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<document id="F09D097F5D207B10F1F0441FB651437B" ID-DOI="10.11646/zootaxa.3652.3.2" ID-GBIF-Dataset="9cb4b047-ae44-419e-898a-eb3bc27c3d15" ID-ISSN="1175-5326" ID-Zenodo-Dep="224020" ID-ZooBank="B334624E-A7F5-4077-BF64-D2AD6DB215D5" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460609979718" checkinUser="plazi" docAuthor="Mattos, Ingrid &amp; Mermudes, José Ricardo M." docDate="2013" docId="03CE87A7FFB06940CFEFFE5DFEAFF822" docLanguage="en" docName="zt03652p342.pdf" docOrigin="Zootaxa 3652 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Paxillus reyesi Mattos &amp; Mermudes, new species" docType="treatment" docVersion="8" lastPageNumber="334" masterDocId="FFF7FFDFFFB76947CF78FFA2FFF8FFF8" masterDocTitle="Synopsis of Paxillus MacLeay, 1819 (Coleoptera: Passalidae): distributional records and descriptions of four new species from Brazil" masterLastPageNumber="342" masterPageNumber="327" pageNumber="334" updateTime="1698328243075" updateUser="plazi">
<mods:mods id="B792DE08384F2A194E1BF610DDDA3BF3" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="783268B9C47371772566D6DF69814254">
<mods:title id="8C903E148DE318555361316767CB19B1">Synopsis of Paxillus MacLeay, 1819 (Coleoptera: Passalidae): distributional records and descriptions of four new species from Brazil</mods:title>
</mods:titleInfo>
<mods:name id="E8E754169A2870A31CA3B7D7D6915C25" type="personal">
<mods:role id="30D43DFC85461ED2A9DFEA242E1EDBB8">
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</mods:role>
<mods:namePart id="01DEF23F79D75AC3C469C4C28B668F4E">Mattos, Ingrid</mods:namePart>
</mods:name>
<mods:name id="9E733CD0C3A364EB5B4F79DBB4CB4A42" type="personal">
<mods:role id="F8F9121404F3FADDB80B94FAA4C28B4C">
<mods:roleTerm id="05940F1F7F1D5D6FB4C57BE05C587E54">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="41E355077628AD616094B071B3B75F71">Mermudes, José Ricardo M.</mods:namePart>
</mods:name>
<mods:typeOfResource id="670020E839FFD9D8925FA5B3637247B8">text</mods:typeOfResource>
<mods:relatedItem id="11CDAC54F5D5D97FF6B19577ADB36143" type="host">
<mods:titleInfo id="F04A69CBC6F0E92C36D2D173D117C99B">
<mods:title id="7BB0D10FD7414529A49CEEB81E4C87A0">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="A884D7639B6A948E0BBC56ACE09FA095">
<mods:date id="3EEEE005DF2C083FE6623D08493085F2">2013</mods:date>
<mods:detail id="BEF06977FE5B017E3899B6813364CC12" type="volume">
<mods:number id="CD56FC6A83C7EE27C2BAB01BE8CA5800">3652</mods:number>
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<mods:detail id="2784E95F6AC1EB0210C0D34901A1F8D3" type="issue">
<mods:number id="056A32476B2BA6C6296EF7561E98AF6E">3</mods:number>
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<mods:classification id="8054BBA09E19DFC7C81872F5BC32C694">journal article</mods:classification>
<mods:identifier id="0A7C74A9FCC5793C90B0F830FAB1CF95" type="DOI">10.11646/zootaxa.3652.3.2</mods:identifier>
<mods:identifier id="8D1E59F025C29AA390F1DFBDF9533B19" type="GBIF-Dataset">9cb4b047-ae44-419e-898a-eb3bc27c3d15</mods:identifier>
<mods:identifier id="91F687C06A7237F19F550D369895506A" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="8A643A95EA823AD919D20B1B83F23506" type="Zenodo-Dep">224020</mods:identifier>
<mods:identifier id="AA1D9D95BB1EADD80BA66CF92EF778F5" type="ZooBank">B334624E-A7F5-4077-BF64-D2AD6DB215D5</mods:identifier>
</mods:mods>
<treatment id="03CE87A7FFB06940CFEFFE5DFEAFF822" ID-DOI="http://doi.org/10.5281/zenodo.6154347" ID-GBIF-Taxon="119568126" ID-Zenodo-Dep="6154347" LSID="urn:lsid:plazi:treatment:03CE87A7FFB06940CFEFFE5DFEAFF822" httpUri="http://treatment.plazi.org/id/03CE87A7FFB06940CFEFFE5DFEAFF822" lastPageNumber="334" pageId="7" pageNumber="334">
<subSubSection id="C37D653AFFB06940CFEFFE5DFED6FDC2" pageId="7" pageNumber="334" type="nomenclature">
<paragraph id="8BD836B1FFB06940CFEFFE5DFD0BFDE1" blockId="7.[151,755,511,570]" box="[151,755,511,537]" pageId="7" pageNumber="334">
<heading id="D09081DDFFB06940CFEFFE5DFD0BFDE1" bold="true" box="[151,755,511,537]" fontSize="11" level="1" pageId="7" pageNumber="334" reason="1">
<emphasis id="B913EAA3FFB06940CFEFFE5DFD0BFDE1" bold="true" box="[151,755,511,537]" pageId="7" pageNumber="334">
<taxonomicName id="4C674D32FFB06940CFEFFE5DFDAFFDE1" authority="Mattos &amp; Mermudes" authorityName="Mattos &amp; Mermudes" box="[151,599,511,537]" class="Insecta" family="Passallidae" genus="Paxillus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="334" phylum="Arthropoda" rank="species" species="reyesi" status="sp. nov.">
<emphasis id="B913EAA3FFB06940CFEFFE5DFEBFFDE1" bold="true" box="[151,327,511,537]" italics="true" pageId="7" pageNumber="334">Paxillus reyesi</emphasis>
Mattos &amp; Mermudes
</taxonomicName>
,
<taxonomicNameLabel id="A22057D8FFB06940CD1BFE5DFD0BFDE1" box="[611,755,511,537]" pageId="7" pageNumber="334" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BD836B1FFB06940CFEFFD80FED6FDC2" blockId="7.[151,755,511,570]" box="[151,302,546,570]" pageId="7" pageNumber="334">
(
<figureCitation id="135C2A34FFB06940CFE7FD80FEDDFDC2" box="[159,293,546,570]" captionStart="FIGURES 12 – 16" captionStartId="8.[151,269,1266,1290]" captionTargetBox="[204,1403,228,1208]" captionTargetId="figure@8.[151,1436,195,1245]" captionTargetPageId="8" captionText="FIGURES 12 – 16. Paxillus reyesi, 12, head and anterior margin of pronotum, dorsal; 13, antennae, dorsal. 14 – 16. Aedeagus: 14, dorsal; 15, lateral; 16, ventral. Scale = 0.5 mm (Figs. 14 – 16 to the same scale)." httpUri="https://zenodo.org/record/224024/files/figure.png" pageId="7" pageNumber="334">Figs. 12–16</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C37D653AFFB06940CFEFFDC8FAC3FA8E" pageId="7" pageNumber="334" type="diagnosis">
<paragraph id="8BD836B1FFB06940CFEFFDC8FCF8FAD6" blockId="7.[151,1437,618,2010]" pageId="7" pageNumber="334">
<emphasis id="B913EAA3FFB06940CFEFFDC8FEE9FD7B" bold="true" box="[151,273,618,643]" pageId="7" pageNumber="334">Diagnosis.</emphasis>
Length
<quantity id="4C9F9B54FFB06940CE08FDC8FE14FD7A" box="[368,492,618,642]" metricMagnitude="-2" metricUnit="m" metricValue="1.95" metricValueMax="2.1" metricValueMin="1.8" pageId="7" pageNumber="334" unit="mm" value="19.5" valueMax="21.0" valueMin="18.0">18–21 mm</quantity>
. Head (
<figureCitation id="135C2A34FFB06940CD3BFDC8FD60FD7A" box="[579,664,618,642]" captionStart="FIGURES 12 – 16" captionStartId="8.[151,269,1266,1290]" captionTargetBox="[204,1403,228,1208]" captionTargetId="figure@8.[151,1436,195,1245]" captionTargetPageId="8" captionText="FIGURES 12 – 16. Paxillus reyesi, 12, head and anterior margin of pronotum, dorsal; 13, antennae, dorsal. 14 – 16. Aedeagus: 14, dorsal; 15, lateral; 16, ventral. Scale = 0.5 mm (Figs. 14 – 16 to the same scale)." httpUri="https://zenodo.org/record/224024/files/figure.png" pageId="7" pageNumber="334">Fig. 12</figureCitation>
): labrum with anterior margin almost concave, sometimes discreetly notched medially; anterior frontal area punctate, punctures shallow and moderately coarse; median frontal area slightly elevated; anterior frontal ridges elevated, almost straight; posterior frontal ridges elevated, straight, and short; medial frontal tubercles robust, slightly larger than inner tubercles; inner tubercles robust, slightly elevated. Lateral frontal area deep and with fine punctures. Central tubercle short, free, more dilated at the base near posterior lateral tubercles. Bridge robust and smooth Lateral post-frontal area smooth and deep. Antennal club (
<figureCitation id="135C2A34FFB06940CA17FCBCFF4EFCA2" captionStart="FIGURES 12 – 16" captionStartId="8.[151,269,1266,1290]" captionTargetBox="[204,1403,228,1208]" captionTargetId="figure@8.[151,1436,195,1245]" captionTargetPageId="8" captionText="FIGURES 12 – 16. Paxillus reyesi, 12, head and anterior margin of pronotum, dorsal; 13, antennae, dorsal. 14 – 16. Aedeagus: 14, dorsal; 15, lateral; 16, ventral. Scale = 0.5 mm (Figs. 14 – 16 to the same scale)." httpUri="https://zenodo.org/record/224024/files/figure.png" pageId="7" pageNumber="334">Fig. 13</figureCitation>
): first antennomere reduced, slightly longer than half of the second antennomere, evidently more narrow. Pronotum: anterior and lateral margins straight; anterior marginal groove short, reaching ¼ of the length of anterior margin, narrow, with fine punctures. Lateral marginal groove narrow with fine punctures; lateral area with irregular and coarse punctures, often fused, moderately dense; scar punctate; median groove almost reaches anterior margin. Scutellum lacking median groove. Prosternal process posteriorly wide, truncate, with or without punctures. Elytra: humerus with tuft of setae (females) or sparse and short setae (males). Mesosternum smooth; scar subequal, half as long as the sides of mesosternum, deep and microsculptured, narrower posteriorly. Metasternum: disc evident, flattened, anterior lateral area finely and densely punctate, with scarce pubescence; posterior lateral area punctate, with dense and shallow punctures, denser near the disc. Mesotibia dorsally pubescent with one medial lateral spine. Metatibiae pubescent and unarmed. Aedeagus (
<figureCitation id="135C2A34FFB06940CDD0FB24FCCAFB66" box="[680,818,1158,1182]" captionStart="FIGURES 12 – 16" captionStartId="8.[151,269,1266,1290]" captionTargetBox="[204,1403,228,1208]" captionTargetId="figure@8.[151,1436,195,1245]" captionTargetPageId="8" captionText="FIGURES 12 – 16. Paxillus reyesi, 12, head and anterior margin of pronotum, dorsal; 13, antennae, dorsal. 14 – 16. Aedeagus: 14, dorsal; 15, lateral; 16, ventral. Scale = 0.5 mm (Figs. 14 – 16 to the same scale)." httpUri="https://zenodo.org/record/224024/files/figure.png" pageId="7" pageNumber="334">Figs. 14–16</figureCitation>
): convex, longer than wide; phallus globular, as long as parameres and phallobase together (lateral view); parameres and phallobase fused. Parameres wider in the median region, slightly more narrow and rounded at apex (lateral view). Phallobase slightly longer than wide, narrowed near phallobase, truncate at apex (lateral view); with sclerotized triangular area (dorsal view); wider than long; ventral distal margin with a small notch (ventral view).
</paragraph>
<paragraph id="8BD836B1FFB06940CFBFFA98FAC3FA8E" blockId="7.[151,1437,618,2010]" pageId="7" pageNumber="334">
<emphasis id="B913EAA3FFB06940CFBFFA98FE5AFAAB" bold="true" box="[199,418,1338,1363]" pageId="7" pageNumber="334">Dimensions (mm).</emphasis>
Total length (from anterior margin of labrum to apex of elytra) 18–21; elytral length (on median line) 11–13; pronotal length (on median line) 4.0–5.5; pronotal width 5.5–6.0; humeral width 6–7.
</paragraph>
</subSubSection>
<subSubSection id="C37D653AFFB06940CFBFFA21FA64F9B6" pageId="7" pageNumber="334" type="materials_examined">
<paragraph id="8BD836B1FFB06940CFBFFA21FA64F9B6" blockId="7.[151,1437,618,2010]" pageId="7" pageNumber="334">
<emphasis id="B913EAA3FFB06940CFBFFA21FE80FA63" bold="true" box="[199,376,1410,1435]" pageId="7" pageNumber="334">
<typeStatus id="54DC8813FFB06940CFBFFA21FEF8FA63" box="[199,256,1411,1435]" pageId="7" pageNumber="334">Type</typeStatus>
material.
</emphasis>
<typeStatus id="54DC8813FFB06940CEFEFA20FE16FA62" box="[390,494,1410,1434]" pageId="7" pageNumber="334" type="holotype">Holotype</typeStatus>
male from
<collectingCountry id="F3707621FFB06940CDF8FA21FD11FA63" box="[640,745,1411,1435]" name="Brazil" pageId="7" pageNumber="334">BRAZIL</collectingCountry>
: Rio Grande do Sul,
<collectingCountry id="F3707621FFB06940CC96FA20FBA9FA62" box="[1006,1105,1410,1434]" name="El Salvador" pageId="7" pageNumber="334">Salvador</collectingCountry>
do Sul,
<specimenCount id="9D61FD38FFB06940CBC6FA20FAF7FA62" box="[1214,1295,1410,1434]" pageId="7" pageNumber="334" type="male">1 male</specimenCount>
(dissected),
<date id="FFD91071FFB06940CFEFFA04FEF7FA46" box="[151,271,1446,1470]" pageId="7" pageNumber="334" value="1974-10-13">13.X.1974</date>
, C. Becker leg., col. MCN 21.193,
<emphasis id="B913EAA3FFB06940CDBFFA04FC45FA46" box="[711,957,1446,1470]" italics="true" pageId="7" pageNumber="334">
Paxillys
<taxonomicName id="4C674D32FFB06940CC56FA04FC45FA46" authority="Fonseca" authorityName="Fonseca" box="[814,957,1446,1470]" class="Insecta" family="Passallidae" genus="Passalus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="334" phylum="Arthropoda" rank="species" species="pentaphyllus">pentaphyllus</taxonomicName>
</emphasis>
Fonseca det. 1988 (MCNZ).
<typeStatus id="54DC8813FFB06940CA5FFA05FA60FA47" box="[1319,1432,1447,1471]" pageId="7" pageNumber="334" type="paratype">Paratypes</typeStatus>
:
<collectingCountry id="F3707621FFB06940CFEFFA69FF07FA1B" box="[151,255,1483,1507]" name="Brazil" pageId="7" pageNumber="334">BRAZIL</collectingCountry>
,
<emphasis id="B913EAA3FFB06940CE74FA69FE48FA1A" box="[268,432,1483,1506]" italics="true" pageId="7" pageNumber="334">Espírito Santo</emphasis>
: Santa Tereza,
<specimenCount id="9D61FD38FFB06940CD1BFA68FD49FA1A" box="[611,689,1482,1506]" pageId="7" pageNumber="334" type="male">1 male</specimenCount>
(dissected),
<date id="FFD91071FFB06940CC3CFA68FC70FA1A" box="[836,904,1482,1506]" pageId="7" pageNumber="334">7.XII.</date>
[19]64, C. Elias leg. (DZUP);
<emphasis id="B913EAA3FFB06940CB9EFA69FA6CFA1A" box="[1254,1428,1483,1506]" italics="true" pageId="7" pageNumber="334">Santa Catarina</emphasis>
:
<collectingCountry id="F3707621FFB06940CFEFFA4CFF11F9FE" box="[151,233,1518,1542]" name="Malta" pageId="7" pageNumber="334">Maratá</collectingCountry>
,
<specimenCount id="9D61FD38FFB06940CF8CFA4CFEB8F9FE" box="[244,320,1518,1542]" pageId="7" pageNumber="334" type="male">1 male</specimenCount>
(dissected),
<date id="FFD91071FFB06940CEA8FA4CFDDAF9FE" box="[464,546,1518,1542]" pageId="7" pageNumber="334" value="1926-02">II.1926</date>
(MAPA);
<emphasis id="B913EAA3FFB06940CDE1FA52FC94F9FE" box="[665,876,1518,1543]" italics="true" pageId="7" pageNumber="334">Rio Grande do Sul</emphasis>
, Alto Feliz,
<specimenCount id="9D61FD38FFB06940CC82FA4CFBA7F9FE" box="[1018,1119,1518,1542]" pageId="7" pageNumber="334" type="female">2 female</specimenCount>
,
<date id="FFD91071FFB06940CB12FA4CFB46F9FE" box="[1130,1214,1518,1542]" pageId="7" pageNumber="334" value="1932-02">II.1932</date>
, (MAPA), Itauba,
<specimenCount id="9D61FD38FFB06940CAF6FA4CFF1EF9D2" pageId="7" pageNumber="334" type="female">1 female</specimenCount>
,
<date id="FFD91071FFB06940CF89F9B0FE97F9D2" box="[241,367,1554,1578]" pageId="7" pageNumber="334" value="1978-04-19">19.IV.1978</date>
, M.H. Galileo leg. (MCNZ); Porto Alegre, Manresa,
<specimenCount id="9D61FD38FFB06940CCB0F9B0FBD4F9D2" box="[968,1068,1554,1578]" pageId="7" pageNumber="334" type="female">1 female</specimenCount>
,
<date id="FFD91071FFB06940CB4FF9B0FB4FF9D2" box="[1079,1207,1554,1578]" pageId="7" pageNumber="334" value="1982-03-29">29.III.1982</date>
, L.
<collectingCountry id="F3707621FFB06940CBA0F9B0FB14F9D2" box="[1240,1260,1554,1578]" name="Iceland" pageId="7" pageNumber="334">O</collectingCountry>
.Ferreira leg.,
<specimenCount id="9D61FD38FFB06940CAF6F9B0FF35F9B6" pageId="7" pageNumber="334" type="male">1 male</specimenCount>
and
<specimenCount id="9D61FD38FFB06940CE7CF994FE90F9B6" box="[260,360,1590,1614]" pageId="7" pageNumber="334" type="female">5 female</specimenCount>
,
<date id="FFD91071FFB06940CE0BF994FE22F9B6" box="[371,474,1590,1614]" pageId="7" pageNumber="334" value="1967-05-04">4.V.1967</date>
, F.R.Meyer leg. (MAPA);
<collectingCountry id="F3707621FFB06940CC7BF995FC77F9B7" box="[771,911,1591,1615]" name="Serbia" pageId="7" pageNumber="334">Montenegro</collectingCountry>
,
<specimenCount id="9D61FD38FFB06940CCE2F994FC06F9B6" box="[922,1022,1590,1614]" pageId="7" pageNumber="334" type="female">2 female</specimenCount>
,
<date id="FFD91071FFB06940CB71F994FB8CF9B6" box="[1033,1140,1590,1614]" pageId="7" pageNumber="334" value="1977-10-06">6.X.1977</date>
, T. Arigony leg. (MCNZ).
</paragraph>
</subSubSection>
<subSubSection id="C37D653AFFB06940CFBFF9F9FCC0F86A" pageId="7" pageNumber="334" type="discussion">
<paragraph id="8BD836B1FFB06940CFBFF9F9FF17F8B2" blockId="7.[151,1437,618,2010]" pageId="7" pageNumber="334">
<emphasis id="B913EAA3FFB06940CFBFF9F9FEC3F98B" bold="true" box="[199,315,1627,1651]" pageId="7" pageNumber="334">Remarks.</emphasis>
<taxonomicName id="4C674D32FFB06940CE3CF9F9FE13F98B" box="[324,491,1627,1651]" class="Insecta" family="Passallidae" genus="Paxillus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="334" phylum="Arthropoda" rank="species" species="reyesi">
<emphasis id="B913EAA3FFB06940CE3CF9F9FE13F98B" box="[324,491,1627,1651]" italics="true" pageId="7" pageNumber="334">Paxillus reyesi</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C674D32FFB06940CDFDF9FEFC7FF98A" authority="Palisot" authorityName="Palisot" box="[645,903,1626,1651]" class="Insecta" family="Passallidae" genus="Passalus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="334" phylum="Arthropoda" rank="species" species="pentaphyllus">
<emphasis id="B913EAA3FFB06940CDFDF9FEFCCAF98B" box="[645,818,1627,1651]" italics="true" pageId="7" pageNumber="334">P. pentaphyllus</emphasis>
Palisot
</taxonomicName>
de Beauvois, 1805,
<taxonomicName id="4C674D32FFB06940CB0CF9FEFA6FF98A" authority="Pangella, 1905" authorityName="Pangella" authorityYear="1905" box="[1140,1431,1626,1651]" class="Insecta" family="Passallidae" genus="Paxillus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="334" phylum="Arthropoda" rank="species" species="borellii">
<emphasis id="B913EAA3FFB06940CB0CF9FEFB1BF98B" box="[1140,1251,1627,1651]" italics="true" pageId="7" pageNumber="334">P. borellii</emphasis>
Pangella, 1905
</taxonomicName>
, and also from
<taxonomicName id="4C674D32FFB06940CE41F922FE61F96E" box="[313,409,1662,1687]" class="Insecta" family="Passallidae" genus="Paxillus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="334" phylum="Arthropoda" rank="species" species="leachi">
<emphasis id="B913EAA3FFB06940CE41F922FE61F96E" box="[313,409,1662,1687]" italics="true" pageId="7" pageNumber="334">P. leachi</emphasis>
</taxonomicName>
by the first antennomere of antennal club reduced (
<figureCitation id="135C2A34FFB06940CCA6F9DCFBC9F96E" box="[990,1073,1662,1686]" captionStart="FIGURES 12 – 16" captionStartId="8.[151,269,1266,1290]" captionTargetBox="[204,1403,228,1208]" captionTargetId="figure@8.[151,1436,195,1245]" captionTargetPageId="8" captionText="FIGURES 12 – 16. Paxillus reyesi, 12, head and anterior margin of pronotum, dorsal; 13, antennae, dorsal. 14 – 16. Aedeagus: 14, dorsal; 15, lateral; 16, ventral. Scale = 0.5 mm (Figs. 14 – 16 to the same scale)." httpUri="https://zenodo.org/record/224024/files/figure.png" pageId="7" pageNumber="334">Fig. 13</figureCitation>
), slightly longer than half of the second antennomere, evidently narrower; but mainly by the characters of aedeagus listed above (
<figureCitation id="135C2A34FFB06940CBABF900FAA5F942" box="[1235,1373,1698,1722]" captionStart="FIGURES 12 – 16" captionStartId="8.[151,269,1266,1290]" captionTargetBox="[204,1403,228,1208]" captionTargetId="figure@8.[151,1436,195,1245]" captionTargetPageId="8" captionText="FIGURES 12 – 16. Paxillus reyesi, 12, head and anterior margin of pronotum, dorsal; 13, antennae, dorsal. 14 – 16. Aedeagus: 14, dorsal; 15, lateral; 16, ventral. Scale = 0.5 mm (Figs. 14 – 16 to the same scale)." httpUri="https://zenodo.org/record/224024/files/figure.png" pageId="7" pageNumber="334">Figs. 14–16</figureCitation>
).
<taxonomicName id="4C674D32FFB06940CFEFF964FEB9F927" box="[151,321,1734,1759]" class="Insecta" family="Passallidae" genus="Paxillus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="334" phylum="Arthropoda" rank="species" species="reyesi">
<emphasis id="B913EAA3FFB06940CFEFF964FEB9F927" box="[151,321,1734,1759]" italics="true" pageId="7" pageNumber="334">Paxillus reyesi</emphasis>
</taxonomicName>
is more similar to
<taxonomicName id="4C674D32FFB06940CD48F96AFCC1F926" authority="Palisot" authorityName="Palisot" box="[560,825,1734,1759]" class="Insecta" family="Passallidae" genus="Passalus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="334" phylum="Arthropoda" rank="species" species="pentaphyllus">
<emphasis id="B913EAA3FFB06940CD48F96AFD19F926" box="[560,737,1734,1759]" italics="true" pageId="7" pageNumber="334">P. pentaphyllus</emphasis>
Palisot
</taxonomicName>
de Beauvois, 1805, which has been the subject of discussion in the literature since Boucher (2006). He questioned the description of the aedeagus of
<taxonomicName id="4C674D32FFB06940CB93F94EFAA0F8FB" box="[1259,1368,1771,1795]" class="Insecta" family="Passallidae" genus="Paxillus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="334" phylum="Arthropoda" rank="species" species="borellii">
<emphasis id="B913EAA3FFB06940CB93F94EFAA0F8FB" box="[1259,1368,1771,1795]" italics="true" pageId="7" pageNumber="334">P. borellii</emphasis>
</taxonomicName>
made by Bührnheim (1978) because it is indeed
<taxonomicName id="4C674D32FFB06940CDEDF8B2FBB5F8DE" authority="Luederwaldt, 1931" authorityName="Luederwaldt" authorityYear="1931" box="[661,1101,1806,1831]" class="Insecta" family="Passallidae" genus="Paxillus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="334" phylum="Arthropoda" rank="species" species="pentaphylloides">
<emphasis id="B913EAA3FFB06940CDEDF8B2FC9EF8DE" box="[661,870,1806,1831]" italics="true" pageId="7" pageNumber="334">P. pentaphylloides</emphasis>
Luederwaldt, 1931
</taxonomicName>
, today synonymized with
<taxonomicName id="4C674D32FFB06940CAFFF8B2FF10F8B3" class="Insecta" family="Passallidae" genus="Paxillus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="334" phylum="Arthropoda" rank="species" species="borellii">
<emphasis id="B913EAA3FFB06940CAFFF8B2FF10F8B3" italics="true" pageId="7" pageNumber="334">P. borellii</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BD836B1FFB06940CFBFF8F4FCC0F86A" blockId="7.[151,1437,618,2010]" pageId="7" pageNumber="334">
Based on comparative studies of the aedeagus,
<taxonomicName id="4C674D32FFB06940CD87F8FAFCA2F897" box="[767,858,1880,1903]" class="Insecta" family="Passallidae" genus="Paxillus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="334" phylum="Arthropoda" rank="species" species="reyes">
<emphasis id="B913EAA3FFB06940CD87F8FAFCA2F897" box="[767,858,1880,1903]" italics="true" pageId="7" pageNumber="334">P. reyes</emphasis>
</taxonomicName>
i is different from the
<taxonomicName id="4C674D32FFB06940CB14F8FAFA8CF896" authority="Palisot" authorityName="Palisot" box="[1132,1396,1878,1903]" class="Insecta" family="Passallidae" genus="Passalus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="334" phylum="Arthropoda" rank="species" species="pentaphyllus">
<emphasis id="B913EAA3FFB06940CB14F8FAFAE4F896" box="[1132,1308,1878,1903]" italics="true" pageId="7" pageNumber="334">P. pentaphyllus</emphasis>
Palisot
</taxonomicName>
de Beauvois, 1805 specimens studied by Boucher (from
<collectingCountry id="F3707621FFB06940CD97F8D8FCD4F86A" box="[751,812,1914,1938]" name="Haiti" pageId="7" pageNumber="334">Haiti</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="C37D653AFFB06940CFBFF83DFEAFF822" pageId="7" pageNumber="334" type="etymology">
<paragraph id="8BD836B1FFB06940CFBFF83DFEAFF822" blockId="7.[151,1437,618,2010]" pageId="7" pageNumber="334">
<emphasis id="B913EAA3FFB06940CFBFF83DFEB6F84F" bold="true" box="[199,334,1951,1975]" pageId="7" pageNumber="334">Etymology.</emphasis>
The name is in honor to Dr. Pedro Reyes-Castillo, who very kindly helped us and encouraged us to study
<taxonomicName id="4C674D32FFB06940CFA2F860FEAAF822" box="[218,338,1986,2010]" class="Insecta" family="Passalidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="334" phylum="Arthropoda" rank="family">Passalidae</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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