417 lines
45 KiB
XML
417 lines
45 KiB
XML
<document ID-CLB-Dataset="21607" ID-DOI="http://dx.doi.org/10.3897/phytokeys.151.50538" ID-GBIF-Dataset="a62873a2-c4e1-4c6d-9baa-5c5fb94e7348" ID-PMC="PMC7329918" ID-Pensoft-Pub="1314-2003-151-107" ID-Pensoft-UUID="42D1EB2DF0545744B92BFCCCB395C45B" ID-PubMed="32655282" ModsDocID="1314-2003-151-107" checkinTime="1593036331120" checkinUser="pensoft" docAuthor="Sylvester, Steven P., Cuta-Alarcon, Lia E., Bravo-Pedraza, William J. & Soreng, Robert J." docDate="2020" docId="4E7CFCB1729C50408D8C94AC584BC6C6" docLanguage="en" docName="PhytoKeys 151: 107-160" docOrigin="PhytoKeys 151" docSource="http://dx.doi.org/10.3897/phytokeys.151.50538" docTitle="Agrostis imberbis Phil. Anales Univ. Chile 94: 11. 1896" docType="treatment" docVersion="6" id="42D1EB2DF0545744B92BFCCCB395C45B" lastPageNumber="107" masterDocId="42D1EB2DF0545744B92BFCCCB395C45B" masterDocTitle="Agrostis and Podagrostis (Agrostidinae, Poaceae) from paramos of Boyaca, Colombia: synoptic taxonomy including a key to Colombian species" masterLastPageNumber="160" masterPageNumber="107" pageNumber="107" updateTime="1732688126400" updateUser="ExternalLinkService">
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<mods:titleInfo id="C7892A48B7AA14E46E6EA6B0EDA56B11">
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<mods:title id="1ED81D1BC7053A8832959AFE1DDB6EF9">Agrostis and Podagrostis (Agrostidinae, Poaceae) from paramos of Boyaca, Colombia: synoptic taxonomy including a key to Colombian species</mods:title>
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<mods:name id="F3EFF4200A412AD818A1DED8B0F6B574" type="personal">
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<mods:roleTerm id="7F7D28D8620B856FA3134B5B26934FED">Author</mods:roleTerm>
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<mods:namePart id="D76996C6424F9EECA9F0A3F32F2507A1">Sylvester, Steven P.</mods:namePart>
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<mods:affiliation id="D64D7F30DD39DA9039F8DCA7B5EF42F2">College of Biology and the Environment, Nanjing Forestry University, Long Pan Road No. 159, Nanjing, 210037, China & Royal Botanic Gardens, Kew, Richmond, Surrey, TW 9 3 AE, UK</mods:affiliation>
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<mods:nameIdentifier id="7EA618C4E921E877B49218992F2D9896" type="orcid">https://orcid.org/0000-0001-5577-8782</mods:nameIdentifier>
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<mods:nameIdentifier id="B68B63FFC9D61E50FB1157C999C91117" type="email">steven_sylvester@hotmail.com</mods:nameIdentifier>
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<mods:roleTerm id="73BC68160923B5CB7C9F6773D82DF7A1">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="0F264C2453ED3A9FCB314ECE8F0D99D3">Cuta-Alarcon, Lia E.</mods:namePart>
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<mods:affiliation id="815176792DEE10465B5B8B24ECBA1315">Grupo Sistematica Biologica, Herbario UPTC, Escuela de Biologia, Facultad de Ciencias, Universidad Pedagogica y Tecnologica de Colombia, Avenida Central del Norte 39 - 115, Tunja-Boyaca, Colombia</mods:affiliation>
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</mods:name>
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<mods:name id="5A869B16C1E0DBF007D6EC42042D9E97" type="personal">
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<mods:roleTerm id="2F02EF8ABDDC0A3FEEAC6C936285A108">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="988E4D50F2402B7A631F5094A3446458">Bravo-Pedraza, William J.</mods:namePart>
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<mods:affiliation id="087519BE273B62C1F7E5BFF5C4F8D56D">Grupo Sistematica Biologica, Herbario UPTC, Escuela de Biologia, Facultad de Ciencias, Universidad Pedagogica y Tecnologica de Colombia, Avenida Central del Norte 39 - 115, Tunja-Boyaca, Colombia</mods:affiliation>
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<mods:name id="4249BF758A90A7D8784FBC1C1C931BC4" type="personal">
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<mods:roleTerm id="C2FE3EE68724E12A66EE4D21196CF120">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="E4E019AA44D0E078D13CA39E8B4E3561">Soreng, Robert J.</mods:namePart>
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<mods:affiliation id="753C2603CEBDC57E3E304D184A76455B">Department of Botany, National Museum of Natural History, Smithsonian Institution, Washington DC 20560, USA</mods:affiliation>
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<mods:nameIdentifier id="D0E686990D85C5091BEBD29714FB2F48" type="orcid">https://orcid.org/0000-0002-8358-4915</mods:nameIdentifier>
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<mods:typeOfResource id="392DD7222841AA0CF45738E51302F9C1">text</mods:typeOfResource>
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<mods:titleInfo id="15460F6EEC1B1955526AB64684CC424D">
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<mods:title id="6820A7D4A7470E840483E7AE8F538BE2">PhytoKeys</mods:title>
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</mods:titleInfo>
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<mods:part id="9E08F54F24E4B2718448F8FE0A31F765">
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<mods:date id="5696950660E1E6B2F28B61EEA50A6062">2020</mods:date>
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<mods:detail id="11D6CB52542BAA740A0610CE43D107F3" type="volume">
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<mods:number id="3F727ED93E850FFA3FD16EDB19685654">151</mods:number>
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</mods:detail>
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<mods:extent id="55DE70945084C10507A8D924DEAEECEA" unit="page">
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<mods:start id="F1834A0E772575DF01A00C85362CA6B4">107</mods:start>
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<mods:end id="411817B9F1BFB90D302D839B67BC3F2B">160</mods:end>
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</mods:extent>
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</mods:relatedItem>
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<mods:location id="B1B80923EA74CFE4968E0DBCE2E97A14">
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<mods:url id="E256DD5D8053A692C8EEF63861E0D0C9">http://dx.doi.org/10.3897/phytokeys.151.50538</mods:url>
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</mods:location>
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<mods:classification id="639D35EE91C62C4D08EBEB941A24DBF3">journal article</mods:classification>
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<mods:identifier id="AE6B27121D06A2B2824C766C236DEABE" type="DOI">http://dx.doi.org/10.3897/phytokeys.151.50538</mods:identifier>
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<mods:identifier id="0868C0353BB201951CFD382A20627B02" type="Pensoft-Pub">1314-2003-151-107</mods:identifier>
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<mods:identifier id="C368F3D895C0B6C322BD59D9400B8FF3" type="Pensoft-UUID">42D1EB2DF0545744B92BFCCCB395C45B</mods:identifier>
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</mods:mods>
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<treatment id="4E7CFCB1729C50408D8C94AC584BC6C6" ID-GBIF-Taxon="164730854" LSID="urn:lsid:plazi:treatment:4E7CFCB1729C50408D8C94AC584BC6C6" httpUri="http://treatment.plazi.org/id/4E7CFCB1729C50408D8C94AC584BC6C6" lastPageNumber="107" pageId="0" pageNumber="107" scope_family="Poaceae">
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<subSubSection id="A7C808B5395826DDDB6DA15FC1FBD4FC" pageId="0" pageNumber="107" type="nomenclature">
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<paragraph id="83AA29A5B577E5AC1F7FA36B779251CF" pageId="0" pageNumber="107">
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<taxonomicName id="10A6F0D405F3B69A58453E5FB0BDFFEA" ID-CoL="B9R3" LSID="4E7CFCB1-729C-5040-8D8C-94AC584BC6C6" authority="Phil. Anales Univ. Chile 94: 11. 1896" authorityName="Phil. Anales Univ. Chile 94: 11." authorityYear="1896" class="Liliopsida" family="Poaceae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis imberbis" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="imberbis">Agrostis cf. imberbis Phil. Anales Univ. Chile 94: 11. 1896</taxonomicName>
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<figureCitation id="1B9B52E93A9F896ECE531F29CB550FD7" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Agrostis cf. imberbis A whole plant B flag ligule and portion of inflorescence C floret, lateral view D portion of inflorescence with close-up of spikelets. Images of Cleef 6821 (US 2785719), courtesy of United States National Herbarium (US)." figureDoi="10.3897/phytokeys.151.50538.figure6" httpUri="https://binary.pensoft.net/fig/424844" pageId="0" pageNumber="107">Fig. 6</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C600CAC2E87BCD7A5589CC7790F89DEC" pageId="0" pageNumber="107" type="reference_group">
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<paragraph id="17774D7DFA77C74E2A86153E23AE8F3F" pageId="0" pageNumber="107">
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=
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<taxonomicName id="B4898BA6846A83C6545EBF50054A7D3E" class="Liliopsida" family="Poaceae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis stenophylla" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="stenophylla">Agrostis stenophylla</taxonomicName>
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Phil., Anales Univ. Chile 94: 10. 1896. Type: Chile.
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<normalizedToken id="4793B856899EFBD92357BC5C9E15B30A" originalValue="Bío-Bío">Bio-Bio</normalizedToken>
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:
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<normalizedToken id="5A497DAC0BA3291DA752F4A2D9BA7759" originalValue="Baños">Banos</normalizedToken>
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de
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<normalizedToken id="6177CD8B4C9DC8D5C99BA9DB97251AAB" originalValue="Chillán">Chillan</normalizedToken>
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, Jan. 1878, Philippi s.n. [
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<normalizedToken id="F0F659A4EA858CB4ECA9F9F3231E53A4" originalValue="“146”">"146"</normalizedToken>
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in Herb. R. A. Phillippi s.d.; "No. 18 (II)" on SGO000000043 isotype] (holotype: SGO-PHIL (SGO-PHIL146); isotypes: BAA (BAA00000030 [image!]), BM (BM000938541 [image!]), SGO (SGO000000043 [image!], SGO000000061 [image!], SGO000000062 [image!]), US (US00156498! fragm. ex SGO-PHIL146), W (W19160040646 [image!])).
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</paragraph>
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<paragraph id="F507487196B7FE296D2045CA174FA12C" pageId="0" pageNumber="107">
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=
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<taxonomicName id="6D017CC1AEBC56311F89CE90401836FC" class="Liliopsida" family="Poaceae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis scotantha" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="scotantha">Agrostis scotantha</taxonomicName>
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Phil., Anales Univ. Chile 94: 16. 1896. Type: Chile.
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<normalizedToken id="D7E90A43DBA091A3BF8E2C62E81F6308" originalValue="Araucanía">Araucania</normalizedToken>
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: La Cueva, Jan. 1887, C. Rahmer s.n. [
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<normalizedToken id="3BA775C98F4CFA130B13D68888DF53B2" originalValue="“183”">"183"</normalizedToken>
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in Herb. R. A. Phillippi s.d.] (holotype: SGO-PHIL (SGO-PHIL183); isotypes: BAA (BAA00000728 [image!] fragm. ex SGO-PHIL183, BAA00000729 [image!] fragm. ex B), SGO (SGO000000055 [image!], SGO000000056 [image!]), US (US00156493! fragm. ex SGO-PHIL183), W (W19160040640 [image!])).
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</paragraph>
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<paragraph id="6DEC966DC2DACBB0D3DAF34641F8C978" pageId="0" pageNumber="107">
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=
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<taxonomicName id="F627372A0CB7C472EAD2D6BAB70EBC6C" class="Liliopsida" family="Poaceae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis moyanoi subsp. var. var. plicatifolia" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="variety" species="moyanoi" subSpecies="var." variety="plicatifolia">Agrostis moyanoi Speg. var. plicatifolia</taxonomicName>
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Speg., Anales Mus. Nac. Buenos Aires 7: 189. 1902. Type: Argentina. Chubut: Corcovado, Oct. 1901, N. Illin 2550 (holotype: LP (LP001364 [image!]; isotypes: BAA (BAA00000023 [image!])).
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</paragraph>
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</subSubSection>
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<subSubSection id="245E0AD84896072C3CD2BFF1E6DAB90D" pageId="0" pageNumber="107" type="materials_examined">
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<paragraph id="8A0168AE51D8CCEE08FB17A00E7D8BB5" pageId="0" pageNumber="107">Type.</paragraph>
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<paragraph id="86B32F2718CB50BEA4E478A89CF9228C" pageId="0" pageNumber="107">
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Chile.
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<normalizedToken id="55C686919E44E6EC7CFA5EAFAADD9808" originalValue="Ñuble">Nuble</normalizedToken>
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: Valle de las Nieblas,
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<normalizedToken id="EC3803BE30B628A374A03603DD70D903" originalValue="Chillán">Chillan</normalizedToken>
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, Jan. 1877, sin col. s.n. [
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<normalizedToken id="99CEBC0D71DD250323ECB0C51776E349" originalValue="“185”">"185"</normalizedToken>
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in Herb. R. A. Phillippi s.d.] (holotype: SGO-PHIL (SGO-PHIL185); isotypes: BAA (BAA00001345 [image!]), BM (BM000938537 [image!]), SGO (SGO000000042 [image!], SGO000000044 [image!], US (US00156439! fragm. ex SGO-PHIL-185), W (W19160040633 [image!])).
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</paragraph>
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</subSubSection>
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<subSubSection id="24D4BCDE04E95B5FCA0B01E997BFF70E" pageId="0" pageNumber="107" type="description">
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<paragraph id="170F96B8B58231593CE3AF6F131EAE53" pageId="0" pageNumber="107">Description.</paragraph>
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<paragraph id="FBBF2F8B4610E2D35471EB5C5135662F" pageId="0" pageNumber="107">
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<emphasis id="B66FF682C7121E3482B7BFB7E8974389" bold="true" pageId="0" pageNumber="107">Perennial herbs</emphasis>
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, laxly to densely tufted, sometimes stooling, with laterally tending or ascending pseudostolons, or rhizomatous with rhizomes ascending or vertical.
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<emphasis id="0011B66EFB6254CD35BCEBF6DEAC8AD4" bold="true" pageId="0" pageNumber="107">Tillers</emphasis>
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intravaginal and extravaginal, with cataphylls usually present.
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<emphasis id="0EA6AB68C40EEE04ED12F04460050CB9" bold="true" pageId="0" pageNumber="107">Culms</emphasis>
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(7-)24-60(-65) cm tall, usually decumbent at their base or slightly creeping and rooting from the nodes, rarely completely erect, firm, with (0-)1-2 nodes exerted at flowering, smooth or scaberulous.
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<emphasis id="BD8F377607279AB42C0C97D7072D1A51" bold="true" pageId="0" pageNumber="107">Leaves</emphasis>
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usually mostly basal in first seasons growth, becoming mostly cauline as plant ages, glabrous, usually scaberulous with undeveloped scabers throughout, sometimes notably scabrous throughout;
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<emphasis id="EFABCA0F482597CA051EAADB70B78588" bold="true" pageId="0" pageNumber="107">ligules</emphasis>
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2.5-6.7(-10) mm long, of upper culm (2.8-)3-6.7(-10) mm long, usually scarious (at least in part), acute to acuminate, sometimes fimbriate, strongly decurrent with the sheath, abaxial surface scabrous;
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<emphasis id="FD0261BA9AC0D446571837B5318A6B8B" bold="true" pageId="0" pageNumber="107">blades</emphasis>
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2.8-9(-11) cm long, 0.5-1 mm wide in diameter as rolled or folded, convolute, involute, or conduplicate, rigid, abaxial surface smooth to scaberulous, adaxial surface lightly scabrous on the veins, apices usually naviculate-acute.
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<emphasis id="DDAA0A23135D7C31792352B722A9A844" bold="true" pageId="0" pageNumber="107">Panicles</emphasis>
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(3-)6-12.5(-20) cm long, (0.5-)2-6 cm wide, open to slightly contracted when young, usually ovoid, slightly to greatly exerted from the basal foliage, lateral branches without spikelets near their base and for a long distance, long and ascending or spreading to becoming divaricate at maturity but not held close to the central inflorescence axis, central axis and panicle branches scaberulous;
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<emphasis id="6C0DDC585A7A5F25FEC142815C0ADB5B" bold="true" pageId="0" pageNumber="107">pedicels</emphasis>
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(0.9-)1.5-3.5 mm long, often slightly longer than their spikelets or sometimes shorter, dilated or not at their apex, scaberulous.
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<emphasis id="88DE521692B206837137CEA32623D842" bold="true" pageId="0" pageNumber="107">Spikelets</emphasis>
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(2.3-)2.5-2.7(-3.5) mm long;
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<emphasis id="2346304C53411470581DCD43A045342D" bold="true" pageId="0" pageNumber="107">glumes</emphasis>
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equal or subequal, similar or lower glume slightly wider and longer than the upper, 1-veined, keels usually scabrous in the distal
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<normalizedToken id="6518B2EF038C5D25CDDA40507759998A" originalValue="½-¾">1/2-3/4</normalizedToken>
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, upper glume sometimes completely smooth, apices acute to acuminate;
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<emphasis id="2829E84A1321B230AEF61BD3D8016614" bold="true" pageId="0" pageNumber="107">floret</emphasis>
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usually 3/4 the length of the glumes or slightly longer;
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<emphasis id="238E12350334697BF216E62794149609" bold="true" pageId="0" pageNumber="107">calluses</emphasis>
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glabrescent, with very sparse short hairs;
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<emphasis id="EF748EB90009C1E941357E69239FCA94" bold="true" pageId="0" pageNumber="107">lemmas</emphasis>
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1.6-1.8(-2.5) mm long, glabrous, lightly scaberulous throughout or just in distal 1/2, sometimes with just undeveloped scabers as pustulate bases, 5-veined, apex obtuse or truncate, denticulate, muticous, mucronate, or sometimes with a short straight awn to 0.5 mm long, inserted in the middle or upper 1/3 of the lemma and not surpassing the glumes, weak and falling easily;
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<emphasis id="23B8F119F625608EB2770FC118B29BDD" bold="true" pageId="0" pageNumber="107">paleas</emphasis>
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usually c. 0.2 mm long, <
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<normalizedToken id="94C4CB6E77DE43B185202F74A72A9701" originalValue="¼">1/4</normalizedToken>
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the length of the lemma;
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<emphasis id="1ACF0E0CCDFE3554D4BF24B3F364F5BF" bold="true" pageId="0" pageNumber="107">rachilla</emphasis>
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absent;
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<emphasis id="476AEF10B6F376C58C96518268FBBB2A" bold="true" pageId="0" pageNumber="107">anthers</emphasis>
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1-1.2(-2) mm long.
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</paragraph>
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<caption id="9846A8EE16D1FBFA5A6D926077D01AFD" doi="10.3897/phytokeys.151.50538.figure6" httpUri="https://binary.pensoft.net/fig/424844" pageId="0" pageNumber="107" start="Figure 6" startId="F6">
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<paragraph id="475BF52172FACA48EB14521B20F44E63" pageId="0" pageNumber="107">
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<emphasis id="12F492118CEC16AE802E66CA16093FAA" bold="true" pageId="0" pageNumber="107">Figure 6.</emphasis>
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<taxonomicName id="5A5771D90B6981C8814432BAEF9B5F3C" class="Liliopsida" family="Poaceae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis imberbis" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="imberbis">Agrostis cf. imberbis</taxonomicName>
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<emphasis id="67B60654DCFA01F71C08AFF886BA2F03" bold="true" pageId="0" pageNumber="107">A</emphasis>
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whole plant
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<emphasis id="7FE196E6BCC5AF417E7B7991DC508B27" bold="true" pageId="0" pageNumber="107">B</emphasis>
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flag ligule and portion of inflorescence
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<emphasis id="08AE1B8E3F81D67CF20D6E3D706778FB" bold="true" pageId="0" pageNumber="107">C</emphasis>
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floret, lateral view
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<emphasis id="03EC48E335C0C2645E8662DA076E4900" bold="true" pageId="0" pageNumber="107">D</emphasis>
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portion of inflorescence with close-up of spikelets. Images of Cleef 6821 (US2785719), courtesy of United States National Herbarium (US).
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection id="7DE54607AE672F6FCF270CEA4ECDB928" pageId="0" pageNumber="107" type="distribution">
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<paragraph id="83172220C4BF4A41C9297E2C63FA5F98" pageId="0" pageNumber="107">Distribution and ecology.</paragraph>
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<paragraph id="72195EDDC12C5E08FFDB717B08FB02D0" pageId="0" pageNumber="107">
|
||
<taxonomicName id="C176605985A6D224E920960CAF97BC6E" class="Liliopsida" family="Poaceae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis imberbis" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="imberbis">Agrostis cf. imberbis</taxonomicName>
|
||
is a new record for Colombia and
|
||
<normalizedToken id="2FCB95BC44972B552C64AAA7B0A68F99" originalValue="páramo">paramo</normalizedToken>
|
||
vegetation in general, and is not found in the most recent checklist for Colombia (
|
||
<bibRefCitation id="F51061BADB60C75A5E28087C5FD821D6" author="Giraldo-Canas, D" editor="Bernal, R" journalOrPublisher="2). Instituto de Ciencias Naturales, Universidad Nacional de Colombia, Bogota D. C." pageId="0" pageNumber="107" pagination="2127 - 2205" refId="B10" refString="Giraldo-Canas, D, Londono, X, Clark, LG, 2016. Poaceae. In: Bernal, R, Gradstein, SR, Celis, M, Eds., Catalogo de plantas y liquenes de Colombia (Vol. 2). Instituto de Ciencias Naturales, Universidad Nacional de Colombia, Bogota D.C.: 2127 - 2205" title="Poaceae." volumeTitle="Catalogo de plantas y liquenes de Colombia (Vol." year="2016">
|
||
<normalizedToken id="4FFAD79FB62B5ABA75B2C4E30322D4A3" originalValue="Giraldo-Cañas">Giraldo-Canas</normalizedToken>
|
||
et al. 2016
|
||
</bibRefCitation>
|
||
), the
|
||
<normalizedToken id="EDAF831F82810060E31EB934EBA0A633" originalValue="páramo">paramo</normalizedToken>
|
||
checklist (
|
||
<bibRefCitation id="62104262A84DF0CF3EF3447E498EAD30" author="Luteyn, JL" journalOrPublisher="Memoirs of the New York Botanical Garden" pageId="0" pageNumber="107" pagination="1 - 278" refId="B20" refString="Luteyn, JL, 1999. Paramos, a checklist of plant diversity, geographical distribution, and botanical literature. Memoirs of the New York Botanical Garden 84: 1 - 278" title="Paramos, a checklist of plant diversity, geographical distribution, and botanical literature." volume="84" year="1999">Luteyn 1999</bibRefCitation>
|
||
) or checklists for Ecuador (
|
||
<bibRefCitation id="B0860FBA4722E86D188ED8825C95EBA5" author="Jorgensen, PM" journalOrPublisher="AAU Reports" pageId="0" pageNumber="107" pagination="1 - 443" refId="B18" refString="Jorgensen, PM, Ulloa-Ulloa, C, 1994. Seed plants of the high Andes of Ecuador: A checklist. AAU Reports 34: 1 - 443" title="Seed plants of the high Andes of Ecuador: A checklist." volume="34" year="1994">
|
||
<normalizedToken id="13D51226BDD4D62E56D6E03BC58A227A" originalValue="Jørgensen">Jorgensen</normalizedToken>
|
||
and Ulloa-Ulloa 1994
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="4E2C285EE58E3EA8D05B6996D3FEC056" author="Jorgensen, PM" journalOrPublisher="Monographs in Systematic Botany from the Missouri Botanical Garden" pageId="0" pageNumber="107" pagination="1 - 1181" refId="B17" refString="Jorgensen, PM, Leon-Yanez, S [Eds], 1999. Catalogue of the Vascular Plants of Ecuador. Monographs in Systematic Botany from the Missouri Botanical Garden 75: 1 - 1181" title="Catalogue of the Vascular Plants of Ecuador." volume="75" year="1999">
|
||
<normalizedToken id="7B79A8F01E60C19F94FBB041B8DEB5B6" originalValue="Jørgensen">Jorgensen</normalizedToken>
|
||
and
|
||
<normalizedToken id="6686F484ADABA2D13611A58AF0C9E69D" originalValue="León-Yánez">Leon-Yanez</normalizedToken>
|
||
1999
|
||
</bibRefCitation>
|
||
), Venezuela (
|
||
<bibRefCitation id="BB856202F541CBB49FC774AB3A6EB3F1" author="Hokche, O" journalOrPublisher="Fundacion Instituto Botanico de Venezuela, Caracas" pageId="0" pageNumber="107" refId="B16" refString="Hokche, O, Berry, PE, Huber, O [Eds], 2008. Nuevo Catalogo de la Flora Vascular de Venezuela. Fundacion Instituto Botanico de Venezuela, Caracas" title="Nuevo Catalogo de la Flora Vascular de Venezuela." year="2008">Hokche et al. 2008</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="96A2350D10AB620666EFC1FA395EDA35" author="Bono, G" editor="Bono, G" journalOrPublisher="Universita degle Studi di Firenze, Firenze" pageId="0" pageNumber="107" pagination="127 - 154" refId="B2" refString="Bono, G, 2010. Poaceae. In: Bono, G, Ed., Catalogo de la Flora y Vegetacion de los Valles de la Vertiente Occidental de los Andes de la Cordillera de Merida. Universita degle Studi di Firenze, Firenze: 127 - 154" title="Poaceae." volumeTitle="Catalogo de la Flora y Vegetacion de los Valles de la Vertiente Occidental de los Andes de la Cordillera de Merida." year="2010">Bono 2010</bibRefCitation>
|
||
), or Costa Rica (
|
||
<bibRefCitation id="55D78C7E4A8C03A666E51FD7BB61E9CA" author="Morales-Quiros, JF" editor="Hammel, BE" journalOrPublisher="Monographs in Systematic Botany from the Missouri Botanical Garden 93 (3)" pageId="0" pageNumber="107" pagination="598 - 821" refId="B22" refString="Morales-Quiros, JF, 2003. Poaceae. In: Hammel, BE, Grayum, MH, Herrera-Mora, C, Zamora-Villalobos, N, Eds., Manual of Plants of Costa Rica. Monographs in Systematic Botany from the Missouri Botanical Garden 93(3): 598 - 821" title="Poaceae." volumeTitle="Manual of Plants of Costa Rica." year="2003">
|
||
<normalizedToken id="3177759940A0D1E9ECDC2012D3EE3C51" originalValue="Morales-Quirós">Morales-Quiros</normalizedToken>
|
||
2003
|
||
</bibRefCitation>
|
||
) which host
|
||
<normalizedToken id="7C16CB8751C87F90AEF41ADBEDE4C439" originalValue="páramo">paramo</normalizedToken>
|
||
vegetation. The species was previously considered to be restricted to Argentina and Chile (
|
||
<bibRefCitation id="3CF1664A72505B9FC567E9D36772E142" author="Rugolo de Agrasar, ZE" editor="Zuloaga, FO" journalOrPublisher="Poaceae: Pooideae (Vol. 3 Tomo 2). IBODA-IMBIV, CONICET, Buenos Aires, Argentina" pageId="0" pageNumber="107" pagination="102 - 125" refId="B30" refString="Rugolo de Agrasar, ZE, 2012. Agrostis L. In: Zuloaga, FO, Rugolo de Agrasar, ZE, Anton, AM, Eds., Flora Argentina. Poaceae: Pooideae (Vol. 3 Tomo 2). IBODA-IMBIV, CONICET, Buenos Aires, Argentina: 102 - 125" title="Agrostis L." volumeTitle="Flora Argentina." year="2012">
|
||
<normalizedToken id="F9979E4523584A73436F2076E7A76F5E" originalValue="Rúgolo">Rugolo</normalizedToken>
|
||
de Agrasar 2012
|
||
</bibRefCitation>
|
||
), but has also been recorded for Bolivia (
|
||
<bibRefCitation id="17B70BA3713F69FEFFD7F03279845565" author="Jorgensen, PM" journalOrPublisher="AAU Reports" pageId="0" pageNumber="107" refId="B19" refString="Jorgensen, PM, Nee, MH, Beck, SG, 2014. . Catalogo de las plantas vasculares de Bolivia, Monographs in systematic botany from the Missouri Botanical Garden 127(1-2): 1-1744. Missouri Botanical Garden Press, St. Louis." title=". Catalogo de las plantas vasculares de Bolivia, Monographs in systematic botany from the Missouri Botanical Garden 127 (1 - 2): 1 - 1744. Missouri Botanical Garden Press, St. Louis." year="2014">
|
||
<normalizedToken id="F0A6F98960E8E112F760077D23011826" originalValue="Jørgensen">Jorgensen</normalizedToken>
|
||
et al. 2014
|
||
</bibRefCitation>
|
||
) and Peru (
|
||
<bibRefCitation id="0CD26BC986F1E6167FC30AF082E62E14" author="Davidse, G" editor="Brako, L" journalOrPublisher="Monographs in Systematic Botany from the Missouri Botanical Garden" pageId="0" pageNumber="107" pagination="926 - 977" refId="B5" refString="Davidse, G, Brako, L, Tovar, O, 1993. Poaceae. In: Brako, L, Zarucchi, J, Eds., Catalogue of the Flowering Plants and Gymnosperms of Peru. Monographs in Systematic Botany from the Missouri Botanical Garden 45: 926 - 977" title="Poaceae." volume="45" volumeTitle="Catalogue of the Flowering Plants and Gymnosperms of Peru." year="1993">Davidse et al. 1993</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="6C6D0F52D5ADD814D75E02FBAB2A286C" author="Soreng, RJ" journalOrPublisher="Contributions from the United States National Herbarium" pageId="0" pageNumber="107" publicationUrl="http://www.tropicos.org/project/" refId="B34" refString="Soreng, RJ, Davidse, G, Peterson, PM, Zuloaga, FO, Judziewicz, EJ, Filgueiras, TS, Morrone, O, 2003 and onwards. . Catalogue of New World Grasses. http://www.tropicos.org/project/" title=". Catalogue of New World Grasses." url="http://www.tropicos.org/project/" year="2003 and onwards">Soreng et al. 2003 and onwards</bibRefCitation>
|
||
; W3TROPICOS-Peru Checklist 2020;
|
||
<normalizedToken id="3431467DD2FB744BBC15D10B4B25827F" originalValue="Óscar">Oscar</normalizedToken>
|
||
Tovar unpubl. data).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B6152D1861815004C8A3834DA0A5EDFA" pageId="0" pageNumber="107" type="materials_examined">
|
||
<paragraph id="31B6A52E5F4DEDD41E0E730585B4F21A" pageId="0" pageNumber="107">Additional specimens examined.</paragraph>
|
||
<paragraph id="5FC5098E7313230E882670C6CE5D8FE2" pageId="0" pageNumber="107">
|
||
<emphasis id="34E46768472DB437274426DF3001142A" bold="true" pageId="0" pageNumber="107">Colombia</emphasis>
|
||
.
|
||
<emphasis id="8D943733B85F05172549D7087B227892" bold="true" pageId="0" pageNumber="107">
|
||
<normalizedToken id="E60594B218B8145CBF4C7A85730F11DC" originalValue="Boyacá">Boyaca</normalizedToken>
|
||
</emphasis>
|
||
: Munic.
|
||
<normalizedToken id="A4968E3DC674CF63A1CAB3342859419B" originalValue="Belén">Belen</normalizedToken>
|
||
,
|
||
<normalizedToken id="DD784620B9FA8D81CD0D2D7E379325B0" originalValue="Páramo">Paramo</normalizedToken>
|
||
de La Rusia, near
|
||
<normalizedToken id="DE630B79EFA134300D0BB2C8BD535AAA" originalValue="Páramo">Paramo</normalizedToken>
|
||
El Consuelo, unprotected private land, somewhat disturbed
|
||
<normalizedToken id="E6D48F09A59D8F717F24384F8596D570" originalValue="páramo">paramo</normalizedToken>
|
||
grazed by horses and rodents, dominated by
|
||
<taxonomicName id="7086E5938E26807FAAD254A791367B37" authorityName="Cuatrec" authorityYear="1973" class="Magnoliopsida" family="Asteraceae" genus="Espeletia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Espeletia boyacensis" order="Asterales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="boyacensis">
|
||
<emphasis id="86CEB4FDACC52863C735CBC27587959D" italics="true" pageId="0" pageNumber="107">Espeletia boyacensis</emphasis>
|
||
</taxonomicName>
|
||
[Cuatrec.],
|
||
<geoCoordinate id="8EC50D7F6CEFF8AC79FD80BE792F08EC" degrees="6.02146" direction="north" orientation="latitude" precision="1" value="6.02146">6.02146N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="07C1EC79EC19B5EB191072C414022881" degrees="72.5718" direction="west" orientation="longitude" precision="5" value="-72.5718">72.5718W</geoCoordinate>
|
||
, 3832 m alt., 22 Nov. 2017, M. Vorontsova 2228 (US); Munic. Chiscas,
|
||
<normalizedToken id="EC97F4192534DCE01BCCF274A919F454" originalValue="Páramo">Paramo</normalizedToken>
|
||
el Penon, Chiscas, borde de bosque de
|
||
<taxonomicName id="95B2B9BA7720B6C0246774D867812CAF" class="Diplopoda" family="Platyrhacidae" genus="Polylepis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polylepis" order="Polydesmida" pageId="0" pageNumber="107" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="575FF73211068E7DB2FD280844DE9CC1" italics="true" pageId="0" pageNumber="107">Polylepis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<geoCoordinate id="F725B1A42EECCC792C0D3D7AACF21BFA" degrees="6.59582" direction="north" orientation="latitude" precision="1" value="6.59582">6.59582N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="0DDD98C9097B6C88850DE8650DB7420D" degrees="72.44284" direction="west" orientation="longitude" precision="1" value="-72.44284">72.44284W</geoCoordinate>
|
||
, 3771 m alt., 5 Mar. 2018, S.P. Sylvester 3159 (K, SI, US); Munic. Chiscas,
|
||
<normalizedToken id="FDD5D8D01BA799D59C8FCB59CEBD0C4A" originalValue="Páramo">Paramo</normalizedToken>
|
||
de Chacaritas, pajonal frailejonal, cercano al cerro de Chacaritas, con fuerte grado de inclinacion,
|
||
<geoCoordinate id="6046439D91520237456B0289B3F6A20F" degrees="6.63108" direction="north" orientation="latitude" precision="1" value="6.63108">6.63108N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="F935056265BD95E38883C5111BB98223" degrees="72.39815" direction="west" orientation="longitude" precision="1" value="-72.39815">72.39815W</geoCoordinate>
|
||
, 4082 m alt., 5 Mar. 2018, S.P. Sylvester 3144 (FMB, K, SI, UPTC, US); Munic. Duitama,
|
||
<normalizedToken id="2B6FFD6C06FD631E4469B0CA75356E69" originalValue="Páramo">Paramo</normalizedToken>
|
||
de La Rusia, NW-N de Duitama, [
|
||
<geoCoordinate id="98CC6AF21B72D78A4C4302C899BC2720" degrees="5.9281" direction="north" orientation="latitude" precision="5" value="5.9281">5.9281N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="423899D461FAE8F919C83FA42001139D" degrees="73.0936" direction="west" orientation="longitude" precision="5" value="-73.0936">73.0936W</geoCoordinate>
|
||
], 3600 m alt., aislada, vertiente seca con
|
||
<taxonomicName id="19C13F499AEA4EBF1351759B9F1B980F" class="Liliopsida" family="Poaceae" genus="Calamagrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Calamagrostis effusa" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="effusa">
|
||
<emphasis id="40CAEB6A1F4D2C8DFA8161630A9C6282" italics="true" pageId="0" pageNumber="107">Calamagrostis effusa</emphasis>
|
||
</taxonomicName>
|
||
[
|
||
<taxonomicName id="33811BA81D9AFC7C29513B79FC92E8ED" genus="Paramochloa" lsidName="Paramochloa effusa" pageId="0" pageNumber="107" rank="species" species="effusa">
|
||
<emphasis id="CBEBF85F90FD54B317B63024323D1544" italics="true" pageId="0" pageNumber="107">Paramochloa effusa</emphasis>
|
||
</taxonomicName>
|
||
] predominante, asociada con
|
||
<taxonomicName id="71EB19169FA1CD963811A856044FE0F3" authorityName="Cuatrec" authorityYear="1973" class="Magnoliopsida" family="Asteraceae" genus="Espeletia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Espeletia boyacensis" order="Asterales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="boyacensis">
|
||
<emphasis id="BF80AE77DE395E127035FE2A25436BD6" italics="true" pageId="0" pageNumber="107">Espeletia boyacensis</emphasis>
|
||
</taxonomicName>
|
||
y
|
||
<taxonomicName id="FB34019885B3EFD1A1DE74D34E30B6C8" authorityName="Ruiz & Pavon" authorityYear="1798" class="Magnoliopsida" family="Rosaceae" genus="Acaena" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Acaena cylindristachya" order="Rosales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="cylindristachya">
|
||
<emphasis id="D18737B4BF3AFE5D7A21B08298FA17D5" italics="true" pageId="0" pageNumber="107">Acaena cylindristachya</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<normalizedToken id="AEBBF511DD1249FF1CA1E4C02F7C8B0D" originalValue="gramínea">graminea</normalizedToken>
|
||
muy abundante, 7 Dec. 1972, A.M. Cleef 6821 (US2785719); Munic. Duitama,
|
||
<normalizedToken id="8E003606BD8BFF738C9DDB0DCCDA6929" originalValue="Páramo">Paramo</normalizedToken>
|
||
de La Rusia,
|
||
<normalizedToken id="5B2AF29B31869BF3CF5935F0FB7621BA" originalValue="vía">via</normalizedToken>
|
||
que conduce a vereda de Avendanos,
|
||
<normalizedToken id="C8A15F19363F972FAD7E22AC12B6705C" originalValue="páramo">paramo</normalizedToken>
|
||
semi-perturbado, pastado por cabras y quemado regularmente hasta hace 1
|
||
<normalizedToken id="B84C6C6411F4B87E0F3FAE5558F9B36D" originalValue="año">ano</normalizedToken>
|
||
,
|
||
<geoCoordinate id="CBA10C48237DF7B84469AAFE556CACB7" degrees="5.95011" direction="north" orientation="latitude" precision="1" value="5.95011">5.95011N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="FBE3437A91D91CFF31EDBE36FF75A46A" degrees="73.09097" direction="west" orientation="longitude" precision="1" value="-73.09097">73.09097W</geoCoordinate>
|
||
, 3795 m alt., 1 Oct. 2017, S.P. Sylvester 3014 (COL, FMB, K, SI, US);
|
||
<geoCoordinate id="E1DC443ACE14C71802DDF0D1DB24EA92" degrees="5.9324667" direction="north" orientation="latitude" precision="1" value="5.9324665">5.9324667N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="F96B15EC99AA24FB1E1E4A57A8E52EFB" degrees="73.0798" direction="west" orientation="longitude" precision="5" value="-73.0798">73.0798W</geoCoordinate>
|
||
, 3726 m alt., 4 Oct. 2017, S.P. Sylvester 3029 (FMB, US); S.P. Sylvester 3031 (K, US); [Munic. El Cocuy and
|
||
<normalizedToken id="190A8A42BCBB65F56D7CAA754660E74B" originalValue="Soatá">Soata</normalizedToken>
|
||
,] between
|
||
<normalizedToken id="5F3FB76B7920BD67E5CA6042FCAA4197" originalValue="Soatá">Soata</normalizedToken>
|
||
and Cocuy,
|
||
<normalizedToken id="A54300C812B84FCAB967A0756A72127F" originalValue="Páramo">Paramo</normalizedToken>
|
||
del Alto del Escobal, [
|
||
<geoCoordinate id="6266069F4B35AA149749997E5CED300E" degrees="6.3514" direction="north" orientation="latitude" precision="5" value="6.3514">6.3514N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="E6242A61126CC570237CBF721C342AFA" degrees="72.5483" direction="west" orientation="longitude" precision="5" value="-72.5483">72.5483W</geoCoordinate>
|
||
], 3750 m alt., 8 Oct 1938, J. Cuatrecasas 1232 (US1772922).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="6531027E9C226465FB9B5FFAA69A8477" pageId="0" pageNumber="107" type="notes">
|
||
<paragraph id="9091805BE63B732D07BE4D9A4B118888" pageId="0" pageNumber="107">Notes.</paragraph>
|
||
<paragraph id="4805C2BCF18588ABE83A9EF10FD53D4E" pageId="0" pageNumber="107">
|
||
Specimens studied from
|
||
<normalizedToken id="88BF94961B022CAE4B3FB51056427B8B" originalValue="Boyacá">Boyaca</normalizedToken>
|
||
match most characteristics of
|
||
<taxonomicName id="046AFCE29322E6D0DFC17E1EDF944C64" lsidName="A. imberbis" pageId="0" pageNumber="107" rank="species" species="imberbis">
|
||
<emphasis id="36F5C4735C30A0DC974DA0F6EFD8CE0F" italics="true" pageId="0" pageNumber="107">A. imberbis</emphasis>
|
||
</taxonomicName>
|
||
apart from sometimes the leaf blade abaxial surfaces being scaberulous, albeit with silica
|
||
<normalizedToken id="AA53B94D673D28CA45735993DC405C5C" originalValue="‘pustules’">'pustules'</normalizedToken>
|
||
present throughout that sometimes developed into short hooks, and spikelets sometimes being shorter, to 2.3 mm long.
|
||
<bibRefCitation id="C89C30FB75DBA778CA04730B898CC385" author="Rugolo de Agrasar, ZE" editor="Zuloaga, FO" journalOrPublisher="Poaceae: Pooideae (Vol. 3 Tomo 2). IBODA-IMBIV, CONICET, Buenos Aires, Argentina" pageId="0" pageNumber="107" pagination="102 - 125" refId="B30" refString="Rugolo de Agrasar, ZE, 2012. Agrostis L. In: Zuloaga, FO, Rugolo de Agrasar, ZE, Anton, AM, Eds., Flora Argentina. Poaceae: Pooideae (Vol. 3 Tomo 2). IBODA-IMBIV, CONICET, Buenos Aires, Argentina: 102 - 125" title="Agrostis L." volumeTitle="Flora Argentina." year="2012">
|
||
<normalizedToken id="4D144B7A0414EFE16B202574BF838F63" originalValue="Rúgolo">Rugolo</normalizedToken>
|
||
de Agrasar (2012)
|
||
</bibRefCitation>
|
||
mentions leaf blades can be smooth in
|
||
<taxonomicName id="6C2595E729C02F20058EEE55910E36A1" lsidName="A. imberbis" pageId="0" pageNumber="107" rank="species" species="imberbis">
|
||
<emphasis id="AD78A39B7BC743EC0A4311DF918F463A" italics="true" pageId="0" pageNumber="107">A. imberbis</emphasis>
|
||
</taxonomicName>
|
||
under exceptional circumstances. Characters that are diagnostic for the species, such as narrow, rolled (convolute or involute) or conduplicate, rigid blades, long acuminate scarious ligules, usually large spikelets> 2.5 mm long, and glabrescent calluses, were all present on the specimens studied from
|
||
<normalizedToken id="F3B0E0F7CE3E7F60BF48C3DC4E39318A" originalValue="Boyacá">Boyaca</normalizedToken>
|
||
. Given its disjunct distribution and slight differences in morphology, coupled with knowledge that other widespread species such as
|
||
<taxonomicName id="75F8725DC31FCE993499A2D0EFA6BBAE" lsidName="A. perennans" pageId="0" pageNumber="107" rank="species" species="perennans">
|
||
<emphasis id="907C6A4CAB47336A4B1F6007F8D772D6" italics="true" pageId="0" pageNumber="107">A. perennans</emphasis>
|
||
</taxonomicName>
|
||
are actually numerous evolutionarily distinct taxa (Konstantin Romaschenko pers. communication), we refer to this species with
|
||
<normalizedToken id="014C5322E1801C8B1619EAFF7E443937" originalValue="‘cf.’">'cf.'</normalizedToken>
|
||
to highlight that it needs to be checked in a molecular framework.
|
||
</paragraph>
|
||
<paragraph id="3D83CAE48C017C4446074FBD8CDD7D23" pageId="0" pageNumber="107">The species has a variable level of panicle congestion depending on the stage of maturity.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="5CB0E89A0DF330ED6AAB7DBC884EDAB3" pageId="0" pageNumber="107" type="similar species">
|
||
<paragraph id="3AB51C6E1B4C7C48257321ED4792EFD8" pageId="0" pageNumber="107">Similar species.</paragraph>
|
||
<paragraph id="249E9FA7A33A571AEEAB76E82CA6D358" pageId="0" pageNumber="107">
|
||
<taxonomicName id="4A1FEEAF0325A980F6518E726BEB27E2" class="Liliopsida" family="Poaceae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis subrepens" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="subrepens">
|
||
<emphasis id="491A83C7D16117F68E0A214C503F96F1" italics="true" pageId="0" pageNumber="107">Agrostis subrepens</emphasis>
|
||
</taxonomicName>
|
||
, a species described from Mexico and whose presence in Colombia is uncertain (see notes on
|
||
<taxonomicName id="5A32529E8BAA194CAE803C373B39E9E5" lsidName="A. subrepens" pageId="0" pageNumber="107" rank="species" species="subrepens">
|
||
<emphasis id="A1DD32FD44C27ABC825151EF49C8CACB" italics="true" pageId="0" pageNumber="107">A. subrepens</emphasis>
|
||
</taxonomicName>
|
||
under 'Excluded
|
||
<normalizedToken id="8AACEDBEF336E6ABCA6D5FDFAAA64DA6" originalValue="species’">species'</normalizedToken>
|
||
at the end of the taxonomic treatment), and
|
||
<taxonomicName id="170CCE53AB4D80C76BCA2C9CA66E42D8" lsidName="A. imberbis" pageId="0" pageNumber="107" rank="species" species="imberbis">A. cf. imberbis</taxonomicName>
|
||
share similar morphologies. This includes the habit being decumbent and stooling, appearing rhizomatous or pseudostoloniferous, blades being convolute and rigid, and panicles usually large and open with spikelets that have a similar shape and size with lemmas lacking notable awns, ligules being strongly decurrent with the sheath and usually scarious (at least in part), and anthers> 1 mm long. While culm and panicle size are generally larger and anthers are often shorter in
|
||
<taxonomicName id="0676154864A38A30D70C50CEF988A92E" lsidName="A. subrepens" pageId="0" pageNumber="107" rank="species" species="subrepens">
|
||
<emphasis id="901B788D44E0807B5BD987C9045A04B0" italics="true" pageId="0" pageNumber="107">A. subrepens</emphasis>
|
||
</taxonomicName>
|
||
, some overlap occurs with
|
||
<taxonomicName id="6777E7805580D867F5E6DF1488796227" lsidName="A. imberbis" pageId="0" pageNumber="107" rank="species" species="imberbis">
|
||
<emphasis id="7545338E63EC01A64C11303C66E7A238" italics="true" pageId="0" pageNumber="107">A. imberbis</emphasis>
|
||
</taxonomicName>
|
||
, with ligule shape and size seeming to be the only solid character to differentiate them.
|
||
<taxonomicName id="975D165BB7C7C609C7259A67F0808095" class="Liliopsida" family="Poaceae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis subrepens" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="subrepens">
|
||
<emphasis id="C91BEF7F9E5699BDFEE080D253A55F7B" italics="true" pageId="0" pageNumber="107">Agrostis subrepens</emphasis>
|
||
</taxonomicName>
|
||
has shorter (<2.2 mm long), obtuse upper culm ligules while
|
||
<taxonomicName id="731B76FE175E6D122141A74998584D85" lsidName="A. imberbis" pageId="0" pageNumber="107" rank="species" species="imberbis">A. cf. imberbis</taxonomicName>
|
||
has acute to acuminate upper culm ligules (2.8-)3-6.7(-10) mm long.
|
||
</paragraph>
|
||
<paragraph id="3B40B842627CD5122D74160C39FFDA3C" pageId="0" pageNumber="107">
|
||
<taxonomicName id="13A7FA885743DADA05B29BC928787DB1" class="Liliopsida" family="Poaceae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis perennans" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="perennans">
|
||
<emphasis id="01145D44D56B8F05CCB4C10903368E73" italics="true" pageId="0" pageNumber="107">Agrostis perennans</emphasis>
|
||
</taxonomicName>
|
||
s.l. can sometimes have involute and rigid basal blades, but the upper culm blades are flat and lax, unlike those of
|
||
<taxonomicName id="D76BAEF1DCF6752DD699F2AC5CCD1F92" lsidName="A. imberbis" pageId="0" pageNumber="107" rank="species" species="imberbis">A. cf. imberbis</taxonomicName>
|
||
, which are rigid and convolute, involute, or strongly conduplicate throughout. The ligules in
|
||
<taxonomicName id="716E78CBD859CB7DCD75E0276300D9AC" lsidName="A. perennans" pageId="0" pageNumber="107" rank="species" species="perennans">
|
||
<emphasis id="605AD059538C46506EF48718E2CC7AB4" italics="true" pageId="0" pageNumber="107">A. perennans</emphasis>
|
||
</taxonomicName>
|
||
s.l. are also usually shorter, with ligules of basal leaves and tillers 0.5-2.5 mm long, while those of
|
||
<taxonomicName id="65BBA8A750A35D0AE7F9B68EA806F17D" lsidName="A. imberbis" pageId="0" pageNumber="107" rank="species" species="imberbis">A. cf. imberbis</taxonomicName>
|
||
are> 2.5 mm long. Ligule apices of
|
||
<taxonomicName id="3E37A55F02B7E025A892C874BD18E515" lsidName="A. perennans" pageId="0" pageNumber="107" rank="species" species="perennans">
|
||
<emphasis id="A4BE321E62124B527CB85A9A66B48A13" italics="true" pageId="0" pageNumber="107">A. perennans</emphasis>
|
||
</taxonomicName>
|
||
s.l. are also truncate or obtuse-triangular, unlike the acute or long acuminate ligules of
|
||
<taxonomicName id="7EB36F834635A6EDFD1090237ABBD937" lsidName="A. imberbis" pageId="0" pageNumber="107" rank="species" species="imberbis">A. cf. imberbis</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="7ADF5E825F0F9AF05AA4267224ECF15E" pageId="0" pageNumber="107">
|
||
<taxonomicName id="7BDA76CADAC2A36EC7E2B3D26784835B" class="Liliopsida" family="Poaceae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis vinealis" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="vinealis">
|
||
<emphasis id="8E6EE39DAFD1D80C689728D7A7741906" italics="true" pageId="0" pageNumber="107">Agrostis vinealis</emphasis>
|
||
</taxonomicName>
|
||
Schreb., a species not recorded for Colombia, but found introduced and naturalized in Argentina and Chile (
|
||
<bibRefCitation id="0C40D87DFF706FBA5878500F85170142" author="Soreng, RJ" editor="Soreng, RJ" journalOrPublisher="Contributions from the United States National Herbarium" pageId="0" pageNumber="107" pagination="42 - 89" refId="B33" refString="Soreng, RJ, Peterson, PM, 2003. Agrostis. In: Soreng, RJ, Peterson, PM, Davidse, G, Judziewicz, EJ, Zuloaga, FO, Filguieras, TS, Morrone, O, Eds., Catalogue of New World Grasses (Poaceae): IV. Subfamily Pooideae. Contributions from the United States National Herbarium 48: 42 - 89" title="Agrostis." volume="48" volumeTitle="Catalogue of New World Grasses (Poaceae): IV. Subfamily Pooideae." year="2003">Soreng and Peterson 2003</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="19A67F0D44F3159024842A38CBC72EC3" author="Rugolo de Agrasar, ZE" editor="Zuloaga, FO" journalOrPublisher="Poaceae: Pooideae (Vol. 3 Tomo 2). IBODA-IMBIV, CONICET, Buenos Aires, Argentina" pageId="0" pageNumber="107" pagination="102 - 125" refId="B30" refString="Rugolo de Agrasar, ZE, 2012. Agrostis L. In: Zuloaga, FO, Rugolo de Agrasar, ZE, Anton, AM, Eds., Flora Argentina. Poaceae: Pooideae (Vol. 3 Tomo 2). IBODA-IMBIV, CONICET, Buenos Aires, Argentina: 102 - 125" title="Agrostis L." volumeTitle="Flora Argentina." year="2012">
|
||
<normalizedToken id="1B616A0034917A23BD785697BE92B548" originalValue="Rúgolo">Rugolo</normalizedToken>
|
||
de Agrasar 2012
|
||
</bibRefCitation>
|
||
), bears similarities in terms of overall habit, being notably rhizomatous and having mainly basal leaves and an exerted open panicle that becomes congested after flowering, as well as other similarities such as the palea being reduced and awns sometimes absent (but usually with a persistent, geniculate and twisted, awn to 4 mm long, inserted near the base of the lemma).
|
||
<taxonomicName id="76AEC81EAB00AECE917363A24A4268B3" class="Liliopsida" family="Poaceae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis vinealis" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="vinealis">
|
||
<emphasis id="96D631EC918AA21D5D618A33216814B2" italics="true" pageId="0" pageNumber="107">Agrostis vinealis</emphasis>
|
||
</taxonomicName>
|
||
usually has leaf blades that are flat towards the base, although sometimes these are involute and setaceous, and scabrous throughout or at least on the adaxial surface. It can also be differentiated by the ligules being often shorter, of the tillers 1-2.5 mm long, of the upper culms 1-4(-5) mm long, with apices rounded or bluntly pointed (vs. of the tillers> 2.5 mm long, of the upper culms (2.8-)3-6.7(-10) mm long, with apices acute to acuminate in
|
||
<taxonomicName id="0BBD51F3E33522B186A5A347437B3975" lsidName="A. imberbis" pageId="0" pageNumber="107" rank="species" species="imberbis">A. cf. imberbis</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |