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<document ID-CLB-Dataset="23665" ID-DOI="http://dx.doi.org/10.3897/alpento.4.49492" ID-GBIF-Dataset="9d8e8024-8c19-4c26-881e-c9ded964a034" ID-Pensoft-Pub="2535-0889-4-1" ID-Pensoft-UUID="E065703019F55C94A3B11E80CC4DBDEA" ID-Zenodo-Dep="3710599" ID-ZooBank="57780613D73C423BA3E0C316DF1B7478" ModsDocID="2535-0889-4-1" checkinTime="1583514546079" checkinUser="pensoft" docAuthor="Baechli, Gerhard &amp; Vilela, Carlos R." docDate="2020" docId="286CDA892A56528CB8AEA6D0FD827FEC" docLanguage="en" docName="AlpEntomol 4: 1-20" docOrigin="Alpine Entomology 4" docPubDate="2020-03-06" docSource="http://dx.doi.org/10.3897/alpento.4.49492" docTitle="Paracacoxenus kaszabi Okada 1973" docType="treatment" docVersion="5" id="E065703019F55C94A3B11E80CC4DBDEA" lastPageNumber="1" masterDocId="E065703019F55C94A3B11E80CC4DBDEA" masterDocTitle="On the identity of Paracacoxenus kaszabi Okada, with the formal description of a new closely related species (Diptera, Drosophilidae)" masterLastPageNumber="20" masterPageNumber="1" pageNumber="1" updateTime="1705951339532" updateUser="ExternalLinkService">
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<mods:title id="E04ABD6B1A6A827061A61C1902F6E8EB">On the identity of Paracacoxenus kaszabi Okada, with the formal description of a new closely related species (Diptera, Drosophilidae)</mods:title>
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<mods:namePart id="986B0844E62C6B8B01E7F229ED210166">Baechli, Gerhard</mods:namePart>
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<mods:affiliation id="0FFE91CB33B056D1B8DF42AF4E70B92F">Zuerich-Irchel, Institut fuer Evolutionsbiologie und Umweltwissenschaften, Zuerich, Switzerland</mods:affiliation>
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<mods:namePart id="F8BB23A66695AE9F36A7381B57EDE9D0">Vilela, Carlos R.</mods:namePart>
<mods:affiliation id="7B29FF4BD26A4F5714E2019123F326EF">Universidade de Sao Paulo, Instituto de Biociencias, Departamento de Genetica e Biologia Evolutiva, Sao Paulo, SP, Brazil</mods:affiliation>
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<treatment id="286CDA892A56528CB8AEA6D0FD827FEC" ID-GBIF-Taxon="162463420" LSID="urn:lsid:plazi:treatment:286CDA892A56528CB8AEA6D0FD827FEC" httpUri="http://treatment.plazi.org/id/286CDA892A56528CB8AEA6D0FD827FEC" lastPageNumber="1" pageId="0" pageNumber="1">
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<taxonomicName id="F188C696113C714A80341C154DD68090" ID-CoL="8D65S" LSID="286CDA89-2A56-528C-B8AE-A6D0FD827FEC" authority="Okada, 1973" authorityName="Okada" authorityYear="1973" class="Insecta" family="Drosophilidae" genus="Paracacoxenus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paracacoxenus kaszabi" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="kaszabi">Paracacoxenus kaszabi Okada, 1973</taxonomicName>
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<figureCitation id="7C498DF66FC8E8DA9022AD107233DF65" captionStart="Figures 13" captionStartId="F1" captionText="Figures 1 - 3. Paracacoxenus kaszabi Okada, ♂ holotype, Central aimak, Ulan-Baator, Nucht im Bogdo ul, Mongolia (HNHM) 1 Labels (mislabelled as a female) and slide with wing and terminalia 2 Photomicrograph of terminalia, dorsal view 3 Right wing, dorsal view, distally optically cut. Photos: Z. Soltesz." figureDoi="10.3897/alpento.4.49492.figures1-3" httpUri="https://binary.pensoft.net/fig/387175" pageId="0" pageNumber="1">Figs 1</figureCitation>
<figureCitation id="45FBBF80602B4475077CE2BE5D0682A5" captionStart="Figures 21, 22" captionStartId="F8" captionText="Figures 21, 22. Paracacoxenus kaszabi Okada, ♂ paratype, Central aimak, Ulan-Baator, Nucht im Bogdo ul, Mongolia (HNHM) 21 Three-branched inner paraphysis, outer paraphysis and aedeagal apodeme, anterior (dorsal) view 22 Three-branched inner paraphysis, outer paraphysis and aedeagal apodeme, oblique posterior view. Abbreviations: a = proximal branch; b = mediodorsal branch; c = distal branch. Scale bar: 0.1 mm." figureDoi="10.3897/alpento.4.49492.figures21-22" httpUri="https://binary.pensoft.net/fig/387182" pageId="0" pageNumber="1">-22</figureCitation>
</paragraph>
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<paragraph id="D78419A5B409F925D357C41CC32A36E0" pageId="0" pageNumber="1">
<taxonomicName id="066C0DF16F86CE6078348F3520AF8944" authorityName="Okada" authorityYear="1973" class="Insecta" family="Drosophilidae" genus="Paracacoxenus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paracacoxenus kaszabi" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="kaszabi">Paracacoxenus kaszabi</taxonomicName>
Okada:
<bibRefCitation id="3D5438191432237D24DE1C3DD0A347AD" author="Okada, T" journalOrPublisher="Annales Historico-Naturales Musei Nationalis Hungarici" pageId="0" pageNumber="1" pagination="271 - 279" refId="B22" refString="Okada, T, 1973. Drosophilidae and Diastatidae from Mongolia (Diptera). Annales Historico-Naturales Musei Nationalis Hungarici 65: 271 - 279" title="Drosophilidae and Diastatidae from Mongolia (Diptera)." volume="65" year="1973">Okada (1973</bibRefCitation>
: 271) [description, figures];
<bibRefCitation id="4DBF6600C53AA2C742094FB0D2F3AB7F" author="Maca, J" journalOrPublisher="Acta Universitatis Carolinae" pageId="0" pageNumber="1" pagination="337 - 342" refId="B14" refString="Maca, J, 1980. Czechoslovak Drosophilidae (Diptera) with micropubescent arista. Acta Universitatis Carolinae 1977: 337 - 342" title="Czechoslovak Drosophilidae (Diptera) with micropubescent arista." volume="1977" year="1980">
<normalizedToken id="24971F58D919687F22AF9E650A57269D" originalValue="Máca">Maca</normalizedToken>
(1980
</bibRefCitation>
: 338) [description, figures, distribution, misidentification, see below];
<bibRefCitation id="FE417D580C82193E5DE01FA3CD724ECB" author="Sidorenko, VS" editor="Sidorenko, VS" journalOrPublisher="Tom VI. Dvukriliye i blokhi. Tshast 2. [Key to the insects of Russian Far East, Vol. VI. Diptera, Siphonaptera. Part 2]. Dalnauka, Vladivostok" pageId="0" pageNumber="1" pagination="211 - 241" refId="B23" refString="Sidorenko, VS, 2001. 112. Sem. Drosophilidae. In: Sidorenko, VS, Kupianskaya, AN, Lelej, AS, Nemkov, PG, Kholin, SK, Eds., Opredelitel nasekomikh dalnego vostoka rossii. Tom VI. Dvukriliye i blokhi. Tshast 2. [Key to the insects of Russian Far East, Vol. VI. Diptera, Siphonaptera. Part 2]. Dalnauka, Vladivostok: 211 - 241" title="112. Sem. Drosophilidae." volumeTitle="Opredelitel nasekomikh dalnego vostoka rossii." year="2001">Sidorenko (2001</bibRefCitation>
: 219) [key].
</paragraph>
<paragraph id="7827E8B08ECD2E6D0556EDA6316A95CE" pageId="0" pageNumber="1">
<taxonomicName id="6435C82FBC1CAE0802F8EDC95F6E9D31" authorityName="Bächli &amp; Vilela" authorityYear="2020" baseAuthorityName="Okada" baseAuthorityYear="1973" class="Insecta" family="Drosophilidae" genus="Cacoxenus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cacoxenus kaszabi" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="kaszabi">Cacoxenus kaszabi</taxonomicName>
(Okada):
<bibRefCitation id="8236E2C08F57FD4BA1B48D365545540C" author="Baechli, G" pageId="0" pageNumber="1" refId="B1" refString="Baechli, G, 1984. Catalog of the types of Drosophilidae in the Hungarian Natural History Museum, Budapest (Diptera). Folia Entomologica Hungarica [Rovartani Kezlemenyek] 45(NS): 27-41." title="Catalog of the types of Drosophilidae in the Hungarian Natural History Museum, Budapest (Diptera). Folia Entomologica Hungarica [Rovartani Kezlemenyek] 45 (NS): 27 - 41." year="1984">
<normalizedToken id="88657EB30C96164114EBB230E67B4532" originalValue="Bächli">Baechli</normalizedToken>
(1984
</bibRefCitation>
: 33) [types];
<bibRefCitation id="176C596E867D5EEA7807D7F7E099EA10" author="Toda, MJ" editor="Takahashi, K" journalOrPublisher="Forestry and Forest Products Research Institute, Sapporo" pageId="0" pageNumber="1" pagination="187 - 195" refId="B25" refString="Toda, MJ, Vinokurov, NN, 1995. Biodiversity in drosophilid communities of cool-temperate and boreal birch forests: a special reference to the vertical distribution within forest. In: Takahashi, K, Osawa, A, Kanazawa, Y, Eds., Proceedings of the Third Symposium on the Joint Siberian Permafrost Studies between Japan and Russia. Forestry and Forest Products Research Institute, Sapporo: 187 - 195" title="Biodiversity in drosophilid communities of cool-temperate and boreal birch forests: a special reference to the vertical distribution within forest." volumeTitle="Proceedings of the Third Symposium on the Joint Siberian Permafrost Studies between Japan and Russia." year="1995">Toda and Vinokurov (1995</bibRefCitation>
: 189) [biology, distribution: Siberia];
<bibRefCitation id="1D9AAC1283D2A9534C1F045EF69045A0" author="Watabe, HA" editor="Takahashi, K" journalOrPublisher="Forestry and Forest Products Research Institute, Sapporo" pageId="0" pageNumber="1" pagination="196 - 200" refId="B31" refString="Watabe, HA, Toda, MJ, Vinokurov, NN, Sidorenko, VS, 1995. A comparative study on drosophilid faunas of east Siberia and neighboring regions. In: Takahashi, K, Osawa, A, Kanazawa, Y, Eds., Proceedings of the Third Symposium on the Joint Siberian Permafrost Studies between Japan and Russia. Forestry and Forest Products Research Institute, Sapporo: 196 - 200" title="A comparative study on drosophilid faunas of east Siberia and neighboring regions." volumeTitle="Proceedings of the Third Symposium on the Joint Siberian Permafrost Studies between Japan and Russia." year="1995">Watabe et al. (1995</bibRefCitation>
: 199) [distribution: Siberia];
<bibRefCitation id="0B4D4FB87CD069F16CBDEE7D4B76F6A2" author="Maca, J" journalOrPublisher="Folia Facultatis Scientiarum Naturalium Universitatis Masarykianae Brunensis, Biologia" pageId="0" pageNumber="1" pagination="429 - 434" refId="B17" refString="Maca, J, Bartak, M, 2001. Drosophilidae. Folia Facultatis Scientiarum Naturalium Universitatis Masarykianae Brunensis, Biologia 105: 429 - 434" title="Drosophilidae." volume="105" year="2001">
<normalizedToken id="BF1B0A637D2DC92EB67F9DFAA5028638" originalValue="Máca">Maca</normalizedToken>
and
<normalizedToken id="37936CD0DE1CC3418476C8ADFA89C8D5" originalValue="Barták">Bartak</normalizedToken>
(2001
</bibRefCitation>
: 431) [distribution, misidentification, see below];
<bibRefCitation id="9596AE15199AAE83DBC034754328FB56" author="Vinokurova, AV" editor="Isaev, AP" journalOrPublisher="IBPK SO RAN, Yakutsk" pageId="0" pageNumber="1" pagination="99 - 106" refId="B30" refString="Vinokurova, AV, 2003. Sostav phauni mukh-drosophilid (Diptera, Drosophilidae) Yakutii [Overal view of the fauna of drosophild flies (Diptera, Drosophilidae) in Yakutia]. In: Isaev, AP, Ed., Entomologiceskie issledovanija v Jakutii: Sbornik naucnych trudov [Entomological investigation in Yakutiya: Collection of scientific papers]. IBPK SO RAN, Yakutsk: 99 - 106" title="Sostav phauni mukh-drosophilid (Diptera, Drosophilidae) Yakutii [Overal view of the fauna of drosophild flies (Diptera, Drosophilidae) in Yakutia]." volumeTitle="Entomologiceskie issledovanija v Jakutii: Sbornik naucnych trudov [Entomological investigation in Yakutiya: Collection of scientific papers]." year="2003">Vinokurova (2003</bibRefCitation>
: 100) [distribution: Siberia].
</paragraph>
<paragraph id="D1C303DD133D4A15C6D3102F51A0D82B" pageId="0" pageNumber="1">
<taxonomicName id="2B7AA785F7FA1C716BFE9B1F1904A727" authorityName="Bächli &amp; Vilela" authorityYear="2020" baseAuthorityName="Okada" baseAuthorityYear="1973" class="Insecta" family="Drosophilidae" genus="Cacoxenus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cacoxenus (Paracacoxenus) kaszabi" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="kaszabi" subGenus="Paracacoxenus">Cacoxenus (Paracacoxenus) kaszabi</taxonomicName>
(Okada):
<bibRefCitation id="4F9F7EB50E6EC544514FEEC8EB535BA2" author="Tsacas, L" journalOrPublisher="Annales de la Societe entomologique de France, Nouvelle Serie" pageId="0" pageNumber="1" pagination="491 - 507" refId="B27" refString="Tsacas, L, Desmier de Chenon, R, 1976. Taxinomie et biogeographie des &quot;genres&quot; Cacoxenus - Paracacoxenus - Gitonides - Gitona (Dipt., Drosophilidae) et biologie d'une nouvelle espece africaine commensale d' Apoidea (Hymenoptera). Annales de la Societe entomologique de France, Nouvelle Serie 12: 491 - 507" title="Taxinomie et biogeographie des &quot; genres &quot; Cacoxenus - Paracacoxenus - Gitonides - Gitona (Dipt., Drosophilidae) et biologie d'une nouvelle espece africaine commensale d' Apoidea (Hymenoptera)." volume="12" year="1976">Tsacas and Desmier de Chenon (1976</bibRefCitation>
: 500) [affiliation];
<bibRefCitation id="01DF3CFBBFC6BD0150333DC52A9DC6ED" author="Wheeler, MR" editor="Ashburner, M" journalOrPublisher="Academic Press, London" pageId="0" pageNumber="1" pagination="1 - 97" refId="B32" refString="Wheeler, MR, 1981. 3a. The Drosophilidae: A Taxonomic Overview. In: Ashburner, M, Carson, HL, Thompson, JN, Eds., The Genetics and Biology of Drosophila. Academic Press, London: 1 - 97" title="3 a. The Drosophilidae: A Taxonomic Overview." volumeTitle="The Genetics and Biology of Drosophila." year="1981">Wheeler (1981</bibRefCitation>
: 23) [affiliation];
<bibRefCitation id="49D491071414AC96C832900610E61A20" author="Baechli, G" editor="Soos, A" journalOrPublisher="Akademiai Kiado, Budapest" pageId="0" pageNumber="1" pagination="186 - 220" refId="B4" refString="Baechli, G, Rocha Pite, MT, 1984. Family Drosophilidae. In: Soos, A, Papp, L, Eds., Catalogue of Palaearctic Diptera 10. Akademiai Kiado, Budapest: 186 - 220" title="Family Drosophilidae." volumeTitle="Catalogue of Palaearctic Diptera 10." year="1984">
<normalizedToken id="AF9C8BB5E90EA1E5019860396A222F17" originalValue="Bächli">Baechli</normalizedToken>
and Rocha
<normalizedToken id="364710D9E75B6B4D8694882ACFA9C30F" originalValue="Pité">Pite</normalizedToken>
(1984
</bibRefCitation>
: 191) [affiliation];
<bibRefCitation id="483BB02F705A7178C8185B740E1DE9EA" DOI="https://doi.org/10.2108/zsj.13.455" author="Toda, MJ" journalOrPublisher="Zoological Science (Zoological Society of Japan)" pageId="0" pageNumber="1" pagination="455 - 477" refId="B26" refString="Toda, MJ, Sidorenko, VS, Watabe, H-A, Kholin, SK, Vinokurov, NN, 1996. A Revision of the Drosophilidae (Diptera) in East Siberia and Russian Far East: Taxonomy and Biogeography. Zoological Science (Zoological Society of Japan) 13: 455 - 477, DOI: https://doi.org/10.2108/zsj.13.455" title="A Revision of the Drosophilidae (Diptera) in East Siberia and Russian Far East: Taxonomy and Biogeography." url="https://doi.org/10.2108/zsj.13.455" volume="13" year="1996">Toda et al. (1996</bibRefCitation>
: 466) [key];
<bibRefCitation id="E55999938F83297AFD928730B25C9FDB" author="Baechli, G" journalOrPublisher="Brill, Leiden" pageId="0" pageNumber="1" refId="B6" refString="Baechli, G, Vilela, CR, Andersson Escher, S, Saura, A, 2004. The Drosophilidae (Diptera) of Fennoscandia and Denmark. Fauna Entomologica Scandinavica, vol. 39. Brill, Leiden" title="The Drosophilidae (Diptera) of Fennoscandia and Denmark. Fauna Entomologica Scandinavica, vol. 39." year="2004">
<normalizedToken id="F33E5BEE3F9619D1E0384682D36ECE76" originalValue="Bächli">Baechli</normalizedToken>
et al. (2004
</bibRefCitation>
: 52) [key];
<bibRefCitation id="0EC531154D69488E20386ED496D8B30B" author="Brake, I" journalOrPublisher="7, Apollo Books, Stenstrup" pageId="0" pageNumber="1" refId="B7" refString="Brake, I, Baechli, G, 2008. Drosophilidae (Diptera). World Catalogue of Insects, vol. 7, Apollo Books, Stenstrup" title="Drosophilidae (Diptera). World Catalogue of Insects, vol." year="2008">
Brake and
<normalizedToken id="307C18E0F55A2DEE74AA8D46E9C0159E" originalValue="Bächli">Baechli</normalizedToken>
(2008
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: 251) [affiliation].
</paragraph>
<paragraph id="4966411FE72E16B50097C16525E88324" pageId="0" pageNumber="1">
<taxonomicName id="4E7F1A273469C62045A14E00524EA621" authorityName="Bächli &amp; Vilela" authorityYear="2020" baseAuthorityName="Okada" baseAuthorityYear="1973" class="Insecta" family="Drosophilidae" genus="Cacoxenus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cacoxenus (Gitonides) kaszabi" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="kaszabi" subGenus="Gitonides">Cacoxenus (Gitonides) kaszabi</taxonomicName>
(Okada):
<bibRefCitation id="835DD29C0A8F69B2457B60F8916710C0" author="Sidorenko, VS" journalOrPublisher="Far Eastern Entomologist" pageId="0" pageNumber="1" pagination="1 - 20" refId="B24" refString="Sidorenko, VS, 2002. Phylogeny of the tribe Steganini Hendel and some related taxa (Diptera, Drosophilidae). Far Eastern Entomologist 111: 1 - 20" title="Phylogeny of the tribe Steganini Hendel and some related taxa (Diptera, Drosophilidae)." volume="111" year="2002">Sidorenko (2002</bibRefCitation>
: 10) [phylogeny].
</paragraph>
</subSubSection>
<subSubSection id="8396DFCEED673D82481675352E37443A" pageId="0" pageNumber="1" type="type locality">
<paragraph id="8B0642EB76349EA335FF98E528D5CA66" pageId="0" pageNumber="1">Type locality.</paragraph>
<paragraph id="FE817284F020432B2A38A0EEC39C6264" pageId="0" pageNumber="1">
MONGOLIA: Central aimak, Ulan-Baator, Nucht im Bogdo ul [
<geoCoordinate id="988806AF453F9535F33E507441A37F0A" degrees="47" direction="north" minutes="54" orientation="latitude" precision="925" value="47.9">47°54N</geoCoordinate>
,
<geoCoordinate id="A9BD4435512C013BD8730BA013B82ACB" degrees="106" direction="east" minutes="54" orientation="longitude" precision="925" value="106.9">106°54E</geoCoordinate>
], 1750m.
</paragraph>
</subSubSection>
<subSubSection id="E09203AD1C244E4F706BA66B1D7772C1" pageId="0" pageNumber="1" type="materials_examined">
<paragraph id="C1D080E3E0191AD3F277FB7AABDF863F" pageId="0" pageNumber="1">Material examined.</paragraph>
<paragraph id="0BF0B82CACD4E2A02944FEEE5FDB7EC5" pageId="0" pageNumber="1">
<materialsCitation id="D7F3E2F46C7017FE5175E49B31B02281" collectingDate="1966-06-10" collectorName="Exp. Dr. Z. Kaszab" country="MONGOLIA" county="Bogdo" determinerName="Vilela" elevation="1750" location="Mongolia" municipality="Nucht" specimenCount="1" specimenCount-male="1" typeStatus="Paratype">
<collectingCountry id="45EFCB1A487F8022FBE63676E3901DE9" name="Mongolia">MONGOLIA</collectingCountry>
:
<specimenCount id="EC11DE97F1BF4744AB9635E1B48463FA" count="1" type="male">1 ♂</specimenCount>
(
<typeStatus id="A6746EFEF24435125DF1B4373F6A0C69" type="paratype">Paratype</typeStatus>
), labelled &quot;
<location id="485A91C873AE53C3747CC61D721D4A84" LSID="urn:lsid:plazi:treatment:286CDA892A56528CB8AEA6D0FD827FEC:485A91C873AE53C3747CC61D721D4A84" country="MONGOLIA" county="Bogdo" municipality="Nucht" name="Mongolia">Mongolia</location>
,
<location id="76E0363FB8401F5CAA880BEB89415A28" LSID="urn:lsid:plazi:treatment:286CDA892A56528CB8AEA6D0FD827FEC:76E0363FB8401F5CAA880BEB89415A28" country="MONGOLIA" county="Bogdo" municipality="Nucht" name="Central">Central</location>
aimak /
<location id="E650BFAC91F0EB1E9C9A7BB1C68AD517" LSID="urn:lsid:plazi:treatment:286CDA892A56528CB8AEA6D0FD827FEC:E650BFAC91F0EB1E9C9A7BB1C68AD517" country="MONGOLIA" county="Bogdo" municipality="Nucht" name="Ulan-Baator">Ulan-Baator</location>
,
<collectingMunicipality id="E5BAFF20B137C6DEEE5BB203CFB0C020">Nucht</collectingMunicipality>
im /
<collectingCounty id="9E0DC6637F9F196389AFEB89D5605F17">Bogdo</collectingCounty>
ul,
<quantity id="5CD99CE7C1A515E6D23ECF763826EFD7" metricMagnitude="3" metricUnit="m" metricValue="1.75" unit="m" value="1750.0">
<elevation id="B7A215CB831E961740EC7794197E992B" metricMagnitude="3" metricUnit="m" metricValue="1.75" unit="m" value="1750.0">1750 m</elevation>
</quantity>
/
<collectorName id="FE1932F1EA0893D6168D2AE32296D0CD">Exp. Dr. Z. Kaszab</collectorName>
, 1966 // Nr. 513 /
<collectingDate id="FC560885E7AC3B296BDDA69A347E0AB4" value="1966-06-10">10.VI.1966</collectingDate>
//
<specimenCount id="3230105B92F0DBC91A24FF1B63633DD8" count="1" type="male"></specimenCount>
//
<typeStatus id="70CA05BCDAED8F15C211F47F55EAAE80" type="paratype">PARATYPE</typeStatus>
[white label] //
<emphasis id="05DBE4D61E6D55D33A764998F96B8776" italics="true" pageId="0" pageNumber="1">Paracacoxenus</emphasis>
/
<emphasis id="71CA05E5EC33D8BAF4CEF379E263CA4F" italics="true" pageId="0" pageNumber="1">kaszabi</emphasis>
Okada /
<normalizedToken id="F825D73A7BBB536134D8D46EE987BC85" originalValue="Bächli">Baechli</normalizedToken>
&amp;
<determinerName id="28658BE5D7A41A131F692ACB30236CA8">Vilela</determinerName>
det. 2017&quot; (HNHM). Photomicrographs of
<typeStatus id="8D1F1BC1A172E2445E99E178D78FF27F" type="holotype">holotype</typeStatus>
slide (HNHM)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="E05E055C1EFD776B3FCC6867CA6BFA25" pageId="0" pageNumber="1" type="diagnosis">
<paragraph id="DB7D9196F97BD1944CE7759C88CDF135" pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph id="73E6A35C09E212A9A8D9260D8815586C" pageId="0" pageNumber="1">
Analysis of some male terminalia sclerites is required to tell this species apart from
<taxonomicName id="6FDC3648CB4537636BAF08E357B83036" authorityName="Bächli &amp; Vilela" authorityYear="2020" class="Insecta" family="Drosophilidae" genus="Paracacoxenus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paracacoxenus macai" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="macai">
<emphasis id="7D8373F19FA9ACDBE0B10FCEB6C5D206" italics="true" pageId="0" pageNumber="1">Paracacoxenus macai</emphasis>
</taxonomicName>
sp. nov. They are as follows: cerci narrow and bearing a longitudinal row of setae adjacent to inner margins (Figs
<figureCitation id="72A95CC81C50E52CD46E814D94FE6D64" captionStart="Figures 1316" captionStartId="F4" captionText="Figures 13 - 16. Paracacoxenus kaszabi Okada, ♂ paratype, Central aimak, Ulan-Baator, Nucht im Bogdo ul, Mongolia (HNHM) 13 Terminalia, dorsal view (in-focus: epandrium, cerci, tips of aedeagus and inner paraphyses) 14 Terminalia, dorsal view (in-focus: all sclerites) 15 Terminalia, posterior view 16 Terminalia, anterior view. Scale bar: 0.1 mm." figureDoi="10.3897/alpento.4.49492.figures13-16" httpUri="https://binary.pensoft.net/fig/387178" pageId="0" pageNumber="1">13</figureCitation>
,
<figureCitation id="71BD488BFA07CD2773C2315E3E49A480" captionStart="Figures 1316" captionStartId="F4" captionText="Figures 13 - 16. Paracacoxenus kaszabi Okada, ♂ paratype, Central aimak, Ulan-Baator, Nucht im Bogdo ul, Mongolia (HNHM) 13 Terminalia, dorsal view (in-focus: epandrium, cerci, tips of aedeagus and inner paraphyses) 14 Terminalia, dorsal view (in-focus: all sclerites) 15 Terminalia, posterior view 16 Terminalia, anterior view. Scale bar: 0.1 mm." figureDoi="10.3897/alpento.4.49492.figures13-16" httpUri="https://binary.pensoft.net/fig/387178" pageId="0" pageNumber="1">14</figureCitation>
), aedeagus ellipsoid-shaped in dorsal and ventral views (Figs
<figureCitation id="7E95A02DF1B2609B23141A3B7F5EBEFB" captionStart="Figures 1316" captionStartId="F4" captionText="Figures 13 - 16. Paracacoxenus kaszabi Okada, ♂ paratype, Central aimak, Ulan-Baator, Nucht im Bogdo ul, Mongolia (HNHM) 13 Terminalia, dorsal view (in-focus: epandrium, cerci, tips of aedeagus and inner paraphyses) 14 Terminalia, dorsal view (in-focus: all sclerites) 15 Terminalia, posterior view 16 Terminalia, anterior view. Scale bar: 0.1 mm." figureDoi="10.3897/alpento.4.49492.figures13-16" httpUri="https://binary.pensoft.net/fig/387178" pageId="0" pageNumber="1">15</figureCitation>
,
<figureCitation id="440847572DE6A69C268234161BBC93CB" captionStart="Figures 1316" captionStartId="F4" captionText="Figures 13 - 16. Paracacoxenus kaszabi Okada, ♂ paratype, Central aimak, Ulan-Baator, Nucht im Bogdo ul, Mongolia (HNHM) 13 Terminalia, dorsal view (in-focus: epandrium, cerci, tips of aedeagus and inner paraphyses) 14 Terminalia, dorsal view (in-focus: all sclerites) 15 Terminalia, posterior view 16 Terminalia, anterior view. Scale bar: 0.1 mm." figureDoi="10.3897/alpento.4.49492.figures13-16" httpUri="https://binary.pensoft.net/fig/387178" pageId="0" pageNumber="1">16</figureCitation>
), inner paraphyses tribranched (Figs
<figureCitation id="2F2521264A8B8C2B1744E3C05B93F9FC" captionStart="Figures 21, 22" captionStartId="F8" captionText="Figures 21, 22. Paracacoxenus kaszabi Okada, ♂ paratype, Central aimak, Ulan-Baator, Nucht im Bogdo ul, Mongolia (HNHM) 21 Three-branched inner paraphysis, outer paraphysis and aedeagal apodeme, anterior (dorsal) view 22 Three-branched inner paraphysis, outer paraphysis and aedeagal apodeme, oblique posterior view. Abbreviations: a = proximal branch; b = mediodorsal branch; c = distal branch. Scale bar: 0.1 mm." figureDoi="10.3897/alpento.4.49492.figures21-22" httpUri="https://binary.pensoft.net/fig/387182" pageId="0" pageNumber="1">21</figureCitation>
,
<figureCitation id="574D0891CF3992739CD4E132CAB09902" captionStart="Figures 21, 22" captionStartId="F8" captionText="Figures 21, 22. Paracacoxenus kaszabi Okada, ♂ paratype, Central aimak, Ulan-Baator, Nucht im Bogdo ul, Mongolia (HNHM) 21 Three-branched inner paraphysis, outer paraphysis and aedeagal apodeme, anterior (dorsal) view 22 Three-branched inner paraphysis, outer paraphysis and aedeagal apodeme, oblique posterior view. Abbreviations: a = proximal branch; b = mediodorsal branch; c = distal branch. Scale bar: 0.1 mm." figureDoi="10.3897/alpento.4.49492.figures21-22" httpUri="https://binary.pensoft.net/fig/387182" pageId="0" pageNumber="1">22</figureCitation>
), and gonopods smooth (Figs
<figureCitation id="F2CDBD2BAB041B532FDD66DB0C29BD2B" captionStart="Figure 17" captionStartId="F5" captionText="Figure 17. Paracacoxenus kaszabi Okada, ♂ paratype, Central aimak, Ulan-Baator, Nucht im Bogdo ul, Mongolia (HNHM); decasternum, hypandrium + gonopods, aedeagus, oblique posterior view. Abbreviations: a = proximal section of decasternum; b and c = distal section of decasternum; d = membranous aedeagus; e = anterior region of hypandrium; f = fused gonopods; g = tip of distal section of decasternum (laterally expanded and fused to dorsal region of aedeagus). Scale bar: 0.1 mm." figureDoi="10.3897/alpento.4.49492.figure17" httpUri="https://binary.pensoft.net/fig/387179" pageId="0" pageNumber="1">17</figureCitation>
,
<figureCitation id="C1E2B208A02C136DD473628CEDCF7F75" captionStart="Figure 18" captionStartId="F6" captionText="Figure 18. Paracacoxenus kaszabi Okada, ♂ paratype, Central aimak, Ulan-Baator, Nucht im Bogdo ul, Mongolia (HNHM); decasternum and hypandrium + gonopods. Abbreviations: a = proximal section of decasternum; b and c = distal section of decasternum; d = tip of distal section of decasternum (laterally expanded); e = hypandrium left arm; f = anterior region of gonopods (fused to each other and to hypandrium); g = posterolateral region of left gonopod. Scale bar: 0.1 mm." figureDoi="10.3897/alpento.4.49492.figure18" httpUri="https://binary.pensoft.net/fig/387180" pageId="0" pageNumber="1">18</figureCitation>
).
</paragraph>
<caption id="CE5553DF6CD8593B2F180357235492B2" ID-Zenodo-Dep="3710601" doi="10.3897/alpento.4.49492.figures1-3" httpUri="https://binary.pensoft.net/fig/387175" pageId="0" pageNumber="1" start="Figures 13" startId="F1">
<paragraph id="7B3ED4E73144BAFC2E53CD477DB57642" pageId="0" pageNumber="1">
<emphasis id="012E2033E7A3B816E3ACF26BA513C4A6" bold="true" pageId="0" pageNumber="1">Figures 1-3.</emphasis>
<taxonomicName id="39EAB1534ED4B10B7892D3D0C0CD3143" authorityName="Okada" authorityYear="1973" class="Insecta" family="Drosophilidae" genus="Paracacoxenus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paracacoxenus kaszabi" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="kaszabi">
<emphasis id="88736D1050CC6065936637A792407DA9" italics="true" pageId="0" pageNumber="1">Paracacoxenus kaszabi</emphasis>
</taxonomicName>
Okada, ♂ holotype, Central aimak, Ulan-Baator, Nucht im Bogdo ul, Mongolia (HNHM)
<emphasis id="30C509BAD1AF1D23018342484048A6DF" bold="true" pageId="0" pageNumber="1">1</emphasis>
Labels (mislabelled as a female) and slide with wing and terminalia
<emphasis id="CE2E1873AEEE64AB4FCB338187048370" bold="true" pageId="0" pageNumber="1">2</emphasis>
Photomicrograph of terminalia, dorsal view
<emphasis id="F4D9084AA57A20702D5BAFAD1DE9053F" bold="true" pageId="0" pageNumber="1">3</emphasis>
Right wing, dorsal view, distally optically cut. Photos: Z.
<normalizedToken id="BFD2DA6927459BBEC6431FA5BA810544" originalValue="Soltèsz">Soltesz</normalizedToken>
.
</paragraph>
</caption>
<caption id="3BBD48B5C78FC10F20C99307014D0A65" ID-Zenodo-Dep="3710617" doi="10.3897/alpento.4.49492.figures4-9" httpUri="https://binary.pensoft.net/fig/387176" pageId="0" pageNumber="1" start="Figures 49" startId="F2">
<paragraph id="63163DE587D59D44215878E2FCCC3CF9" pageId="0" pageNumber="1">
<emphasis id="357AA9623D890CBE2E95963226CF1058" bold="true" pageId="0" pageNumber="1">Figures 4-9.</emphasis>
<taxonomicName id="0B4D2E17FD51121931228F9137A625A2" authorityName="Okada" authorityYear="1973" class="Insecta" family="Drosophilidae" genus="Paracacoxenus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paracacoxenus kaszabi" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="kaszabi">
<emphasis id="05A6760DF91B93C5DEC57E1FF9F979DD" italics="true" pageId="0" pageNumber="1">Paracacoxenus kaszabi</emphasis>
</taxonomicName>
Okada, ♂ paratype, Central aimak, Ulan-Baator, Nucht im Bogdo ul, Mongolia (HNHM)
<emphasis id="D6BE4285EFBA3F2EFFAD0FC7411B042F" bold="true" pageId="0" pageNumber="1">4</emphasis>
Habitus, left lateral view
<emphasis id="35C33A0BD9BCE70FE9C00DA7DFCBBCC0" bold="true" pageId="0" pageNumber="1">5</emphasis>
Habitus, oblique dorsal
<emphasis id="7E0C8CCBFC3F8E556554CC1B0997E3F4" bold="true" pageId="0" pageNumber="1">6</emphasis>
Habitus, dorsal
<emphasis id="07F9E75A93EB61429818C3FD7AA329D8" bold="true" pageId="0" pageNumber="1">7</emphasis>
Head close-up, left lateral
<emphasis id="6046F3415BF3B5EAFFA6171B86C2660E" bold="true" pageId="0" pageNumber="1">8</emphasis>
Head close-up, dorsal
<emphasis id="C0024CFFD3C8B399E5C1AB51C1B6A778" bold="true" pageId="0" pageNumber="1">9</emphasis>
Terminalia close-up, posterior. Scale bars: 0.5 mm (
<emphasis id="BE54B2E47D93C5E9D65059058F35B86E" bold="true" pageId="0" pageNumber="1">7-9</emphasis>
); 1.0 mm (
<emphasis id="4537A8D5D52C32E157C8D82F6AE7430D" bold="true" pageId="0" pageNumber="1">4-6</emphasis>
).
</paragraph>
</caption>
<caption id="879614423CF9D24C24C804598EF18B1A" ID-Zenodo-Dep="3710619" doi="10.3897/alpento.4.49492.figures10-12" httpUri="https://binary.pensoft.net/fig/387177" pageId="0" pageNumber="1" start="Figures 1012" startId="F3">
<paragraph id="1B797D524A68F5BFC4058B59B7F5EF53" pageId="0" pageNumber="1">
<emphasis id="A8581C70D3406EE4645622A150178A21" bold="true" pageId="0" pageNumber="1">Figures 10-12.</emphasis>
<taxonomicName id="A23C95DF7BE726D3671B3039CE1919DA" authorityName="Okada" authorityYear="1973" class="Insecta" family="Drosophilidae" genus="Paracacoxenus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paracacoxenus kaszabi" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="kaszabi">
<emphasis id="21445CA688B62591F7EEF76FB745F2BD" italics="true" pageId="0" pageNumber="1">Paracacoxenus kaszabi</emphasis>
</taxonomicName>
Okada, ♂ paratype, Central aimak, Ulan-Baator, Nucht im Bogdo ul, Mongolia (HNHM)
<emphasis id="E7D8570D2E88CE6A7FC822AB63BFE981" bold="true" pageId="0" pageNumber="1">10</emphasis>
Terminalia, oblique dorsal view
<emphasis id="23C1B3B93C97F4921C6FDEA1AEBA4E81" bold="true" pageId="0" pageNumber="1">11</emphasis>
Terminalia, left lateral view
<emphasis id="0F187535A69595F733D1F26803FD39B0" bold="true" pageId="0" pageNumber="1">12</emphasis>
Terminalia, oblique ventral view. Scale bar: 0.1 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="9FAAFD995087C69ED50266783752B2CD" pageId="0" pageNumber="1" type="description">
<paragraph id="F631084E1961A9DD643C2A2886DEEAA0" pageId="0" pageNumber="1">Description.</paragraph>
<paragraph id="2E1741EB1823FF94B9F60E56B856532F" pageId="0" pageNumber="1">
♂.
<emphasis id="52D6603A41F8C7FA65DE81989B816963" bold="true" pageId="0" pageNumber="1">Head</emphasis>
: Frons (Figs
<figureCitation id="F6C25231C9BF773F352210E89F17D953" captionStart="Figures 49" captionStartId="F2" captionText="Figures 4 - 9. Paracacoxenus kaszabi Okada, ♂ paratype, Central aimak, Ulan-Baator, Nucht im Bogdo ul, Mongolia (HNHM) 4 Habitus, left lateral view 5 Habitus, oblique dorsal 6 Habitus, dorsal 7 Head close-up, left lateral 8 Head close-up, dorsal 9 Terminalia close-up, posterior. Scale bars: 0.5 mm (7 - 9); 1.0 mm (4 - 6)." figureDoi="10.3897/alpento.4.49492.figures4-9" httpUri="https://binary.pensoft.net/fig/387176" pageId="0" pageNumber="1">6</figureCitation>
,
<figureCitation id="C86787F76CAA7454D0D25BF8FC39CCCF" captionStart="Figures 49" captionStartId="F2" captionText="Figures 4 - 9. Paracacoxenus kaszabi Okada, ♂ paratype, Central aimak, Ulan-Baator, Nucht im Bogdo ul, Mongolia (HNHM) 4 Habitus, left lateral view 5 Habitus, oblique dorsal 6 Habitus, dorsal 7 Head close-up, left lateral 8 Head close-up, dorsal 9 Terminalia close-up, posterior. Scale bars: 0.5 mm (7 - 9); 1.0 mm (4 - 6)." figureDoi="10.3897/alpento.4.49492.figures4-9" httpUri="https://binary.pensoft.net/fig/387176" pageId="0" pageNumber="1">8</figureCitation>
) centrally velvety black; all setae black. Frontal length 0.38 mm; frontal index about 1.00, top to bottom width ratio about 1.19. Frontal triangle not distinguishable; ocellar triangle greyish-black, microtrichose, about 40 % of frontal length. Orbital plates greyish-black, microtrichose, about 80 % of frontal length. Orbital setae arranged almost in a row, distance of or3 to or1 about 157 % of or3 to vtm, or1 / or3 ratio about 1.09, or2 / or1 ratio about 0.83; vt index about 1.00, postocellar setae broken or invisible; ocellar setae about 57 %, of frontal length, vibrissa (Fig.
<figureCitation id="4CCD834DEAE0E877664F20AC7C24E1D4" captionStart="Figures 49" captionStartId="F2" captionText="Figures 4 - 9. Paracacoxenus kaszabi Okada, ♂ paratype, Central aimak, Ulan-Baator, Nucht im Bogdo ul, Mongolia (HNHM) 4 Habitus, left lateral view 5 Habitus, oblique dorsal 6 Habitus, dorsal 7 Head close-up, left lateral 8 Head close-up, dorsal 9 Terminalia close-up, posterior. Scale bars: 0.5 mm (7 - 9); 1.0 mm (4 - 6)." figureDoi="10.3897/alpento.4.49492.figures4-9" httpUri="https://binary.pensoft.net/fig/387176" pageId="0" pageNumber="1">7</figureCitation>
) about 50 % of frontal length. Face black. Carina narrow, slightly prominent. Cheek index about 9. Eye index about 1.26. Occiput black. Antenna black. Flagellomere 1 roundish, length to width ratio about 1.00. Arista microtrichose (Fig.
<figureCitation id="963DD8DF1E46377BD0FC3D5B02CF7CFE" captionStart="Figures 49" captionStartId="F2" captionText="Figures 4 - 9. Paracacoxenus kaszabi Okada, ♂ paratype, Central aimak, Ulan-Baator, Nucht im Bogdo ul, Mongolia (HNHM) 4 Habitus, left lateral view 5 Habitus, oblique dorsal 6 Habitus, dorsal 7 Head close-up, left lateral 8 Head close-up, dorsal 9 Terminalia close-up, posterior. Scale bars: 0.5 mm (7 - 9); 1.0 mm (4 - 6)." figureDoi="10.3897/alpento.4.49492.figures4-9" httpUri="https://binary.pensoft.net/fig/387176" pageId="0" pageNumber="1">7</figureCitation>
). Proboscis black. Palpus short, black, with 2 more prominent apical setae.
<emphasis id="685D8B547CDED2D3A9B20DD7D88FEA30" bold="true" pageId="0" pageNumber="1">Thorax</emphasis>
dull black, length about 1.19 mm, 8 rows of acrostichal setae (Figs
<figureCitation id="BBCE313C575E7BD3F5B0536C240603F7" captionStart="Figures 49" captionStartId="F2" captionText="Figures 4 - 9. Paracacoxenus kaszabi Okada, ♂ paratype, Central aimak, Ulan-Baator, Nucht im Bogdo ul, Mongolia (HNHM) 4 Habitus, left lateral view 5 Habitus, oblique dorsal 6 Habitus, dorsal 7 Head close-up, left lateral 8 Head close-up, dorsal 9 Terminalia close-up, posterior. Scale bars: 0.5 mm (7 - 9); 1.0 mm (4 - 6)." figureDoi="10.3897/alpento.4.49492.figures4-9" httpUri="https://binary.pensoft.net/fig/387176" pageId="0" pageNumber="1">4-6</figureCitation>
), one prominent h. Anterior and posterior dorsocentral setae close together, transverse distance about 6x of longitudinal distance; dc index = 0.38. Scutellar setae nearly equidistant; basal ones divergent; apical scut broken. Haltere whitish. Legs black, knees slightly paler.
<emphasis id="FC87A715CFF973C7EBFAEC3B24E4E867" bold="true" pageId="0" pageNumber="1">Wing</emphasis>
hyaline, damaged and partly folded, length 1.40 mm (Fig.
<figureCitation id="AFAB2715586D8ABC6D6BFABA874B987B" captionStart="Figures 13" captionStartId="F1" captionText="Figures 1 - 3. Paracacoxenus kaszabi Okada, ♂ holotype, Central aimak, Ulan-Baator, Nucht im Bogdo ul, Mongolia (HNHM) 1 Labels (mislabelled as a female) and slide with wing and terminalia 2 Photomicrograph of terminalia, dorsal view 3 Right wing, dorsal view, distally optically cut. Photos: Z. Soltesz." figureDoi="10.3897/alpento.4.49492.figures1-3" httpUri="https://binary.pensoft.net/fig/387175" pageId="0" pageNumber="1">3</figureCitation>
from holotype).
<emphasis id="6694806F81DB781F58ECC23396D73D33" bold="true" pageId="0" pageNumber="1">Abdomen</emphasis>
dull black (Figs
<figureCitation id="0648D0430BF96AA0DA9469842F9D33F4" captionStart="Figures 49" captionStartId="F2" captionText="Figures 4 - 9. Paracacoxenus kaszabi Okada, ♂ paratype, Central aimak, Ulan-Baator, Nucht im Bogdo ul, Mongolia (HNHM) 4 Habitus, left lateral view 5 Habitus, oblique dorsal 6 Habitus, dorsal 7 Head close-up, left lateral 8 Head close-up, dorsal 9 Terminalia close-up, posterior. Scale bars: 0.5 mm (7 - 9); 1.0 mm (4 - 6)." figureDoi="10.3897/alpento.4.49492.figures4-9" httpUri="https://binary.pensoft.net/fig/387176" pageId="0" pageNumber="1">4</figureCitation>
,
<figureCitation id="73988E696C9A46DF319909910AA3F1EE" captionStart="Figures 49" captionStartId="F2" captionText="Figures 4 - 9. Paracacoxenus kaszabi Okada, ♂ paratype, Central aimak, Ulan-Baator, Nucht im Bogdo ul, Mongolia (HNHM) 4 Habitus, left lateral view 5 Habitus, oblique dorsal 6 Habitus, dorsal 7 Head close-up, left lateral 8 Head close-up, dorsal 9 Terminalia close-up, posterior. Scale bars: 0.5 mm (7 - 9); 1.0 mm (4 - 6)." figureDoi="10.3897/alpento.4.49492.figures4-9" httpUri="https://binary.pensoft.net/fig/387176" pageId="0" pageNumber="1">9</figureCitation>
). Syntergite 6 + 7 (Fig.
<figureCitation id="A59FF8CB7BAD9EEA2E62E90A6AA44489" captionStart="Figures 49" captionStartId="F2" captionText="Figures 4 - 9. Paracacoxenus kaszabi Okada, ♂ paratype, Central aimak, Ulan-Baator, Nucht im Bogdo ul, Mongolia (HNHM) 4 Habitus, left lateral view 5 Habitus, oblique dorsal 6 Habitus, dorsal 7 Head close-up, left lateral 8 Head close-up, dorsal 9 Terminalia close-up, posterior. Scale bars: 0.5 mm (7 - 9); 1.0 mm (4 - 6)." figureDoi="10.3897/alpento.4.49492.figures4-9" httpUri="https://binary.pensoft.net/fig/387176" pageId="0" pageNumber="1">9</figureCitation>
) without a claw-like extension.
<emphasis id="F95DE8088B220689D7248B943D207318" bold="true" pageId="0" pageNumber="1">Terminalia</emphasis>
♂ (Figs
<figureCitation id="62A4E07AF55C0ADB36CA46647B054C58" captionStart="Figures 13" captionStartId="F1" captionText="Figures 1 - 3. Paracacoxenus kaszabi Okada, ♂ holotype, Central aimak, Ulan-Baator, Nucht im Bogdo ul, Mongolia (HNHM) 1 Labels (mislabelled as a female) and slide with wing and terminalia 2 Photomicrograph of terminalia, dorsal view 3 Right wing, dorsal view, distally optically cut. Photos: Z. Soltesz." figureDoi="10.3897/alpento.4.49492.figures1-3" httpUri="https://binary.pensoft.net/fig/387175" pageId="0" pageNumber="1">2</figureCitation>
[holotype], 9-22): Epandrium microtrichose (Fig.
<figureCitation id="BF9FC2F2181BBDC8572160DFF389387D" captionStart="Figures 1316" captionStartId="F4" captionText="Figures 13 - 16. Paracacoxenus kaszabi Okada, ♂ paratype, Central aimak, Ulan-Baator, Nucht im Bogdo ul, Mongolia (HNHM) 13 Terminalia, dorsal view (in-focus: epandrium, cerci, tips of aedeagus and inner paraphyses) 14 Terminalia, dorsal view (in-focus: all sclerites) 15 Terminalia, posterior view 16 Terminalia, anterior view. Scale bar: 0.1 mm." figureDoi="10.3897/alpento.4.49492.figures13-16" httpUri="https://binary.pensoft.net/fig/387178" pageId="0" pageNumber="1">14</figureCitation>
), except for the anteroventral region, with no ventral, and 1-2 dorsal setae, ventral lobe fused to surstylus. Cercus narrow [wide in
<taxonomicName id="76A316CF6C3E307906821E74E5F90C0A" authorityName="Bächli &amp; Vilela" authorityYear="2020" class="Insecta" family="Drosophilidae" genus="Paracacoxenus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paracacoxenus macai" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="macai">
<emphasis id="258FD8C2C5047845CC8563DAB2DFFECE" italics="true" pageId="0" pageNumber="1">Paracacoxenus macai</emphasis>
</taxonomicName>
sp. nov.], ventrally positioned, laterally connected to epandrium by membranous tissue, covered with only 8 setae [18-21 in
<taxonomicName id="4F65AFB33D84A6B85BBCE55FD79C6728" class="Hexapoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="P. macai" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="macai">
<emphasis id="6FC476576396EB40555FA5E5C75B1328" italics="true" pageId="0" pageNumber="1">P. macai</emphasis>
</taxonomicName>
sp. nov.] organized in a longitudinal row (Figs
<figureCitation id="38343E5E36187EA1FFDEDD085AE23DF1" captionStart="Figures 1316" captionStartId="F4" captionText="Figures 13 - 16. Paracacoxenus kaszabi Okada, ♂ paratype, Central aimak, Ulan-Baator, Nucht im Bogdo ul, Mongolia (HNHM) 13 Terminalia, dorsal view (in-focus: epandrium, cerci, tips of aedeagus and inner paraphyses) 14 Terminalia, dorsal view (in-focus: all sclerites) 15 Terminalia, posterior view 16 Terminalia, anterior view. Scale bar: 0.1 mm." figureDoi="10.3897/alpento.4.49492.figures13-16" httpUri="https://binary.pensoft.net/fig/387178" pageId="0" pageNumber="1">13</figureCitation>
,
<figureCitation id="71CCC4F39CBA5CD507D1E124E532AB01" captionStart="Figures 1316" captionStartId="F4" captionText="Figures 13 - 16. Paracacoxenus kaszabi Okada, ♂ paratype, Central aimak, Ulan-Baator, Nucht im Bogdo ul, Mongolia (HNHM) 13 Terminalia, dorsal view (in-focus: epandrium, cerci, tips of aedeagus and inner paraphyses) 14 Terminalia, dorsal view (in-focus: all sclerites) 15 Terminalia, posterior view 16 Terminalia, anterior view. Scale bar: 0.1 mm." figureDoi="10.3897/alpento.4.49492.figures13-16" httpUri="https://binary.pensoft.net/fig/387178" pageId="0" pageNumber="1">14</figureCitation>
) adjacent to inner margin, microtrichose, without ventral lobe. Surstylus without prensisetae, bearing 3 outer setae (Figs
<figureCitation id="E84AC84C6C4C1FECCD60446BB90E1633" captionStart="Figures 1012" captionStartId="F3" captionText="Figures 10 - 12. Paracacoxenus kaszabi Okada, ♂ paratype, Central aimak, Ulan-Baator, Nucht im Bogdo ul, Mongolia (HNHM) 10 Terminalia, oblique dorsal view 11 Terminalia, left lateral view 12 Terminalia, oblique ventral view. Scale bar: 0.1 mm." figureDoi="10.3897/alpento.4.49492.figures10-12" httpUri="https://binary.pensoft.net/fig/387177" pageId="0" pageNumber="1">12</figureCitation>
,
<figureCitation id="80940609C51F8DAD14B73B052EB8F722" captionStart="Figures 1316" captionStartId="F4" captionText="Figures 13 - 16. Paracacoxenus kaszabi Okada, ♂ paratype, Central aimak, Ulan-Baator, Nucht im Bogdo ul, Mongolia (HNHM) 13 Terminalia, dorsal view (in-focus: epandrium, cerci, tips of aedeagus and inner paraphyses) 14 Terminalia, dorsal view (in-focus: all sclerites) 15 Terminalia, posterior view 16 Terminalia, anterior view. Scale bar: 0.1 mm." figureDoi="10.3897/alpento.4.49492.figures13-16" httpUri="https://binary.pensoft.net/fig/387178" pageId="0" pageNumber="1">13</figureCitation>
,
<figureCitation id="FC5BC4EE853906A6C5C91F2E22AF973F" captionStart="Figures 19, 20" captionStartId="F7" captionText="Figures 19, 20. Paracacoxenus kaszabi Okada, ♂ paratype, Central aimak, Ulan-Baator, Nucht im Bogdo ul, Mongolia (HNHM) 19 Aedeagal apodeme, left inner paraphysis, left outer paraphysis, membranous aedeagus and posterior ejaculatory duct, left lateral view 20 Left surstylus, decasternum, hypandrium, gonopod, aedeagus and posterior ejaculatory duct, lateral view. Scale bar: 0.1 mm." figureDoi="10.3897/alpento.4.49492.figures19-20" httpUri="https://binary.pensoft.net/fig/387181" pageId="0" pageNumber="1">20</figureCitation>
) [2 in
<taxonomicName id="62FA5E2E51B91FE08665170A0798D9A2" class="Hexapoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="P. macai" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="macai">
<emphasis id="C045EA2D8AC5A68686C38A073D23171B" italics="true" pageId="0" pageNumber="1">P. macai</emphasis>
</taxonomicName>
sp. nov.] on central membranous area and no inner seta, not microtrichose, completely fused to ventral lobe of epandrium which is slightly microtrichose in the fusion area (Figs
<figureCitation id="1D9DB7756652AE0C134D31F5245C3E75" captionStart="Figures 1316" captionStartId="F4" captionText="Figures 13 - 16. Paracacoxenus kaszabi Okada, ♂ paratype, Central aimak, Ulan-Baator, Nucht im Bogdo ul, Mongolia (HNHM) 13 Terminalia, dorsal view (in-focus: epandrium, cerci, tips of aedeagus and inner paraphyses) 14 Terminalia, dorsal view (in-focus: all sclerites) 15 Terminalia, posterior view 16 Terminalia, anterior view. Scale bar: 0.1 mm." figureDoi="10.3897/alpento.4.49492.figures13-16" httpUri="https://binary.pensoft.net/fig/387178" pageId="0" pageNumber="1">13</figureCitation>
,
<figureCitation id="6DB23EAEBA68773379E984B9AE3AE480" captionStart="Figures 1316" captionStartId="F4" captionText="Figures 13 - 16. Paracacoxenus kaszabi Okada, ♂ paratype, Central aimak, Ulan-Baator, Nucht im Bogdo ul, Mongolia (HNHM) 13 Terminalia, dorsal view (in-focus: epandrium, cerci, tips of aedeagus and inner paraphyses) 14 Terminalia, dorsal view (in-focus: all sclerites) 15 Terminalia, posterior view 16 Terminalia, anterior view. Scale bar: 0.1 mm." figureDoi="10.3897/alpento.4.49492.figures13-16" httpUri="https://binary.pensoft.net/fig/387178" pageId="0" pageNumber="1">14</figureCitation>
). Decasternum divided into two articulate sections connected by a membranous strip, folded over one another (Figs
<figureCitation id="BCBEC36B07AEFA62FAA21142AF047A60" captionStart="Figure 17" captionStartId="F5" captionText="Figure 17. Paracacoxenus kaszabi Okada, ♂ paratype, Central aimak, Ulan-Baator, Nucht im Bogdo ul, Mongolia (HNHM); decasternum, hypandrium + gonopods, aedeagus, oblique posterior view. Abbreviations: a = proximal section of decasternum; b and c = distal section of decasternum; d = membranous aedeagus; e = anterior region of hypandrium; f = fused gonopods; g = tip of distal section of decasternum (laterally expanded and fused to dorsal region of aedeagus). Scale bar: 0.1 mm." figureDoi="10.3897/alpento.4.49492.figure17" httpUri="https://binary.pensoft.net/fig/387179" pageId="0" pageNumber="1">17</figureCitation>
,
<figureCitation id="937B544C17777AAC5534893B157E8301" captionStart="Figure 18" captionStartId="F6" captionText="Figure 18. Paracacoxenus kaszabi Okada, ♂ paratype, Central aimak, Ulan-Baator, Nucht im Bogdo ul, Mongolia (HNHM); decasternum and hypandrium + gonopods. Abbreviations: a = proximal section of decasternum; b and c = distal section of decasternum; d = tip of distal section of decasternum (laterally expanded); e = hypandrium left arm; f = anterior region of gonopods (fused to each other and to hypandrium); g = posterolateral region of left gonopod. Scale bar: 0.1 mm." figureDoi="10.3897/alpento.4.49492.figure18" httpUri="https://binary.pensoft.net/fig/387180" pageId="0" pageNumber="1">18</figureCitation>
); proximal section flattened, anteromedially somewhat projected posterad, medially slightly concave, distally fused to surstyli and laterally connected to hypandrium arms by membranous tissue (Figs
<figureCitation id="8E1B3D63837EC3917FA567B41E9BAB03" captionStart="Figures 1316" captionStartId="F4" captionText="Figures 13 - 16. Paracacoxenus kaszabi Okada, ♂ paratype, Central aimak, Ulan-Baator, Nucht im Bogdo ul, Mongolia (HNHM) 13 Terminalia, dorsal view (in-focus: epandrium, cerci, tips of aedeagus and inner paraphyses) 14 Terminalia, dorsal view (in-focus: all sclerites) 15 Terminalia, posterior view 16 Terminalia, anterior view. Scale bar: 0.1 mm." figureDoi="10.3897/alpento.4.49492.figures13-16" httpUri="https://binary.pensoft.net/fig/387178" pageId="0" pageNumber="1">16</figureCitation>
,
<figureCitation id="9C29D59912DB33904C2336C039672B8D" captionStart="Figure 17" captionStartId="F5" captionText="Figure 17. Paracacoxenus kaszabi Okada, ♂ paratype, Central aimak, Ulan-Baator, Nucht im Bogdo ul, Mongolia (HNHM); decasternum, hypandrium + gonopods, aedeagus, oblique posterior view. Abbreviations: a = proximal section of decasternum; b and c = distal section of decasternum; d = membranous aedeagus; e = anterior region of hypandrium; f = fused gonopods; g = tip of distal section of decasternum (laterally expanded and fused to dorsal region of aedeagus). Scale bar: 0.1 mm." figureDoi="10.3897/alpento.4.49492.figure17" httpUri="https://binary.pensoft.net/fig/387179" pageId="0" pageNumber="1">17</figureCitation>
); distal section [ventral process of
<bibRefCitation id="7A77867B8063ACCC5887F6B7310676E0" author="Grimaldi, DA" journalOrPublisher="American Museum of Natural History" pageId="0" pageNumber="1" pagination="1 - 139" refId="B9" refString="Grimaldi, DA, 1990. A phylogenetic, revised classification of genera in the Drosophilidae (Diptera). American Museum of Natural History 197: 1 - 139" title="A phylogenetic, revised classification of genera in the Drosophilidae (Diptera)." volume="197" year="1990">Grimaldi (1990)</bibRefCitation>
: 76, 77 (fig. 424), 78] V-shaped at rest (Figs
<figureCitation id="BBB87005619324AC0BDA4FD45C8953A3" captionStart="Figure 17" captionStartId="F5" captionText="Figure 17. Paracacoxenus kaszabi Okada, ♂ paratype, Central aimak, Ulan-Baator, Nucht im Bogdo ul, Mongolia (HNHM); decasternum, hypandrium + gonopods, aedeagus, oblique posterior view. Abbreviations: a = proximal section of decasternum; b and c = distal section of decasternum; d = membranous aedeagus; e = anterior region of hypandrium; f = fused gonopods; g = tip of distal section of decasternum (laterally expanded and fused to dorsal region of aedeagus). Scale bar: 0.1 mm." figureDoi="10.3897/alpento.4.49492.figure17" httpUri="https://binary.pensoft.net/fig/387179" pageId="0" pageNumber="1">17b</figureCitation>
,
<figureCitation id="ED98EA28E080058DF32FDB1703B86A38" captionStart="Figure 17" captionStartId="F5" captionText="Figure 17. Paracacoxenus kaszabi Okada, ♂ paratype, Central aimak, Ulan-Baator, Nucht im Bogdo ul, Mongolia (HNHM); decasternum, hypandrium + gonopods, aedeagus, oblique posterior view. Abbreviations: a = proximal section of decasternum; b and c = distal section of decasternum; d = membranous aedeagus; e = anterior region of hypandrium; f = fused gonopods; g = tip of distal section of decasternum (laterally expanded and fused to dorsal region of aedeagus). Scale bar: 0.1 mm." figureDoi="10.3897/alpento.4.49492.figure17" httpUri="https://binary.pensoft.net/fig/387179" pageId="0" pageNumber="1">17c</figureCitation>
,
<figureCitation id="8977CE3C1E6217EDC4ABBBAB3F09AD8C" captionStart="Figures 19, 20" captionStartId="F7" captionText="Figures 19, 20. Paracacoxenus kaszabi Okada, ♂ paratype, Central aimak, Ulan-Baator, Nucht im Bogdo ul, Mongolia (HNHM) 19 Aedeagal apodeme, left inner paraphysis, left outer paraphysis, membranous aedeagus and posterior ejaculatory duct, left lateral view 20 Left surstylus, decasternum, hypandrium, gonopod, aedeagus and posterior ejaculatory duct, lateral view. Scale bar: 0.1 mm." figureDoi="10.3897/alpento.4.49492.figures19-20" httpUri="https://binary.pensoft.net/fig/387181" pageId="0" pageNumber="1">20</figureCitation>
), proximally bifid, distally flattened, slightly bifid and expanded laterad, marginally non-crested (Figs
<figureCitation id="6918E28F5FB832B9ACAAF6335851F72D" captionStart="Figure 17" captionStartId="F5" captionText="Figure 17. Paracacoxenus kaszabi Okada, ♂ paratype, Central aimak, Ulan-Baator, Nucht im Bogdo ul, Mongolia (HNHM); decasternum, hypandrium + gonopods, aedeagus, oblique posterior view. Abbreviations: a = proximal section of decasternum; b and c = distal section of decasternum; d = membranous aedeagus; e = anterior region of hypandrium; f = fused gonopods; g = tip of distal section of decasternum (laterally expanded and fused to dorsal region of aedeagus). Scale bar: 0.1 mm." figureDoi="10.3897/alpento.4.49492.figure17" httpUri="https://binary.pensoft.net/fig/387179" pageId="0" pageNumber="1">17g</figureCitation>
,
<figureCitation id="68E6FABF9620848F4E306FB71901A4AD" captionStart="Figure 18" captionStartId="F6" captionText="Figure 18. Paracacoxenus kaszabi Okada, ♂ paratype, Central aimak, Ulan-Baator, Nucht im Bogdo ul, Mongolia (HNHM); decasternum and hypandrium + gonopods. Abbreviations: a = proximal section of decasternum; b and c = distal section of decasternum; d = tip of distal section of decasternum (laterally expanded); e = hypandrium left arm; f = anterior region of gonopods (fused to each other and to hypandrium); g = posterolateral region of left gonopod. Scale bar: 0.1 mm." figureDoi="10.3897/alpento.4.49492.figure18" httpUri="https://binary.pensoft.net/fig/387180" pageId="0" pageNumber="1">18d</figureCitation>
) [widely expanded and marginally crested in
<taxonomicName id="2C6ED9BD14052835CE8020B82007D705" class="Hexapoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="P. macai" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="macai">
<emphasis id="BA995F9E7757E2260D38422D81F10285" italics="true" pageId="0" pageNumber="1">P. macai</emphasis>
</taxonomicName>
sp. nov.] and fused to dorsomedian region of aedeagus. Hypandrium somewhat square-shaped in posterior view (Figs
<figureCitation id="766CBA59AC6FF195ECB4BF778D3151D7" captionStart="Figures 13" captionStartId="F1" captionText="Figures 1 - 3. Paracacoxenus kaszabi Okada, ♂ holotype, Central aimak, Ulan-Baator, Nucht im Bogdo ul, Mongolia (HNHM) 1 Labels (mislabelled as a female) and slide with wing and terminalia 2 Photomicrograph of terminalia, dorsal view 3 Right wing, dorsal view, distally optically cut. Photos: Z. Soltesz." figureDoi="10.3897/alpento.4.49492.figures1-3" httpUri="https://binary.pensoft.net/fig/387175" pageId="0" pageNumber="1">2</figureCitation>
[holotype], 15, 16, 18), as long as epandrium, anterior margin slightly convex and expanded laterad; posterior hypandrium process absent, &quot;dorsal arch&quot; (see
<bibRefCitation id="60FA0669786CADDFFCF72C4756D1EF0A" author="Baechli, G" journalOrPublisher="Brill, Leiden" pageId="0" pageNumber="1" refId="B6" refString="Baechli, G, Vilela, CR, Andersson Escher, S, Saura, A, 2004. The Drosophilidae (Diptera) of Fennoscandia and Denmark. Fauna Entomologica Scandinavica, vol. 39. Brill, Leiden" title="The Drosophilidae (Diptera) of Fennoscandia and Denmark. Fauna Entomologica Scandinavica, vol. 39." year="2004">
<normalizedToken id="CD7D73C264C2DDA3B67B586494E23AF5" originalValue="Bächli">Baechli</normalizedToken>
et al. 2004
</bibRefCitation>
: 14) formed by a complex, two-sectioned decasternum (Figs
<figureCitation id="52229F7A61274F10746FA4EDE265D48B" captionStart="Figure 17" captionStartId="F5" captionText="Figure 17. Paracacoxenus kaszabi Okada, ♂ paratype, Central aimak, Ulan-Baator, Nucht im Bogdo ul, Mongolia (HNHM); decasternum, hypandrium + gonopods, aedeagus, oblique posterior view. Abbreviations: a = proximal section of decasternum; b and c = distal section of decasternum; d = membranous aedeagus; e = anterior region of hypandrium; f = fused gonopods; g = tip of distal section of decasternum (laterally expanded and fused to dorsal region of aedeagus). Scale bar: 0.1 mm." figureDoi="10.3897/alpento.4.49492.figure17" httpUri="https://binary.pensoft.net/fig/387179" pageId="0" pageNumber="1">17a-c</figureCitation>
,
<figureCitation id="87459F63BF2C6975CF8DC3E218143ED3" captionStart="Figures 19, 20" captionStartId="F7" captionText="Figures 19, 20. Paracacoxenus kaszabi Okada, ♂ paratype, Central aimak, Ulan-Baator, Nucht im Bogdo ul, Mongolia (HNHM) 19 Aedeagal apodeme, left inner paraphysis, left outer paraphysis, membranous aedeagus and posterior ejaculatory duct, left lateral view 20 Left surstylus, decasternum, hypandrium, gonopod, aedeagus and posterior ejaculatory duct, lateral view. Scale bar: 0.1 mm." figureDoi="10.3897/alpento.4.49492.figures19-20" httpUri="https://binary.pensoft.net/fig/387181" pageId="0" pageNumber="1">20</figureCitation>
); gonopods fused to each other and to anterior very end of hypandrium, devoid of warts [conspicuously warted in
<taxonomicName id="2671280734CC359A15BCB951D2EB0CF6" class="Hexapoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="P. macai" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="macai">
<emphasis id="1A40C9BD5A4EA9849CF349143B5BD51F" italics="true" pageId="0" pageNumber="1">P. macai</emphasis>
</taxonomicName>
sp. nov.], bearing 0-1 lateral setula (Figs
<figureCitation id="213DB90BFA3E6975675F90FC8D96D8D0" captionStart="Figure 17" captionStartId="F5" captionText="Figure 17. Paracacoxenus kaszabi Okada, ♂ paratype, Central aimak, Ulan-Baator, Nucht im Bogdo ul, Mongolia (HNHM); decasternum, hypandrium + gonopods, aedeagus, oblique posterior view. Abbreviations: a = proximal section of decasternum; b and c = distal section of decasternum; d = membranous aedeagus; e = anterior region of hypandrium; f = fused gonopods; g = tip of distal section of decasternum (laterally expanded and fused to dorsal region of aedeagus). Scale bar: 0.1 mm." figureDoi="10.3897/alpento.4.49492.figure17" httpUri="https://binary.pensoft.net/fig/387179" pageId="0" pageNumber="1">17</figureCitation>
,
<figureCitation id="12F6D1296AD105E81FCAB3D5E1761730" captionStart="Figure 18" captionStartId="F6" captionText="Figure 18. Paracacoxenus kaszabi Okada, ♂ paratype, Central aimak, Ulan-Baator, Nucht im Bogdo ul, Mongolia (HNHM); decasternum and hypandrium + gonopods. Abbreviations: a = proximal section of decasternum; b and c = distal section of decasternum; d = tip of distal section of decasternum (laterally expanded); e = hypandrium left arm; f = anterior region of gonopods (fused to each other and to hypandrium); g = posterolateral region of left gonopod. Scale bar: 0.1 mm." figureDoi="10.3897/alpento.4.49492.figure18" httpUri="https://binary.pensoft.net/fig/387180" pageId="0" pageNumber="1">18</figureCitation>
,
<figureCitation id="179CC877D22839E6D9A75F12503A2382" captionStart="Figures 19, 20" captionStartId="F7" captionText="Figures 19, 20. Paracacoxenus kaszabi Okada, ♂ paratype, Central aimak, Ulan-Baator, Nucht im Bogdo ul, Mongolia (HNHM) 19 Aedeagal apodeme, left inner paraphysis, left outer paraphysis, membranous aedeagus and posterior ejaculatory duct, left lateral view 20 Left surstylus, decasternum, hypandrium, gonopod, aedeagus and posterior ejaculatory duct, lateral view. Scale bar: 0.1 mm." figureDoi="10.3897/alpento.4.49492.figures19-20" httpUri="https://binary.pensoft.net/fig/387181" pageId="0" pageNumber="1">20</figureCitation>
). Aedeagus reduced to a membranous dorsoventrally flattened bag (Fig.
<figureCitation id="FB6C2466CFFF0AAD6B962ACF4071A090" captionStart="Figure 17" captionStartId="F5" captionText="Figure 17. Paracacoxenus kaszabi Okada, ♂ paratype, Central aimak, Ulan-Baator, Nucht im Bogdo ul, Mongolia (HNHM); decasternum, hypandrium + gonopods, aedeagus, oblique posterior view. Abbreviations: a = proximal section of decasternum; b and c = distal section of decasternum; d = membranous aedeagus; e = anterior region of hypandrium; f = fused gonopods; g = tip of distal section of decasternum (laterally expanded and fused to dorsal region of aedeagus). Scale bar: 0.1 mm." figureDoi="10.3897/alpento.4.49492.figure17" httpUri="https://binary.pensoft.net/fig/387179" pageId="0" pageNumber="1">17</figureCitation>
), ellipsoid-shaped [triangle-shaped in
<taxonomicName id="7BD05D8185838EC47764364E22B84AEB" class="Hexapoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="P. macai" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="macai">
<emphasis id="A7A3A97769F4601E8A34EE28A6951263" italics="true" pageId="0" pageNumber="1">P. macai</emphasis>
</taxonomicName>
sp. nov.] in dorsal and ventral views, microtrichose, and dorsally fused to the dorsoventrally flattened end of distal section of decasternum. Two pairs of paraphyses (Figs
<figureCitation id="1EDEBD39CE3EC9D0D43AC6CBEDE022DA" captionStart="Figures 19, 20" captionStartId="F7" captionText="Figures 19, 20. Paracacoxenus kaszabi Okada, ♂ paratype, Central aimak, Ulan-Baator, Nucht im Bogdo ul, Mongolia (HNHM) 19 Aedeagal apodeme, left inner paraphysis, left outer paraphysis, membranous aedeagus and posterior ejaculatory duct, left lateral view 20 Left surstylus, decasternum, hypandrium, gonopod, aedeagus and posterior ejaculatory duct, lateral view. Scale bar: 0.1 mm." figureDoi="10.3897/alpento.4.49492.figures19-20" httpUri="https://binary.pensoft.net/fig/387181" pageId="0" pageNumber="1">19</figureCitation>
,
<figureCitation id="D2C18D2F07375243671048210CAACCED" captionStart="Figures 21, 22" captionStartId="F8" captionText="Figures 21, 22. Paracacoxenus kaszabi Okada, ♂ paratype, Central aimak, Ulan-Baator, Nucht im Bogdo ul, Mongolia (HNHM) 21 Three-branched inner paraphysis, outer paraphysis and aedeagal apodeme, anterior (dorsal) view 22 Three-branched inner paraphysis, outer paraphysis and aedeagal apodeme, oblique posterior view. Abbreviations: a = proximal branch; b = mediodorsal branch; c = distal branch. Scale bar: 0.1 mm." figureDoi="10.3897/alpento.4.49492.figures21-22" httpUri="https://binary.pensoft.net/fig/387182" pageId="0" pageNumber="1">21</figureCitation>
,
<figureCitation id="6FC8E05AC06D3C1CD95FF30F1946AF5F" captionStart="Figures 21, 22" captionStartId="F8" captionText="Figures 21, 22. Paracacoxenus kaszabi Okada, ♂ paratype, Central aimak, Ulan-Baator, Nucht im Bogdo ul, Mongolia (HNHM) 21 Three-branched inner paraphysis, outer paraphysis and aedeagal apodeme, anterior (dorsal) view 22 Three-branched inner paraphysis, outer paraphysis and aedeagal apodeme, oblique posterior view. Abbreviations: a = proximal branch; b = mediodorsal branch; c = distal branch. Scale bar: 0.1 mm." figureDoi="10.3897/alpento.4.49492.figures21-22" httpUri="https://binary.pensoft.net/fig/387182" pageId="0" pageNumber="1">22</figureCitation>
). Outer paraphysis laterally flattened and tube-shaped, subapically enlarged [triangle-shaped in
<taxonomicName id="16A6139E12AB07D8A57FFB83A022823D" class="Hexapoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="P. macai" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="macai">
<emphasis id="3F98914602AAB9F307548BA513EFFDCF" italics="true" pageId="0" pageNumber="1">P. macai</emphasis>
</taxonomicName>
sp. nov.], and turned ventrad in lateral view, with two parallel dorsal rows of ca. 4-5 outer setulae per row, articulated both with lateral inner margin (gonopods) of hypandrium medially, and with posterolateral region of aedeagal apodeme (Fig.
<figureCitation id="7B62C609876EC24F176A5288045F41B1" captionStart="Figures 19, 20" captionStartId="F7" captionText="Figures 19, 20. Paracacoxenus kaszabi Okada, ♂ paratype, Central aimak, Ulan-Baator, Nucht im Bogdo ul, Mongolia (HNHM) 19 Aedeagal apodeme, left inner paraphysis, left outer paraphysis, membranous aedeagus and posterior ejaculatory duct, left lateral view 20 Left surstylus, decasternum, hypandrium, gonopod, aedeagus and posterior ejaculatory duct, lateral view. Scale bar: 0.1 mm." figureDoi="10.3897/alpento.4.49492.figures19-20" httpUri="https://binary.pensoft.net/fig/387181" pageId="0" pageNumber="1">19</figureCitation>
). Inner paraphysis slightly shorter than outer paraphysis (Figs
<figureCitation id="9B153619863410DDD511D3864C6E5422" captionStart="Figures 1012" captionStartId="F3" captionText="Figures 10 - 12. Paracacoxenus kaszabi Okada, ♂ paratype, Central aimak, Ulan-Baator, Nucht im Bogdo ul, Mongolia (HNHM) 10 Terminalia, oblique dorsal view 11 Terminalia, left lateral view 12 Terminalia, oblique ventral view. Scale bar: 0.1 mm." figureDoi="10.3897/alpento.4.49492.figures10-12" httpUri="https://binary.pensoft.net/fig/387177" pageId="0" pageNumber="1">11</figureCitation>
,
<figureCitation id="4388682EBBE2AEC318E3B5BCC9307834" captionStart="Figures 1012" captionStartId="F3" captionText="Figures 10 - 12. Paracacoxenus kaszabi Okada, ♂ paratype, Central aimak, Ulan-Baator, Nucht im Bogdo ul, Mongolia (HNHM) 10 Terminalia, oblique dorsal view 11 Terminalia, left lateral view 12 Terminalia, oblique ventral view. Scale bar: 0.1 mm." figureDoi="10.3897/alpento.4.49492.figures10-12" httpUri="https://binary.pensoft.net/fig/387177" pageId="0" pageNumber="1">12</figureCitation>
), strongly sclerotized, bare, mediolaterally fused to aedeagal apodeme, tribranched (Figs
<figureCitation id="AAAA80BFD66AC9375D38025F92B34DF7" captionStart="Figures 21, 22" captionStartId="F8" captionText="Figures 21, 22. Paracacoxenus kaszabi Okada, ♂ paratype, Central aimak, Ulan-Baator, Nucht im Bogdo ul, Mongolia (HNHM) 21 Three-branched inner paraphysis, outer paraphysis and aedeagal apodeme, anterior (dorsal) view 22 Three-branched inner paraphysis, outer paraphysis and aedeagal apodeme, oblique posterior view. Abbreviations: a = proximal branch; b = mediodorsal branch; c = distal branch. Scale bar: 0.1 mm." figureDoi="10.3897/alpento.4.49492.figures21-22" httpUri="https://binary.pensoft.net/fig/387182" pageId="0" pageNumber="1">21a-c</figureCitation>
,
<figureCitation id="E6AE7D104B274DACF9CDDDE1B9C9878A" captionStart="Figures 21, 22" captionStartId="F8" captionText="Figures 21, 22. Paracacoxenus kaszabi Okada, ♂ paratype, Central aimak, Ulan-Baator, Nucht im Bogdo ul, Mongolia (HNHM) 21 Three-branched inner paraphysis, outer paraphysis and aedeagal apodeme, anterior (dorsal) view 22 Three-branched inner paraphysis, outer paraphysis and aedeagal apodeme, oblique posterior view. Abbreviations: a = proximal branch; b = mediodorsal branch; c = distal branch. Scale bar: 0.1 mm." figureDoi="10.3897/alpento.4.49492.figures21-22" httpUri="https://binary.pensoft.net/fig/387182" pageId="0" pageNumber="1">22a-c</figureCitation>
) [heptabranched in
<taxonomicName id="FD82EDD3FB2A721FBE2E40F0010305A2" class="Hexapoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="P. macai" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="macai">
<emphasis id="A7C0765814EABA48FBBBE9DCBE8DA52B" italics="true" pageId="0" pageNumber="1">P. macai</emphasis>
</taxonomicName>
sp. nov.]: distal branch long, apically sharp, slightly turned dorsad and inwards, submediodorsal branch wide, triangle-shaped, pointed laterad and dorsad, proximal branches somewhat rectangle-shaped, blunt, pointed dorsalwards and distally connected to each other by a membranous strip embracing submedian region of distal section of decasternum (Figs
<figureCitation id="2FD7BD559E9A5AA2A3B89B89D645918B" captionStart="Figures 1012" captionStartId="F3" captionText="Figures 10 - 12. Paracacoxenus kaszabi Okada, ♂ paratype, Central aimak, Ulan-Baator, Nucht im Bogdo ul, Mongolia (HNHM) 10 Terminalia, oblique dorsal view 11 Terminalia, left lateral view 12 Terminalia, oblique ventral view. Scale bar: 0.1 mm." figureDoi="10.3897/alpento.4.49492.figures10-12" httpUri="https://binary.pensoft.net/fig/387177" pageId="0" pageNumber="1">10</figureCitation>
,
<figureCitation id="1391F91B80EDE51796C99E3B8C1A0A30" captionStart="Figures 21, 22" captionStartId="F8" captionText="Figures 21, 22. Paracacoxenus kaszabi Okada, ♂ paratype, Central aimak, Ulan-Baator, Nucht im Bogdo ul, Mongolia (HNHM) 21 Three-branched inner paraphysis, outer paraphysis and aedeagal apodeme, anterior (dorsal) view 22 Three-branched inner paraphysis, outer paraphysis and aedeagal apodeme, oblique posterior view. Abbreviations: a = proximal branch; b = mediodorsal branch; c = distal branch. Scale bar: 0.1 mm." figureDoi="10.3897/alpento.4.49492.figures21-22" httpUri="https://binary.pensoft.net/fig/387182" pageId="0" pageNumber="1">21</figureCitation>
). Aedeagal apodeme shorter than outer paraphysis (Fig.
<figureCitation id="7547FC4BF0968B9DF2A787C698A36FE5" captionStart="Figures 19, 20" captionStartId="F7" captionText="Figures 19, 20. Paracacoxenus kaszabi Okada, ♂ paratype, Central aimak, Ulan-Baator, Nucht im Bogdo ul, Mongolia (HNHM) 19 Aedeagal apodeme, left inner paraphysis, left outer paraphysis, membranous aedeagus and posterior ejaculatory duct, left lateral view 20 Left surstylus, decasternum, hypandrium, gonopod, aedeagus and posterior ejaculatory duct, lateral view. Scale bar: 0.1 mm." figureDoi="10.3897/alpento.4.49492.figures19-20" httpUri="https://binary.pensoft.net/fig/387181" pageId="0" pageNumber="1">19</figureCitation>
), dorsoventrally flattened, strongly sclerotized, laterally fused to anterior region of inner paraphysis (Fig.
<figureCitation id="877FB8A1549C75256B4537938FD4C408" captionStart="Figures 21, 22" captionStartId="F8" captionText="Figures 21, 22. Paracacoxenus kaszabi Okada, ♂ paratype, Central aimak, Ulan-Baator, Nucht im Bogdo ul, Mongolia (HNHM) 21 Three-branched inner paraphysis, outer paraphysis and aedeagal apodeme, anterior (dorsal) view 22 Three-branched inner paraphysis, outer paraphysis and aedeagal apodeme, oblique posterior view. Abbreviations: a = proximal branch; b = mediodorsal branch; c = distal branch. Scale bar: 0.1 mm." figureDoi="10.3897/alpento.4.49492.figures21-22" httpUri="https://binary.pensoft.net/fig/387182" pageId="0" pageNumber="1">21</figureCitation>
). Ventral rod apparently absent, probably turned backwards and fused to posteroventral region of aedeagal apodeme (Fig.
<figureCitation id="3B374B67EA7C44F8DD7676C6CBA7D484" captionStart="Figures 1012" captionStartId="F3" captionText="Figures 10 - 12. Paracacoxenus kaszabi Okada, ♂ paratype, Central aimak, Ulan-Baator, Nucht im Bogdo ul, Mongolia (HNHM) 10 Terminalia, oblique dorsal view 11 Terminalia, left lateral view 12 Terminalia, oblique ventral view. Scale bar: 0.1 mm." figureDoi="10.3897/alpento.4.49492.figures10-12" httpUri="https://binary.pensoft.net/fig/387177" pageId="0" pageNumber="1">11</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="3D332D242BA510BDBA2DA150F9F6E6D1" pageId="0" pageNumber="1" type="distribution">
<paragraph id="CC378109863A110168424C069855670F" pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph id="B8D431BF7C48F10F1D0C9C6B399EAC42" pageId="0" pageNumber="1">
Mongolia:
<bibRefCitation id="988CE954B7DB9D3AB9482B2D605440EE" author="Okada, T" journalOrPublisher="Annales Historico-Naturales Musei Nationalis Hungarici" pageId="0" pageNumber="1" pagination="271 - 279" refId="B22" refString="Okada, T, 1973. Drosophilidae and Diastatidae from Mongolia (Diptera). Annales Historico-Naturales Musei Nationalis Hungarici 65: 271 - 279" title="Drosophilidae and Diastatidae from Mongolia (Diptera)." volume="65" year="1973">Okada (1973)</bibRefCitation>
: 271. Russia, Siberia:
<bibRefCitation id="D77D1A6945937108ACF4B67850080591" author="Toda, MJ" editor="Takahashi, K" journalOrPublisher="Forestry and Forest Products Research Institute, Sapporo" pageId="0" pageNumber="1" pagination="187 - 195" refId="B25" refString="Toda, MJ, Vinokurov, NN, 1995. Biodiversity in drosophilid communities of cool-temperate and boreal birch forests: a special reference to the vertical distribution within forest. In: Takahashi, K, Osawa, A, Kanazawa, Y, Eds., Proceedings of the Third Symposium on the Joint Siberian Permafrost Studies between Japan and Russia. Forestry and Forest Products Research Institute, Sapporo: 187 - 195" title="Biodiversity in drosophilid communities of cool-temperate and boreal birch forests: a special reference to the vertical distribution within forest." volumeTitle="Proceedings of the Third Symposium on the Joint Siberian Permafrost Studies between Japan and Russia." year="1995">Toda and Vinokurov (1995)</bibRefCitation>
: 189;
<bibRefCitation id="DCE340EBF2AAB0D6C2399D99776FF0CA" author="Watabe, HA" editor="Takahashi, K" journalOrPublisher="Forestry and Forest Products Research Institute, Sapporo" pageId="0" pageNumber="1" pagination="196 - 200" refId="B31" refString="Watabe, HA, Toda, MJ, Vinokurov, NN, Sidorenko, VS, 1995. A comparative study on drosophilid faunas of east Siberia and neighboring regions. In: Takahashi, K, Osawa, A, Kanazawa, Y, Eds., Proceedings of the Third Symposium on the Joint Siberian Permafrost Studies between Japan and Russia. Forestry and Forest Products Research Institute, Sapporo: 196 - 200" title="A comparative study on drosophilid faunas of east Siberia and neighboring regions." volumeTitle="Proceedings of the Third Symposium on the Joint Siberian Permafrost Studies between Japan and Russia." year="1995">Watabe et al. (1995)</bibRefCitation>
: 199;
<bibRefCitation id="E80138E9BE9B4CDAF47B1B9EFBC82720" author="Vinokurova, AV" editor="Isaev, AP" journalOrPublisher="IBPK SO RAN, Yakutsk" pageId="0" pageNumber="1" pagination="99 - 106" refId="B30" refString="Vinokurova, AV, 2003. Sostav phauni mukh-drosophilid (Diptera, Drosophilidae) Yakutii [Overal view of the fauna of drosophild flies (Diptera, Drosophilidae) in Yakutia]. In: Isaev, AP, Ed., Entomologiceskie issledovanija v Jakutii: Sbornik naucnych trudov [Entomological investigation in Yakutiya: Collection of scientific papers]. IBPK SO RAN, Yakutsk: 99 - 106" title="Sostav phauni mukh-drosophilid (Diptera, Drosophilidae) Yakutii [Overal view of the fauna of drosophild flies (Diptera, Drosophilidae) in Yakutia]." volumeTitle="Entomologiceskie issledovanija v Jakutii: Sbornik naucnych trudov [Entomological investigation in Yakutiya: Collection of scientific papers]." year="2003">Vinokurova (2003)</bibRefCitation>
: 100.
</paragraph>
<caption id="B57CD1147DA52C799CD008086738A95C" ID-Zenodo-Dep="3710621" doi="10.3897/alpento.4.49492.figures13-16" httpUri="https://binary.pensoft.net/fig/387178" pageId="0" pageNumber="1" start="Figures 1316" startId="F4">
<paragraph id="A4630A646BBE05A226237AF07DCAA2B4" pageId="0" pageNumber="1">
<emphasis id="A306778F5EFE644CA8EB5DBEEFC9DDEE" bold="true" pageId="0" pageNumber="1">Figures 13-16.</emphasis>
<taxonomicName id="A54C5F72DB96A511B55CAD08C0AE895A" authorityName="Okada" authorityYear="1973" class="Insecta" family="Drosophilidae" genus="Paracacoxenus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paracacoxenus kaszabi" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="kaszabi">
<emphasis id="A0BDD54EFC43F9F5C2DEC1E18F19CFB5" italics="true" pageId="0" pageNumber="1">Paracacoxenus kaszabi</emphasis>
</taxonomicName>
Okada, ♂ paratype, Central aimak, Ulan-Baator, Nucht im Bogdo ul, Mongolia (HNHM)
<emphasis id="689841269A6032AD7DEF07FC7C7BEA76" bold="true" pageId="0" pageNumber="1">13</emphasis>
Terminalia, dorsal view (in-focus: epandrium, cerci, tips of aedeagus and inner paraphyses)
<emphasis id="3CEE5AFFB4E10674C35B006871167B7D" bold="true" pageId="0" pageNumber="1">14</emphasis>
Terminalia, dorsal view (in-focus: all sclerites)
<emphasis id="AB9B3E927BE2C03D02E78790EA1865A6" bold="true" pageId="0" pageNumber="1">15</emphasis>
Terminalia, posterior view
<emphasis id="81D4E39F0CDDFCE393ACC1AFF2C406ED" bold="true" pageId="0" pageNumber="1">16</emphasis>
Terminalia, anterior view. Scale bar: 0.1 mm.
</paragraph>
</caption>
<caption id="6E23264401DC51258559A02D67D11356" ID-Zenodo-Dep="3710623" doi="10.3897/alpento.4.49492.figure17" httpUri="https://binary.pensoft.net/fig/387179" pageId="0" pageNumber="1" start="Figure 17" startId="F5">
<paragraph id="C3D7D68403E8504778567AC20B0DBB79" pageId="0" pageNumber="1">
<emphasis id="F0BDA6FEAD96CCE783CCDF6528C40C24" bold="true" pageId="0" pageNumber="1">Figure 17.</emphasis>
<taxonomicName id="AF4ECD272814DA2B5F1B43FAFB2225BC" authorityName="Okada" authorityYear="1973" class="Insecta" family="Drosophilidae" genus="Paracacoxenus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paracacoxenus kaszabi" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="kaszabi">
<emphasis id="6CFAF300214AF31144DCE70559953698" italics="true" pageId="0" pageNumber="1">Paracacoxenus kaszabi</emphasis>
</taxonomicName>
Okada, ♂ paratype, Central aimak, Ulan-Baator, Nucht im Bogdo ul, Mongolia (HNHM); decasternum, hypandrium + gonopods, aedeagus, oblique posterior view. Abbreviations: a = proximal section of decasternum; b and c = distal section of decasternum; d = membranous aedeagus; e = anterior region of hypandrium; f = fused gonopods; g = tip of distal section of decasternum (laterally expanded and fused to dorsal region of aedeagus). Scale bar: 0.1 mm.
</paragraph>
</caption>
<caption id="0F39494E75CACE007C27145C8E571361" ID-Zenodo-Dep="3710625" doi="10.3897/alpento.4.49492.figure18" httpUri="https://binary.pensoft.net/fig/387180" pageId="0" pageNumber="1" start="Figure 18" startId="F6">
<paragraph id="88C11933BEB161BD627D8863569FDC53" pageId="0" pageNumber="1">
<emphasis id="C64A37D8D6740F48CBA64E6839DF091D" bold="true" pageId="0" pageNumber="1">Figure 18.</emphasis>
<taxonomicName id="2E86B8FA38DA290F81AE6540D9F79C23" authorityName="Okada" authorityYear="1973" class="Insecta" family="Drosophilidae" genus="Paracacoxenus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paracacoxenus kaszabi" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="kaszabi">
<emphasis id="245044EE5A2DB52A3B816A5D59DEBE68" italics="true" pageId="0" pageNumber="1">Paracacoxenus kaszabi</emphasis>
</taxonomicName>
Okada, ♂ paratype, Central aimak, Ulan-Baator, Nucht im Bogdo ul, Mongolia (HNHM); decasternum and hypandrium + gonopods. Abbreviations: a = proximal section of decasternum; b and c = distal section of decasternum; d = tip of distal section of decasternum (laterally expanded); e = hypandrium left arm; f = anterior region of gonopods (fused to each other and to hypandrium); g = posterolateral region of left gonopod. Scale bar: 0.1 mm.
</paragraph>
</caption>
<caption id="1B91176CACFADFD2A3CC0FCE094F5450" ID-Zenodo-Dep="3710627" doi="10.3897/alpento.4.49492.figures19-20" httpUri="https://binary.pensoft.net/fig/387181" pageId="0" pageNumber="1" start="Figures 19, 20" startId="F7">
<paragraph id="6DF4B5C53665875E47EE1BA1DAA08BD6" pageId="0" pageNumber="1">
<emphasis id="8E83918D174D792502E052B3761435AB" bold="true" pageId="0" pageNumber="1">Figures 19, 20.</emphasis>
<taxonomicName id="7EC24D13B351EC33D80E5623871AF8AF" authorityName="Okada" authorityYear="1973" class="Insecta" family="Drosophilidae" genus="Paracacoxenus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paracacoxenus kaszabi" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="kaszabi">
<emphasis id="EF7053F3BE16985B86F08284A7E356E2" italics="true" pageId="0" pageNumber="1">Paracacoxenus kaszabi</emphasis>
</taxonomicName>
Okada, ♂ paratype, Central aimak, Ulan-Baator, Nucht im Bogdo ul, Mongolia (HNHM)
<emphasis id="9667341F21125B91069EB4B4D4225C5C" bold="true" pageId="0" pageNumber="1">19</emphasis>
Aedeagal apodeme, left inner paraphysis, left outer paraphysis, membranous aedeagus and posterior ejaculatory duct, left lateral view
<emphasis id="FE08A3F63E45E92F2980F09013537E16" bold="true" pageId="0" pageNumber="1">20</emphasis>
Left surstylus, decasternum, hypandrium, gonopod, aedeagus and posterior ejaculatory duct, lateral view. Scale bar: 0.1 mm.
</paragraph>
</caption>
<caption id="37216D506DDA3805762DDEBF0CDEF347" ID-Zenodo-Dep="3710629" doi="10.3897/alpento.4.49492.figures21-22" httpUri="https://binary.pensoft.net/fig/387182" pageId="0" pageNumber="1" start="Figures 21, 22" startId="F8">
<paragraph id="84E78AD48F17F65DD1BCE992BA9D5138" pageId="0" pageNumber="1">
<emphasis id="8CF012531BAEEC95F9697E88090E2B0D" bold="true" pageId="0" pageNumber="1">Figures 21, 22.</emphasis>
<taxonomicName id="51D638CD282051E35C78CE13F3264FE1" authorityName="Okada" authorityYear="1973" class="Insecta" family="Drosophilidae" genus="Paracacoxenus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paracacoxenus kaszabi" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="kaszabi">
<emphasis id="AD327522A282680F9FC81DA519A450DE" italics="true" pageId="0" pageNumber="1">Paracacoxenus kaszabi</emphasis>
</taxonomicName>
Okada, ♂ paratype, Central aimak, Ulan-Baator, Nucht im Bogdo ul, Mongolia (HNHM)
<emphasis id="CA265929E41AA86EB2475B987A3D0B4B" bold="true" pageId="0" pageNumber="1">21</emphasis>
Three-branched inner paraphysis, outer paraphysis and aedeagal apodeme, anterior (dorsal) view
<emphasis id="D9870F9D508D031A2965013DA56AD1C6" bold="true" pageId="0" pageNumber="1">22</emphasis>
Three-branched inner paraphysis, outer paraphysis and aedeagal apodeme, oblique posterior view. Abbreviations: a = proximal branch; b = mediodorsal branch; c = distal branch. Scale bar: 0.1 mm.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>