228 lines
27 KiB
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228 lines
27 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.949.51269" ID-GBIF-Dataset="e4d2c425-4596-487c-8c4f-b09330b48203" ID-PMC="PMC7381721" ID-Pensoft-Pub="1313-2970-949-1" ID-Pensoft-UUID="303D64593B3B550DBB7CE01AA547409D" ID-PubMed="32765176" ID-ZooBank="93BFA448BB7343CDBC4EA86090DA63D5" ModsDocID="1313-2970-949-1" checkinTime="1594832489974" checkinUser="pensoft" docAuthor="Salata, Sebastian & Fisher, Brian L." docDate="2020" docId="306B1012E807513380E1A0347C50C614" docLanguage="en" docName="ZooKeys 949: 1-185" docOrigin="ZooKeys 949" docSource="http://dx.doi.org/10.3897/zookeys.949.51269" docTitle="Pheidole dasos Salata & Fisher 2020, sp. nov." docType="treatment" docUuid="CAD761EC-FFAC-4EC6-B80E-F4C0720AF757" docUuidSource="ZooBank" docVersion="4" id="303D64593B3B550DBB7CE01AA547409D" lastPageNumber="1" masterDocId="303D64593B3B550DBB7CE01AA547409D" masterDocTitle="Taxonomic revision of the Pheidole sikorae species group (Hymenoptera, Formicidae) from Madagascar" masterLastPageNumber="185" masterPageNumber="1" pageNumber="1" updateTime="1668169097828" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Taxonomic revision of the Pheidole sikorae species group (Hymenoptera, Formicidae) from Madagascar</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Salata, Sebastian</mods:namePart>
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<mods:affiliation>Department of Entomology, California Academy of Sciences, San Francisco, CA 94118, USA</mods:affiliation>
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<mods:nameIdentifier type="orcid">https://orcid.org/0000-0003-0811-2309</mods:nameIdentifier>
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<mods:nameIdentifier type="email">sdsalata@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Fisher, Brian L.</mods:namePart>
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<mods:affiliation>Department of Entomology, California Academy of Sciences, San Francisco, CA 94118, USA</mods:affiliation>
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<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-4653-3270</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2020</mods:date>
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<mods:detail type="volume">
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<mods:number>949</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>185</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.949.51269</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.949.51269</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-949-1</mods:identifier>
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<mods:identifier type="ZooBank">93BFA448BB7343CDBC4EA86090DA63D5</mods:identifier>
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<mods:identifier type="Pensoft-UUID">303D64593B3B550DBB7CE01AA547409D</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="165090191" LSID="urn:lsid:zoobank.org:act:CAD761EC-FFAC-4EC6-B80E-F4C0720AF757" httpUri="http://treatment.plazi.org/id/306B1012E807513380E1A0347C50C614" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName LSID="306B1012-E807-5133-80E1-A0347C50C614" authority="Salata & Fisher, 2020" authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole dasos" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="dasos" status="sp. nov.">Pheidole dasos</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Pheidole dasos sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923284) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0212441)." figureDoi="10.3897/zookeys.949.51269.figure28" httpUri="https://binary.pensoft.net/fig/432197" pageId="0" pageNumber="1">Figs 28A-F</figureCitation>
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<figureCitation captionStart="Figure 63" captionStartId="F63" captionText="Figure 63. Major worker, hypostomal teeth. Pheidole alina sp. nov. (A). Pheidole ambohimanga sp. nov. (B). Pheidole analavelona sp. nov. (C). Pheidole andohahela sp. nov. (D). Pheidole anomala sp. nov. (E). Pheidole anosyenne sp. nov. (F). Pheidole antranohofa sp. nov. (G). Pheidole beanka sp. nov. (H). Pheidole befotaka sp. nov. (I). Pheidole dasos sp. nov. (J). Pheidole flavominuta sp. nov. (K). Pheidole gracilis sp. nov. (L). Pheidole haboka sp. nov. (M). Pheidole havoana sp. nov. (N). Pheidole hazo sp. nov. (O). Pheidole itremo sp. nov. (P). Pheidole joffreville sp. nov. (Q). Pheidole kely sp. nov. (R). Pheidole lavasoa sp. nov. (S). Pheidole litigiosa Forel (T). Pheidole mahamavo sp. nov. (U). Pheidole mainty sp. nov. (V). Pheidole mamiratra sp. nov. (W). Pheidole manantenina sp. nov. (X)." figureDoi="10.3897/zookeys.949.51269.figure63" httpUri="https://binary.pensoft.net/fig/432232" pageId="0" pageNumber="1">, 63J</figureCitation>
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<figureCitation captionStart="Figure 65" captionStartId="F65" captionText="Figure 65. Distribution. Pheidole alina sp. nov. (A). Pheidole ambohimanga sp. nov. (B). Pheidole analavelona sp. nov. (C). Pheidole andohahela sp. nov. (D). Pheidole anomala sp. nov. (E). Pheidole anosyenne sp. nov. (F). Pheidole antranohofa sp. nov. (G). Pheidole beanka sp. nov. (H). Pheidole befotaka sp. nov. (I). Pheidole dasos sp. nov. (J). Pheidole flavominuta sp. nov. (K). Pheidole gracilis sp. nov. (L). Pheidole haboka sp. nov. (M). Pheidole havoana sp. nov. (N). Pheidole hazo sp. nov. (O). Pheidole itremo sp. nov. (P). Pheidole joffreville sp. nov. (Q). Pheidole kely sp. nov. (R). Pheidole lavasoa sp. nov. (S). Pheidole litigiosa Forel (T)." figureDoi="10.3897/zookeys.949.51269.figure65" httpUri="https://binary.pensoft.net/fig/432234" pageId="0" pageNumber="1">, 65J</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="type material">
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<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Holotype.</emphasis>
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Madagascar. • 1 major worker; Antsiranana; Makirovana forest;
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<geoCoordinate degrees="14.104" direction="south" orientation="latitude" precision="55" value="-14.104">-14.104</geoCoordinate>
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,
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<geoCoordinate degrees="50.03574" direction="east" orientation="longitude" precision="1" value="50.03574">50.03574</geoCoordinate>
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; alt. 225 m; 4 May 2011; B. L. Fisher et al. leg.; rainforest, ex rotten log; BLF27046; CASENT0212441 (CASC).
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Paratype.</emphasis>
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• 1w.; same data as for holotype, CASENT0923284 (CASC).
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</paragraph>
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<caption doi="10.3897/zookeys.949.51269.figure28" httpUri="https://binary.pensoft.net/fig/432197" pageId="0" pageNumber="1" start="Figure 28" startId="F28">
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Figure 28.</emphasis>
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<taxonomicName authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole dasos" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="dasos">
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<emphasis italics="true" pageId="0" pageNumber="1">Pheidole dasos</emphasis>
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</taxonomicName>
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sp. nov., full-face view (
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<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
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), profile (
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<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
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), and dorsal view (
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<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
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) of paratype minor worker (CASENT0923284) and full-face view (
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<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
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), profile (
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<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
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), and dorsal view (
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<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
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) of holotype major worker (CASENT0212441).
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="materials_examined">
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<paragraph pageId="0" pageNumber="1">Other material.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Madagascar. -
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<emphasis bold="true" pageId="0" pageNumber="1">Antsiranana</emphasis>
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: •1w., 1s.; Makirovana forest;
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<geoCoordinate degrees="14.17066" direction="south" orientation="latitude" precision="1" value="-14.17066">-14.17066</geoCoordinate>
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,
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<geoCoordinate degrees="49.95409" direction="east" orientation="longitude" precision="1" value="49.95409">49.95409</geoCoordinate>
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; alt. 415 m; 29 Apr 2011; B. L. Fisher et al. leg.; rainforest, ex rotten log; BLF26619 (CASC).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="diagnosis">
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<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Moderately large species.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Major workers.</emphasis>
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Head in full-face view sub-oval, not widening posteriorly, with anterior and posterior sides relatively straight, in lateral view sub-oval; ventral and dorsal faces convex; sides of the head with moderately dense, moderately long, subdecumbent to suberect pilosity; medial part of frons with thick, interrupted, sparse, and longitudinal rugae with indistinctly foveolate interspaces; anterolateral sides of frons with sparse, thick, and longitudinal rugae, posterolateral sides with rugae more irregular, interspaces with sparse foveolae; occipital lobes, and area posterolateral from eyes without smooth notches; scape, when laid back, exceeding the midlength of head by two-fifths of its length; inner hypostomal teeth distinct, large, closely spaced, triangular, with rounded apex directed upward; outer hypostomal teeth lobe-like, narrower, and approximately as high as inner teeth; inner and outer hypostomal teeth closely spaced and not connected by concavity; mesosoma rugofoveolate; dorsal promesonotum with additional thick and transverse rugae; lateral sides of pronotum with smooth notches; gaster smooth; body orange.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Minor workers.</emphasis>
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Head foveolate; vertex and median frons with fading sculpture; area posterolateral from eyes smooth; antennal sockets with few indistinct, curved outward rugae and foveolate interspaces; scape, when laid back, exceeding the posterior head margin by one-third of its length; promesonotum low and moderately long; promesonotal groove absent; propodeal spines small and triangular; mesosoma predominantly smooth; lateral sides with very sparse and indistinct, thin rugulae; body brown.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">Description.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Major workers.</emphasis>
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Measurements (
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<emphasis italics="true" pageId="0" pageNumber="1">N</emphasis>
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= 2): HL: 1.1, 1.14; HW: 1.04, 1.12; SL: 0.79, 0.81; EL: 0.16, 0.14; WL: 0.98, 1.05; PSL: 0.14, 0.15; MTL: 0.72, 0.77; PNW: 0.4, 0.45; PTW: 0.13, 0.13; PPW: 0.29, 0.31; CI: 105.9, 101.4; SI: 76.2, 72.6; PSLI: 12.4, 13.5; PPI: 45.8, 41.7; PNI: 38.8, 39.9; MTI: 68.8, 68.6.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Head.</emphasis>
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In full-face view sub-oval, not widening posteriorly, with anterior and posterior sides relatively straight (Fig.
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<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Pheidole dasos sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923284) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0212441)." figureDoi="10.3897/zookeys.949.51269.figure28" httpUri="https://binary.pensoft.net/fig/432197" pageId="0" pageNumber="1">28B</figureCitation>
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). In lateral view sub-oval; ventral and dorsal faces convex; inner hypostomal teeth visible. Sides of the head with moderately dense, moderately long, subdecumbent to suberect pilosity; whole head with dense, long, decumbent to erect pilosity. Medial part of frons with thick, interrupted, sparse, and longitudinal rugae with indistinctly foveolate interspaces; anterolateral sides of frons with sparse, thick, and longitudinal rugae, posterolateral sides with rugae more irregular, interspaces with sparse foveolae. Occipital lobes with slightly thinner, sparse, and irregular rugae and smooth to indistinctly foveolate interspaces. Area posterolateral from eyes with denser and thinner longitudinal rugae with rugofoveolate interspaces. Gena with relatively sparse and thick, longitudinal rugae with distinctly rugofoveolate interspaces. Centre of clypeus smooth and shiny, lateral sides with indistinct rugulae; median notch present, moderately wide, and shallow; median longitudinal carina absent; lateral longitudinal carinae absent. Scape, when laid back, exceeding the midlength of head by two-fifths of its length; pilosity subdecumbent to erect (Fig.
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<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Pheidole dasos sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923284) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0212441)." figureDoi="10.3897/zookeys.949.51269.figure28" httpUri="https://binary.pensoft.net/fig/432197" pageId="0" pageNumber="1">28B, D</figureCitation>
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). Inner hypostomal teeth distinct, large, closely spaced, triangular, with rounded apex directed upward; outer hypostomal teeth lobe-like, narrower than and approximately as high as inner teeth; inner and outer hypostomal teeth closely spaced and not connected by concavity (Fig.
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<figureCitation captionStart="Figure 63" captionStartId="F63" captionText="Figure 63. Major worker, hypostomal teeth. Pheidole alina sp. nov. (A). Pheidole ambohimanga sp. nov. (B). Pheidole analavelona sp. nov. (C). Pheidole andohahela sp. nov. (D). Pheidole anomala sp. nov. (E). Pheidole anosyenne sp. nov. (F). Pheidole antranohofa sp. nov. (G). Pheidole beanka sp. nov. (H). Pheidole befotaka sp. nov. (I). Pheidole dasos sp. nov. (J). Pheidole flavominuta sp. nov. (K). Pheidole gracilis sp. nov. (L). Pheidole haboka sp. nov. (M). Pheidole havoana sp. nov. (N). Pheidole hazo sp. nov. (O). Pheidole itremo sp. nov. (P). Pheidole joffreville sp. nov. (Q). Pheidole kely sp. nov. (R). Pheidole lavasoa sp. nov. (S). Pheidole litigiosa Forel (T). Pheidole mahamavo sp. nov. (U). Pheidole mainty sp. nov. (V). Pheidole mamiratra sp. nov. (W). Pheidole manantenina sp. nov. (X)." figureDoi="10.3897/zookeys.949.51269.figure63" httpUri="https://binary.pensoft.net/fig/432232" pageId="0" pageNumber="1">63J</figureCitation>
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).
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Mesosoma.</emphasis>
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In lateral view, promesonotum short, angular and moderately low, posterior mesonotum moderately steep, mesonotal process indistinct, tubercle-like; promesonotal groove absent; metanotal groove absent; propodeal spines short, with wide base and acute apex; humeral area produced (Fig.
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<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Pheidole dasos sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923284) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0212441)." figureDoi="10.3897/zookeys.949.51269.figure28" httpUri="https://binary.pensoft.net/fig/432197" pageId="0" pageNumber="1">28D</figureCitation>
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). Surface shiny and rugofoveolate; dorsal promesonotum with additional thick and transverse rugae; lateral sides of pronotum with smooth notches. Pilosity moderately dense, long, and erect (Fig.
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<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Pheidole dasos sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923284) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0212441)." figureDoi="10.3897/zookeys.949.51269.figure28" httpUri="https://binary.pensoft.net/fig/432197" pageId="0" pageNumber="1">28D, F</figureCitation>
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).
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Petiole.</emphasis>
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Shiny with dense foveolae; node finely foveolate, triangular, with rounded and thick apex, in rear view node dorsoventrally slightly convex; pilosity moderately sparse and erect (Fig.
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<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Pheidole dasos sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923284) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0212441)." figureDoi="10.3897/zookeys.949.51269.figure28" httpUri="https://binary.pensoft.net/fig/432197" pageId="0" pageNumber="1">28D, F</figureCitation>
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).
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Postpetiole.</emphasis>
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Shiny and foveolate; dorsum with reduced sculpture and smooth notch; in dorsal view oval, lateral margins medially with two dentate projections; pilosity long, moderately sparse and erect (Fig.
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<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Pheidole dasos sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923284) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0212441)." figureDoi="10.3897/zookeys.949.51269.figure28" httpUri="https://binary.pensoft.net/fig/432197" pageId="0" pageNumber="1">28D, F</figureCitation>
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).
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Gaster.</emphasis>
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Shiny and smooth; pilosity moderately dense, long and erect (Fig.
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<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Pheidole dasos sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923284) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0212441)." figureDoi="10.3897/zookeys.949.51269.figure28" httpUri="https://binary.pensoft.net/fig/432197" pageId="0" pageNumber="1">28D, F</figureCitation>
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).
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Colour.</emphasis>
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Orange with yellowish legs (Fig.
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<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Pheidole dasos sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923284) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0212441)." figureDoi="10.3897/zookeys.949.51269.figure28" httpUri="https://binary.pensoft.net/fig/432197" pageId="0" pageNumber="1">28D, F</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">Description.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Minor workers.</emphasis>
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Measurements (
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<emphasis italics="true" pageId="0" pageNumber="1">N</emphasis>
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= 2): HL: 0.56, 0.61; HW: 0.46, 0.49; SL: 0.71, 0.74; EL: 0.1, 0.1; WL: 0.74, 0.79; PSL: 0.08, 0.09; MTL: 0.57, 0.59; PNW: 0.31, 0.32; PTW: 0.09, 0.07; PPW: 0.12, 0.14; CI: 122.0, 122.7; SI: 153.6, 150.9; PSLI: 13.8, 15.4; PPI: 74.3, 52.9; PNI: 67.5, 65.7; MTI: 123.3, 120.5.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Head.</emphasis>
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Cephalic margin slightly convex (Fig.
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<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Pheidole dasos sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923284) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0212441)." figureDoi="10.3897/zookeys.949.51269.figure28" httpUri="https://binary.pensoft.net/fig/432197" pageId="0" pageNumber="1">28A</figureCitation>
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). Pilosity relatively sparse, moderately long, subdecumbent to erect. Sculpture shiny and foveolate; vertex and median frons with fading sculpture; area posterolateral from eyes smooth; antennal sockets with few indistinct, curved outward rugae and foveolate interspaces. Clypeus with median longitudinal carina absent; two lateral longitudinal carinae absent. Scape, when laid back, exceeding the posterior head margin by one-third of its length; pilosity dense, subdecumbent to erect (Fig.
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<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Pheidole dasos sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923284) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0212441)." figureDoi="10.3897/zookeys.949.51269.figure28" httpUri="https://binary.pensoft.net/fig/432197" pageId="0" pageNumber="1">28A, C</figureCitation>
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).
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Mesosoma.</emphasis>
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In lateral view, promesonotum low and moderately long, arched; promesonotal groove absent; metanotal groove distinct; propodeal spines small, triangular (Fig.
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<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Pheidole dasos sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923284) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0212441)." figureDoi="10.3897/zookeys.949.51269.figure28" httpUri="https://binary.pensoft.net/fig/432197" pageId="0" pageNumber="1">28C</figureCitation>
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). Sculpture shiny and predominantly smooth; lateral sides with very sparse and indistinct, thin rugulae. Pilosity very sparse, moderately long, and erect (Fig.
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<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Pheidole dasos sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923284) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0212441)." figureDoi="10.3897/zookeys.949.51269.figure28" httpUri="https://binary.pensoft.net/fig/432197" pageId="0" pageNumber="1">28C, E</figureCitation>
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).
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Gaster.</emphasis>
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With sparse, erect pilosity (Fig.
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<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Pheidole dasos sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923284) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0212441)." figureDoi="10.3897/zookeys.949.51269.figure28" httpUri="https://binary.pensoft.net/fig/432197" pageId="0" pageNumber="1">28C, E</figureCitation>
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).
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Colour.</emphasis>
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Brown, legs and gaster brighter (Fig.
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<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Pheidole dasos sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923284) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0212441)." figureDoi="10.3897/zookeys.949.51269.figure28" httpUri="https://binary.pensoft.net/fig/432197" pageId="0" pageNumber="1">28C, E</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="etymology">
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<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="1">Greek for forest in reference to habitat preferences of the species.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="biology_ecology">
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<paragraph pageId="0" pageNumber="1">Biology.</paragraph>
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<paragraph pageId="0" pageNumber="1">The species was collected between 225-415 m in elevation, in rainforest. Nests were located in rotten logs.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="comments">
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<paragraph pageId="0" pageNumber="1">Comments.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole dasos" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="dasos">
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<emphasis italics="true" pageId="0" pageNumber="1">Pheidole dasos</emphasis>
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</taxonomicName>
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sp. nov., described from Makirovana forest in Antsiranana, is most similar to
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<taxonomicName lsidName="P. hazo" pageId="0" pageNumber="1" rank="species" species="hazo">
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<emphasis italics="true" pageId="0" pageNumber="1">P. hazo</emphasis>
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</taxonomicName>
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sp. nov., known from the vicinity of Andranoma in Antananarivo, and
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<taxonomicName lsidName="P. sparsa" pageId="0" pageNumber="1" rank="species" species="sparsa">
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<emphasis italics="true" pageId="0" pageNumber="1">P. sparsa</emphasis>
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</taxonomicName>
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sp. nov., recorded so far only from Bemanevika in Mahajanga. Majors of
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<taxonomicName lsidName="P. dasos" pageId="0" pageNumber="1" rank="species" species="dasos">
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<emphasis italics="true" pageId="0" pageNumber="1">P. dasos</emphasis>
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</taxonomicName>
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sp. nov. can be distinguished from both taxa by medial part of frons with moderately dense rugae with interspaces with indistinct foveolae, and pronotum predominantly sculptured with smooth notches on lateral sides; minors of
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<taxonomicName lsidName="P. dasos" pageId="0" pageNumber="1" rank="species" species="dasos">
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<emphasis italics="true" pageId="0" pageNumber="1">P. dasos</emphasis>
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</taxonomicName>
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sp. nov. can be distinguished from
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<taxonomicName lsidName="P. sparsa" pageId="0" pageNumber="1" rank="species" species="sparsa">
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<emphasis italics="true" pageId="0" pageNumber="1">P. sparsa</emphasis>
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</taxonomicName>
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sp. nov. and
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<taxonomicName lsidName="P. hazo" pageId="0" pageNumber="1" rank="species" species="hazo">
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<emphasis italics="true" pageId="0" pageNumber="1">P. hazo</emphasis>
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</taxonomicName>
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sp. nov. by brown body colouration, longer scape exceeding the posterior head margin by one-third of its length, and predominantly smooth mesosoma with very sparse and indistinct, thin rugulae on lateral sides.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |