275 lines
36 KiB
XML
275 lines
36 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.463.8499" ID-PMC="PMC4294300" ID-Pensoft-Pub="1313-2970-463-21" ID-Pensoft-UUID="3E5CFFC6FFC91628FFF6E355951DFFF8" ID-PubMed="25589861" ID-Zenodo-Dep="578654" ID-ZooBank="71C1AE92B0CE4D26B1E5076E480180AD" ModsDocID="1313-2970-463-21" checkinTime="1451244962410" checkinUser="pensoft" docAuthor="Solodovnikov, Igor A., Zamotajlov, Alexandr S., Fominykh, Dmitriy D. & Titarenko, Andrey Y." docDate="2014" docId="0F50EB992CFD5483847B7E6066C59AF3" docLanguage="en" docName="ZooKeys 463: 21-56" docOrigin="ZooKeys 463" docPubDate="2014-12-12" docSource="http://dx.doi.org/10.3897/zookeys.463.8499" docTitle="Carabus (Archiplectes) pseudopshuensis Zamotajlov 1991" docType="treatment" docVersion="3" id="3E5CFFC6FFC91628FFF6E355951DFFF8" lastPageNumber="43" masterDocId="3E5CFFC6FFC91628FFF6E355951DFFF8" masterDocTitle="Contribution to the knowledge of the Carabus (Archiplectes) satyrus Kurnakov, 1962, species complex in Abkhazia (Coleoptera, Carabidae, Carabini)" masterLastPageNumber="56" masterPageNumber="21" pageNumber="38" updateTime="1668159733340" updateUser="ExternalLinkService">
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<mods:titleInfo>
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<mods:title>Contribution to the knowledge of the Carabus (Archiplectes) satyrus Kurnakov, 1962, species complex in Abkhazia (Coleoptera, Carabidae, Carabini)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Solodovnikov, Igor A.</mods:namePart>
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<mods:affiliation>Vitebsk State P. M. Masherov University, Moskovskyi prospect, 33, Vitebsk 210038, Belarus</mods:affiliation>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Zamotajlov, Alexandr S.</mods:namePart>
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<mods:affiliation>Kuban State Agrarian University, Kalinin St. 13, Krasnodar 350044, Russia</mods:affiliation>
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<mods:nameIdentifier type="email">a_zamotajlov@mail.ru</mods:nameIdentifier>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Fominykh, Dmitriy D.</mods:namePart>
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<mods:affiliation>Russian Entomological Society, Kuban Branch, Kalinin St. 13, Krasnodar 350044, Russia</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Titarenko, Andrey Y.</mods:namePart>
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<mods:affiliation>Publicly Traded Company " Morpho Absoloni ", Bolotnikovskaya St. 5 / 3, Moscow 117556, Russia</mods:affiliation>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
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<mods:part>
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<mods:date>2014</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2014-12-12</mods:number>
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<mods:detail type="volume">
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<mods:number>463</mods:number>
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<mods:extent unit="page">
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<mods:start>21</mods:start>
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<mods:url>http://dx.doi.org/10.3897/zookeys.463.8499</mods:url>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.463.8499</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-463-21</mods:identifier>
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<subSection lastPageId="22" lastPageNumber="43" pageId="17" pageNumber="38" type="taxonomy">
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<treatment ID-GBIF-Taxon="182234823" LSID="urn:lsid:plazi:treatment:0F50EB992CFD5483847B7E6066C59AF3" httpUri="http://treatment.plazi.org/id/0F50EB992CFD5483847B7E6066C59AF3" lastPageId="22" lastPageNumber="43" pageId="17" pageNumber="38">
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<subSubSection pageId="17" pageNumber="38" type="nomenclature">
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<paragraph pageId="17" pageNumber="38">
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<taxonomicName LSID="0F50EB99-2CFD-5483-847B-7E6066C59AF3" authority="Zamotajlov, 1991" authorityName="Zamotajlov" authorityYear="1991" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) pseudopshuensis" order="Coleoptera" pageId="17" pageNumber="38" phylum="Arthropoda" rank="species" species="pseudopshuensis" subGenus="Archiplectes">Carabus (Archiplectes) pseudopshuensis Zamotajlov, 1991</taxonomicName>
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Distribution of morphometric characters in males of Carabus (Archiplectes) besleticus resheviensis (River Reshevie), Carabus (Archiplectes) pseudopshuensis (River Reshevie), and Carabus (Archiplectes) juenthneri juenthneri (Village Pskhu env.) constructed using discriminant analysis based on 8 parameters." pageId="17" pageNumber="38">Figs 5</figureCitation>
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Distribution of morphometric characters in females of Carabus (Archiplectes) besleticus resheviensis (River Reshevie), Carabus (Archiplectes) pseudopshuensis (River Reshevie), and Carabus (Archiplectes) juenthneri juenthneri (Village Pskhu env.) constructed using discriminant analysis based on 8 parameters." pageId="17" pageNumber="38">, 6</figureCitation>
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<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Distribution map of Carabus (Archiplectes) satyrus, Carabus (Archiplectes) besleticus, and Carabus (Archiplectes) pseudopshuensis in Abkhazia." pageId="17" pageNumber="38">, 7</figureCitation>
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<figureCitation captionStart="Figures 8–15" captionStartId="F8" captionText="Figures 8 - 15. Fully inflated endophallus preparation and apical lamella of aedeagus of Carabus (Archiplectes) satyrus and Carabus (Archiplectes) pseudopshuensis. 8 - 11 Carabus (Archiplectes) satyrus 12 - 15 Carabus (Archiplectes) pseudopshuensis." pageId="17" pageNumber="38">, 12-15</figureCitation>
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<figureCitation captionStart="Figures 78–81" captionStartId="F19" captionText="Figures 78 - 81. Dorsal habitus, Carabus (Archiplectes) pseudopshuensis. 78 - 79 Abkhazia, Village Pskhu env 80 - 81 Abkhazia, N slopes of Bzybian Mt. Range, Village Serebryanyi env." pageId="17" pageNumber="38">, 78-89</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="18" pageNumber="39" type="reference_group">
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<paragraph pageId="18" pageNumber="39">
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<taxonomicName class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) satyrus subsp. pseudopshuensis" order="Coleoptera" pageId="18" pageNumber="39" phylum="Arthropoda" rank="subSpecies" species="satyrus" subGenus="Archiplectes" subSpecies="pseudopshuensis">
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<pageBreakToken pageId="18" pageNumber="39" start="start">Carabus</pageBreakToken>
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(Archiplectes) satyrus pseudopshuensis
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</taxonomicName>
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Zamotajlov, 1991: 36 ("Environs of Pskhu");
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<bibRefCitation author="Bousquet, Y" journalOrPublisher="Catalogue of Palaearctic Coleoptera" pageId="23" pageNumber="44" pagination="118 - 201" refId="B2" refString="Bousquet, Y, Brezina, B, Davies, A, Farkac, J, Smetana, A, 2003. Tribe Carabini Latreille, 1802. Catalogue of Palaearctic Coleoptera Vol. 1: 118 - 201" title="Tribe Carabini Latreille, 1802." volume="Vol. 1" year="2003">Bousquet et al. 2003</bibRefCitation>
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: 132;
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<bibRefCitation author="Retezar, I" journalOrPublisher="Mesterprint Kft., Budapest" pageId="23" pageNumber="44" refId="B8" refString="Retezar, I, 2008. The Carabus of Abkhazia, Caucasus (Coleoptera, Carabidae): Iconography, genital morphology, systematics and faunistics. Mesterprint Kft., Budapest" title="The Carabus of Abkhazia, Caucasus (Coleoptera, Carabidae): Iconography, genital morphology, systematics and faunistics." year="2008">
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<normalizedToken originalValue="Retezár">Retezar</normalizedToken>
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2008
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</bibRefCitation>
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: 41;
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</paragraph>
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<paragraph pageId="18" pageNumber="39">
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<taxonomicName class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Tribax) satyrus subsp. pseudopshuensis" order="Coleoptera" pageId="18" pageNumber="39" phylum="Arthropoda" rank="subSpecies" species="satyrus" subGenus="Tribax" subSpecies="pseudopshuensis">Carabus (Tribax) satyrus pseudopshuensis</taxonomicName>
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:
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<bibRefCitation author="Deuve, T" journalOrPublisher="Pensoft Publishers, Sofia-Moscow" pageId="23" pageNumber="44" refId="B3" refString="Deuve, T, 2004. Illustrated Catalogue of the Genus Carabus of the World (Coleoptera: Carabidae). Pensoft Publishers, Sofia-Moscow" title="Illustrated Catalogue of the Genus Carabus of the World (Coleoptera: Carabidae)." year="2004">Deuve 2004</bibRefCitation>
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: 275.
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</paragraph>
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<paragraph pageId="18" pageNumber="39">
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<taxonomicName authorityName="Zamotajlov" authorityYear="1991" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) pseudopshuensis" order="Coleoptera" pageId="18" pageNumber="39" phylum="Arthropoda" rank="species" species="pseudopshuensis" subGenus="Archiplectes">Carabus (Archiplectes) pseudopshuensis</taxonomicName>
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:
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<bibRefCitation author="Fominykh, DD" journalOrPublisher="Galanika, St. Petersburg" pageId="23" pageNumber="44" refId="B4" refString="Fominykh, DD, Zamotajlov, AS, 2012. Contribution to the knowledge of the Carabus subgenus Archiplectes (Coleoptera, Carabidae) of the middle course of Bzyb River. XIV Congress of the Russian Entomological Society. Saint Petersburg, August 27 - September 1, 2012. Materials of the Congress. Galanika, St. Petersburg" title="Contribution to the knowledge of the Carabus subgenus Archiplectes (Coleoptera, Carabidae) of the middle course of Bzyb River. XIV Congress of the Russian Entomological Society. Saint Petersburg, August 27 - September 1, 2012. Materials of the Congress." year="2012">Fominykh and Zamotajlov 2012</bibRefCitation>
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: 444;
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<bibRefCitation author="Retezar, I" journalOrPublisher="Folia Entomologica Hungarica" pageId="23" pageNumber="44" pagination="1 - 6" refId="B9" refString="Retezar, I, 2013. Taxonomic and faunistic studies on the subgenus Carabus (Archiplectes) Gottwald, 1982 in Abkhazia (Coleoptera: Carabidae). Folia Entomologica Hungarica 74: 1 - 6" title="Taxonomic and faunistic studies on the subgenus Carabus (Archiplectes) Gottwald, 1982 in Abkhazia (Coleoptera: Carabidae)." volume="74" year="2013">
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<normalizedToken originalValue="Retezár">Retezar</normalizedToken>
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2013
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</bibRefCitation>
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: 4.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="21" lastPageNumber="42" pageId="18" pageNumber="39" type="comparative material examined">
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<paragraph pageId="18" pageNumber="39">Comparative material examined.</paragraph>
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<paragraph lastPageId="21" lastPageNumber="42" pageId="18" pageNumber="39">
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3351 specimens were examined (135 specimens measured, 60 male genitalia preparations studied): 2 males, 4 females, Abkhazia, Sukhum Distr., left bank of River Aguripsta near outfall of River Belaya, beech forest
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<pageBreakToken pageId="19" pageNumber="40" start="start">site</pageBreakToken>
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, 800-1000 m, 11.V-27.VI.2010, leg. D. Fominykh, I. Solodovnikov (cZAM, cFDD, cSIA); 5 males, 5 females, Abkhazia, Sukhum Distr., left bank of River Aguripsta near Mt. Svyataya, 800 m, 10.V-28.VI.2010, leg. D. Fominykh, I. Solodovnikov (cZAM, cFDD); 4 males, 2 females, Abkhazia, Sukhum Distr., vicinities of Village Pskhu, S slope of Mt. Svyataya, beech forest site, 650-700 m, 10.V-28.VI.2010, leg. I. Solodovnikov, D. Fominykh, (cSIA); 21 males, 25 females, Abkhazia, Sukhum Distr., SE of Village Pskhu, hamlet Bitaga, Mt. Bzybskaya, 550-650 m, beech forest site, 12.V-30.VI.2010, leg. I. Solodovnikov, D. Fominykh, (cSIA); 547 males, 763 females, Abkhazia, Sukhum Distr., left bank of River Aguripsta near Village Pskhu, W slopes of Mt. Chibiskha, 700 m, 25.IV-04.VII.2013, leg. D. Fominykh (cZAM, cFDD, cSAA, cBAS); 48 males, 62 females, Abkhazia, Sukhum Distr., right bank of River Bzyb, S slopes of Mt. Chibiskha, 550 m, 25.IV-04.VII.2013, leg. D. Fominykh (cZAM, cFDD); 5 males, 4 females, Abkhazia, Sukhum Distr., right bank of River Bzyb, S slopes of Mt. Chibiskha upstream hamlet Bitaga, karst, beech forest site, 600-620 m,
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<geoCoordinate degrees="43" direction="north" minutes="21" orientation="latitude" precision="1" seconds="29.36" value="43.358154">N43°21'29.36"</geoCoordinate>
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/
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<geoCoordinate degrees="040" direction="east" minutes="49" orientation="longitude" precision="1" seconds="58.89" value="40.833023">E040°49'58.89"</geoCoordinate>
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, 25.IV-04.VII.2013, leg. D. Fominykh (cSIA); 6 males, 6 females, Abkhazia, Sukhum Distr., right bank of River Baul near Village Sanchara, 1030 m,
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<geoCoordinate degrees="43" direction="north" minutes="24,9" orientation="latitude" precision="92" value="43.415">N43°24,9'</geoCoordinate>
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,
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<geoCoordinate degrees="040" direction="east" minutes="51,4" orientation="longitude" precision="92" value="40.856667">E040°51,4'</geoCoordinate>
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, VII.2010, leg. I.
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<normalizedToken originalValue="Retezár">Retezar</normalizedToken>
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(cZAM, cFDD); 2 males, 2 females, Abkhazia, Sukhum Distr., vicinities of Village Sanchara, River Baul, beech forest site, 20.VII.1991, leg. M.N. Maksimenkov (cSIA); 2 females, same locality, 800 m, 10.VIII.1989, leg. M.N. Maksimenkov (cSIA); 2 males, Village Sanchara, VI.1991, leg. M.N. Maksimenkov (cSIA); 1 male, Abkhazia, Sukhum Distr., Pskhu, 10.IX.1991, local collector (cSIA); 3 males, 4 females, Abkhazia, Sukhum Distr., Bzybian Mt. Range, upper reaches of RRiver West Gumista, Dou
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<pageBreakToken pageId="20" pageNumber="41" start="start">Pass</pageBreakToken>
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, 1300 m, 30.IV-19.VII.2012, leg. D. Fominykh (cFDD); 10 males, 13 females, Abkhazia, Sukhum Distr., N slopes of Bzybian Mt. Range near Bzyb valley, 1000 m, 30.IV.2012, leg. D. Fominykh (cZAM, cFDD, cSAA); 40 males, 30 females,
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<pageBreakToken pageId="21" pageNumber="42" start="start">Abkhazia</pageBreakToken>
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, Sukhum Distr., N slopes of Bzybian Mt. Range near Bzyb valley, 800 m, 30.IV-19.VII.2012, leg. D. Fominykh (cZAM, cFDD, cSIA, cSAA); 351 males, 449 females, Abkhazia, Sukhum Distr., N slopes of Bzybian Mt. Range, right bank of River Reshevie, 700-1000 m, 1.V-20.VII.2012, leg. D. Fominykh (cZAM, cFDD, cSAA, cBAS); 2 males, 5 females, Abkhazia, Sukhum Distr., left bank of River Bzyb, right bank of River Reshevie, beech, maple, fern forest site, 670-720 m, 13-14.V.2010, leg. I. Solodovnikov (cSIA); 9 males, 18 females, same locality, 13.V-02.VII.2010, leg. I. Solodovnikov, D. Fominykh (cSIA, cPIG, cKVM); 14 males, 16 females, same locality, 04.V-20.VII.2012, leg. D. Fominykh, A. Safronov (cSIA); 386 males, 516 females, Abkhazia, Sukhum Distr., N Slopes of Bzybian Mt. Range near Village Serebryanyi, 500-600 m, 25.IV-05.VII.2013, leg. D. Fominykh (cZAM, cFDD); 5 males, 8 females, Abkhazia, Sukhum Distr., vicinities of Village Pskhu, hamlet Serebryanyi, left bank of River Bzyb, karst, beech forest site, 600-620 m,
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<geoCoordinate degrees="43" direction="north" minutes="21" orientation="latitude" precision="1" seconds="52.90" value="43.364693">N43°21'52.90"</geoCoordinate>
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/
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<geoCoordinate degrees="040" direction="east" minutes="48" orientation="longitude" precision="1" seconds="52.90" value="40.814693">E040°48'52.90"</geoCoordinate>
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, 25.IV-05.VII.2013, leg. D. Fominykh (cSIA).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="21" pageNumber="42" type="description">
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<paragraph pageId="21" pageNumber="42">Description.</paragraph>
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<paragraph pageId="21" pageNumber="42">Medium-sized form, length 26.5-34.0 mm, males 28.0-32.0 mm, females 28.5-34.0 mm long. Underside black, dorsum normally with bright metallic lustre, green, dark blue, bronze, crimson, violet, less often black, with transitional color forms in males, dichromatic and trichromatic individuals being also known, females matte. Body slender.</paragraph>
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<paragraph pageId="21" pageNumber="42">
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Head not inflated. Pronotum variable in shape, from transverse to subquadrate. PW/PL of individuals from the right bank of River Bzyb hardly depends upon altitude in males and constitute constantly 1.32-1.39, in females, upon decrease of altitude, PW/PL varies from 1.38 at 700 m to 1.48 at 1100 m. Hind angles slightly protruding backwards. In the left-bank populations, pronotum of nearly the same proportions or less transverse, particularly in females. PW/PL varies there in males of three studied populations from 1.26 to 1.39, and in females - from 1.34 to 1.42, reaching extreme value of 1.23 in the population, inhabiting environs of the hamlet Serebryanyi. Elytra oblong-ovate in the right-bank populations, also vary in shape from distinctly elongate to almost ovate. EL/EW gradually varies in males depending upon altitude decrease from 1.89 to 1.69, in females such correlations are not obviously revealed and irregularly varies from 1.69 to 1.76. Elytral sculpture varies from almost smooth to coarsely granular, with strongly pronounced punctation in the middle interspaces. EL/EW is more or less stable in the left-bank populations and measures 1.66-1.70 in males and 1.60-1.75 in females. Elytral sculpture granular, interspaces regular, nearly straight, some male specimens possess somewhat confused elytral sculpture. The main morphometric measurements are presented in Table
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<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Morphometric characteristic of males (n = 67) and females (n = 68) of Carabus (Archiplectes) pseudopshuensis." httpUri="http://table.plazi.org/id/0C7DFCF5F18DFCF33E2586198364B05A" pageId="21" pageNumber="42" tableUuid="0C7DFCF5F18DFCF33E2586198364B05A">1</tableCitation>
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.
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</paragraph>
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<paragraph pageId="21" pageNumber="42">
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The structure of endophallus testifies to the assignment of this form to the
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<taxonomicName class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="reitteri" order="Coleoptera" pageId="21" pageNumber="42" phylum="Arthropoda" rank="species" species="reitteri">
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<emphasis italics="true" pageId="21" pageNumber="42">reitteri</emphasis>
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</taxonomicName>
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-group. It is easily distinguishable from
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<taxonomicName authorityName="Kurnakov" authorityYear="1962" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) satyrus" order="Coleoptera" pageId="21" pageNumber="42" phylum="Arthropoda" rank="species" species="satyrus" subGenus="Archiplectes">Carabus (Archiplectes) satyrus</taxonomicName>
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,
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<taxonomicName authorityName="Kurnakov" authorityYear="1972" baseAuthorityName="besleticus Kurnakov" baseAuthorityYear="1972" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) besleticus" order="Coleoptera" pageId="21" pageNumber="42" phylum="Arthropoda" rank="species" species="besleticus" subGenus="Archiplectes">Carabus (Archiplectes) besleticus</taxonomicName>
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, and
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<taxonomicName baseAuthorityName="Reitter" baseAuthorityYear="1899" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) juenthneri" order="Coleoptera" pageId="21" pageNumber="42" phylum="Arthropoda" rank="species" species="juenthneri" subGenus="Archiplectes">Carabus (Archiplectes) juenthneri</taxonomicName>
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by the shape of the dorsal appendix, which is less elongate and possesses a more rectangular form.
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</paragraph>
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<caption pageId="21" pageNumber="42" start="Figures 78–81" startId="F19">
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<paragraph pageId="21" pageNumber="42">
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<emphasis bold="true" pageId="21" pageNumber="42">Figures 78-81.</emphasis>
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Dorsal habitus,
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<taxonomicName authorityName="Zamotajlov" authorityYear="1991" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) pseudopshuensis" order="Coleoptera" pageId="21" pageNumber="42" phylum="Arthropoda" rank="species" species="pseudopshuensis" subGenus="Archiplectes">Carabus (Archiplectes) pseudopshuensis</taxonomicName>
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.
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<emphasis bold="true" pageId="21" pageNumber="42">78-79</emphasis>
|
||
Abkhazia, Village Pskhu env
|
||
<emphasis bold="true" pageId="21" pageNumber="42">80-81</emphasis>
|
||
Abkhazia, N slopes of Bzybian Mt. Range, Village Serebryanyi env.
|
||
</paragraph>
|
||
</caption>
|
||
<caption pageId="21" pageNumber="42" start="Figures 82–89" startId="F20">
|
||
<paragraph pageId="21" pageNumber="42">
|
||
<emphasis bold="true" pageId="21" pageNumber="42">Figures 82-89.</emphasis>
|
||
Dorsal habitus,
|
||
<taxonomicName authorityName="Zamotajlov" authorityYear="1991" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) pseudopshuensis" order="Coleoptera" pageId="21" pageNumber="42" phylum="Arthropoda" rank="species" species="pseudopshuensis" subGenus="Archiplectes">Carabus (Archiplectes) pseudopshuensis</taxonomicName>
|
||
.
|
||
<emphasis bold="true" pageId="21" pageNumber="42">82-83, 88-89</emphasis>
|
||
Abkhazia, N slopes of Bzybian Mt. Range, Village Serebryanyi env
|
||
<emphasis bold="true" pageId="21" pageNumber="42">84</emphasis>
|
||
Abkhazia, Village Sanchara env
|
||
<emphasis bold="true" pageId="21" pageNumber="42">85</emphasis>
|
||
Abkhazia, Village Pskhu env., Mt. Svyataya
|
||
<emphasis bold="true" pageId="21" pageNumber="42">86-87</emphasis>
|
||
Abkhazia, Village Pskhu env.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="22" lastPageNumber="43" pageId="21" pageNumber="42" type="differential diagnosis">
|
||
<paragraph pageId="21" pageNumber="42">Differential diagnosis.</paragraph>
|
||
<paragraph lastPageId="22" lastPageNumber="43" pageId="21" pageNumber="42">
|
||
Strongly resembles
|
||
<taxonomicName authority="juenthneri" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) juenthneri subsp. juenthneri" order="Coleoptera" pageId="22" pageNumber="43" phylum="Arthropoda" rank="subSpecies" species="juenthneri" subGenus="Archiplectes" subSpecies="juenthneri">
|
||
<pageBreakToken pageId="22" pageNumber="43" start="start">Carabus</pageBreakToken>
|
||
(Archiplectes) juenthneri juenthneri
|
||
</taxonomicName>
|
||
, which occurs sympatrically in some localities on the right bank of River Bzyb; however, it differs in smaller body size, more elongate and ovate body, and also in the structure of the male genitalia. Upstream, in the valley of River Aguripsta,
|
||
<taxonomicName authority="juenthneri" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) juenthneri subsp. juenthneri" order="Coleoptera" pageId="22" pageNumber="43" phylum="Arthropoda" rank="subSpecies" species="juenthneri" subGenus="Archiplectes" subSpecies="juenthneri">Carabus (Archiplectes) juenthneri juenthneri</taxonomicName>
|
||
completely substitutes this taxon. On the left bank of River Bzyb it also occurs sympatrically with
|
||
<taxonomicName authorityName="Solodovnikov, Zamotajlov & Fominykh" authorityYear="2014" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) besleticus subsp. resheviensis" order="Coleoptera" pageId="22" pageNumber="43" phylum="Arthropoda" rank="subSpecies" species="besleticus" subGenus="Archiplectes" subSpecies="resheviensis">Carabus (Archiplectes) besleticus resheviensis</taxonomicName>
|
||
from which it differs in considerably smaller body size and different male genitalia. The authors have succeeded in collecting in such localities few transitional individuals, apparently being hybrid forms. Morphometric characters are illustrated in Figures
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Distribution of morphometric characters in males of Carabus (Archiplectes) besleticus resheviensis (River Reshevie), Carabus (Archiplectes) pseudopshuensis (River Reshevie), and Carabus (Archiplectes) juenthneri juenthneri (Village Pskhu env.) constructed using discriminant analysis based on 8 parameters." pageId="22" pageNumber="43">5</figureCitation>
|
||
-
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Distribution of morphometric characters in females of Carabus (Archiplectes) besleticus resheviensis (River Reshevie), Carabus (Archiplectes) pseudopshuensis (River Reshevie), and Carabus (Archiplectes) juenthneri juenthneri (Village Pskhu env.) constructed using discriminant analysis based on 8 parameters." pageId="22" pageNumber="43">6</figureCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="22" pageNumber="43" type="distribution">
|
||
<paragraph pageId="22" pageNumber="43">Distribution.</paragraph>
|
||
<paragraph pageId="22" pageNumber="43">
|
||
Populates the left bank of the River Aguripsta from the confluence of River Belaya (its left tributary) up to its outfall, and also right board of River Bzyb, upstream confluence of River Aguripsta, also occurs at northern slopes of Bzybian Mt. Range, reaching its watershed at 1300 m, where dwells together with
|
||
<taxonomicName authorityName="Kurnakov" authorityYear="1962" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) apollo subsp. tenebricosus" order="Coleoptera" pageId="22" pageNumber="43" phylum="Arthropoda" rank="subSpecies" species="apollo" subGenus="Archiplectes" subSpecies="tenebricosus">Carabus (Archiplectes) apollo tenebricosus</taxonomicName>
|
||
, however it has not been found at its southern slopes, where apparently it is completely substituted by the latter.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="22" pageNumber="43" type="habitat">
|
||
<paragraph pageId="22" pageNumber="43">Habitat.</paragraph>
|
||
<paragraph pageId="22" pageNumber="43">
|
||
Inhabits mainly beech and fir-beech mixed forests, developed on rich ground litter, and also petrous forest taluses at karstic landforms. The population density is rather high in its known geographic range, except for the middle flow of River Aguripsta (natural border of its distribution area), the maximum density of imago is recorded for the altitude interval of 600-900 m. Activity of imago proceeds from April to July, solitary individuals are sometimes observed at the end of summer or the middle of September. The following forms of
|
||
<taxonomicName authorityName="Linnaeus" authorityYear="1758" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus" order="Coleoptera" pageId="22" pageNumber="43" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="22" pageNumber="43">Carabus</emphasis>
|
||
</taxonomicName>
|
||
occur sympatrically on the left bank of River Bzyb:
|
||
<taxonomicName authorityName="Solodovnikov, Zamotajlov & Fominykh" authorityYear="2014" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) besleticus subsp. resheviensis" order="Coleoptera" pageId="22" pageNumber="43" phylum="Arthropoda" rank="subSpecies" species="besleticus" subGenus="Archiplectes" subSpecies="resheviensis">Carabus (Archiplectes) besleticus resheviensis</taxonomicName>
|
||
,
|
||
<taxonomicName authorityName="Kurnakov" authorityYear="1962" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) apollo subsp. tenebricosus" order="Coleoptera" pageId="22" pageNumber="43" phylum="Arthropoda" rank="subSpecies" species="apollo" subGenus="Archiplectes" subSpecies="tenebricosus">Carabus (Archiplectes) apollo tenebricosus</taxonomicName>
|
||
,
|
||
<taxonomicName authority="apschuanus" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Tribax) apschuanus subsp. apschuanus" order="Coleoptera" pageId="22" pageNumber="43" phylum="Arthropoda" rank="subSpecies" species="apschuanus" subGenus="Tribax" subSpecies="apschuanus">Carabus (Tribax) apschuanus apschuanus</taxonomicName>
|
||
,
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Tribax) constantinovi subsp. otcharensis" order="Coleoptera" pageId="22" pageNumber="43" phylum="Arthropoda" rank="subSpecies" species="constantinovi" subGenus="Tribax" subSpecies="otcharensis">Carabus (Tribax) constantinovi otcharensis</taxonomicName>
|
||
,
|
||
<taxonomicName authority="circassicus" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Tribax) circassicus subsp. circassicus" order="Coleoptera" pageId="22" pageNumber="43" phylum="Arthropoda" rank="subSpecies" species="circassicus" subGenus="Tribax" subSpecies="circassicus">Carabus (Tribax) circassicus circassicus</taxonomicName>
|
||
(natio
|
||
<taxonomicName class="Insecta" family="Carabidae" form="abasinus" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="22" pageNumber="43" phylum="Arthropoda" rank="form">
|
||
<emphasis italics="true" pageId="22" pageNumber="43">abasinus</emphasis>
|
||
</taxonomicName>
|
||
),
|
||
<taxonomicName authorityName="Mandl" authorityYear="1955" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Microplectes) argonautarum subsp. reischitzi" order="Coleoptera" pageId="22" pageNumber="43" phylum="Arthropoda" rank="subSpecies" species="argonautarum" subGenus="Microplectes" subSpecies="reischitzi">Carabus (Microplectes) argonautarum reischitzi</taxonomicName>
|
||
,
|
||
<taxonomicName authorityName="Motschulsky" authorityYear="1840" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Megodontus) septemcarinatus" order="Coleoptera" pageId="22" pageNumber="43" phylum="Arthropoda" rank="species" species="septemcarinatus" subGenus="Megodontus">Carabus (Megodontus) septemcarinatus</taxonomicName>
|
||
,
|
||
<taxonomicName authorityName="Fischer von Waldheim" authorityYear="1823" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Eucarabus) cumanus" order="Coleoptera" pageId="22" pageNumber="43" phylum="Arthropoda" rank="species" species="cumanus" subGenus="Eucarabus">Carabus (Eucarabus) cumanus</taxonomicName>
|
||
Fischer von Waldheim, 1823,
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Sphodristocarabus) armeniacus subsp. dvorschaki" order="Coleoptera" pageId="22" pageNumber="43" phylum="Arthropoda" rank="subSpecies" species="armeniacus" subGenus="Sphodristocarabus" subSpecies="dvorschaki">Carabus (Sphodristocarabus) armeniacus dvorschaki</taxonomicName>
|
||
, and
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Procerus) caucasicus subsp. colchicus" order="Coleoptera" pageId="22" pageNumber="43" phylum="Arthropoda" rank="subSpecies" species="caucasicus" subGenus="Procerus" subSpecies="colchicus">Carabus (Procerus) caucasicus colchicus</taxonomicName>
|
||
; on the right bank of River Bzyb:
|
||
<taxonomicName authority="juenthneri" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) juenthneri subsp. juenthneri" order="Coleoptera" pageId="22" pageNumber="43" phylum="Arthropoda" rank="subSpecies" species="juenthneri" subGenus="Archiplectes" subSpecies="juenthneri">Carabus (Archiplectes) juenthneri juenthneri</taxonomicName>
|
||
,
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Tribax) apschuanus subsp. schoeni" order="Coleoptera" pageId="22" pageNumber="43" phylum="Arthropoda" rank="subSpecies" species="apschuanus" subGenus="Tribax" subSpecies="schoeni">Carabus (Tribax) apschuanus schoeni</taxonomicName>
|
||
<normalizedToken originalValue="Novotný">Novotny</normalizedToken>
|
||
&
|
||
<normalizedToken originalValue="Voříšek">Vorisek</normalizedToken>
|
||
, 1988,
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Tribax) constantinovi subsp. otcharensis" order="Coleoptera" pageId="22" pageNumber="43" phylum="Arthropoda" rank="subSpecies" species="constantinovi" subGenus="Tribax" subSpecies="otcharensis">Carabus (Tribax) constantinovi otcharensis</taxonomicName>
|
||
,
|
||
<taxonomicName authority="circassicus" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Tribax) circassicus subsp. circassicus" order="Coleoptera" pageId="22" pageNumber="43" phylum="Arthropoda" rank="subSpecies" species="circassicus" subGenus="Tribax" subSpecies="circassicus">Carabus (Tribax) circassicus circassicus</taxonomicName>
|
||
(natio
|
||
<taxonomicName class="Insecta" family="Carabidae" form="abasinus" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="22" pageNumber="43" phylum="Arthropoda" rank="form">
|
||
<emphasis italics="true" pageId="22" pageNumber="43">abasinus</emphasis>
|
||
</taxonomicName>
|
||
),
|
||
<taxonomicName authorityName="Mandl" authorityYear="1955" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Microplectes) argonautarum subsp. reischitzi" order="Coleoptera" pageId="22" pageNumber="43" phylum="Arthropoda" rank="subSpecies" species="argonautarum" subGenus="Microplectes" subSpecies="reischitzi">Carabus (Microplectes) argonautarum reischitzi</taxonomicName>
|
||
,
|
||
<taxonomicName authorityName="Motschulsky" authorityYear="1840" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Megodontus) septemcarinatus" order="Coleoptera" pageId="22" pageNumber="43" phylum="Arthropoda" rank="species" species="septemcarinatus" subGenus="Megodontus">Carabus (Megodontus) septemcarinatus</taxonomicName>
|
||
,
|
||
<taxonomicName authorityName="Fischer von Waldheim" authorityYear="1823" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Eucarabus) cumanus" order="Coleoptera" pageId="22" pageNumber="43" phylum="Arthropoda" rank="species" species="cumanus" subGenus="Eucarabus">Carabus (Eucarabus) cumanus</taxonomicName>
|
||
,
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Pachycarabus) imitator subsp. katharinae" order="Coleoptera" pageId="22" pageNumber="43" phylum="Arthropoda" rank="subSpecies" species="imitator" subGenus="Pachycarabus" subSpecies="katharinae">Carabus (Pachycarabus) imitator katharinae</taxonomicName>
|
||
Reitter, 1896,
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Sphodristocarabus) armeniacus subsp. janthinus" order="Coleoptera" pageId="22" pageNumber="43" phylum="Arthropoda" rank="subSpecies" species="armeniacus" subGenus="Sphodristocarabus" subSpecies="janthinus">Carabus (Sphodristocarabus) armeniacus janthinus</taxonomicName>
|
||
Ganglbauer, 1887, and
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Procerus) caucasicus subsp. colchicus" order="Coleoptera" pageId="22" pageNumber="43" phylum="Arthropoda" rank="subSpecies" species="caucasicus" subGenus="Procerus" subSpecies="colchicus">Carabus (Procerus) caucasicus colchicus</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</subSection>
|
||
</document> |