treatments-xml/data/96/2D/87/962D87B3FFB39C466383FC284B34FAA9.xml

<document id="129084D9A50B93032329DDF5B6E193B7" ID-DOI="10.5281/zenodo.5277501" ID-ISSN="0253-116X" ID-Zenodo-Dep="5277501" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629969536188" checkinUser="marcus" docAuthor="Esser, Jens &amp; R, Afrotropical" docDate="2019" docId="962D87B3FFB39C466383FC284B34FAA9" docLanguage="en" docName="LinzerbiolBeitr.2019.51.2.915-928.pdf" docOrigin="Linzer biologische Beiträge 51 (2)" docStyle="DocumentStyle:24E5EB772D355DE2EE23B1E8D07853E0.9:LinzerbiolBeitr.2006-.journal_article" docStyleId="24E5EB772D355DE2EE23B1E8D07853E0" docStyleName="LinzerbiolBeitr.2006-.journal_article" docStyleVersion="9" docTitle="Mnioticus gugheanus Esser &amp; R 2019, nov.sp." docType="treatment" docVersion="3" lastPageNumber="918" masterDocId="6A14FFCBFFB19C456314FF814852FF90" masterDocTitle="Revision of Mnioticus COOMBS &amp; WOODROFFE, 1962 (Coleoptera, Cryptophagidae)" masterLastPageNumber="928" masterPageNumber="915" pageNumber="917" updateTime="1699132627152" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="8582E1C5AF77D62AAD66CF1EC31EE977">Revision of Mnioticus COOMBS &amp; WOODROFFE, 1962 (Coleoptera, Cryptophagidae)</mods:title>
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<taxonomicName id="D9844D26FFB39C476383FC284928FC2F" authority="Esser &amp; R, 2019" authorityName="Esser &amp; R" authorityYear="2019" box="[151,378,937,959]" class="Insecta" family="Cryptophagidae" genus="Mnioticus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="917" phylum="Arthropoda" rank="species" species="gugheanus" status="sp. nov.">
<emphasis id="2CF0EAB7FFB39C476383FC284928FC2F" bold="true" box="[151,378,937,959]" italics="true" pageId="2" pageNumber="917">Mnioticus gugheanus</emphasis>
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(
<figureCitation id="86BF2A20FFB39C4762CFFC284A4CFC2F" box="[475,542,937,959]" captionStart="Figs 9-12" captionStartId="12.[151,191,1483,1505]" captionTargetBox="[159,1029,205,1437]" captionTargetId="figure-41@12.[153,1061,196,1465]" captionTargetPageId="12" captionText="Figs 9-12: (9) Mnioticus holmi nov.sp., dark specimen, paratype, female; (10) Mnioticus karisimbianus nov.sp., paratype, female; (11) Mnioticus gugheanus nov.sp. Gughe Mts., paratype, male; (12) Mnioticus kinangopianus nov.sp. Aberdare, paratype, female." figureDoi="http://doi.org/10.5281/zenodo.5277509" httpUri="https://zenodo.org/record/5277509/files/figure.png" pageId="2" pageNumber="917">fig. 11</figureCitation>
)
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</paragraph>
<paragraph id="1E3B36A5FFB39C476383FC4E4B77FBCC" blockId="2.[151,1059,975,1117]" pageId="2" pageNumber="917">
<materialsCitation id="AEEC3CF8FFB39C476383FC4E4AC3FB84" collectingDate="1948-12-22" collectionCode="BMNH" country="Ethiopia" county="Lobelia" elevation="3200" location="Gamo Prov." municipality="Mt. Gughe" pageId="2" pageNumber="917" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
T y p e m a t e r i a l: 
<typeStatus id="C13F8807FFB39C476288FC4E49BFFC75" box="[412,493,975,997]" pageId="2" pageNumber="917" type="holotype">Holotype</typeStatus>
<specimenCount id="0882FD2CFFB39C4762E8FC4E4A42FC74" box="[508,528,975,996]" pageId="2" pageNumber="917" type="male">♂</specimenCount>
: &quot; 
<collectingCountry id="66937635FFB39C47613DFC4E4A2AFC75" box="[553,632,975,997]" name="Ethiopia" pageId="2" pageNumber="917">Ethiopia</collectingCountry>
: 1948-1949, Hugh Scott, B.M. 1949-184 / Abyssinia: 
<location id="1B5B607EFFB39C47623EFC6649C8FC6C" LSID="urn:lsid:plazi:treatment:962D87B3FFB39C466383FC284B34FAA9:1B5B607EFFB39C47623EFC6649C8FC6C" box="[298,410,999,1020]" country="Ethiopia" county="Lobelia" municipality="Mt. Gughe" name="Gamo Prov." pageId="2" pageNumber="917">Gamo Prov.</location>
, 
<collectingMunicipality id="FE5FACDFFFB39C4762B1FC664A5BFC6D" box="[421,521,999,1021]" pageId="2" pageNumber="917">Mt. Gughé</collectingMunicipality>
, ca. 
<quantity id="D97C9B40FFB39C476123FC664AD9FC6D" box="[567,651,999,1021]" metricMagnitude="3" metricUnit="m" metricValue="3.2004" pageId="2" pageNumber="917" unit="ft" value="10500.0">
<elevation id="95A9D196FFB39C476123FC664AD9FC6D" box="[567,651,999,1021]" metricMagnitude="3" metricUnit="m" metricValue="3.2004" pageId="2" pageNumber="917" unit="ft" value="10500.0">10,500 ft</elevation>
</quantity>
, 
<date id="6A3A1065FFB39C476183FC664B50FC6C" box="[663,770,999,1020]" pageId="2" pageNumber="917" value="1948-12-22">
<collectingDate id="7A7EE98DFFB39C476183FC664B50FC6C" box="[663,770,999,1020]" pageId="2" pageNumber="917" value="1948-12-22">22.XII.1948</collectingDate>
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/ from dying stem and fruiting inflorescence of 
<collectingCounty id="F75A4E29FFB39C47624CFC7E49C9FB85" box="[344,411,1023,1045]" pageId="2" pageNumber="917">Lobelia</collectingCounty>
rhynchopetalum&quot; [
<collectionCode id="7895AE60FFB39C476157FC7F4ADEFB84" box="[579,652,1022,1044]" pageId="2" pageNumber="917">BMNH</collectionCode>
]
</materialsCitation>
. 
<materialsCitation id="AEEC3CF8FFB39C47618FFC7E498AFBBC" collectingDate="1948-12-22" collectionCode="BMNH" country="Ethiopia" county="Lobelia" elevation="3200" location="Gamo Prov." municipality="Mt. Gughe" pageId="2" pageNumber="917" specimenCount="37" typeStatus="paratype">
<typeStatus id="C13F8807FFB39C47618FFC7E4AACFB84" box="[667,766,1023,1044]" pageId="2" pageNumber="917" type="paratype">Paratypes</typeStatus>
: 
<specimenCount id="0882FD2CFFB39C47601CFC7E4B14FB84" box="[776,838,1023,1044]" pageId="2" pageNumber="917" type="generic">37 ex.</specimenCount>
with the same data as the holotype [
<collectionCode id="7895AE60FFB39C476258FB9649C9FBBC" box="[332,411,1047,1068]" pageId="2" pageNumber="917">BMNH</collectionCode>
, cES]
</materialsCitation>
; 
<materialsCitation id="AEEC3CF8FFB39C4762F0FB964B73FBCC" collectingDate="1948-12-21" collectionCode="BMNH" country="Ethiopia" elevation="3353" location="Gamo Prov." municipality="Mt. Gughe" pageId="2" pageNumber="917" specimenCount="6" typeStatus="paratype">
<specimenCount id="0882FD2CFFB39C4762F0FB964A47FBBC" box="[484,533,1047,1069]" pageId="2" pageNumber="917" type="generic">6 ex.</specimenCount>
&quot; 
<collectingCountry id="66937635FFB39C476130FB964A2EFBBD" box="[548,636,1047,1069]" name="Ethiopia" pageId="2" pageNumber="917">Ethiopia</collectingCountry>
: 1948-1949, Hugh Scott, B.M. 1949-184 / Abyssinia: 
<location id="1B5B607EFFB39C47622CFBAE49E5FBD4" LSID="urn:lsid:plazi:treatment:962D87B3FFB39C466383FC284B34FAA9:1B5B607EFFB39C47622CFBAE49E5FBD4" box="[312,439,1071,1092]" country="Ethiopia" municipality="Mt. Gughe" name="Gamo Prov." pageId="2" pageNumber="917">Gamo Prov.</location>
, 
<collectingMunicipality id="FE5FACDFFFB39C4762D0FBAE4A64FBD5" box="[452,566,1071,1093]" pageId="2" pageNumber="917">Mt. Gughé</collectingMunicipality>
, over 
<quantity id="D97C9B40FFB39C47616FFBAE4A88FBD5" box="[635,730,1071,1093]" metricMagnitude="3" metricUnit="m" metricValue="3.3528" pageId="2" pageNumber="917" unit="ft" value="11000.0">
<elevation id="95A9D196FFB39C47616FFBAE4A88FBD5" box="[635,730,1071,1093]" metricMagnitude="3" metricUnit="m" metricValue="3.3528" pageId="2" pageNumber="917" unit="ft" value="11000.0">11,000 ft</elevation>
</quantity>
, 
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<collectingDate id="7A7EE98DFFB39C4761FCFBAE4B0DFBD4" box="[744,863,1071,1092]" pageId="2" pageNumber="917" value="1948-12-21">21.XII.1948</collectingDate>
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/ among decaying stems and leaves of Kniphofia on south-west slope&quot; [
<collectionCode id="7895AE60FFB39C4761DDFBC74B49FBCC" box="[713,795,1094,1116]" pageId="2" pageNumber="917">BMNH</collectionCode>
]
</materialsCitation>
.
</paragraph>
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E t y m o l o g y: Named after the Mt. Gughé (
<collectingCountry id="66937635FFB39C47619BFBEA4AB8FB11" box="[655,746,1131,1153]" name="Ethiopia" pageId="2" pageNumber="917">Ethiopia</collectingCountry>
), where the 
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material was collected.
</paragraph>
<paragraph id="1E3B36A5FFB39C476383FB2E4B65FA79" blockId="2.[151,1059,1198,1513]" pageId="2" pageNumber="917">
D e s c r i p t i o n: Male, 
<quantity id="D97C9B40FFB39C4762A8FB2F4A5EFB54" box="[444,524,1198,1220]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="2" pageNumber="917" unit="mm" value="1.9">1.9 mm</quantity>
, reddish-brown, shiny, elytra with a semi-circular black spot (except suture, base and apex). Membranous wings absent. Legs and antennae slender, tarsi 5-segmented, hind tarsi 4-segmented (in female 5-segmented). Antennae 11-segmented with a 3-segmented club. Segments 4 to 8 more or less globular, segments 1 to 3 slightly longer than broad. Segments 9 and 10 moderately transverse, 11 dropshaped and asymmetrical. Eyes large with fine ommatidia, puncture on head fine and sparse. Pronotum also with fine and sparse puncture. Pronotum convex and moderately transverse (1.5 times broader than long), with sides rounded, more narrowed to base in the posterior third and minute serration. Basal furrow flat, basal fovea deep. Anterior margin slightly concave, posterior margin slightly convex. Elytra elongate, puncture stronger and less sparse than on pronotum. Humeral callus absent.
</paragraph>
<paragraph id="1E3B36A5FFB39C466383FA784957FECE" blockId="2.[151,1059,1529,1580]" lastBlockId="3.[151,1058,211,350]" lastPageId="3" lastPageNumber="918" pageId="2" pageNumber="917">
C o m m e n t: Resembles in general shape to the most species except 
<taxonomicName id="D9844D26FFB39C4760BDFA784977F9BC" authority="(SCOTT, 1936)" baseAuthorityName="SCOTT" baseAuthorityYear="1936" class="Insecta" family="Cryptophagidae" genus="Mnioticus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="917" phylum="Arthropoda" rank="species" species="gibbinsi">
<emphasis id="2CF0EAB7FFB39C4760BDFA784C70F99E" box="[937,1058,1529,1550]" italics="true" pageId="2" pageNumber="917">M. gibbinsi</emphasis>
(
<bibRefCitation id="7A154B54FFB39C47638BF997494CF9BC" author="SCOTT H" box="[159,286,1558,1580]" pageId="2" pageNumber="917" pagination="235 - 245" refId="ref6631" refString="SCOTT H. (1936): Coleoptera associated with the giant Lobelias and arborescent Senecios of eastern Africa. - Journal of the Linnean Society London 39: 235 - 245; 252 - 281." type="journal article" year="1936">SCOTT, 1936</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicName id="D9844D26FFB39C47623AF9964A68F9BC" authority="(SCOTT, 1936)" baseAuthorityName="SCOTT" baseAuthorityYear="1936" box="[302,570,1558,1580]" class="Insecta" family="Cryptophagidae" genus="Mnioticus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="917" phylum="Arthropoda" rank="species" species="hancocki">
<emphasis id="2CF0EAB7FFB39C47623AF99649F5F9BB" box="[302,423,1558,1580]" italics="true" pageId="2" pageNumber="917">M. hancocki</emphasis>
(
<bibRefCitation id="7A154B54FFB39C4762A0F9974A61F9BC" author="SCOTT H" box="[436,563,1558,1580]" pageId="2" pageNumber="917" pagination="235 - 245" refId="ref6631" refString="SCOTT H. (1936): Coleoptera associated with the giant Lobelias and arborescent Senecios of eastern Africa. - Journal of the Linnean Society London 39: 235 - 245; 252 - 281." type="journal article" year="1936">SCOTT, 1936</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicName id="D9844D26FFB39C476150F9964B16F9BC" authority="(SCOTT, 1936)" baseAuthorityName="SCOTT" baseAuthorityYear="1936" box="[580,836,1558,1580]" class="Insecta" family="Cryptophagidae" genus="Mnioticus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="917" phylum="Arthropoda" rank="species" species="jeanneli">
<emphasis id="2CF0EAB7FFB39C476150F9964AE1F9BB" box="[580,691,1558,1580]" italics="true" pageId="2" pageNumber="917">M. jeanneli</emphasis>
(
<bibRefCitation id="7A154B54FFB39C4761D4F9974B6FF9BC" author="SCOTT H" box="[704,829,1558,1580]" pageId="2" pageNumber="917" pagination="235 - 245" refId="ref6631" refString="SCOTT H. (1936): Coleoptera associated with the giant Lobelias and arborescent Senecios of eastern Africa. - Journal of the Linnean Society London 39: 235 - 245; 252 - 281." type="journal article" year="1936">SCOTT, 1936</bibRefCitation>
)
</taxonomicName>
and 
<taxonomicName id="D9844D26FFB39C476067F9964C71F9BB" authority="Esser &amp; R, 2019" authorityName="Esser &amp; R" authorityYear="2019" box="[883,1059,1558,1580]" class="Insecta" family="Cryptophagidae" genus="Mnioticus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="917" phylum="Arthropoda" rank="species" species="kinangopianus" status="sp. nov.">
<emphasis id="2CF0EAB7FFB39C476067F9964C71F9BB" box="[883,1059,1558,1580]" italics="true" pageId="2" pageNumber="917">M. kinangopianus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="37C357CCFFB29C466383FF5548B3FF79" box="[151,225,212,233]" pageId="3" pageNumber="918" rank="species">nov.sp.</taxonomicNameLabel>
but normally separated by its dark spots on elytra (spot is of variable in size but well circumscribed). The also Ethiopian species 
<taxonomicName id="D9844D26FFB29C46619EFF704B81FE97" authority="(SCOTT, 1936)" baseAuthorityName="SCOTT" baseAuthorityYear="1936" box="[650,979,241,263]" class="Insecta" family="Cryptophagidae" genus="Mnioticus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="918" phylum="Arthropoda" rank="species" species="rhynchopetali">
<emphasis id="2CF0EAB7FFB29C46619EFF704B6BFE96" box="[650,825,241,262]" italics="true" pageId="3" pageNumber="918">M. rhynchopetali</emphasis>
(
<bibRefCitation id="7A154B54FFB29C46605DFF704B9EFE97" author="SCOTT H" box="[841,972,241,263]" pageId="3" pageNumber="918" pagination="235 - 245" refId="ref6631" refString="SCOTT H. (1936): Coleoptera associated with the giant Lobelias and arborescent Senecios of eastern Africa. - Journal of the Linnean Society London 39: 235 - 245; 252 - 281." type="journal article" year="1936">SCOTT, 1936</bibRefCitation>
)
</taxonomicName>
(
<figureCitation id="86BF2A20FFB29C4660F6FF704C48FE97" box="[994,1050,241,263]" captionStart="Figs 1-4" captionStartId="10.[151,191,1517,1539]" captionTargetBox="[161,1061,211,1506]" captionTargetId="figure-46@10.[153,1061,197,1507]" captionTargetPageId="10" captionText="Figs 1-4: (1) Mnioticus gibbinsi (SCOTT, 1936), male; (2) Mnioticus hancocki (SCOTT, 1936), male; (3) Mnioticus rhynchopetali (SCOTT, 1936), holotype, male; (4) Mnioticus rougemonti ESSER, 2014, paratype, male." figureDoi="http://doi.org/10.5281/zenodo.5277503" httpUri="https://zenodo.org/record/5277503/files/figure.png" pageId="3" pageNumber="918">fig. 3</figureCitation>
) differs by its longer legs and more slender antennae. 
<taxonomicName id="D9844D26FFB29C4661A6FE8E4B9CFEB3" authority="ESSER, 2014" authorityName="ESSER" authorityYear="2014" box="[690,974,270,292]" class="Insecta" family="Cryptophagidae" genus="Mnioticus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="918" phylum="Arthropoda" rank="species" species="rougemonti">
<emphasis id="2CF0EAB7FFB29C4661A6FE8E4B1BFEB4" box="[690,841,271,292]" italics="true" pageId="3" pageNumber="918">M. rougemonti</emphasis>
<bibRefCitation id="7A154B54FFB29C466044FE8E4B9CFEB3" author="ESSER J" box="[848,974,270,292]" pageId="3" pageNumber="918" pagination="247 - 250" refId="ref6498" refString="ESSER J. (2014): Mnioticus rougemonti nov. sp. von Mnioticus COOMBS &amp; WOODROFFE, 1962 aus Athiopien (Coleoptera: Cryptophagidae). - Entomologische Zeitschrift 124 / 4: 247 - 250." type="journal article" year="2014">ESSER, 2014</bibRefCitation>
</taxonomicName>
(
<figureCitation id="86BF2A20FFB29C4660C9FE8F4C44FEB4" box="[989,1046,270,292]" captionStart="Figs 1-4" captionStartId="10.[151,191,1517,1539]" captionTargetBox="[161,1061,211,1506]" captionTargetId="figure-46@10.[153,1061,197,1507]" captionTargetPageId="10" captionText="Figs 1-4: (1) Mnioticus gibbinsi (SCOTT, 1936), male; (2) Mnioticus hancocki (SCOTT, 1936), male; (3) Mnioticus rhynchopetali (SCOTT, 1936), holotype, male; (4) Mnioticus rougemonti ESSER, 2014, paratype, male." figureDoi="http://doi.org/10.5281/zenodo.5277503" httpUri="https://zenodo.org/record/5277503/files/figure.png" pageId="3" pageNumber="918">fig. 4</figureCitation>
), described also from 
<collectingCountry id="66937635FFB29C466267FEAA4999FED1" box="[371,459,299,321]" name="Ethiopia" pageId="3" pageNumber="918">Ethiopia</collectingCountry>
, has smaller eyes and antennae segments 9 and 10 less transverse.
</paragraph>
<paragraph id="1E3B36A5FFB29C466383FEEF4A2CFE31" blockId="3.[151,1058,366,417]" pageId="3" pageNumber="918">Pale specimens have only a small dark spot, which is enlarged in darker specimens. Very dark specimens have also a darkened pronotum.</paragraph>
<paragraph id="1E3B36A5FFB29C466383FE4D4C72FD97" blockId="3.[151,483,459,481]" lastBlockId="3.[151,1059,495,735]" pageId="3" pageNumber="918">
<emphasis id="2CF0EAB7FFB29C466383FE4D49C4FE71" bold="true" box="[151,406,459,481]" pageId="3" pageNumber="918">
<taxonomicName id="D9844D26FFB29C466383FE4D4916FE71" authority="Esser &amp; R, 2019" authorityName="Esser &amp; R" authorityYear="2019" box="[151,324,459,481]" class="Insecta" family="Cryptophagidae" genus="Mnioticus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="918" phylum="Arthropoda" rank="species" species="holmi" status="sp. nov.">
<emphasis id="2CF0EAB7FFB29C466383FE4D4916FE71" bold="true" box="[151,324,459,481]" italics="true" pageId="3" pageNumber="918">Mnioticus holmi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="37C357CCFFB29C46625EFE4D49C4FE71" box="[330,406,460,481]" pageId="3" pageNumber="918" rank="species">nov.sp.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="86BF2A20FFB29C4662B0FE4D4989FE72" box="[420,475,460,482]" captionStart="Figs 9-12" captionStartId="12.[151,191,1483,1505]" captionTargetBox="[159,1029,205,1437]" captionTargetId="figure-41@12.[153,1061,196,1465]" captionTargetPageId="12" captionText="Figs 9-12: (9) Mnioticus holmi nov.sp., dark specimen, paratype, female; (10) Mnioticus karisimbianus nov.sp., paratype, female; (11) Mnioticus gugheanus nov.sp. Gughe Mts., paratype, male; (12) Mnioticus kinangopianus nov.sp. Aberdare, paratype, female." figureDoi="http://doi.org/10.5281/zenodo.5277509" httpUri="https://zenodo.org/record/5277509/files/figure.png" pageId="3" pageNumber="918">fig. 9</figureCitation>
) T y p e m a t e r i a l: 
<typeStatus id="C13F8807FFB29C46629AFE7349B2FD98" box="[398,480,498,520]" pageId="3" pageNumber="918" type="holotype">Holotype</typeStatus>
<specimenCount id="0882FD2CFFB29C4662F3FE7349A8FD97" box="[487,506,498,519]" pageId="3" pageNumber="918" type="male">♂</specimenCount>
: &quot;EAK Ruwenzori, Kitandara, 
<date id="6A3A1065FFB29C466007FE734B24FD97" box="[787,886,498,519]" pageId="3" pageNumber="918" value="1948-04-10">10.IV.1948</date>
, Å. Holm Nr. 93 /
</paragraph>
<paragraph id="1E3B36A5FFB29C4663D3FD8B49D2FDEF" blockId="3.[151,1059,495,735]" pageId="3" pageNumber="918">
<materialsCitation id="AEEC3CF8FFB29C4663D3FD8B49EAFDA7" collectingDate="1948-12-22" collectionCode="NHRM" country="Ethiopia" county="Lobelia" elevation="3850" location="Gamo Prov." municipality="Mt. Gughe" pageId="3" pageNumber="918" specimenCount="6" specimenCount-female="5" specimenCount-male="1" typeStatus="paratype">
<quantity id="D97C9B40FFB29C4663D3FD8B495EFD8F" box="[199,268,522,544]" metricMagnitude="3" metricUnit="m" metricValue="3.85" pageId="3" pageNumber="918" unit="m" value="3850.0">
<elevation id="95A9D196FFB29C4663D3FD8B495EFD8F" box="[199,268,522,544]" metricMagnitude="3" metricUnit="m" metricValue="3.85" pageId="3" pageNumber="918" unit="m" value="3850.0">3850 m</elevation>
</quantity>
, Sålln mossa i Senecio/Lobelia-skog.&quot; [
<collectionCode id="7895AE60FFB29C466177FD884AFEFD8F" box="[611,684,521,543]" country="Sweden" httpUri="http://grbio.org/cool/jrkv-fynr" name="Naturhistoriska Rijkmuseet" pageId="3" pageNumber="918">NHRM</collectionCode>
]. 
<typeStatus id="C13F8807FFB29C4661ADFD8B4B43FD8F" box="[697,785,522,543]" pageId="3" pageNumber="918" type="paratype">Paratypes</typeStatus>
: 
<specimenCount id="0882FD2CFFB29C466003FD8B4B6BFD8F" box="[791,825,522,543]" pageId="3" pageNumber="918" type="male">1 ♂</specimenCount>
, 
<specimenCount id="0882FD2CFFB29C466056FD8B4B31FD8F" box="[834,867,522,544]" pageId="3" pageNumber="918" type="female">5 ♀</specimenCount>
with the same data as the holotype [
<collectionCode id="7895AE60FFB29C46622FFDA349D0FDA7" box="[315,386,546,567]" country="Sweden" httpUri="http://grbio.org/cool/jrkv-fynr" name="Naturhistoriska Rijkmuseet" pageId="3" pageNumber="918">NHRM</collectionCode>
, cES]
</materialsCitation>
; 
<materialsCitation id="AEEC3CF8FFB29C4662D4FDA3492EFDEF" collectingDate="1963-07-19" collectingDateMax="1963-07-22" collectingDateMin="1963-07-19" collectionCode="RMAC, RMCA" collectorName="M. - J. Celis" country="Democratic Republic of the Congo" elevation="3" location="Congo" pageId="3" pageNumber="918" specimenCount="4" specimenCount-female="4" typeStatus="paratype">
<specimenCount id="0882FD2CFFB29C4662D4FDA349B0FDA7" box="[448,482,546,567]" pageId="3" pageNumber="918" type="female">3 ♀</specimenCount>
&quot; 
<collectingCountry id="66937635FFB29C4662FBFDA34A7CFDA7" box="[495,558,546,567]" name="Democratic Republic of the Congo" pageId="3" pageNumber="918">Congo</collectingCountry>
: P. N. A., 
<date id="6A3A1065FFB29C46619CFDA34B44FDA8" box="[648,790,546,568]" pageId="3" pageNumber="918" value="1963-07-21" valueMax="1963-07-22" valueMin="1963-07-21">
<collectingDate id="7A7EE98DFFB29C46619CFDA34B44FDA8" box="[648,790,546,568]" pageId="3" pageNumber="918" value="1963-07-21" valueMax="1963-07-22" valueMin="1963-07-21">21-22-VII-1963</collectingDate>
</date>
, RP. 
<collectorName id="B3715373FFB29C466050FDA34BF7FDA8" box="[836,933,546,568]" pageId="3" pageNumber="918">M.-J. Célis</collectorName>
2129 / Massif Ruwenzori, Mahungu, r. dr. Katsambu, 
<quantity id="D97C9B40FFB29C466130FDBB4A3DFDDF" box="[548,623,570,591]" metricMagnitude="0" metricUnit="m" metricValue="3.28" pageId="3" pageNumber="918" unit="m" value="3.28">
<elevation id="95A9D196FFB29C466130FDBB4A3DFDDF" box="[548,623,570,591]" metricMagnitude="0" metricUnit="m" metricValue="3.28" pageId="3" pageNumber="918" unit="m" value="3.28">3.280 m</elevation>
</quantity>
, terreau » [
<collectionCode id="7895AE60FFB29C4661C4FDBB4B47FDDF" box="[720,789,570,591]" pageId="3" pageNumber="918">RMAC</collectionCode>
, cES], 
<specimenCount id="0882FD2CFFB29C466040FDBB4B25FDDF" box="[852,887,570,591]" pageId="3" pageNumber="918" type="female">1 ♀</specimenCount>
« 
<collectingCountry id="66937635FFB29C46609AFDBB4B9EFDDF" box="[910,972,570,591]" name="Democratic Republic of the Congo" pageId="3" pageNumber="918">Congo</collectingCountry>
: P. N. A., 
<date id="6A3A1065FFB29C4663D0FDD34967FDF8" box="[196,309,594,616]" pageId="3" pageNumber="918" value="1963-07-19">
<collectingDate id="7A7EE98DFFB29C4663D0FDD34967FDF8" box="[196,309,594,616]" pageId="3" pageNumber="918" value="1963-07-19">19-VII-1963</collectingDate>
</date>
, RP. 
<collectorName id="B3715373FFB29C466273FDD34998FDF8" box="[359,458,594,616]" pageId="3" pageNumber="918">M.-J. Célis</collectorName>
2135 / Massif Ruwenzori, Mahungu, camp des porteurs, 
<quantity id="D97C9B40FFB29C4660C7FDD34C4DFDF7" box="[979,1055,594,616]" metricMagnitude="0" metricUnit="m" metricValue="3.25" pageId="3" pageNumber="918" unit="m" value="3.25">
<elevation id="95A9D196FFB29C4660C7FDD34C4DFDF7" box="[979,1055,594,616]" metricMagnitude="0" metricUnit="m" metricValue="3.25" pageId="3" pageNumber="918" unit="m" value="3.25">3.250 m</elevation>
</quantity>
, Ericetum » [
<collectionCode id="7895AE60FFB29C46623BFDE84925FDEF" box="[303,375,617,639]" country="Belgium" httpUri="http://grbio.org/cool/ee3x-fpgk" name="Royal Museum for Central Africa" pageId="3" pageNumber="918" type="Museum">RMCA</collectionCode>
]
</materialsCitation>
.
</paragraph>
<paragraph id="1E3B36A5FFB29C466383FD0E4C71FD51" blockId="3.[151,1059,495,735]" pageId="3" pageNumber="918">
E t y m o l o g y: Named after Åke Holm (1909-1989) who collected the 
<typeStatus id="C13F8807FFB29C466089FD0F4BA4FD34" box="[925,1014,654,676]" pageId="3" pageNumber="918" type="holotype">holotype</typeStatus>
and a few 
<typeStatus id="C13F8807FFB29C4663CCFD2D496EFD51" box="[216,316,684,705]" pageId="3" pageNumber="918" type="paratype">paratypes</typeStatus>
, and also a lot of 
<taxonomicName id="D9844D26FFB29C4662E3FD2A4ACBFD51" box="[503,665,683,705]" class="Insecta" family="Cryptophagidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="918" phylum="Arthropoda" rank="family">Cryptophagidae</taxonomicName>
from several mountain ranges in East
</paragraph>
<paragraph id="1E3B36A5FFB29C466383FD48488DFD4F" blockId="3.[151,1059,495,735]" box="[151,223,713,735]" pageId="3" pageNumber="918">Africa.</paragraph>
<paragraph id="1E3B36A5FFB29C466383FD6E4901FB11" blockId="3.[151,1059,750,1153]" pageId="3" pageNumber="918">
D e s c r i p t i o n: Male, 
<quantity id="D97C9B40FFB29C4662AEFD6F4A5FFC94" box="[442,525,750,772]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="3" pageNumber="918" unit="mm" value="1.9">1.9 mm</quantity>
, reddish-brown, elytra pale (except suture), underside dark, posteriorly paler, shiny. Covered with whitish hairs, more or less (on sides of elytra) decumbent. Membranous wings absent. Legs and antennae slender, tarsi 5- segmented, anterior tarsomeres 1-3 widened (only in male), antennae 11-segmented with a 3-segmented club. Segment 3 elongated and pear-shaped, segment 2 somewhat shorter and broader. Segment 1 stout, segments 4 to 8 more or less long as broad. Segments 9 and 10 moderately transverse, 11 fusiform and asymmetrical. Eyes moderately large with coarse ommatidia, puncture on head fine and sparse. Pronotum moderately transverse (1,4 times broader than long), broadest in the middle, sides narrowed anteriorly and posteriorly, convex, before posterior angles abruptly concave. Sides with fine crenulation. Anterior margin nearly straight, basal margin slightly rounded backwards in the medium half. Basal furrow moderate, basal fovea short. Puncture on pronotum fine and sparse. Elytra elongated, without any features. Puncture distinctly denser and stronger than on pronotum.
</paragraph>
<paragraph id="1E3B36A5FFB29C466383FB104B34FAA9" blockId="3.[151,1059,1169,1337]" pageId="3" pageNumber="918">
C o m m e n t: Not well distinguishable from 
<taxonomicName id="D9844D26FFB29C466163FB104A9DFB36" authorityName="Esser &amp; R" authorityYear="2019" box="[631,719,1169,1190]" class="Insecta" family="Cryptophagidae" genus="Mnioticus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="918" phylum="Arthropoda" rank="species" species="scotti" status="nom. nov.">
<emphasis id="2CF0EAB7FFB29C466163FB104ADEFB36" box="[631,652,1169,1190]" italics="true" pageId="3" pageNumber="918">M</emphasis>
. 
<emphasis id="2CF0EAB7FFB29C46618CFB104A9DFB36" box="[664,719,1169,1190]" italics="true" pageId="3" pageNumber="918">scotti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="37C357CCFFB29C4661C2FB104B64FB36" box="[726,822,1169,1190]" pageId="3" pageNumber="918">nom.nov.</taxonomicNameLabel>
except by the origin of specimens. 
<taxonomicName id="D9844D26FFB29C466205FB2E493FFB54" authorityName="Esser &amp; R" authorityYear="2019" box="[273,365,1199,1220]" class="Insecta" family="Cryptophagidae" genus="Mnioticus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="918" phylum="Arthropoda" rank="species" species="scotti" status="nom. nov.">
<emphasis id="2CF0EAB7FFB29C466205FB2E4974FB54" box="[273,294,1199,1220]" italics="true" pageId="3" pageNumber="918">M</emphasis>
. 
<emphasis id="2CF0EAB7FFB29C466222FB2E493FFB54" box="[310,365,1199,1220]" italics="true" pageId="3" pageNumber="918">scotti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="37C357CCFFB29C466263FB2E4985FB54" box="[375,471,1199,1220]" pageId="3" pageNumber="918">nom.nov.</taxonomicNameLabel>
is known by 
<specimenCount id="0882FD2CFFB29C466179FB2E4AB9FB54" box="[621,747,1198,1220]" pageId="3" pageNumber="918" type="female">two females</specimenCount>
(
<typeStatus id="C13F8807FFB29C4661E8FB2F4B04FB54" box="[764,854,1198,1220]" pageId="3" pageNumber="918" type="holotype">holotype</typeStatus>
and one additional specimen). For distinguishing the two species the discovery of male of 
<taxonomicName id="D9844D26FFB29C466061FB4D4B9CFB71" authorityName="Esser &amp; R" authorityYear="2019" box="[885,974,1228,1249]" class="Insecta" family="Cryptophagidae" genus="Mnioticus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="918" phylum="Arthropoda" rank="species" species="scotti">
<emphasis id="2CF0EAB7FFB29C466061FB4D4BD8FB71" box="[885,906,1228,1249]" italics="true" pageId="3" pageNumber="918">M</emphasis>
. 
<emphasis id="2CF0EAB7FFB29C466083FB4D4B9CFB71" box="[919,974,1228,1249]" italics="true" pageId="3" pageNumber="918">scotti</emphasis>
</taxonomicName>
is helpful. Also similar to 
<taxonomicName id="D9844D26FFB29C46627FFB684A7BFB6E" authority="Esser &amp; R, 2019" authorityName="Esser &amp; R" authorityYear="2019" box="[363,553,1257,1278]" class="Insecta" family="Cryptophagidae" genus="Mnioticus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="918" phylum="Arthropoda" rank="species" species="ruwenzoriensis" status="sp. nov.">
<emphasis id="2CF0EAB7FFB29C46627FFB6849D2FB6E" box="[363,384,1257,1278]" italics="true" pageId="3" pageNumber="918">M</emphasis>
. 
<emphasis id="2CF0EAB7FFB29C466284FB684A7BFB6E" box="[400,553,1257,1278]" italics="true" pageId="3" pageNumber="918">ruwenzoriensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="37C357CCFFB29C466127FB684A2FFB6E" box="[563,637,1257,1278]" pageId="3" pageNumber="918" rank="species">nov.sp.</taxonomicNameLabel>
but pronotum with puncture and shiny. Sides of pronotum concave in the posterior part. The 
<taxonomicName id="D9844D26FFB29C4661D5FA864B27FA8B" authority="Esser &amp; R, 2019" authorityName="Esser &amp; R" authorityYear="2019" box="[705,885,1286,1308]" class="Insecta" family="Cryptophagidae" genus="Mnioticus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="918" phylum="Arthropoda" rank="species" species="karisimbianus" status="sp. nov.">
<emphasis id="2CF0EAB7FFB29C4661D5FA864A84FA8C" box="[705,726,1287,1308]" italics="true" pageId="3" pageNumber="918">M</emphasis>
. 
<emphasis id="2CF0EAB7FFB29C4661F1FA874B27FA8B" box="[741,885,1286,1307]" italics="true" pageId="3" pageNumber="918">karisimbianus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="37C357CCFFB29C466068FA864B94FA8C" box="[892,966,1287,1308]" pageId="3" pageNumber="918" rank="species">nov.sp.</taxonomicNameLabel>
(
<figureCitation id="86BF2A20FFB29C4660C1FA874C48FA8C" box="[981,1050,1286,1308]" captionStart="Figs 9-12" captionStartId="12.[151,191,1483,1505]" captionTargetBox="[159,1029,205,1437]" captionTargetId="figure-41@12.[153,1061,196,1465]" captionTargetPageId="12" captionText="Figs 9-12: (9) Mnioticus holmi nov.sp., dark specimen, paratype, female; (10) Mnioticus karisimbianus nov.sp., paratype, female; (11) Mnioticus gugheanus nov.sp. Gughe Mts., paratype, male; (12) Mnioticus kinangopianus nov.sp. Aberdare, paratype, female." figureDoi="http://doi.org/10.5281/zenodo.5277509" httpUri="https://zenodo.org/record/5277509/files/figure.png" pageId="3" pageNumber="918">fig. 10</figureCitation>
) with elytra more stout is so far not known from Ruwenzori Mountains.
</paragraph>
</subSubSection>
</treatment>
</document>