9AE0091F3320E7C4E2927ED5">2006</mods:date>
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<mods:identifier id="7F7A7E541781140DA24AE1A3AE651985" type="DOI">10.5281/zenodo.174029</mods:identifier>
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<treatment id="7F54080DFFECFF86D56ADBE7FC3DFA9B" ID-DOI="http://doi.org/10.5281/zenodo.6258179" ID-GBIF-Taxon="119350243" ID-Zenodo-Dep="6258179" LSID="urn:lsid:plazi:treatment:7F54080DFFECFF86D56ADBE7FC3DFA9B" httpUri="http://treatment.plazi.org/id/7F54080DFFECFF86D56ADBE7FC3DFA9B" lastPageId="2" lastPageNumber="371" pageId="1" pageNumber="370">
<subSubSection id="BFE7EA90FFECFF85D56ADBE7FD25F91D" box="[264,679,1742,1768]" pageId="1" pageNumber="370" type="nomenclature">
<paragraph id="F742B91BFFECFF85D56ADBE7FD25F91D" blockId="1.[264,679,1742,1768]" box="[264,679,1742,1768]" pageId="1" pageNumber="370">
<heading id="AC0A0E77FFECFF85D56ADBE7FD25F91D" bold="true" box="[264,679,1742,1768]" fontSize="11" level="1" pageId="1" pageNumber="370" reason="1">
<emphasis id="C5896509FFECFF85D56ADBE7FD25F91D" bold="true" box="[264,679,1742,1768]" pageId="1" pageNumber="370">
<taxonomicName id="30FDC298FFECFF85D56ADBE7FE49F91D" box="[264,459,1742,1768]" class="Insecta" family="Terebellidae" genus="Laena" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="370" phylum="Arthropoda" rank="species" species="kenyirica" status="sp. nov.">
<emphasis id="C5896509FFECFF85D56ADBE7FE49F91D" bold="true" box="[264,459,1742,1768]" italics="true" pageId="1" pageNumber="370">Laena kenyirica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DEBAD872FFECFF85D5B6DBE7FD94F91D" box="[468,534,1742,1768]" pageId="1" pageNumber="370" rank="species">n. sp.</taxonomicNameLabel>
(
<figureCitation id="6FC6A59EFFECFF85D647DBE7FDF0F91D" box="[549,626,1742,1768]" captionStart="FIGURES 1 8" captionStartId="7.[264,382,1734,1758]" captionTargetBox="[289,1307,328,1705]" captionTargetId="figure@7.[285,1310,326,1715]" captionTargetPageId="7" captionText="FIGURES 1 8. Dorsal view of Laena species. 1 Laena champasaka n. sp., Ψ holotype CSBC, Laos. 2 Laena gialaica n. sp., Ψ holotype CSBC, Vietnam. 3 Laena inthanonica n. sp., ♂ holotype MHNG, Thailand. 4 Laena jocheni n. sp., ♂ holotype SMNS, Laos. 5 Laena kenyirica n. sp., ♂ holotype SMNS, Malaysia. 6 Laena khaolaka n. sp., ♂ holotype SMNS, Thailand. 7 Laena kiyoshii n. sp., ♂ holotype SMNS, Laos. 8 Laena kurbatovi n. sp., Ψ holotype HNHM, Myanmar / Burma." httpUri="https://zenodo.org/record/174030/files/figure.png" pageId="1" pageNumber="370">Figs 5</figureCitation>
,
<figureCitation id="6FC6A59EFFECFF85D61DDBE7FD1CF91D" box="[639,670,1742,1768]" captionStart="FIGURES 17 26" captionStartId="12.[264,382,1382,1406]" captionTargetBox="[346,1246,319,1358]" captionTargetId="figure@12.[342,1249,316,1361]" captionTargetPageId="12" captionText="FIGURES 17 26. Aedeagus of Laena species. 17 Laena inthanonica n. sp., ♂ holotype MHNG, Thailand. 18 Laena jocheni n. sp., ♂ holotype SMNS, Laos. 19 Laena kenyirica n. sp., ♂ holotype SMNS, Malaysia. 20 Laena khaolaka n. sp., ♂ holotype SMNS, Thailand. 21 Laena kiyoshii n. sp., ♂ holotype SMNS, Laos. 22 Laena moguntia n. sp., ♂ holotype CKAO, Vietnam. 23 Laena schulzi n. sp., ♂ holotype SMNS, Malaysia. 24 Laena rolandi n. sp., ♂ holotype CRGT, Thailand. 25 Laena tamdaoensis, ♂ holotype NSMT, Vietnam. 26 Laena vietnamica, non­type ♂ HNHM, Vietnam." httpUri="https://zenodo.org/record/174032/files/figure.png" pageId="1" pageNumber="370">19</figureCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="BFE7EA90FFECFF86D56ADA30FCE4FEC3" lastPageId="2" lastPageNumber="371" pageId="1" pageNumber="370" type="materials_examined">
<paragraph id="F742B91BFFECFF85D56ADA30FC03F8AE" blockId="1.[264,1323,1817,1883]" pageId="1" pageNumber="370">
<typeStatus id="284607B9FFECFF85D56ADA30FEF8F8C6" box="[264,378,1817,1843]" pageId="1" pageNumber="370" type="holotype">
<emphasis id="C5896509FFECFF85D56ADA30FEF8F8C6" bold="true" box="[264,378,1817,1843]" pageId="1" pageNumber="370">Holotype</emphasis>
</typeStatus>
(♂):
<collectingCountry id="8FEAF98BFFECFF85D5D5DA30FDA7F8C6" box="[439,549,1817,1843]" name="Malaysia" pageId="1" pageNumber="370">Malaysia</collectingCountry>
, Lake Kenyir,
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SW Dam,
<quantity id="300514FEFFECFF85D7F1DA30FC5FF8C6" box="[915,989,1817,1843]" metricMagnitude="4" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="370" unit="km" value="50.0">50 km</quantity>
SW Kuala Terengganu,
<quantity id="300514FEFFECFF85D09DDA30FEA3F8AE" metricMagnitude="2" metricUnit="m" metricValue="3.5" pageId="1" pageNumber="370" unit="m" value="350.0">350 m</quantity>
,
<date id="83439FDBFFECFF85D54CDA68FE66F8AE" box="[302,484,1857,1883]" pageId="1" pageNumber="370" value="2001-07-07" valueMax="2001-07-12" valueMin="2001-07-07">7.12.VII.2001</date>
, leg. A. Schulz &amp; K. Vock,
<collectionCode id="91EC21DEFFECFF85D74BDA68FCFFF8AE" LSID="urn:lsid:biocol.org:col:34840" box="[809,893,1857,1883]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="1" pageNumber="370">SMNS</collectionCode>
.
</paragraph>
<paragraph id="F742B91BFFEFFF86D55ADC35FCE4FEC3" blockId="2.[264,1324,284,1390]" box="[312,870,284,310]" pageId="2" pageNumber="371">
<typeStatus id="284607B9FFEFFF86D55ADC35FE2CFEC3" box="[312,430,284,310]" pageId="2" pageNumber="371" type="paratype">Paratypes</typeStatus>
: Same data as
<typeStatus id="284607B9FFEFFF86D63BDC35FD40FEC3" box="[601,706,284,310]" pageId="2" pageNumber="371" type="holotype">holotype</typeStatus>
, 2 ex.
<collectionCode id="91EC21DEFFEFFF86D76DDC35FCE1FEC3" LSID="urn:lsid:biocol.org:col:34840" box="[783,867,284,310]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="2" pageNumber="371">SMNS</collectionCode>
.
</paragraph>
</subSubSection>
<subSubSection id="BFE7EA90FFEFFF86D55ADC6AFB7EFEAB" box="[312,1276,323,350]" pageId="2" pageNumber="371" type="etymology">
<paragraph id="F742B91BFFEFFF86D55ADC6AFB7EFEAB" blockId="2.[264,1324,284,1390]" box="[312,1276,323,350]" pageId="2" pageNumber="371">
<emphasis id="C5896509FFEFFF86D55ADC6AFE4AFEA8" bold="true" box="[312,456,323,349]" pageId="2" pageNumber="371">Etymology:</emphasis>
Named after Lake Kenyir, in whose vicinity the
<typeStatus id="284607B9FFEFFF86D067DC6DFBC1FEAB" box="[1029,1091,324,350]" pageId="2" pageNumber="371">types</typeStatus>
were collected.
</paragraph>
</subSubSection>
<subSubSection id="BFE7EA90FFEFFF86D55ADC42FD67FCE3" pageId="2" pageNumber="371" type="diagnosis">
<paragraph id="F742B91BFFEFFF86D55ADC42FD67FCE3" blockId="2.[264,1324,284,1390]" pageId="2" pageNumber="371">
<emphasis id="C5896509FFEFFF86D55ADC42FE38FE70" bold="true" box="[312,442,363,389]" pageId="2" pageNumber="371">Diagnosis:</emphasis>
The species can be recognized by the lack of prominent eyes, distinctly crenulate lateral margins of the pronotum, rows of elytral punctures without setation, distinctly convex elytral intervals, distinctly crenulate lateral intervals in the humeral region, and aedeagus with joint subquadrate parameres.
<taxonomicName id="30FDC298FFEFFF86D7FCDCCDFBDEFE08" box="[926,1116,484,509]" class="Insecta" family="Terebellidae" genus="Laena" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="371" phylum="Arthropoda" rank="species" species="kenyirica" status="sp. nov.">
<emphasis id="C5896509FFEFFF86D7FCDCCDFBDEFE08" box="[926,1116,484,509]" italics="true" pageId="2" pageNumber="371">Laena kenyirica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DEBAD872FFEFFF86D006DCCAFB25FE08" box="[1124,1191,483,509]" pageId="2" pageNumber="371" rank="species">
<emphasis id="C5896509FFEFFF86D006DCCAFB25FE08" bold="true" box="[1124,1191,483,509]" pageId="2" pageNumber="371">n. sp.</emphasis>
</taxonomicNameLabel>
runs in the key of the Malaysian species (
<bibRefCitation id="936CC4EAFFEFFF86D6C2DF25FCF1FDD3" author="Schawaller" box="[672,883,524,550]" pageId="2" pageNumber="384" refString="Schawaller, W. (1995) Neue Laena - Arten (Coleoptera: Tenebrionidae) aus Malaysia. Stuttgarter Beitrage zur Naturkunde, (A) 523, 1 - 16." type="journal article" year="1995">Schawaller 1995</bibRefCitation>
) to the species
<taxonomicName id="30FDC298FFEFFF86D031DF25FE57FDBB" authority="Schawaller, 1995" authorityName="Schawaller" authorityYear="1995" class="Insecta" family="Terebellidae" genus="Laena" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="371" phylum="Arthropoda" rank="species" species="gentingica">
<emphasis id="C5896509FFEFFF86D031DF25FAA9FDD0" box="[1107,1323,524,549]" italics="true" pageId="2" pageNumber="371">Laena gentingica</emphasis>
<bibRefCitation id="936CC4EAFFEFFF86D56ADF1DFE57FDBB" author="Schawaller" box="[264,469,564,590]" pageId="2" pageNumber="384" refString="Schawaller, W. (1995) Neue Laena - Arten (Coleoptera: Tenebrionidae) aus Malaysia. Stuttgarter Beitrage zur Naturkunde, (A) 523, 1 - 16." type="journal article" year="1995">Schawaller, 1995</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="30FDC298FFEFFF86D670DF1DFC73FDBB" authority="Schawaller, 1995" authorityName="Schawaller" authorityYear="1995" box="[530,1009,564,590]" class="Insecta" family="Terebellidae" genus="Laena" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="371" phylum="Arthropoda" rank="species" species="brinchangensis">
<emphasis id="C5896509FFEFFF86D670DF1DFC9AFDB8" box="[530,792,564,589]" italics="true" pageId="2" pageNumber="371">Laena brinchangensis</emphasis>
<bibRefCitation id="936CC4EAFFEFFF86D743DF1DFC73FDBB" author="Schawaller" box="[801,1009,564,590]" pageId="2" pageNumber="384" refString="Schawaller, W. (1995) Neue Laena - Arten (Coleoptera: Tenebrionidae) aus Malaysia. Stuttgarter Beitrage zur Naturkunde, (A) 523, 1 - 16." type="journal article" year="1995">Schawaller, 1995</bibRefCitation>
</taxonomicName>
; both with a smaller body size, distinctly prominent eyes, a different shape of the pronotum, a different dorsal punctation and setation, and a different shape of the aedeagus with joint triangular parameres.
<taxonomicName id="30FDC298FFEFFF86D5F0DF85FDD1FD30" box="[402,595,684,709]" class="Insecta" family="Terebellidae" genus="Laena" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="371" phylum="Arthropoda" rank="species" species="kenyirica" status="sp. nov.">
<emphasis id="C5896509FFEFFF86D5F0DF85FDD1FD30" box="[402,595,684,709]" italics="true" pageId="2" pageNumber="371">Laena kenyirica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DEBAD872FFEFFF86D63FDF85FD20FD33" box="[605,674,684,710]" pageId="2" pageNumber="371" rank="species">
<emphasis id="C5896509FFEFFF86D63FDF85FD20FD33" bold="true" box="[605,674,684,710]" pageId="2" pageNumber="371">n. sp.</emphasis>
</taxonomicNameLabel>
differs from the syntopic
<taxonomicName id="30FDC298FFEFFF86D783DF85FB07FD30" box="[993,1157,684,709]" class="Insecta" family="Terebellidae" genus="Laena" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="371" phylum="Arthropoda" rank="species" species="schulzi" status="sp. nov.">
<emphasis id="C5896509FFEFFF86D783DF85FB07FD30" box="[993,1157,684,709]" italics="true" pageId="2" pageNumber="371">Laena schulzi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DEBAD872FFEFFF86D0F2DF85FB57FD33" box="[1168,1237,684,710]" pageId="2" pageNumber="371" rank="species">
<emphasis id="C5896509FFEFFF86D0F2DF85FB57FD33" bold="true" box="[1168,1237,684,710]" pageId="2" pageNumber="371">n. sp.</emphasis>
</taxonomicNameLabel>
by the bigger body size, by a different shape of pronotum and elytra, by distinct dorsal setation and by a different shape of the aedeagus.
</paragraph>
</subSubSection>
<subSubSection id="BFE7EA90FFEFFF86D55ADE0DFD03FB53" pageId="2" pageNumber="371" type="description">
<paragraph id="F742B91BFFEFFF86D55ADE0DFD03FB53" blockId="2.[264,1324,284,1390]" pageId="2" pageNumber="371">
<emphasis id="C5896509FFEFFF86D55ADE0DFE50FCCB" bold="true" box="[312,466,804,830]" pageId="2" pageNumber="371">Description:</emphasis>
Body length 3.84.8 mm. Eyes not prominent. Pronotum (
<figureCitation id="6FC6A59EFFEFFF86D0F9DE0DFB66FCCB" box="[1179,1252,804,830]" captionStart="FIGURES 1 8" captionStartId="7.[264,382,1734,1758]" captionTargetBox="[289,1307,328,1705]" captionTargetId="figure@7.[285,1310,326,1715]" captionTargetPageId="7" captionText="FIGURES 1 8. Dorsal view of Laena species. 1 Laena champasaka n. sp., Ψ holotype CSBC, Laos. 2 Laena gialaica n. sp., Ψ holotype CSBC, Vietnam. 3 Laena inthanonica n. sp., ♂ holotype MHNG, Thailand. 4 Laena jocheni n. sp., ♂ holotype SMNS, Laos. 5 Laena kenyirica n. sp., ♂ holotype SMNS, Malaysia. 6 Laena khaolaka n. sp., ♂ holotype SMNS, Thailand. 7 Laena kiyoshii n. sp., ♂ holotype SMNS, Laos. 8 Laena kurbatovi n. sp., Ψ holotype HNHM, Myanmar / Burma." httpUri="https://zenodo.org/record/174030/files/figure.png" pageId="2" pageNumber="371">Fig. 5</figureCitation>
) with large punctures, distance between 0.53 diameters, most punctures bearing long erect seta; surface flat, shining; lateral margin distinctly bordered, crenulate; propleura with similar punctation but without setation. Elytra (
<figureCitation id="6FC6A59EFFEFFF86D681DEB5FCA9FC43" box="[739,811,924,950]" captionStart="FIGURES 1 8" captionStartId="7.[264,382,1734,1758]" captionTargetBox="[289,1307,328,1705]" captionTargetId="figure@7.[285,1310,326,1715]" captionTargetPageId="7" captionText="FIGURES 1 8. Dorsal view of Laena species. 1 Laena champasaka n. sp., Ψ holotype CSBC, Laos. 2 Laena gialaica n. sp., Ψ holotype CSBC, Vietnam. 3 Laena inthanonica n. sp., ♂ holotype MHNG, Thailand. 4 Laena jocheni n. sp., ♂ holotype SMNS, Laos. 5 Laena kenyirica n. sp., ♂ holotype SMNS, Malaysia. 6 Laena khaolaka n. sp., ♂ holotype SMNS, Thailand. 7 Laena kiyoshii n. sp., ♂ holotype SMNS, Laos. 8 Laena kurbatovi n. sp., Ψ holotype HNHM, Myanmar / Burma." httpUri="https://zenodo.org/record/174030/files/figure.png" pageId="2" pageNumber="371">Fig. 5</figureCitation>
) with complete rows of punctures without striae, punctures equal in size to pronotal punctures, without setae; intervals with row of fine punctures each bearing long, erect seta; all intervals convex, shining, external intervals near humeral region serrate by punctures; interval
<quantity id="300514FEFFEFFF86D7B0D93DFB83FBDB" box="[978,1025,1044,1070]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="2" pageNumber="371" unit="in" value="7.0">7 in</quantity>
the humeral region with indistinct setiferous pore, interval 9 with 4 indistinct setiferous pores. Femora in both sexes without distinct tooth or other modification. Tibiae in males without distinct sexual character. Aedeagus see
<figureCitation id="6FC6A59EFFEFFF86D645D9A5FDFCFB53" box="[551,638,1164,1190]" captionStart="FIGURES 17 26" captionStartId="12.[264,382,1382,1406]" captionTargetBox="[346,1246,319,1358]" captionTargetId="figure@12.[342,1249,316,1361]" captionTargetPageId="12" captionText="FIGURES 17 26. Aedeagus of Laena species. 17 Laena inthanonica n. sp., ♂ holotype MHNG, Thailand. 18 Laena jocheni n. sp., ♂ holotype SMNS, Laos. 19 Laena kenyirica n. sp., ♂ holotype SMNS, Malaysia. 20 Laena khaolaka n. sp., ♂ holotype SMNS, Thailand. 21 Laena kiyoshii n. sp., ♂ holotype SMNS, Laos. 22 Laena moguntia n. sp., ♂ holotype CKAO, Vietnam. 23 Laena schulzi n. sp., ♂ holotype SMNS, Malaysia. 24 Laena rolandi n. sp., ♂ holotype CRGT, Thailand. 25 Laena tamdaoensis, ♂ holotype NSMT, Vietnam. 26 Laena vietnamica, non­type ♂ HNHM, Vietnam." httpUri="https://zenodo.org/record/174032/files/figure.png" pageId="2" pageNumber="371">Fig. 19</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="BFE7EA90FFEFFF86D55AD99DFC3DFA9B" pageId="2" pageNumber="371" type="discussion">
<paragraph id="F742B91BFFEFFF86D55AD99DFC3DFA9B" blockId="2.[264,1324,284,1390]" pageId="2" pageNumber="371">
<emphasis id="C5896509FFEFFF86D55AD99DFE31FB3B" bold="true" box="[312,435,1204,1230]" pageId="2" pageNumber="371">Remarks:</emphasis>
<taxonomicName id="30FDC298FFEFFF86D5DED99DFDFEFB38" box="[444,636,1204,1229]" class="Insecta" family="Terebellidae" genus="Laena" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="371" phylum="Arthropoda" rank="species" species="kenyirica" status="sp. nov.">
<emphasis id="C5896509FFEFFF86D5DED99DFDFEFB38" box="[444,636,1204,1229]" italics="true" pageId="2" pageNumber="371">Laena kenyirica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DEBAD872FFEFFF86D6E7D99DFD4BFB3B" box="[645,713,1204,1230]" pageId="2" pageNumber="371" rank="species">
<emphasis id="C5896509FFEFFF86D6E7D99DFD4BFB3B" bold="true" box="[645,713,1204,1230]" pageId="2" pageNumber="371">n. sp.</emphasis>
</taxonomicNameLabel>
(and the syntopic
<taxonomicName id="30FDC298FFEFFF86D7CBD99DFBCCFB38" box="[937,1102,1204,1229]" class="Insecta" family="Terebellidae" genus="Laena" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="371" phylum="Arthropoda" rank="species" species="schulzi" status="sp. nov.">
<emphasis id="C5896509FFEFFF86D7CBD99DFBCCFB38" box="[937,1102,1204,1229]" italics="true" pageId="2" pageNumber="371">Laena schulzi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DEBAD872FFEFFF86D034D99DFB18FB3B" box="[1110,1178,1204,1230]" pageId="2" pageNumber="371" rank="species">
<emphasis id="C5896509FFEFFF86D034D99DFB18FB3B" bold="true" box="[1110,1178,1204,1230]" pageId="2" pageNumber="371">n. sp.</emphasis>
</taxonomicNameLabel>
, see below) populates lowland forest in the northeastern part of the Malayan Peninsula, thus living disjunctly from the
<typeStatus id="284607B9FFEFFF86D590D82DFDA7FAEB" box="[498,549,1284,1310]" pageId="2" pageNumber="371">type</typeStatus>
localities of all the other known Malaysian species in the central mountain ranges above
<quantity id="300514FEFFEFFF86D642D805FDFEFAB3" box="[544,636,1324,1350]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="371" unit="m" value="1000.0">1000 m</quantity>
. This different distributional and biological patterns might be considered as further arguments for the specific validity.
</paragraph>
</subSubSection>
</treatment>
</document>