161 lines
19 KiB
XML
161 lines
19 KiB
XML
<document id="3EA4BE66976B40DEF314135BF08C75E5" ID-CLB-Dataset="277216" ID-DOI="http://doi.org/10.5281/zenodo.10135885" ID-GBIF-Dataset="482f3dde-6113-4733-945d-ae301578ea12" ID-ISSN="1211-8788" ID-Zenodo-Dep="10135885" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1699970497318" checkinUser="juliana" docAuthor="Löbl, Ivan" docDate="2023" docId="7626AB1DFFE8FFF4E8978C8C7168F9EA" docLanguage="en" docName="ActaMusMoravSciBiol.108.1-2.1-34.pdf" docOrigin="Acta Musei Moraviae, Scientiae biologicae 108 (1 - 2)" docTitle="Scaphisoma bryanti Löbl 2023, sp. nov." docType="treatment" docVersion="5" lastPageNumber="10" masterDocId="8A1FD365FFE0FFFDE916887C7240FFFA" masterDocTitle="On the Scaphidiinae (Coleoptera: Staphylinidae) of Java, Indonesia" masterLastPageNumber="34" masterPageNumber="1" pageNumber="9" updateTime="1700051061570" updateUser="ExternalLinkService">
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<mods:mods id="EF723B21481499794D02853E07524AC1" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="0CD995851E38F8AEB79BCE6E6F174236">
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<mods:title id="E938BFEE1092F28894F5C9683C1B8091">On the Scaphidiinae (Coleoptera: Staphylinidae) of Java, Indonesia</mods:title>
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</mods:titleInfo>
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<mods:name id="372C3F47BE245DDB7CA4594AF8498685" type="personal">
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<mods:roleTerm id="9445B1CDC2D6782565675493209D1D25">Author</mods:roleTerm>
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<mods:namePart id="1708A1839291A26E4C925CDBB0A8449B">Löbl, Ivan</mods:namePart>
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<mods:affiliation id="47E22EF8380252EA0C8747B0775A2E7E">Museum dhistoire naturelle, C. P. 6434, CH 1211 Genève 6, Switzerland</mods:affiliation>
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<mods:nameIdentifier id="E36F0B83F3F599EE6E99082BCB646F83" type="email">email: Ivan.lobl@bluewin.ch</mods:nameIdentifier>
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<mods:typeOfResource id="0572641568E1BB2A587F9C77FEC40BD6">text</mods:typeOfResource>
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<mods:title id="82E1B14A9C491EC7929B5EB7A61495D8">Acta Musei Moraviae, Scientiae biologicae</mods:title>
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</mods:titleInfo>
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<mods:part id="F4D2014AA9CA9DE2C500840EEF2DC4F5">
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<mods:date id="BFC667DF6D83E327AAF2167B42AE6ECC">2023</mods:date>
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<mods:number id="0D2ECD5ABFFC25B97AFE7E9AA0194145">2023-11-01</mods:number>
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<mods:detail id="71A17919C4AF5BFF3460F9D0D9A6611F" type="volume">
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<mods:number id="7B83A67A0F45341865549927A5874F40">108</mods:number>
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<mods:number id="D9A7A1001F4766E0DBDF2A81CB656749">1 - 2</mods:number>
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<mods:start id="3F23E51C374B1C3F2A7E40DB2EC5F2F4">1</mods:start>
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<mods:classification id="B06E202F7DCFBE0259FBF65D8DC7D40B">journal article</mods:classification>
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<mods:identifier id="E46920031C4AF5556F299514E48438F6" type="CLB-Dataset">277216</mods:identifier>
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<mods:identifier id="AE5D05E42E2098090C6F3ADA7609256F" type="DOI">http://doi.org/10.5281/zenodo.10135885</mods:identifier>
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<mods:identifier id="951CE8C495BFACE46DB8FCAEF81AA256" type="GBIF-Dataset">482f3dde-6113-4733-945d-ae301578ea12</mods:identifier>
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<mods:identifier id="C1BA8497A0B0843B740AB2162E1C34A9" type="ISSN">1211-8788</mods:identifier>
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<mods:identifier id="534D82541790D8D963F0B0DEFF62FC6C" type="Zenodo-Dep">10135885</mods:identifier>
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</mods:mods>
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<treatment id="7626AB1DFFE8FFF4E8978C8C7168F9EA" ID-GBIF-Taxon="217574520" LSID="urn:lsid:plazi:treatment:7626AB1DFFE8FFF4E8978C8C7168F9EA" httpUri="http://treatment.plazi.org/id/7626AB1DFFE8FFF4E8978C8C7168F9EA" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">
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<subSubSection id="B6954980FFE8FFF5E8978C8C70F9FAF3" box="[385,697,1264,1289]" pageId="8" pageNumber="9" type="nomenclature">
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<paragraph id="FE301A0BFFE8FFF5E8978C8C70F9FAF3" blockId="8.[385,697,1264,1289]" box="[385,697,1264,1289]" pageId="8" pageNumber="9">
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<heading id="A578AD67FFE8FFF5E8978C8C70F9FAF3" bold="true" box="[385,697,1264,1289]" fontSize="10" level="2" pageId="8" pageNumber="9" reason="2">
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<emphasis id="CCFBC619FFE8FFF5E8978C8C70F9FAF3" bold="true" box="[385,697,1264,1289]" pageId="8" pageNumber="9">
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<taxonomicName id="398F6188FFE8FFF5E8978C8C701CFAF2" authority="Löbl, 2023" authorityName="Löbl" authorityYear="2023" box="[385,604,1264,1288]" class="Insecta" family="Staphylinidae" genus="Scaphisoma" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="bryanti" status="sp. nov.">
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<emphasis id="CCFBC619FFE8FFF5E8978C8C701CFAF2" bold="true" box="[385,604,1264,1288]" italics="true" pageId="8" pageNumber="9">Scaphisoma bryanti</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="D7C87B62FFE8FFF5EB758C8D70F9FAF3" box="[611,697,1265,1289]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="B6954980FFE8FFF5EAFA8C8D761BFAF3" box="[1004,1115,1265,1289]" pageId="8" pageNumber="9" type="description">
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<paragraph id="FE301A0BFFE8FFF5EAFA8C8D761BFAF3" blockId="8.[1004,1115,1265,1289]" box="[1004,1115,1265,1289]" pageId="8" pageNumber="9">
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(
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<figureCitation id="66B4068EFFE8FFF5EAE58C8D7613FAF3" box="[1011,1107,1265,1289]" captionStart="Figs 8-13" captionStartId="11.[304,342,1687,1706]" captionTargetBox="[351,1196,395,1653]" captionTargetId="figure-83@11.[339,1215,390,1657]" captionTargetPageId="11" captionText="Figs 8-13. Male genitalia. 8 Scaphisoma bryanti sp. nov., aedeagus in dorsal view, scale = 0.2 mm. 9 Ditto, in ventral view, without proximal part of basal bulb, scale = 0.1 mm. 10, 11 Scaphisoma drescheri sp. nov., aedeagus in dorsal and lateral views, scale = 0.1 mm. 12, 13 Scaphisoma multispinosum sp. nov., aedeagus in dorsal and lateral views, scale = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10135889" httpUri="https://zenodo.org/record/10135889/files/figure.png" pageId="8" pageNumber="9">Figs 8, 9</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="B6954980FFE8FFF5E85A8D5C70D1FAB6" pageId="8" pageNumber="9" type="materials_examined">
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<paragraph id="FE301A0BFFE8FFF5E85A8D5C70D1FAB6" blockId="8.[332,1280,1312,1357]" pageId="8" pageNumber="9">
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<emphasis id="CCFBC619FFE8FFF5E85A8D5C73BFFAC9" bold="true" box="[332,511,1312,1331]" pageId="8" pageNumber="9">Material examined.</emphasis>
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<materialsCitation id="4EE71056FFE8FFF5EB128D5C762AFAC9" ID-GBIF-Occurrence="4438213359" box="[516,1130,1312,1331]" collectingDate="2009-04-10" collectionCode="NHML" collectorName="G. E. Bryant" location="Kawah Manoek" pageId="8" pageNumber="9" specimenCount="1" specimenCount-male="1" stateProvince="Java" typeStatus="holotype">
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<typeStatus id="2134A4A9FFE8FFF5EB128D5C7013FAC9" box="[516,595,1312,1331]" pageId="8" pageNumber="9">Holotype</typeStatus>
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<specimenCount id="E889D182FFE8FFF5EB4C8D5C70C6FAC9" box="[602,646,1312,1331]" pageId="8" pageNumber="9" type="male">male</specimenCount>
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,
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<location id="FB504CD0FFE8FFF5EB998D5C7159FAC9" LSID="urn:lsid:plazi:treatment:7626AB1DFFE8FFF4E8978C8C7168F9EA:FB504CD0FFE8FFF5EB998D5C7159FAC9" box="[655,793,1312,1331]" name="Kawah Manoek" pageId="8" pageNumber="9" stateProvince="Java">Kawah Manoek</location>
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,
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<collectingRegion id="3C4BD4E9FFE8FFF5EA348D5C710AFAC9" box="[802,842,1312,1331]" pageId="8" pageNumber="9">Java</collectingRegion>
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,
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<collectorName id="537A7FDDFFE8FFF5EA458D5C7184FAC9" box="[851,964,1312,1331]" pageId="8" pageNumber="9">G. E. Bryant</collectorName>
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.
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<date id="8A313CCBFFE8FFF5EADA8D5C7650FAC9" box="[972,1040,1312,1331]" pageId="8" pageNumber="9" value="2009-04-10">
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<collectingDate id="9A75C523FFE8FFF5EADA8D5C7650FAC9" box="[972,1040,1312,1331]" pageId="8" pageNumber="9" value="2009-04-10">10.iv.09</collectingDate>
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</date>
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(
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<collectionCode id="989E82CEFFE8FFF5ED0B8D5C7621FAC9" box="[1053,1121,1312,1331]" collectionName="Libya, Tripoli, Natural History Museum" pageId="8" pageNumber="9">NHML</collectionCode>
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).
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</materialsCitation>
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<materialsCitation id="4EE71056FFE8FFF5ED798D5C70D1FAB6" ID-GBIF-Occurrence="4438213328" collectingDate="2009-04-10" collectionCode="MHNG" collectorName="G. E. Bryant" location="Kawah Manoek" pageId="8" pageNumber="9" specimenCount="1" specimenCount-female="1" stateProvince="Java" typeStatus="paratype">
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<typeStatus id="2134A4A9FFE8FFF5ED798D5C76FAFAC9" box="[1135,1210,1312,1331]" pageId="8" pageNumber="9">Paratype</typeStatus>
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<specimenCount id="E889D182FFE8FFF5EDD68D5C76BCFAC9" box="[1216,1276,1312,1331]" pageId="8" pageNumber="9" type="female">female</specimenCount>
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, with same data as holotype (
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<collectionCode id="989E82CEFFE8FFF5EB578D4570C7FAB6" box="[577,647,1337,1356]" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle" pageId="8" pageNumber="9">MHNG</collectionCode>
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).
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="B6954980FFE8FFF4E85A8D1E70F5FB1B" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9" type="description">
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<paragraph id="FE301A0BFFE8FFF4E85A8D1E717DFB7A" blockId="8.[332,1280,1376,1754]" lastBlockId="9.[304,1252,393,1552]" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">
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<emphasis id="CCFBC619FFE8FFF5E85A8D1E7397FA80" bold="true" box="[332,471,1378,1402]" pageId="8" pageNumber="9">Description.</emphasis>
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Length 2.92
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<quantity id="3977B7EEFFE8FFF5EB7E8D1C7091FA80" box="[616,721,1376,1402]" metricMagnitude="-3" metricUnit="m" metricValue="2.95" pageId="8" pageNumber="9" unit="mm" value="2.95">2.95 mm</quantity>
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, width 1.65
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<quantity id="3977B7EEFFE8FFF5EA4E8D1C7182FA80" box="[856,962,1376,1402]" metricMagnitude="-3" metricUnit="m" metricValue="1.67" pageId="8" pageNumber="9" unit="mm" value="1.67">1.67 mm</quantity>
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. Head and most of pronotum ochraceous. Basal margin of pronotum black; black area expanded mesally and nearly reaching pronotal midlength, forming notched spot. Elytron ochraceous on prevailing surface, black or blackish along basal and lateral margins, and along suture up to light apical seventh. Dark lateral band expanding anterior of lateral midlength to form small transverse band nearly reaching elytral midwidth. Margin of dark basal area irregular, denticulate and expanding along sutural stria to form wide band reaching somewhat posterior of elytral midlength. Small light brown spot present on subapical elytral area. Epipleuron blackish. Hypomeron ochraceous with narrowly darkened basal and ventral margins. Venter of mesothorax and metathorax black. Ventrite I blackish on prevailing surface, becoming lighter apicad. Following ventrites yellowish. Appendages ochraceous to yellowish. Length/width ratios of antennomeres
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<date id="8A313CCBFFE8FFF5EA678EBF7194F920" box="[881,980,1730,1754]" pageId="8" pageNumber="9" value="2010-03-16">III 16/10</date>
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:
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<date id="8A313CCBFFE8FFF5EAC98EBF765FF920" box="[991,1055,1730,1754]" pageId="8" pageNumber="9" value="1940-04">IV 40</date>
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/9:
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<date id="8A313CCBFFE8FFF5ED568EBF7637F920" box="[1088,1143,1730,1754]" pageId="8" pageNumber="9" value="1960-05">V 60</date>
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/9:
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<date id="8A313CCBFFE8FFF5ED8E8EBF7699F920" box="[1176,1241,1730,1754]" pageId="8" pageNumber="9" value="1968-06">VI 68</date>
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/11:
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<date id="8A313CCBFFE9FFF4E82689F5733EFE5B" box="[304,382,393,417]" pageId="9" pageNumber="10" value="1965-07">VII 65</date>
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/15:
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<date id="8A313CCBFFE9FFF4E8A789F57047FE5B" box="[433,519,393,417]" pageId="9" pageNumber="10" value="1955-08">VIII 55</date>
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/10:
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<date id="8A313CCBFFE9FFF4EB2D89F570C1FE5B" box="[571,641,393,417]" pageId="9" pageNumber="10" value="1968-09">IX 68</date>
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/15:
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<date id="8A313CCBFFE9FFF4EBA289F570B0FE5B" box="[692,752,393,417]" pageId="9" pageNumber="10" value="1957-10">X 57</date>
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/15: XI: 73/17. Pronotum and elytra lacking microsculpture. Pronotum with rounded lateral margins, lateral carinae visible in dorsal view, lateral margin stria impunctate, discal punctation extremely fine, hardly visible at 100 times magnification. Tip of scutellum exposed, triangular. Elytron rather strongly narrowed apicad, with lateral margin oblique posterior of basal third; lateral margin carina exposed in dorsal view; lateral margin stria punctate; apical margin rounded; inner apical angle situated posterior of level of outer apical angle; apical crenulation present; sutural margin not raised; sutural stria shallow, starting at or posterior of level of scutellum, gradually converging to suture, adsutural area flat, near scutellum about 0.08
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<quantity id="3977B7EEFFE9FFF4E8738ADB738CFD3A" box="[357,460,679,705]" metricMagnitude="-5" metricUnit="m" metricValue="9.0" pageId="9" pageNumber="10" unit="mm" value="0.09">0.09 mm</quantity>
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wide, very finely punctate; discal punctation much coarser than that on pronotum, punctures not clearly delimited, puncture intervals mostly about two to three times as large as puncture diameters. Hind wings fully developed. Hypomeron smooth. Mesanepisternum extremely finely and sparsely punctate, lacking microsculpture. Mesepimeron well five times as long as wide, nearly twice as long as interval between its tip and mesocoxa. Metaventrite with strigulate microsculpture, very finely and sparsely punctate; median area of metaventrite convex, lacking impressions or stria; antecoxal puncture rows present, poorly distinct. Submesocoxal areas about
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<quantity id="3977B7EEFFE9FFF4ED168BF57620FC5A" box="[1024,1120,905,929]" metricMagnitude="-5" metricUnit="m" metricValue="4.0" pageId="9" pageNumber="10" unit="mm" value="0.04">0.04 mm</quantity>
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long, about as long as seventh of shortest interval to metacoxa; submesocoxal lines nearly parallel, very finely punctate. Metanepisternum flat, lacking microsculpture, slightly narrowing anteriad, with suture straight except at rounded anterior angle. Protibiae and metatibiae straight, mesotibiae curved. Ventrites with strigulate microsculpture and punctation as fine and sparse as than on metaventrite. Ventrite I with submetacoxal areas 0.06
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<quantity id="3977B7EEFFE9FFF4ED688C5B76A2FBBA" box="[1150,1250,1063,1089]" metricMagnitude="-5" metricUnit="m" metricValue="7" pageId="9" pageNumber="10" unit="mm" value="0.07">0.07 mm</quantity>
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long, about as long as fourth of shortest interval to apical margin of ventrite, submetacoxal lines convex, very finely punctate.
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</paragraph>
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<paragraph id="FE301A0BFFE9FFF4E8738CF570F5FB1B" blockId="9.[304,1252,393,1552]" pageId="9" pageNumber="10">
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<emphasis id="CCFBC619FFE9FFF4E8738CF573E3FB5B" box="[357,419,1161,1185]" italics="true" pageId="9" pageNumber="10">Male.</emphasis>
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Protarsomeres and mesotarsomeres I to III distinctly widened, narrower than apex of protibia. Apical margin of ventrite VI subangulate, lacking distinct lobe. Aedeagus (
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<figureCitation id="66B4068EFFE9FFF4E8BE8CB57048FB1B" box="[424,520,1225,1249]" captionStart="Figs 8-13" captionStartId="11.[304,342,1687,1706]" captionTargetBox="[351,1196,395,1653]" captionTargetId="figure-83@11.[339,1215,390,1657]" captionTargetPageId="11" captionText="Figs 8-13. Male genitalia. 8 Scaphisoma bryanti sp. nov., aedeagus in dorsal view, scale = 0.2 mm. 9 Ditto, in ventral view, without proximal part of basal bulb, scale = 0.1 mm. 10, 11 Scaphisoma drescheri sp. nov., aedeagus in dorsal and lateral views, scale = 0.1 mm. 12, 13 Scaphisoma multispinosum sp. nov., aedeagus in dorsal and lateral views, scale = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10135889" httpUri="https://zenodo.org/record/10135889/files/figure.png" pageId="9" pageNumber="10">Figs 8, 9</figureCitation>
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)
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<quantity id="3977B7EEFFE9FFF4EB018CB57036FB1A" box="[535,630,1225,1249]" metricMagnitude="-3" metricUnit="m" metricValue="1.01" pageId="9" pageNumber="10" unit="mm" value="1.01">1.01 mm</quantity>
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long.
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</paragraph>
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</subSubSection>
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<subSubSection id="B6954980FFE9FFF4E8268C8D7380FAD3" pageId="9" pageNumber="10" type="etymology">
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<paragraph id="FE301A0BFFE9FFF4E8268C8D7380FAD3" blockId="9.[304,1252,393,1552]" pageId="9" pageNumber="10">
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<emphasis id="CCFBC619FFE9FFF4E8268C8D73F0FAF3" bold="true" box="[304,432,1265,1289]" pageId="9" pageNumber="10">Etymology.</emphasis>
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The species is named in honour of its collector, Gilbert Ernest Bryant (1878 1965).
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</paragraph>
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</subSubSection>
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<subSubSection id="B6954980FFE9FFF4E8268D447168F9EA" pageId="9" pageNumber="10" type="diagnosis">
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<paragraph id="FE301A0BFFE9FFF4E8268D447168F9EA" blockId="9.[304,1252,393,1552]" pageId="9" pageNumber="10">
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<emphasis id="CCFBC619FFE9FFF4E8268D44706FFAAA" bold="true" box="[304,559,1336,1360]" pageId="9" pageNumber="10">Differential diagnosis.</emphasis>
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The conspicuous colour pattern of the pronotum of this new species is similar to that of
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<taxonomicName id="398F6188FFE9FFF4EB598D247177FA8A" baseAuthorityName="Pic" baseAuthorityYear="1915" box="[591,823,1368,1392]" class="Insecta" family="Staphylinidae" genus="Scaphisoma" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="testaceomaculatum">
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<emphasis id="CCFBC619FFE9FFF4EB598D247177FA8A" box="[591,823,1368,1392]" italics="true" pageId="9" pageNumber="10">S. testaceomaculatum</emphasis>
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</taxonomicName>
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, but the elytra are light on a much larger surface. The latter species differs notably by the elytra with sutural striae deep and much more converging to the suture, and the adsutural areas coarsely punctate. Unlike
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<taxonomicName id="398F6188FFE9FFF4EDD98DE473BEFA2A" baseAuthorityName="Pic" baseAuthorityYear="1915" class="Insecta" family="Staphylinidae" genus="Scaphisoma" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="testaceomaculatum">
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<emphasis id="CCFBC619FFE9FFF4EDD98DE473BEFA2A" italics="true" pageId="9" pageNumber="10">S. testaceomaculatum</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="398F6188FFE9FFF4EB1B8DC47038FA2A" authorityName="Löbl" authorityYear="2023" box="[525,632,1464,1488]" class="Insecta" family="Staphylinidae" genus="Scaphisoma" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="bryanti">
|
||
<emphasis id="CCFBC619FFE9FFF4EB1B8DC47038FA2A" box="[525,632,1464,1488]" italics="true" pageId="9" pageNumber="10">S. bryanti</emphasis>
|
||
</taxonomicName>
|
||
has a symmetrical aedeagus and is a members of the
|
||
<taxonomicName id="398F6188FFE9FFF4EDD98DC47339FA0A" authorityName="Motschulsky" authorityYear="1863" class="Insecta" family="Staphylinidae" genus="Scaphisoma" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="pictum">
|
||
<emphasis id="CCFBC619FFE9FFF4EDD98DC47339FA0A" italics="true" pageId="9" pageNumber="10">S. pictum</emphasis>
|
||
</taxonomicName>
|
||
group. It may be distinguished from other species of the group by its large body size and the colour of the pronotum and elytra.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |