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<mods:title id="9C1E9795E2FAEBCB5C5B609AEFB84FF4">A review of the genus Bifidocoelotes Wang, 2002, with the description of three new species from China (Araneae, Agelenidae)</mods:title>
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<treatment id="03CE87C2D705FF8BB798FDC9FA3CFB42" ID-DOI="http://doi.org/10.5281/zenodo.7469747" ID-GBIF-Taxon="207883698" ID-Zenodo-Dep="7469747" LSID="urn:lsid:plazi:treatment:03CE87C2D705FF8BB798FDC9FA3CFB42" httpUri="http://treatment.plazi.org/id/03CE87C2D705FF8BB798FDC9FA3CFB42" lastPageId="2" lastPageNumber="181" pageId="1" pageNumber="180">
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<paragraph id="8BD836D4D705FF88B798FDC9FD93FD74" blockId="1.[151,563,618,645]" box="[151,563,618,645]" pageId="1" pageNumber="180">
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Genus
<taxonomicName id="4C674D57D705FF88B7E1FDC9FD93FD74" ID-CoL="62G7F" ID-ENA="1983079" authority="Wang, 2002" authorityName="Wang" authorityYear="2002" box="[238,563,618,645]" class="Arachnida" family="Agelenidae" genus="Bifidocoelotes" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="180" phylum="Arthropoda" rank="genus">
<emphasis id="B913EAC6D705FF88B7E1FDC9FE39FD75" bold="true" box="[238,409,618,645]" italics="true" pageId="1" pageNumber="180">Bifidocoelotes</emphasis>
<bibRefCitation id="EFF64B25D705FF88B6AFFDC8FD93FD74" author="Wang, X. P." box="[416,563,618,645]" pageId="1" pageNumber="180" pagination="1 - 150" refId="ref6635" refString="Wang, X. P. (2002) A generic-level revision of the spider subfamily Coelotinae (Araneae, Amaurobiidae). Bulletin of the American Museum of Natural History, 269, 1 - 150. https: // doi. org / 10.1206 / 0003 - 0090 (2002) 269 &lt;0001: AGLROT&gt; 2.0. CO; 2" type="journal article" year="2002">Wang, 2002</bibRefCitation>
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<typeStatus id="54DC8876D705FF88B798FD0CFF6CFD37" box="[151,204,687,711]" pageId="1" pageNumber="180">Type</typeStatus>
species:
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<taxonomicName id="4C674D57D705FF88B623FD0CFD3FFD37" ID-CoL="99CNM" authority="Li &amp; Blick, 2020" authorityName=": Li &amp; Blick" authorityYear="2020" box="[300,671,687,711]" class="Arachnida" family="Agelenidae" genus="Bifidocoelotes" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="180" phylum="Arthropoda" rank="species" species="tsoi">
<emphasis id="B913EAC6D705FF88B623FD0CFE48FD37" box="[300,488,687,711]" italics="true" pageId="1" pageNumber="180">Bifidocoelotes tsoi</emphasis>
<bibRefCitation id="EFF64B25D705FF88B6E1FD0CFD3FFD37" author="Li, S. Q. &amp; Blick, T." box="[494,671,687,711]" pageId="1" pageNumber="180" pagination="118" refId="ref6452" refString="Li, S. Q. &amp; Blick, T. (2020) Bifidocoelotes tsoi, a new name to replace Bifidocoelotes bifidus (Araneae, Agelenidae). Acta Arachnologica Sinica, 29 (2), 118. https: // doi. org / 10.3969 / j. issn. 1005 - 9628.2020.02.009" type="journal article" year="2020">Li &amp; Blick, 2020</bibRefCitation>
</taxonomicName>
.
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<paragraph id="8BD836D4D705FF88B798FD56FC36FBDF" blockId="1.[151,1437,757,1072]" pageId="1" pageNumber="180">
<emphasis id="B913EAC6D705FF88B798FD56FEB4FCFF" bold="true" box="[151,276,757,783]" pageId="1" pageNumber="180">Diagnosis:</emphasis>
Females can be recognized by the combination of the following characteristics: a single epigynal tooth pale in color and variably keratinized (this kind of tooth is often difficult to see) which originates on the anterior margin of the epigyne, is at least half the length of epigyne, and has a bifurcated tip of varying degrees (
<figureCitation id="135C2A51D705FF88B217FC9EFA2FFCA7" box="[1304,1423,829,855]" captionStart="FIGURE 1" captionStartId="1.[151,250,1666,1690]" captionTargetBox="[163,1418,1102,1635]" captionTargetId="figure-290@1.[151,1436,1093,1640]" captionTargetPageId="1" captionText="FIGURE 1. The epigynes of six Bifidocoelotes spp., ventral view. A. Bifidocoelotes obscurus (after Zhou et al. 2017); B. Bifidocoelotes primus (after Zhou et al. 2017); C. Bifidocoelotes tsoi (after Wang et al. 2001); D. Bifidocoelotes mammiformis sp. nov.; E. Bifidocoelotes elongatus sp. nov.; F. Bifidocoelotes quadratus sp. nov.. Scale bar: 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.7456721" httpUri="https://zenodo.org/record/7456721/files/figure.png" pageId="1" pageNumber="180">Fig. 1AF</figureCitation>
); the presence of a spermathecal head, and the different configuration of spermathecae and copulatory ducts. Males can be recognized by the combination of the following characteristics: a more or less bifid conductor; a long and slender embolus originating proximally; an extremely long cymbial furrow; a relatively small and spoon-like median apophysis; a broad retrolateral tibial apophysis; a lateral tibial apophysis present and a finger-like patellar apophysis (
<figureCitation id="135C2A51D705FF88B61AFC52FE34FBFB" box="[277,404,1009,1035]" captionStart="FIGURE 2" captionStartId="2.[151,250,435,459]" captionTargetBox="[151,1432,181,408]" captionTargetId="figure-31@2.[151,1436,181,412]" captionTargetPageId="2" captionText="FIGURE 2. The distal parts of conductors of four Bifidocoelotes spp., ventral view.A. Bifidocoelotes mammiformis sp. nov.; B. Bifidocoelotes tsoi (after Wang et al. 2001); C. Bifidocoelotes primus (after Zhou et al. 2017); D. Bifidocoelotes obscurus (after Zhou et al. 2017). Scale bars: A, B, D = 0.1 mm; C = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.7456725" httpUri="https://zenodo.org/record/7456725/files/figure.png" pageId="1" pageNumber="180">Figs 2AD</figureCitation>
,
<figureCitation id="135C2A51D705FF88B691FC52FE41FBFB" box="[414,481,1009,1035]" captionStart="FIGURE 4" captionStartId="5.[151,250,1878,1902]" captionTargetBox="[228,1357,181,1854]" captionTargetId="figure-19@5.[228,1360,181,1854]" captionTargetPageId="5" captionText="FIGURE 4. Bifidocoelotes mammiformis sp. nov., holotype male. A. Habitus, dorsal view; B. Same, ventral view; C. Left palp, prolateral view; D. Same, ventral view; E. Same, retrolateral view. Abbreviations: CF, cymbial furrow; CL, conductor lamella; Co, conductor; Em, embolus; LTA, lateral tibial apophysis; MA, median apophysis; PA, patellar apophysis; RTA, retrolateral tibial apophysis. Scale bars: A, B = 1 mm; CE = 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.7456731" httpUri="https://zenodo.org/record/7456731/files/figure.png" pageId="1" pageNumber="180">4CE</figureCitation>
;
<bibRefCitation id="EFF64B25D705FF88B6E4FC52FD04FBFB" author="Zhou, Y. C. &amp; Yuen, Y. L. &amp; Zhang, Z. S." box="[491,676,1009,1036]" pageId="1" pageNumber="180" pagination="429 - 436" refId="ref6836" refString="Zhou, Y. C., Yuen, Y. L. &amp; Zhang, Z. S. (2017) A new Bifidocoelotes species, with the description of the male of B. primus from Hong Kong, China (Agelenidae: Coelotinae). Zootaxa, 4232 (3), 429 - 436. https: // doi. org / 10.11646 / zootaxa. 4232.3.11" type="journal article" year="2017">
Zhou
<emphasis id="B913EAC6D705FF88B523FC51FDFCFBFB" box="[556,604,1009,1035]" italics="true" pageId="1" pageNumber="180">et al</emphasis>
. 2017
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: figs 1CE, 2EG, 3CE, 4EG, 5;
<bibRefCitation id="EFF64B25D705FF88B321FC51FB4DFBFB" author="Wang, X. P. &amp; Tso, I. M. &amp; Wu, H. Y." box="[1070,1261,1009,1036]" pageId="1" pageNumber="180" pagination="127 - 133" refId="ref6755" refString="Wang, X. P., Tso, I. M. &amp; Wu, H. Y. (2001) Three new Coelotes species (Araneae: Amaurobiidae) from Taiwan. Zoological Studies, 40, 127 - 133." type="journal article" year="2001">
Wang
<emphasis id="B913EAC6D705FF88B37AFC51FB05FBFB" box="[1141,1189,1009,1035]" italics="true" pageId="1" pageNumber="180">et al</emphasis>
. 2001
</bibRefCitation>
: figs 3, 4). Both sexes have three promarginal and two retromarginal cheliceral teeth.
</paragraph>
</subSubSection>
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<paragraph id="8BD836D4D705FF88B798F921FB0AF911" blockId="1.[151,1436,1666,1762]" pageId="1" pageNumber="180">
<emphasis id="B913EAC6D705FF88B798F921FEB6F96A" bold="true" box="[151,278,1666,1690]" pageId="1" pageNumber="180">FIGURE 1.</emphasis>
The epigynes of six
<taxonomicName id="4C674D57D705FF88B6F6F921FD60F969" box="[505,704,1666,1690]" class="Arachnida" family="Agelenidae" genus="Bifidocoelotes" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="180" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B913EAC6D705FF88B6F6F921FD2AF96A" box="[505,650,1666,1690]" italics="true" pageId="1" pageNumber="180">Bifidocoelotes</emphasis>
spp.
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, ventral view. A.
<taxonomicName id="4C674D57D705FF88B473F921FBD2F96A" authorityName="Zhou, Yuen &amp; Zhang" authorityYear="2017" box="[892,1138,1666,1690]" class="Arachnida" family="Agelenidae" genus="Bifidocoelotes" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="180" phylum="Arthropoda" rank="species" species="obscurus">
<emphasis id="B913EAC6D705FF88B473F921FBD2F96A" box="[892,1138,1666,1690]" italics="true" pageId="1" pageNumber="180">Bifidocoelotes obscurus</emphasis>
</taxonomicName>
(after
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Zhou
<emphasis id="B913EAC6D705FF88B3F5F920FA8AF96A" box="[1274,1322,1666,1690]" italics="true" pageId="1" pageNumber="180">et al</emphasis>
. 2017
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); B.
<taxonomicName id="4C674D57D705FF88B798F905FED5F94E" authorityName="MRDH" baseAuthorityName="Fox" baseAuthorityYear="1937" box="[151,373,1702,1726]" class="Arachnida" family="Agelenidae" genus="Bifidocoelotes" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="180" phylum="Arthropoda" rank="species" species="primus">
<emphasis id="B913EAC6D705FF88B798F905FED5F94E" box="[151,373,1702,1726]" italics="true" pageId="1" pageNumber="180">Bifidocoelotes primus</emphasis>
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(after
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Zhou
<emphasis id="B913EAC6D705FF88B6F9F904FD83F94E" box="[502,547,1702,1726]" italics="true" pageId="1" pageNumber="180">et al</emphasis>
. 2017
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); C.
<taxonomicName id="4C674D57D705FF88B599F905FCF3F94E" authorityName=": Li &amp; Blick" authorityYear="2020" box="[662,851,1702,1726]" class="Arachnida" family="Agelenidae" genus="Bifidocoelotes" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="180" phylum="Arthropoda" rank="species" species="tsoi">
<emphasis id="B913EAC6D705FF88B599F905FCF3F94E" box="[662,851,1702,1726]" italics="true" pageId="1" pageNumber="180">Bifidocoelotes tsoi</emphasis>
</taxonomicName>
(after
<bibRefCitation id="EFF64B25D705FF88B499F905FBE8F94E" author="Wang, X. P. &amp; Tso, I. M. &amp; Wu, H. Y." box="[918,1096,1702,1726]" pageId="1" pageNumber="180" pagination="127 - 133" refId="ref6755" refString="Wang, X. P., Tso, I. M. &amp; Wu, H. Y. (2001) Three new Coelotes species (Araneae: Amaurobiidae) from Taiwan. Zoological Studies, 40, 127 - 133." type="journal article" year="2001">
Wang
<emphasis id="B913EAC6D705FF88B4D8F904FBA4F94E" box="[983,1028,1702,1726]" italics="true" pageId="1" pageNumber="180">et al</emphasis>
. 2001
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); D.
<taxonomicName id="4C674D57D705FF88B376F905FA3CF94E" authority="Liao &amp; Wang &amp; Yin &amp; Xu, 2022" authorityName="Liao &amp; Wang &amp; Yin &amp; Xu" authorityYear="2022" box="[1145,1436,1702,1726]" class="Arachnida" family="Agelenidae" genus="Bifidocoelotes" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="180" phylum="Arthropoda" rank="species" species="mammiformis" status="sp. nov.">
<emphasis id="B913EAC6D705FF88B376F905FA3CF94E" box="[1145,1436,1702,1726]" italics="true" pageId="1" pageNumber="180">Bifidocoelotes mammiformis</emphasis>
</taxonomicName>
<emphasis id="B913EAC6D705FF88B798F969FF48F912" bold="true" box="[151,232,1738,1762]" pageId="1" pageNumber="180">
<taxonomicNameLabel id="A22057BDD705FF88B798F969FF48F912" box="[151,232,1738,1762]" pageId="1" pageNumber="180" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; E.
<taxonomicName id="4C674D57D705FF88B61DF969FDACF912" authority="Liao &amp; Wang &amp; Yin &amp; Xu, 2022" authorityName="Liao &amp; Wang &amp; Yin &amp; Xu" authorityYear="2022" box="[274,524,1738,1762]" class="Arachnida" family="Agelenidae" genus="Bifidocoelotes" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="180" phylum="Arthropoda" rank="species" species="elongatus" status="sp. nov.">
<emphasis id="B913EAC6D705FF88B61DF969FDACF912" box="[274,524,1738,1762]" italics="true" pageId="1" pageNumber="180">Bifidocoelotes elongatus</emphasis>
</taxonomicName>
<emphasis id="B913EAC6D705FF88B51DF969FDC2F912" bold="true" box="[530,610,1738,1762]" pageId="1" pageNumber="180">
<taxonomicNameLabel id="A22057BDD705FF88B51DF969FDC2F912" box="[530,610,1738,1762]" pageId="1" pageNumber="180" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; F.
<taxonomicName id="4C674D57D705FF88B586F969FC27F912" authority="Liao &amp; Wang &amp; Yin &amp; Xu, 2022" authorityName="Liao &amp; Wang &amp; Yin &amp; Xu" authorityYear="2022" box="[649,903,1738,1762]" class="Arachnida" family="Agelenidae" genus="Bifidocoelotes" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="180" phylum="Arthropoda" rank="species" species="quadratus" status="sp. nov.">
<emphasis id="B913EAC6D705FF88B586F969FC27F912" box="[649,903,1738,1762]" italics="true" pageId="1" pageNumber="180">Bifidocoelotes quadratus</emphasis>
</taxonomicName>
<emphasis id="B913EAC6D705FF88B482F969FC7EF912" bold="true" box="[909,990,1738,1762]" pageId="1" pageNumber="180">
<taxonomicNameLabel id="A22057BDD705FF88B482F969FC7EF912" box="[909,990,1738,1762]" pageId="1" pageNumber="180" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
. Scale bar: 0.3 mm.
</paragraph>
</caption>
<subSubSection id="C37D655FD705FF88B7C8F8B3FE88F884" pageId="1" pageNumber="180" type="synonymic_list">
<paragraph id="8BD836D4D705FF88B7C8F8B3FE88F884" blockId="1.[151,1437,1808,1943]" pageId="1" pageNumber="180">
<emphasis id="B913EAC6D705FF88B7C8F8B3FEC9F8DA" bold="true" box="[199,361,1808,1834]" pageId="1" pageNumber="180">Composition:</emphasis>
Six species,
<taxonomicName id="4C674D57D705FF88B6F9F8B3FCA8F8DA" authority="Liao &amp; Wang &amp; Yin &amp; Xu, 2022" authorityName="Liao &amp; Wang &amp; Yin &amp; Xu" authorityYear="2022" box="[502,776,1808,1834]" class="Arachnida" family="Agelenidae" genus="Bifidocoelotes" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="180" phylum="Arthropoda" rank="species" species="elongatus" status="sp. nov.">
<emphasis id="B913EAC6D705FF88B6F9F8B3FCA8F8DA" box="[502,776,1808,1834]" italics="true" pageId="1" pageNumber="180">Bifidocoelotes elongatus</emphasis>
</taxonomicName>
<emphasis id="B913EAC6D705FF88B401F8B3FCC5F8DA" bold="true" box="[782,869,1808,1834]" pageId="1" pageNumber="180">
<taxonomicNameLabel id="A22057BDD705FF88B401F8B3FCC5F8DA" box="[782,869,1808,1834]" pageId="1" pageNumber="180" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<specimenCount id="9D61FD5DD705FF88B479F8B3FC29F8DA" box="[886,905,1808,1834]" pageId="1" pageNumber="180" type="female"></specimenCount>
),
<taxonomicName id="4C674D57D705FF88B491F8B2FBF6F8DA" authority="Liao &amp; Wang &amp; Yin &amp; Xu, 2022" authorityName="Liao &amp; Wang &amp; Yin &amp; Xu" authorityYear="2022" box="[926,1110,1808,1834]" class="Arachnida" family="Agelenidae" genus="Bifidocoelotes" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="180" phylum="Arthropoda" rank="species" species="mammiformis" status="sp. nov.">
<emphasis id="B913EAC6D705FF88B491F8B2FBF6F8DA" box="[926,1110,1808,1834]" italics="true" pageId="1" pageNumber="180">B. mammiformis</emphasis>
</taxonomicName>
<emphasis id="B913EAC6D705FF88B354F8B3FB13F8DA" bold="true" box="[1115,1203,1808,1834]" pageId="1" pageNumber="180">
<taxonomicNameLabel id="A22057BDD705FF88B354F8B3FB13F8DA" box="[1115,1203,1808,1834]" pageId="1" pageNumber="180" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<specimenCount id="9D61FD5DD705FF88B3CBF8B3FB77F8DA" box="[1220,1239,1808,1834]" pageId="1" pageNumber="180" type="female"></specimenCount>
, ♁),
<taxonomicName id="4C674D57D705FF88B202F8B2FA3DF8DA" authority="Liao &amp; Wang &amp; Yin &amp; Xu, 2022" authorityName="Liao &amp; Wang &amp; Yin &amp; Xu" authorityYear="2022" box="[1293,1437,1808,1834]" class="Arachnida" family="Agelenidae" genus="Bifidocoelotes" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="180" phylum="Arthropoda" rank="species" species="quadratus" status="sp. nov.">
<emphasis id="B913EAC6D705FF88B202F8B2FA3DF8DA" box="[1293,1437,1808,1834]" italics="true" pageId="1" pageNumber="180">B. quadratus</emphasis>
</taxonomicName>
<emphasis id="B913EAC6D705FF88B798F897FF51F8BE" bold="true" box="[151,241,1844,1870]" pageId="1" pageNumber="180">
<taxonomicNameLabel id="A22057BDD705FF88B798F897FF51F8BE" box="[151,241,1844,1870]" pageId="1" pageNumber="180" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<specimenCount id="9D61FD5DD705FF88B60CF897FEB6F8BE" box="[259,278,1844,1870]" pageId="1" pageNumber="180" type="female"></specimenCount>
);
<taxonomicName id="4C674D57D705FF88B621F896FD4DF8BE" authority="Zhou, Yuen &amp; Zhang, 2017" authorityName="Zhou, Yuen &amp; Zhang" authorityYear="2017" box="[302,749,1844,1871]" class="Arachnida" family="Agelenidae" genus="Bifidocoelotes" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="180" phylum="Arthropoda" rank="species" species="obscurus">
<emphasis id="B913EAC6D705FF88B621F896FE12F8BE" box="[302,434,1844,1870]" italics="true" pageId="1" pageNumber="180">B. obscurus</emphasis>
<bibRefCitation id="EFF64B25D705FF88B6B5F897FD4DF8BE" author="Zhou, Y. C. &amp; Yuen, Y. L. &amp; Zhang, Z. S." box="[442,749,1844,1871]" pageId="1" pageNumber="180" pagination="429 - 436" refId="ref6836" refString="Zhou, Y. C., Yuen, Y. L. &amp; Zhang, Z. S. (2017) A new Bifidocoelotes species, with the description of the male of B. primus from Hong Kong, China (Agelenidae: Coelotinae). Zootaxa, 4232 (3), 429 - 436. https: // doi. org / 10.11646 / zootaxa. 4232.3.11" type="journal article" year="2017">Zhou, Yuen &amp; Zhang, 2017</bibRefCitation>
</taxonomicName>
(
<specimenCount id="9D61FD5DD705FF88B5F0F897FCB2F8BE" box="[767,786,1844,1870]" pageId="1" pageNumber="180" type="female"></specimenCount>
, ♁),
<taxonomicName id="4C674D57D705FF88B442F896FBE5F8BE" authority="(Fox, 1937)" authorityName="MRDH" baseAuthorityName="Fox" baseAuthorityYear="1937" box="[845,1093,1844,1870]" class="Arachnida" family="Agelenidae" genus="Bifidocoelotes" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="180" phylum="Arthropoda" rank="species" species="primus">
<emphasis id="B913EAC6D705FF88B442F896FC1AF8BE" box="[845,954,1845,1870]" italics="true" pageId="1" pageNumber="180">B. primus</emphasis>
(
<bibRefCitation id="EFF64B25D705FF88B4C6F897FB9DF8BE" author="Fox, I." box="[969,1085,1844,1870]" pageId="1" pageNumber="180" pagination="1 - 9" refId="ref6378" refString="Fox, I. (1937) New species and records of Chinese spiders. American Museum Novitates, 907, 1 - 9." type="journal article" year="1937">Fox, 1937</bibRefCitation>
)
</taxonomicName>
(
<specimenCount id="9D61FD5DD705FF88B358F897FBCAF8BE" box="[1111,1130,1844,1870]" pageId="1" pageNumber="180" type="female"></specimenCount>
, ♁) and
<taxonomicName id="4C674D57D705FF88B3C2F896FF6FF882" authority="Li &amp; Blick, 2020" authorityName=": Li &amp; Blick" authorityYear="2020" class="Arachnida" family="Agelenidae" genus="Bifidocoelotes" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="180" phylum="Arthropoda" rank="species" species="tsoi">
<emphasis id="B913EAC6D705FF88B3C2F896FAB4F8BE" box="[1229,1300,1845,1870]" italics="true" pageId="1" pageNumber="180">B. tsoi</emphasis>
<bibRefCitation id="EFF64B25D705FF88B213F897FF6FF882" author="Li, S. Q. &amp; Blick, T." pageId="1" pageNumber="180" pagination="118" refId="ref6452" refString="Li, S. Q. &amp; Blick, T. (2020) Bifidocoelotes tsoi, a new name to replace Bifidocoelotes bifidus (Araneae, Agelenidae). Acta Arachnologica Sinica, 29 (2), 118. https: // doi. org / 10.3969 / j. issn. 1005 - 9628.2020.02.009" type="journal article" year="2020">Li &amp; Blick, 2020</bibRefCitation>
</taxonomicName>
(
<specimenCount id="9D61FD5DD705FF88B7EDF8FBFF55F882" box="[226,245,1880,1906]" pageId="1" pageNumber="180" type="female"></specimenCount>
, ♁).
</paragraph>
</subSubSection>
<subSubSection id="C37D655FD705FF88B7C8F8DFFBAAF866" box="[199,1034,1916,1943]" pageId="1" pageNumber="180" type="distribution">
<paragraph id="8BD836D4D705FF88B7C8F8DFFBAAF866" blockId="1.[151,1437,1808,1943]" box="[199,1034,1916,1943]" pageId="1" pageNumber="180">
<emphasis id="B913EAC6D705FF88B7C8F8DFFEC3F866" bold="true" box="[199,355,1916,1942]" pageId="1" pageNumber="180">Distribution:</emphasis>
<collectingCountry id="F3707644D705FF88B665F8DFFE0DF866" box="[362,429,1916,1942]" name="China" pageId="1" pageNumber="180">China</collectingCountry>
(
<collectingRegion id="49A3F836D705FF88B6B4F8DFFDE2F866" box="[443,578,1916,1942]" country="China" name="Guangdong" pageId="1" pageNumber="180">Guangdong</collectingRegion>
,
<collectingRegion id="49A3F836D705FF88B541F8DFFD11F866" box="[590,689,1916,1942]" country="China" name="Guangxi" pageId="1" pageNumber="180">Guangxi</collectingRegion>
,
<collectingCountry id="F3707644D705FF88B5B3F8DEFCE2F867" box="[700,834,1917,1943]" name="Hong Kong" pageId="1" pageNumber="180">Hong Kong</collectingCountry>
, Taiwan) (
<figureCitation id="135C2A51D705FF88B4B7F8DFFC5CF866" box="[952,1020,1916,1942]" captionStart="FIGURE 7" captionStartId="10.[151,250,1002,1026]" captionTargetBox="[151,1436,181,978]" captionTargetId="figure-31@10.[151,1436,181,978]" captionTargetPageId="10" captionText="FIGURE 7. Collection localities for Bifidocoelotes in China. 1. B. mammiformis sp. nov.; 2. B. elongatus sp. nov.; 3. B. quadratus sp. nov.; 4. B. tsoi; 5. B. obscurus; 6. B. primus." figureDoi="http://doi.org/10.5281/zenodo.7456741" httpUri="https://zenodo.org/record/7456741/files/figure.png" pageId="1" pageNumber="180">Fig. 7</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF18665CD706FF8BB798FE10FCB9FDE3" ID-DOI="http://doi.org/10.5281/zenodo.7456725" ID-Zenodo-Dep="7456725" httpUri="https://zenodo.org/record/7456725/files/figure.png" pageId="2" pageNumber="181" startId="2.[151,250,435,459]" targetBox="[151,1432,181,408]" targetPageId="2">
<paragraph id="8BD836D4D706FF8BB798FE10FCB9FDE3" blockId="2.[151,1436,435,532]" pageId="2" pageNumber="181">
<emphasis id="B913EAC6D706FF8BB798FE10FEB2FE3B" bold="true" box="[151,274,435,459]" pageId="2" pageNumber="181">FIGURE 2.</emphasis>
The distal parts of conductors of four
<taxonomicName id="4C674D57D706FF8BB59EFE10FCF4FE3B" box="[657,852,435,459]" class="Arachnida" family="Agelenidae" genus="Bifidocoelotes" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="181" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B913EAC6D706FF8BB59EFE10FC82FE3B" box="[657,802,435,459]" italics="true" pageId="2" pageNumber="181">Bifidocoelotes</emphasis>
spp.
</taxonomicName>
, ventral view. A.
<taxonomicName id="4C674D57D706FF8BB30EFE10FA82FE3C" authority="Liao &amp; Wang &amp; Yin &amp; Xu, 2022" authorityName="Liao &amp; Wang &amp; Yin &amp; Xu" authorityYear="2022" box="[1025,1314,435,460]" class="Arachnida" family="Agelenidae" genus="Bifidocoelotes" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="181" phylum="Arthropoda" rank="species" species="mammiformis" status="sp. nov.">
<emphasis id="B913EAC6D706FF8BB30EFE10FA82FE3C" box="[1025,1314,435,460]" italics="true" pageId="2" pageNumber="181">Bifidocoelotes mammiformis</emphasis>
</taxonomicName>
<emphasis id="B913EAC6D706FF8BB227FE17FAD8FE3C" bold="true" box="[1320,1400,436,460]" pageId="2" pageNumber="181">
<taxonomicNameLabel id="A22057BDD706FF8BB227FE17FAD8FE3C" box="[1320,1400,436,460]" pageId="2" pageNumber="181" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; B.
<taxonomicName id="4C674D57D706FF8BB798FE74FEF3FE1F" authorityName=": Li &amp; Blick" authorityYear="2020" box="[151,339,471,495]" class="Arachnida" family="Agelenidae" genus="Bifidocoelotes" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="181" phylum="Arthropoda" rank="species" species="tsoi">
<emphasis id="B913EAC6D706FF8BB798FE74FEF3FE1F" box="[151,339,471,495]" italics="true" pageId="2" pageNumber="181">Bifidocoelotes tsoi</emphasis>
</taxonomicName>
(after
<bibRefCitation id="EFF64B25D706FF8BB69AFE7BFDE5FE00" author="Wang, X. P. &amp; Tso, I. M. &amp; Wu, H. Y." box="[405,581,471,496]" pageId="2" pageNumber="181" pagination="127 - 133" refId="ref6755" refString="Wang, X. P., Tso, I. M. &amp; Wu, H. Y. (2001) Three new Coelotes species (Araneae: Amaurobiidae) from Taiwan. Zoological Studies, 40, 127 - 133." type="journal article" year="2001">
Wang
<emphasis id="B913EAC6D706FF8BB6DAFE7BFDA8FE1F" box="[469,520,471,495]" italics="true" pageId="2" pageNumber="181">et al.</emphasis>
2001
</bibRefCitation>
); C.
<taxonomicName id="4C674D57D706FF8BB57CFE74FCF0FE1F" authorityName="MRDH" baseAuthorityName="Fox" baseAuthorityYear="1937" box="[627,848,471,495]" class="Arachnida" family="Agelenidae" genus="Bifidocoelotes" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="181" phylum="Arthropoda" rank="species" species="primus">
<emphasis id="B913EAC6D706FF8BB57CFE74FCF0FE1F" box="[627,848,471,495]" italics="true" pageId="2" pageNumber="181">Bifidocoelotes primus</emphasis>
</taxonomicName>
(after
<bibRefCitation id="EFF64B25D706FF8BB49CFE74FB9DFE00" author="Zhou, Y. C. &amp; Yuen, Y. L. &amp; Zhang, Z. S." box="[915,1085,471,496]" pageId="2" pageNumber="181" pagination="429 - 436" refId="ref6836" refString="Zhou, Y. C., Yuen, Y. L. &amp; Zhang, Z. S. (2017) A new Bifidocoelotes species, with the description of the male of B. primus from Hong Kong, China (Agelenidae: Coelotinae). Zootaxa, 4232 (3), 429 - 436. https: // doi. org / 10.11646 / zootaxa. 4232.3.11" type="journal article" year="2017">
Zhou
<emphasis id="B913EAC6D706FF8BB4C1FE7BFC5AFE1F" box="[974,1018,471,495]" italics="true" pageId="2" pageNumber="181">et al</emphasis>
. 2017
</bibRefCitation>
); D.
<taxonomicName id="4C674D57D706FF8BB362FE74FAFFFE1F" authorityName="Zhou, Yuen &amp; Zhang" authorityYear="2017" box="[1133,1375,471,495]" class="Arachnida" family="Agelenidae" genus="Bifidocoelotes" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="181" phylum="Arthropoda" rank="species" species="obscurus">
<emphasis id="B913EAC6D706FF8BB362FE74FAFFFE1F" box="[1133,1375,471,495]" italics="true" pageId="2" pageNumber="181">Bifidocoelotes obscurus</emphasis>
</taxonomicName>
(after
<bibRefCitation id="EFF64B25D706FF8BB798FE58FEE3FDE4" author="Zhou, Y. C. &amp; Yuen, Y. L. &amp; Zhang, Z. S." box="[151,323,507,532]" pageId="2" pageNumber="181" pagination="429 - 436" refId="ref6836" refString="Zhou, Y. C., Yuen, Y. L. &amp; Zhang, Z. S. (2017) A new Bifidocoelotes species, with the description of the male of B. primus from Hong Kong, China (Agelenidae: Coelotinae). Zootaxa, 4232 (3), 429 - 436. https: // doi. org / 10.11646 / zootaxa. 4232.3.11" type="journal article" year="2017">
Zhou
<emphasis id="B913EAC6D706FF8BB7DCFE5FFEA0FDE3" box="[211,256,507,531]" italics="true" pageId="2" pageNumber="181">et al</emphasis>
. 2017
</bibRefCitation>
). Scale bars: A, B, D = 0.1 mm; C = 0.05 mm.
</paragraph>
</caption>
<subSubSection id="C37D655FD706FF8BB798FDC5FD88FD8F" box="[151,552,613,640]" pageId="2" pageNumber="181" type="multiple">
<paragraph id="8BD836D4D706FF8BB798FDC5FD88FD8F" blockId="2.[151,552,613,640]" box="[151,552,613,640]" pageId="2" pageNumber="181">
<heading id="D09081B8D706FF8BB798FDC5FD88FD8F" bold="true" box="[151,552,613,640]" fontSize="11" level="1" pageId="2" pageNumber="181" reason="1">
<emphasis id="B913EAC6D706FF8BB798FDC5FD88FD8F" bold="true" box="[151,552,613,640]" pageId="2" pageNumber="181">
Key to the
<taxonomicName id="4C674D57D706FF8BB610FDC6FD88FD8F" box="[287,552,613,640]" class="Arachnida" family="Agelenidae" genus="Bifidocoelotes" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="181" phylum="Arthropoda" rank="species" species="undefined-1">
<emphasis id="B913EAC6D706FF8BB610FDC6FE6AFD70" bold="true" box="[287,458,613,640]" italics="true" pageId="2" pageNumber="181">Bifidocoelotes</emphasis>
species
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37D655FD706FF8BB798FD08FA3CFB42" pageId="2" pageNumber="181" type="key">
<key id="FFF661E5D706FF8BB798FD08FA3CFB42" pageId="2" pageNumber="181">
<keyStep id="30932E71D706FF8BB798FD08FA3CFD2E" pageId="2" pageNumber="181">
<paragraph id="8BD836D4D706FF8BB798FD08FA3CFD30" blockId="2.[151,1437,683,1203]" box="[151,1436,683,705]" pageId="2" pageNumber="181">
<keyLead id="309695E1D706FF8BB798FD08FA3CFD30" box="[151,1436,683,705]" pageId="2" pageNumber="181">1. Male............................................................................................... 2</keyLead>
</paragraph>
<paragraph id="8BD836D4D706FF8BB798FD6BFA3CFD2E" blockId="2.[151,1437,683,1203]" box="[151,1436,712,734]" pageId="2" pageNumber="181">
<keyLead id="309695E1D706FF8BB798FD6BFA3CFD2E" box="[151,1436,712,734]" pageId="2" pageNumber="181">- Female............................................................................................. 5</keyLead>
</paragraph>
</keyStep>
<keyStep id="30932E71D706FF8BB798FD46FA3CFCE8" pageId="2" pageNumber="181">
<paragraph id="8BD836D4D706FF8BB798FD46FA3DFD0B" blockId="2.[151,1437,683,1203]" box="[151,1437,741,763]" pageId="2" pageNumber="181">
<keyLead id="309695E1D706FF8BB798FD46FA3DFD0B" box="[151,1437,741,763]" pageId="2" pageNumber="181">
2. Conductor slightly bifurcated terminally (
<figureCitation id="135C2A51D706FF8BB57DFD46FD1EFD0A" box="[626,702,741,763]" captionStart="FIGURE 2" captionStartId="2.[151,250,435,459]" captionTargetBox="[151,1432,181,408]" captionTargetId="figure-31@2.[151,1436,181,412]" captionTargetPageId="2" captionText="FIGURE 2. The distal parts of conductors of four Bifidocoelotes spp., ventral view.A. Bifidocoelotes mammiformis sp. nov.; B. Bifidocoelotes tsoi (after Wang et al. 2001); C. Bifidocoelotes primus (after Zhou et al. 2017); D. Bifidocoelotes obscurus (after Zhou et al. 2017). Scale bars: A, B, D = 0.1 mm; C = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.7456725" httpUri="https://zenodo.org/record/7456725/files/figure.png" pageId="2" pageNumber="181">Fig. 2C</figureCitation>
;
<bibRefCitation id="EFF64B25D706FF8BB5C7FD46FCCAFD0A" author="Zhou, Y. C. &amp; Yuen, Y. L. &amp; Zhang, Z. S." box="[712,874,741,763]" pageId="2" pageNumber="181" pagination="429 - 436" refId="ref6836" refString="Zhou, Y. C., Yuen, Y. L. &amp; Zhang, Z. S. (2017) A new Bifidocoelotes species, with the description of the male of B. primus from Hong Kong, China (Agelenidae: Coelotinae). Zootaxa, 4232 (3), 429 - 436. https: // doi. org / 10.11646 / zootaxa. 4232.3.11" type="journal article" year="2017">
Zhou
<emphasis id="B913EAC6D706FF8BB40FFD45FC8BFD0A" box="[768,811,741,763]" italics="true" pageId="2" pageNumber="181">et al</emphasis>
. 2017
</bibRefCitation>
: fig. 5)................................
<taxonomicName id="4C674D57D706FF8BB230FD45FA3DFD0B" ID-CoL="LT4L" authorityName="MRDH" baseAuthorityName="Fox" baseAuthorityYear="1937" box="[1343,1437,742,763]" class="Arachnida" family="Agelenidae" genus="Bifidocoelotes" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="181" phylum="Arthropoda" rank="species" species="primus">
<emphasis id="B913EAC6D706FF8BB230FD45FA3DFD0B" box="[1343,1437,742,763]" italics="true" pageId="2" pageNumber="181">B. primus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BD836D4D706FF8BB798FCA0FA3CFCE8" blockId="2.[151,1437,683,1203]" box="[151,1436,771,793]" pageId="2" pageNumber="181">
<keyLead id="309695E1D706FF8BB798FCA0FA3CFCE8" box="[151,1436,771,793]" pageId="2" pageNumber="181">
- Conductor strongly bifurcated terminally (
<figureCitation id="135C2A51D706FF8BB576FCA0FD5DFCE8" box="[633,765,771,793]" captionStart="FIGURE 2" captionStartId="2.[151,250,435,459]" captionTargetBox="[151,1432,181,408]" captionTargetId="figure-31@2.[151,1436,181,412]" captionTargetPageId="2" captionText="FIGURE 2. The distal parts of conductors of four Bifidocoelotes spp., ventral view.A. Bifidocoelotes mammiformis sp. nov.; B. Bifidocoelotes tsoi (after Wang et al. 2001); C. Bifidocoelotes primus (after Zhou et al. 2017); D. Bifidocoelotes obscurus (after Zhou et al. 2017). Scale bars: A, B, D = 0.1 mm; C = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.7456725" httpUri="https://zenodo.org/record/7456725/files/figure.png" pageId="2" pageNumber="181">Fig. 2A, B, D</figureCitation>
)..................................................... 3
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30932E71D706FF8BB798FC83FA3CFCA2" pageId="2" pageNumber="181">
<paragraph id="8BD836D4D706FF8BB798FC83FA3DFCC6" blockId="2.[151,1437,683,1203]" box="[151,1437,800,822]" pageId="2" pageNumber="181">
<keyLead id="309695E1D706FF8BB798FC83FA3DFCC6" box="[151,1437,800,822]" pageId="2" pageNumber="181">
3. The two branches of conductor subequal in length (
<figureCitation id="135C2A51D706FF8BB5DFFC83FCB9FCC5" box="[720,793,800,822]" captionStart="FIGURE 2" captionStartId="2.[151,250,435,459]" captionTargetBox="[151,1432,181,408]" captionTargetId="figure-31@2.[151,1436,181,412]" captionTargetPageId="2" captionText="FIGURE 2. The distal parts of conductors of four Bifidocoelotes spp., ventral view.A. Bifidocoelotes mammiformis sp. nov.; B. Bifidocoelotes tsoi (after Wang et al. 2001); C. Bifidocoelotes primus (after Zhou et al. 2017); D. Bifidocoelotes obscurus (after Zhou et al. 2017). Scale bars: A, B, D = 0.1 mm; C = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.7456725" httpUri="https://zenodo.org/record/7456725/files/figure.png" pageId="2" pageNumber="181">Fig. 2B</figureCitation>
)..............................................
<taxonomicName id="4C674D57D706FF8BB250FC82FA3DFCC6" ID-CoL="99CNM" authorityName=": Li &amp; Blick" authorityYear="2020" box="[1375,1437,801,822]" class="Arachnida" family="Agelenidae" genus="Bifidocoelotes" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="181" phylum="Arthropoda" rank="species" species="tsoi">
<emphasis id="B913EAC6D706FF8BB250FC82FA3DFCC6" box="[1375,1437,801,822]" italics="true" pageId="2" pageNumber="181">B. tsoi</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BD836D4D706FF8BB798FC9DFA3CFCA2" blockId="2.[151,1437,683,1203]" box="[151,1436,829,851]" pageId="2" pageNumber="181">
<keyLead id="309695E1D706FF8BB798FC9DFA3CFCA2" box="[151,1436,829,851]" pageId="2" pageNumber="181">
- One branch of conductor clearly longer than the other (
<figureCitation id="135C2A51D706FF8BB5FCFC9EFCFBFCA2" box="[755,859,829,851]" captionStart="FIGURE 2" captionStartId="2.[151,250,435,459]" captionTargetBox="[151,1432,181,408]" captionTargetId="figure-31@2.[151,1436,181,412]" captionTargetPageId="2" captionText="FIGURE 2. The distal parts of conductors of four Bifidocoelotes spp., ventral view.A. Bifidocoelotes mammiformis sp. nov.; B. Bifidocoelotes tsoi (after Wang et al. 2001); C. Bifidocoelotes primus (after Zhou et al. 2017); D. Bifidocoelotes obscurus (after Zhou et al. 2017). Scale bars: A, B, D = 0.1 mm; C = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.7456725" httpUri="https://zenodo.org/record/7456725/files/figure.png" pageId="2" pageNumber="181">Fig. 2A, D</figureCitation>
)............................................. 4
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30932E71D706FF8BB798FCF8FA3DFC7D" pageId="2" pageNumber="181">
<paragraph id="8BD836D4D706FF8BB798FCF8FA3CFC80" blockId="2.[151,1437,683,1203]" box="[151,1436,859,881]" pageId="2" pageNumber="181">
<keyLead id="309695E1D706FF8BB798FCF8FA3CFC80" box="[151,1436,859,881]" pageId="2" pageNumber="181">
4. The proximal branch of conductor smaller and shorter than the other (
<figureCitation id="135C2A51D706FF8BB48FFCF8FC6CFC80" box="[896,972,859,881]" captionStart="FIGURE 2" captionStartId="2.[151,250,435,459]" captionTargetBox="[151,1432,181,408]" captionTargetId="figure-31@2.[151,1436,181,412]" captionTargetPageId="2" captionText="FIGURE 2. The distal parts of conductors of four Bifidocoelotes spp., ventral view.A. Bifidocoelotes mammiformis sp. nov.; B. Bifidocoelotes tsoi (after Wang et al. 2001); C. Bifidocoelotes primus (after Zhou et al. 2017); D. Bifidocoelotes obscurus (after Zhou et al. 2017). Scale bars: A, B, D = 0.1 mm; C = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.7456725" httpUri="https://zenodo.org/record/7456725/files/figure.png" pageId="2" pageNumber="181">Fig. 2A</figureCitation>
)................
<taxonomicName id="4C674D57D706FF8BB3A4FCF8FAEAFC80" authority="Liao &amp; Wang &amp; Yin &amp; Xu, 2022" authorityName="Liao &amp; Wang &amp; Yin &amp; Xu" authorityYear="2022" box="[1195,1354,859,880]" class="Arachnida" family="Agelenidae" genus="Bifidocoelotes" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="181" phylum="Arthropoda" rank="species" species="mammiformis" status="sp. nov.">
<emphasis id="B913EAC6D706FF8BB3A4FCF8FAEAFC80" box="[1195,1354,859,880]" italics="true" pageId="2" pageNumber="181">B. mammiformis</emphasis>
</taxonomicName>
<emphasis id="B913EAC6D706FF8BB25FFCF8FA3CFC80" bold="true" box="[1360,1436,859,880]" pageId="2" pageNumber="181">
<taxonomicNameLabel id="A22057BDD706FF8BB25FFCF8FA3CFC80" box="[1360,1436,859,880]" pageId="2" pageNumber="181" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BD836D4D706FF8BB798FCDBFA3DFC7D" blockId="2.[151,1437,683,1203]" box="[151,1437,888,910]" pageId="2" pageNumber="181">
<keyLead id="309695E1D706FF8BB798FCDBFA3DFC7D" box="[151,1437,888,910]" pageId="2" pageNumber="181">
- The proximal branch of conductor stronger and longer than the other (
<figureCitation id="135C2A51D706FF8BB48CFCDBFC6EFC7D" box="[899,974,888,910]" captionStart="FIGURE 2" captionStartId="2.[151,250,435,459]" captionTargetBox="[151,1432,181,408]" captionTargetId="figure-31@2.[151,1436,181,412]" captionTargetPageId="2" captionText="FIGURE 2. The distal parts of conductors of four Bifidocoelotes spp., ventral view.A. Bifidocoelotes mammiformis sp. nov.; B. Bifidocoelotes tsoi (after Wang et al. 2001); C. Bifidocoelotes primus (after Zhou et al. 2017); D. Bifidocoelotes obscurus (after Zhou et al. 2017). Scale bars: A, B, D = 0.1 mm; C = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.7456725" httpUri="https://zenodo.org/record/7456725/files/figure.png" pageId="2" pageNumber="181">Fig. 2D</figureCitation>
)...........................
<taxonomicName id="4C674D57D706FF8BB224FCDAFA3DFC7D" ID-CoL="LT4K" authorityName="Zhou, Yuen &amp; Zhang" authorityYear="2017" box="[1323,1437,888,910]" class="Arachnida" family="Agelenidae" genus="Bifidocoelotes" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="181" phylum="Arthropoda" rank="species" species="obscurus">
<emphasis id="B913EAC6D706FF8BB224FCDAFA3DFC7D" box="[1323,1437,888,910]" italics="true" pageId="2" pageNumber="181">B. obscurus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30932E71D706FF8BB798FC36FA3CFC38" pageId="2" pageNumber="181">
<paragraph id="8BD836D4D706FF8BB798FC36FA3CFC5B" blockId="2.[151,1437,683,1203]" box="[151,1436,917,939]" pageId="2" pageNumber="181">
<keyLead id="309695E1D706FF8BB798FC36FA3CFC5B" box="[151,1436,917,939]" pageId="2" pageNumber="181">
5. Epigynal tooth only slightly bifurcated (
<figureCitation id="135C2A51D706FF8BB56CFC36FD69FC5A" box="[611,713,917,939]" captionStart="FIGURE 1" captionStartId="1.[151,250,1666,1690]" captionTargetBox="[163,1418,1102,1635]" captionTargetId="figure-290@1.[151,1436,1093,1640]" captionTargetPageId="1" captionText="FIGURE 1. The epigynes of six Bifidocoelotes spp., ventral view. A. Bifidocoelotes obscurus (after Zhou et al. 2017); B. Bifidocoelotes primus (after Zhou et al. 2017); C. Bifidocoelotes tsoi (after Wang et al. 2001); D. Bifidocoelotes mammiformis sp. nov.; E. Bifidocoelotes elongatus sp. nov.; F. Bifidocoelotes quadratus sp. nov.. Scale bar: 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.7456721" httpUri="https://zenodo.org/record/7456721/files/figure.png" pageId="2" pageNumber="181">Fig. 1A, E</figureCitation>
).......................................................... 6
</keyLead>
</paragraph>
<paragraph id="8BD836D4D706FF8BB798FC10FA3CFC38" blockId="2.[151,1437,683,1203]" box="[151,1436,947,969]" pageId="2" pageNumber="181">
<keyLead id="309695E1D706FF8BB798FC10FA3CFC38" box="[151,1436,947,969]" pageId="2" pageNumber="181">
- Epigynal tooth deeply bifurcated (bifurcated ca. 1/3 to 1/2 the length of the epigynal tooth) (
<figureCitation id="135C2A51D706FF8BB357FC10FB78FC38" box="[1112,1240,947,969]" captionStart="FIGURE 1" captionStartId="1.[151,250,1666,1690]" captionTargetBox="[163,1418,1102,1635]" captionTargetId="figure-290@1.[151,1436,1093,1640]" captionTargetPageId="1" captionText="FIGURE 1. The epigynes of six Bifidocoelotes spp., ventral view. A. Bifidocoelotes obscurus (after Zhou et al. 2017); B. Bifidocoelotes primus (after Zhou et al. 2017); C. Bifidocoelotes tsoi (after Wang et al. 2001); D. Bifidocoelotes mammiformis sp. nov.; E. Bifidocoelotes elongatus sp. nov.; F. Bifidocoelotes quadratus sp. nov.. Scale bar: 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.7456721" httpUri="https://zenodo.org/record/7456721/files/figure.png" pageId="2" pageNumber="181">Fig. 1BD, F</figureCitation>
).............. 7
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30932E71D706FF8BB798FC73FA3CFBF2" pageId="2" pageNumber="181">
<paragraph id="8BD836D4D706FF8BB798FC73FA3DFC15" blockId="2.[151,1437,683,1203]" box="[151,1437,976,998]" pageId="2" pageNumber="181">
<keyLead id="309695E1D706FF8BB798FC73FA3DFC15" box="[151,1437,976,998]" pageId="2" pageNumber="181">
6. Spermathecal head mammillary (
<bibRefCitation id="EFF64B25D706FF8BB52FFC73FD64FC15" author="Zhou, Y. C. &amp; Yuen, Y. L. &amp; Zhang, Z. S." box="[544,708,976,998]" pageId="2" pageNumber="181" pagination="429 - 436" refId="ref6836" refString="Zhou, Y. C., Yuen, Y. L. &amp; Zhang, Z. S. (2017) A new Bifidocoelotes species, with the description of the male of B. primus from Hong Kong, China (Agelenidae: Coelotinae). Zootaxa, 4232 (3), 429 - 436. https: // doi. org / 10.11646 / zootaxa. 4232.3.11" type="journal article" year="2017">
Zhou
<emphasis id="B913EAC6D706FF8BB555FC72FD24FC15" box="[602,644,976,998]" italics="true" pageId="2" pageNumber="181">et al</emphasis>
. 2017
</bibRefCitation>
: figs 1B, 2D).......................................
<taxonomicName id="4C674D57D706FF8BB224FC72FA3DFC15" ID-CoL="LT4K" authorityName="Zhou, Yuen &amp; Zhang" authorityYear="2017" box="[1323,1437,976,998]" class="Arachnida" family="Agelenidae" genus="Bifidocoelotes" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="181" phylum="Arthropoda" rank="species" species="obscurus">
<emphasis id="B913EAC6D706FF8BB224FC72FA3DFC15" box="[1323,1437,976,998]" italics="true" pageId="2" pageNumber="181">B. obscurus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BD836D4D706FF8BB798FC4DFA3CFBF2" blockId="2.[151,1437,683,1203]" box="[151,1436,1005,1027]" pageId="2" pageNumber="181">
<keyLead id="309695E1D706FF8BB798FC4DFA3CFBF2" box="[151,1436,1005,1027]" pageId="2" pageNumber="181">
- Spermathecal head thumb-shaped (
<figureCitation id="135C2A51D706FF8BB53CFC4EFDDEFBF2" box="[563,638,1005,1027]" captionStart="FIGURE 3" captionStartId="3.[151,250,1870,1894]" captionTargetBox="[151,1436,345,1837]" captionTargetId="figure-76@3.[151,1436,345,1847]" captionTargetPageId="3" captionText="FIGURE 3. Bifidocoelotes elongatus sp. nov., holotype female. A. Habitus, dorsal view; B. Same, ventral view; C. Epigyne, ventral view; D. Same, dorsal view. Abbreviations: CD, copulatory duct; CO, copulatory opening; ET, epigynal tooth; FD, fertilization duct; SH, spermathecal head; Sp, spermatheca. Scale bars: A, B = 1 mm; C, D = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7456729" httpUri="https://zenodo.org/record/7456729/files/figure.png" pageId="2" pageNumber="181">Fig. 3D</figureCitation>
)................................................
<taxonomicName id="4C674D57D706FF8BB3DDFC4DFAE9FBF2" authority="Liao &amp; Wang &amp; Yin &amp; Xu, 2022" authorityName="Liao &amp; Wang &amp; Yin &amp; Xu" authorityYear="2022" box="[1234,1353,1005,1027]" class="Arachnida" family="Agelenidae" genus="Bifidocoelotes" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="181" phylum="Arthropoda" rank="species" species="elongatus" status="sp. nov.">
<emphasis id="B913EAC6D706FF8BB3DDFC4DFAE9FBF2" box="[1234,1353,1005,1027]" italics="true" pageId="2" pageNumber="181">B. elongatus</emphasis>
</taxonomicName>
<emphasis id="B913EAC6D706FF8BB25FFC4EFA3CFBF2" bold="true" box="[1360,1436,1005,1026]" pageId="2" pageNumber="181">
<taxonomicNameLabel id="A22057BDD706FF8BB25FFC4EFA3CFBF2" box="[1360,1436,1005,1026]" pageId="2" pageNumber="181" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30932E71D706FF8BB798FBA8FA3CFBCD" pageId="2" pageNumber="181">
<paragraph id="8BD836D4D706FF8BB798FBA8FA3CFBD0" blockId="2.[151,1437,683,1203]" box="[151,1436,1035,1057]" pageId="2" pageNumber="181">
<keyLead id="309695E1D706FF8BB798FBA8FA3CFBD0" box="[151,1436,1035,1057]" pageId="2" pageNumber="181">
7. Atrium extremely large, quadrate (
<figureCitation id="135C2A51D706FF8BB521FBA8FDD7FBD0" box="[558,631,1035,1057]" captionStart="FIGURE 1" captionStartId="1.[151,250,1666,1690]" captionTargetBox="[163,1418,1102,1635]" captionTargetId="figure-290@1.[151,1436,1093,1640]" captionTargetPageId="1" captionText="FIGURE 1. The epigynes of six Bifidocoelotes spp., ventral view. A. Bifidocoelotes obscurus (after Zhou et al. 2017); B. Bifidocoelotes primus (after Zhou et al. 2017); C. Bifidocoelotes tsoi (after Wang et al. 2001); D. Bifidocoelotes mammiformis sp. nov.; E. Bifidocoelotes elongatus sp. nov.; F. Bifidocoelotes quadratus sp. nov.. Scale bar: 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.7456721" httpUri="https://zenodo.org/record/7456721/files/figure.png" pageId="2" pageNumber="181">Figs 1F</figureCitation>
,
<figureCitation id="135C2A51D706FF8BB58DFBA8FD3FFBD1" box="[642,671,1035,1057]" captionStart="FIGURE 6" captionStartId="9.[151,250,1943,1967]" captionTargetBox="[183,1404,181,1910]" captionTargetId="figure-19@9.[183,1404,181,1920]" captionTargetPageId="9" captionText="FIGURE 6. Bifidocoelotes quadratus sp. nov., holotype female. A. Habitus, dorsal view; B. Same, ventral view; C. Epigyne, ventral view; D. Same, dorsal view. Abbreviations: At, atrium; CD, copulatory duct; ET, epigynal tooth; FD, fertilization duct; SH, spermathecal head; Sp, spermatheca. Scale bars: A, B = 1 mm; C, D = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7456739" httpUri="https://zenodo.org/record/7456739/files/figure.png" pageId="2" pageNumber="181">6C</figureCitation>
)............................................
<taxonomicName id="4C674D57D706FF8BB3C1FBA8FAE9FBD0" authority="Liao &amp; Wang &amp; Yin &amp; Xu, 2022" authorityName="Liao &amp; Wang &amp; Yin &amp; Xu" authorityYear="2022" box="[1230,1353,1035,1056]" class="Arachnida" family="Agelenidae" genus="Bifidocoelotes" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="181" phylum="Arthropoda" rank="species" species="quadratus" status="sp. nov.">
<emphasis id="B913EAC6D706FF8BB3C1FBA8FAE9FBD0" box="[1230,1353,1035,1056]" italics="true" pageId="2" pageNumber="181">B. quadratus</emphasis>
</taxonomicName>
<emphasis id="B913EAC6D706FF8BB25FFBA8FA3CFBD0" bold="true" box="[1360,1436,1035,1056]" pageId="2" pageNumber="181">
<taxonomicNameLabel id="A22057BDD706FF8BB25FFBA8FA3CFBD0" box="[1360,1436,1035,1056]" pageId="2" pageNumber="181" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BD836D4D706FF8BB798FB8BFA3CFBCD" blockId="2.[151,1437,683,1203]" box="[151,1436,1064,1086]" pageId="2" pageNumber="181">
<keyLead id="309695E1D706FF8BB798FB8BFA3CFBCD" box="[151,1436,1064,1086]" pageId="2" pageNumber="181">
- Atrium relatively small or absent (
<figureCitation id="135C2A51D706FF8BB523FB8BFD33FBCD" box="[556,659,1064,1086]" captionStart="FIGURE 1" captionStartId="1.[151,250,1666,1690]" captionTargetBox="[163,1418,1102,1635]" captionTargetId="figure-290@1.[151,1436,1093,1640]" captionTargetPageId="1" captionText="FIGURE 1. The epigynes of six Bifidocoelotes spp., ventral view. A. Bifidocoelotes obscurus (after Zhou et al. 2017); B. Bifidocoelotes primus (after Zhou et al. 2017); C. Bifidocoelotes tsoi (after Wang et al. 2001); D. Bifidocoelotes mammiformis sp. nov.; E. Bifidocoelotes elongatus sp. nov.; F. Bifidocoelotes quadratus sp. nov.. Scale bar: 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.7456721" httpUri="https://zenodo.org/record/7456721/files/figure.png" pageId="2" pageNumber="181">Fig. 1BD</figureCitation>
).............................................................. 8
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30932E71D706FF8BB798FBE6FA3CFB88" pageId="2" pageNumber="181">
<paragraph id="8BD836D4D706FF8BB798FBE6FA3DFBAB" blockId="2.[151,1437,683,1203]" box="[151,1437,1093,1115]" pageId="2" pageNumber="181">
<keyLead id="309695E1D706FF8BB798FBE6FA3DFBAB" box="[151,1437,1093,1115]" pageId="2" pageNumber="181">
8. Spermathecae longitudinally extended (
<bibRefCitation id="EFF64B25D706FF8BB551FBE6FCA7FBAA" author="Wang, X. P. &amp; Tso, I. M. &amp; Wu, H. Y." box="[606,775,1093,1115]" pageId="2" pageNumber="181" pagination="127 - 133" refId="ref6755" refString="Wang, X. P., Tso, I. M. &amp; Wu, H. Y. (2001) Three new Coelotes species (Araneae: Amaurobiidae) from Taiwan. Zoological Studies, 40, 127 - 133." type="journal article" year="2001">
Wang
<emphasis id="B913EAC6D706FF8BB592FBE5FD67FBAA" box="[669,711,1093,1115]" italics="true" pageId="2" pageNumber="181">et al</emphasis>
. 2001
</bibRefCitation>
: figs 1, 2)........................................
<taxonomicName id="4C674D57D706FF8BB250FBE5FA3DFBAB" ID-CoL="99CNM" authorityName=": Li &amp; Blick" authorityYear="2020" box="[1375,1437,1094,1115]" class="Arachnida" family="Agelenidae" genus="Bifidocoelotes" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="181" phylum="Arthropoda" rank="species" species="tsoi">
<emphasis id="B913EAC6D706FF8BB250FBE5FA3DFBAB" box="[1375,1437,1094,1115]" italics="true" pageId="2" pageNumber="181">B. tsoi</emphasis>
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</paragraph>
<paragraph id="8BD836D4D706FF8BB798FBC0FA3CFB88" blockId="2.[151,1437,683,1203]" box="[151,1436,1123,1145]" pageId="2" pageNumber="181">
<keyLead id="309695E1D706FF8BB798FBC0FA3CFB88" box="[151,1436,1123,1145]" pageId="2" pageNumber="181">
- Spermathecae more or less globular (
<figureCitation id="135C2A51D706FF8BB548FBC0FD34FB89" box="[583,660,1123,1145]" captionStart="FIGURE 5" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[216,1369,181,1911]" captionTargetId="figure-31@6.[216,1371,181,1920]" captionTargetPageId="6" captionText="FIGURE 5. Bifidocoelotes mammiformis sp. nov., paratype female.A. Habitus, dorsal view; B. Same, ventral view; C. Epigyne, ventral view; D. Same, dorsal view. Abbreviations: CD, copulatory duct; CO, copulatory opening; ET, epigynal tooth; FD, fertilization duct; SH, spermathecal head; Sp, spermatheca. Scale bars: A, B = 1 mm; C, D = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7456735" httpUri="https://zenodo.org/record/7456735/files/figure.png" pageId="2" pageNumber="181">Fig. 5D</figureCitation>
;
<bibRefCitation id="EFF64B25D706FF8BB591FBC0FCE0FB88" author="Zhou, Y. C. &amp; Yuen, Y. L. &amp; Zhang, Z. S." box="[670,832,1123,1145]" pageId="2" pageNumber="181" pagination="429 - 436" refId="ref6836" refString="Zhou, Y. C., Yuen, Y. L. &amp; Zhang, Z. S. (2017) A new Bifidocoelotes species, with the description of the male of B. primus from Hong Kong, China (Agelenidae: Coelotinae). Zootaxa, 4232 (3), 429 - 436. https: // doi. org / 10.11646 / zootaxa. 4232.3.11" type="journal article" year="2017">
Zhou
<emphasis id="B913EAC6D706FF8BB5D9FBC0FCA1FB88" box="[726,769,1123,1144]" italics="true" pageId="2" pageNumber="181">et al</emphasis>
. 2017
</bibRefCitation>
: figs 3B, 4D)...................................... 9
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30932E71D706FF8BB798FB23FA3CFB42" pageId="2" pageNumber="181">
<paragraph id="8BD836D4D706FF8BB798FB23FA3DFB66" blockId="2.[151,1437,683,1203]" box="[151,1437,1152,1174]" pageId="2" pageNumber="181">
<keyLead id="309695E1D706FF8BB798FB23FA3DFB66" box="[151,1437,1152,1174]" pageId="2" pageNumber="181">
9. Spermathecae long finger-shaped, almost touching each other (
<bibRefCitation id="EFF64B25D706FF8BB431FB23FC42FB65" author="Zhou, Y. C. &amp; Yuen, Y. L. &amp; Zhang, Z. S." box="[830,994,1152,1174]" pageId="2" pageNumber="181" pagination="429 - 436" refId="ref6836" refString="Zhou, Y. C., Yuen, Y. L. &amp; Zhang, Z. S. (2017) A new Bifidocoelotes species, with the description of the male of B. primus from Hong Kong, China (Agelenidae: Coelotinae). Zootaxa, 4232 (3), 429 - 436. https: // doi. org / 10.11646 / zootaxa. 4232.3.11" type="journal article" year="2017">
Zhou
<emphasis id="B913EAC6D706FF8BB477FB22FC03FB65" box="[888,931,1152,1174]" italics="true" pageId="2" pageNumber="181">et al</emphasis>
. 2017
</bibRefCitation>
: figs 3B, 4D).................
<taxonomicName id="4C674D57D706FF8BB230FB22FA3DFB66" ID-CoL="LT4L" authorityName="MRDH" baseAuthorityName="Fox" baseAuthorityYear="1937" box="[1343,1437,1153,1174]" class="Arachnida" family="Agelenidae" genus="Bifidocoelotes" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="181" phylum="Arthropoda" rank="species" species="primus">
<emphasis id="B913EAC6D706FF8BB230FB22FA3DFB66" box="[1343,1437,1153,1174]" italics="true" pageId="2" pageNumber="181">B. primus</emphasis>
</taxonomicName>
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</paragraph>
<paragraph id="8BD836D4D706FF8BB798FB3DFA3CFB42" blockId="2.[151,1437,683,1203]" box="[151,1436,1181,1203]" pageId="2" pageNumber="181">
<keyLead id="309695E1D706FF8BB798FB3DFA3CFB42" box="[151,1436,1181,1203]" pageId="2" pageNumber="181">
- Spermathecae small ball-shaped and separated far from each other (
<figureCitation id="135C2A51D706FF8BB463FB3EFC17FB43" box="[876,951,1181,1203]" captionStart="FIGURE 5" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[216,1369,181,1911]" captionTargetId="figure-31@6.[216,1371,181,1920]" captionTargetPageId="6" captionText="FIGURE 5. Bifidocoelotes mammiformis sp. nov., paratype female.A. Habitus, dorsal view; B. Same, ventral view; C. Epigyne, ventral view; D. Same, dorsal view. Abbreviations: CD, copulatory duct; CO, copulatory opening; ET, epigynal tooth; FD, fertilization duct; SH, spermathecal head; Sp, spermatheca. Scale bars: A, B = 1 mm; C, D = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7456735" httpUri="https://zenodo.org/record/7456735/files/figure.png" pageId="2" pageNumber="181">Fig. 5D</figureCitation>
)..................
<taxonomicName id="4C674D57D706FF8BB3A4FB3DFAEAFB42" authority="Liao &amp; Wang &amp; Yin &amp; Xu, 2022" authorityName="Liao &amp; Wang &amp; Yin &amp; Xu" authorityYear="2022" box="[1195,1354,1181,1203]" class="Arachnida" family="Agelenidae" genus="Bifidocoelotes" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="181" phylum="Arthropoda" rank="species" species="mammiformis" status="sp. nov.">
<emphasis id="B913EAC6D706FF8BB3A4FB3DFAEAFB42" box="[1195,1354,1181,1203]" italics="true" pageId="2" pageNumber="181">B. mammiformis</emphasis>
</taxonomicName>
<emphasis id="B913EAC6D706FF8BB25FFB3EFA3CFB42" bold="true" box="[1360,1436,1181,1202]" pageId="2" pageNumber="181">
<taxonomicNameLabel id="A22057BDD706FF8BB25FFB3EFA3CFB42" box="[1360,1436,1181,1202]" pageId="2" pageNumber="181" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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