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<document id="4E5CF81899CCA55BB5CB9311BA35E539" ID-CLB-Dataset="99512" ID-DOI="10.5252/geodiversitas2022v44a23" ID-GBIF-Dataset="1818f0c9-dbee-4fa4-b69a-ad6d3712913b" ID-ISSN="1638-9395" ID-Zenodo-Dep="6819412" ID-ZooBank="urn:lsid:zoobank.org:pub:999E7A6A-3781-4746-8519-62E1010D3880" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1657550474273" checkinUser="felipe" docAuthor="Klopfstein, Seraina" docDate="2022" docId="FC1A686EFFAFFFF68F1042DDFE480901" docLanguage="en" docName="Geodiversitas.44.23.645-664.pdf.imf" docOrigin="Geodiversitas 44 (23)" docStyle="DocumentStyle:F830B10FF475E64C1F1601E3B32DDC00.4:Geodiversitas.2018-.journal_article" docStyleId="F830B10FF475E64C1F1601E3B32DDC00" docStyleName="Geodiversitas.2018-.journal_article" docStyleVersion="4" docTitle="Xanthopimpla ciboisae Klopfstein 2022, n. sp." docType="treatment" docUuid="412A6A93-542C-4326-83F6-BFBC29358A4D" docUuidSource="ZooBank" docVersion="9" lastPageNumber="660" masterDocId="00231016FFBFFFE78E394169FF940D6F" masterDocTitle="High diversity of pimpline parasitoid wasps (Hymenoptera, Ichneumonidae, Pimplinae) from the lowermost Eocene Fur Formation (Denmark)" masterLastPageNumber="664" masterPageNumber="645" pageNumber="659" updateTime="1699358479615" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:title id="BA6A6302E97DCA66FEB9043EF9F6BCE8">High diversity of pimpline parasitoid wasps (Hymenoptera, Ichneumonidae, Pimplinae) from the lowermost Eocene Fur Formation (Denmark)</mods:title>
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<mods:namePart id="8BF607C036B4ED2E12779E66806D08DB">Klopfstein, Seraina</mods:namePart>
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<mods:title id="85C1E1071ED88135CC5B8E31D8FA132A">Geodiversitas</mods:title>
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<mods:date id="5FF768AD0C6E575449C453BAF8EDE7A6">2022</mods:date>
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<mods:number id="C4A72C7E05CD6CFBE674B88E3CFF955A">2022-07-07</mods:number>
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<treatment id="FC1A686EFFAFFFF68F1042DDFE480901" ID-DOI="http://doi.org/10.5281/zenodo.6857742" ID-GBIF-Taxon="197821812" ID-Zenodo-Dep="6857742" LSID="urn:lsid:zoobank.org:act:412A6A93-542C-4326-83F6-BFBC29358A4D" httpUri="http://treatment.plazi.org/id/FC1A686EFFAFFFF68F1042DDFE480901" lastPageId="17" lastPageNumber="660" pageId="16" pageNumber="659">
<subSubSection id="3CA98AF3FFAFFFF78F1042DDFDF60EA0" box="[297,610,948,975]" pageId="16" pageNumber="659" type="nomenclature">
<paragraph id="740CD978FFAFFFF78F1042DDFDF60EA0" blockId="16.[297,610,948,975]" box="[297,610,948,975]" pageId="16" pageNumber="659">
<heading id="2F446E14FFAFFFF78F1042DDFDF60EA0" box="[297,610,948,975]" centered="true" fontSize="11" level="2" pageId="16" pageNumber="659" reason="2">
<taxonomicName id="B3B3A2FBFFAFFFF78F1042DDFDB40EA0" authority="Klopfstein, 2022" authorityName="Klopfstein" authorityYear="2022" box="[297,544,948,975]" class="Insecta" family="Ichneumonidae" genus="Xanthopimpla" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="659" phylum="Arthropoda" rank="species" species="ciboisae" status="sp. nov.">
<emphasis id="46C7056AFFAFFFF78F1042DDFDB40EA0" bold="true" box="[297,544,948,975]" italics="true" pageId="16" pageNumber="659">Xanthopimpla ciboisae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5DF4B811FFAFFFF78C1E42DCFDF60EA0" box="[551,610,949,975]" pageId="16" pageNumber="659" rank="species">n. sp.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="3CA98AF3FFAFFFF78FAE42BCFC930947" pageId="16" pageNumber="659" type="description">
<paragraph id="740CD978FFAFFFF78FAE42BCFE600E80" blockId="16.[407,500,980,1007]" box="[407,500,980,1007]" pageId="16" pageNumber="659">
(
<figureCitation id="EC88C5FDFFAFFFF78F9942BDFE7F0E80" box="[416,491,980,1007]" captionStart="FIG" captionStartId="16.[132,143,779,796]" captionTargetBox="[132,1444,215,737]" captionTargetId="figure-633@16.[698,1267,213,738]" captionTargetPageId="16" captionText="FIG. 10. — Holotype of Xanthopimpla ciboisae n. sp. (FUR-10046), photograph and interpretative drawing, where dotted lines represent uncertain and/or interpolated interpretations. Scale bar: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6819434" httpUri="https://zenodo.org/record/6819434/files/figure.png" pageId="16" pageNumber="659">Fig. 10</figureCitation>
)
</paragraph>
<paragraph id="740CD978FFAFFFF78EBD457EFC930947" blockId="16.[132,775,1047,1064]" box="[132,775,1047,1064]" pageId="16" pageNumber="659">
<uri id="0022D57AFFAFFFF78EBD457EFC930947" box="[132,775,1047,1064]" pageId="16" pageNumber="659">
urn:lsid:zoobank.org:act:
<uuid id="0015E3ADFFAFFFF78F65457EFC930947" box="[348,775,1047,1064]" pageId="16" pageNumber="659">412A6A93-542C-4326-83F6-BFBC29358A4D</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="3CA98AF3FFAFFFF78EBD4535FC9C091C" box="[132,776,1115,1139]" pageId="16" pageNumber="659" type="materials_examined">
<paragraph id="740CD978FFAFFFF78EBD4535FC9C091C" blockId="16.[132,776,1115,1139]" box="[132,776,1115,1139]" pageId="16" pageNumber="659">
TYPE MATERIAL. —
<materialsCitation id="C4DBD325FFAFFFF78F074532FC9C091C" ID-GBIF-Occurrence="3863921315" box="[318,776,1115,1139]" collectionCode="FUR" collectorName="M. Breiner" country="Denmark" location="Skive Kommune" municipality="Jutland" pageId="16" pageNumber="659" specimenCode="FUR-10046" specimenCount="1" specimenCount-female="1" stateProvince="Limfjord" typeStatus="holotype">
<typeStatus id="AB0867DAFFAFFFF78F074532FE0D091C" box="[318,409,1115,1139]" pageId="16" pageNumber="659" type="holotype">Holotype</typeStatus>
<specimenCount id="62B512F1FFAFFFF78FA74532FE74091C" box="[414,480,1115,1139]" pageId="16" pageNumber="659" type="female">female</specimenCount>
:
<specimenCode id="24157103FFAFFFF78FDE4532FDCA091C" box="[487,606,1115,1139]" pageId="16" pageNumber="659">FUR-10046</specimenCode>
(leg.
<collectorName id="D946BCAEFFAFFFF78CA94535FD69091C" box="[656,765,1115,1139]" pageId="16" pageNumber="659">M. Breiner</collectorName>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="3CA98AF3FFAFFFF78EBA45E4FE0F082B" pageId="16" pageNumber="659" type="etymology">
<paragraph id="740CD978FFAFFFF78EBA45E4FE0F082B" blockId="16.[131,775,1165,1348]" pageId="16" pageNumber="659">ETYMOLOGY. — This wasp is dedicated to Alice Cibois, to honour her dedication as president of the Swiss Systematics Society and her creative and innovative efforts to promote our craft among the general public. She especially drew attention to the ongoing discovery and naming of new species through the curated list “New Species Swiss Made”, which year after year feature dozens to hundred new species across the tree of life.</paragraph>
</subSubSection>
<subSubSection id="3CA98AF3FFAFFFF78EBD4437FF5F08B3" pageId="16" pageNumber="659" type="materials_examined">
<paragraph id="740CD978FFAFFFF78EBD4437FDC008FF" blockId="16.[132,777,1373,1424]" pageId="16" pageNumber="659">
<typeStatus id="AB0867DAFFAFFFF78EBD4437FF22081A" box="[132,182,1374,1397]" pageId="16" pageNumber="659">TYPE</typeStatus>
HORIZON AND LOCALITY. —
<collectingCountry id="0CA499E8FFAFFFF78FD14434FDDF081A" box="[488,587,1373,1397]" name="Denmark" pageId="16" pageNumber="659">Denmark</collectingCountry>
,
<collectingMunicipality id="94684302FFAFFFF78C6E4434FD37081A" box="[599,675,1373,1397]" pageId="16" pageNumber="659">Jutland</collectingMunicipality>
,
<collectingRegion id="B677179AFFAFFFF78C894434FC9D081A" box="[688,777,1373,1397]" pageId="16" pageNumber="659">Limfjord</collectingRegion>
region,
<location id="716C8FA3FFAFFFF78EF64411FEE108E0" LSID="urn:lsid:plazi:treatment:FC1A686EFFAFFFF68F1042DDFE480901:716C8FA3FFAFFFF78EF64411FEE108E0" box="[207,373,1400,1424]" country="Denmark" municipality="Jutland" name="Skive Kommune" pageId="16" pageNumber="659" stateProvince="Limfjord">Skive Kommune</location>
, Fur. Silstrup member.
</paragraph>
<paragraph id="740CD978FFAFFFF78EBD44C3FF5F08B3" blockId="16.[132,775,1450,1500]" pageId="16" pageNumber="659">
MATERIAL EXAMINED. —
<materialsCitation id="C4DBD325FFAFFFF78FB944C3FF5F08B3" ID-GBIF-Occurrence="3863921310" collectionCode="FUR" pageId="16" pageNumber="659" specimenCode="FUR- 14684" specimenCount="1">
Tentatively placed in this species:
<specimenCode id="24157103FFAFFFF78CF744C3FF5308B3" pageId="16" pageNumber="659">FUR- 14684</specimenCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="3CA98AF3FFAFFFF78EBD449FFE890A84" pageId="16" pageNumber="659" type="diagnosis">
<paragraph id="740CD978FFAFFFF78EBD449FFE290B8D" blockId="16.[131,775,1526,2027]" pageId="16" pageNumber="659">
DIAGNOSIS. — Taxonomic placement: while many of the characters that can be used to diagnose
<taxonomicName id="B3B3A2FBFFAFFFF78FD34779FDC50B47" box="[490,593,1552,1576]" pageId="16" pageNumber="647" rank="subFamily" subFamily="Pimplinae">Pimplinae</taxonomicName>
are well visible in this specimen (
<tableCitation id="3931ECC3FFAFFFF78F1B4742FEFB0B2C" box="[290,367,1579,1603]" captionStart="TABLE" captionStartId="18.[132,143,219,236]" captionTargetPageId="18" captionText="TABLE 1. — Overview of characters supporting placement in Pimplinae. Brackets indicate uncertain interpretation or visibility of only some aspect of a character. 1, forewig with areolet triangular or oblique-quadrate; 2, forewing vein 2m-cu bowed outwards, with two bullae (in brackets if bullae not visible); 3, first tergite attached broadly to propodeum, parallel-sided or somewhat converging towards base; usually 1.0-2.0 × longer than wide; 4, spiracle of first tergite in front of middle; first sternite shorter than half the length of tergite; 5, ovipositor clearly protruding from metasomal apex, often long; if shorter than metasoma, then usually rather robust; 6, ventral valve of ovipositor with teeth, dorsal valve without a notch; 7, claws with basal lobe (present in many extant Pimplinae, especially Ephialtini); 8, unique character combination present that allows association with a particular genus in Pimplinae." httpUri="http://table.plazi.org/id/20CC89F0FFADFFF58EBD41B2FBBC0C31" pageId="16" pageNumber="659" tableUuid="20CC89F0FFADFFF58EBD41B2FBBC0C31">Table 1</tableCitation>
), the most unequivocal evidence stems from generic placement: an association with
<taxonomicName id="B3B3A2FBFFAFFFF78C58472CFD7B0B32" authorityName="Saussure" authorityYear="1892" box="[609,751,1605,1629]" class="Insecta" family="Ichneumonidae" genus="Xanthopimpla" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="659" phylum="Arthropoda" rank="genus">
<emphasis id="46C7056AFFAFFFF78C58472CFD7B0B32" box="[609,751,1605,1629]" italics="true" pageId="16" pageNumber="659">Xanthopimpla</emphasis>
</taxonomicName>
is evidenced by the colour pattern (especially the paired dark marks at least on tergite 3, robust legs, very short hind wing 1-Cu compared to cu-a, stout metasoma, and robust ovipositor. Forewing vein 2m-cu furthermore shows a zig-zag pattern that is very rare outside that genus.
</paragraph>
<paragraph id="740CD978FFAFFFF78EBD478CFE890A84" blockId="16.[131,775,1526,2027]" pageId="16" pageNumber="659">
Species diagnosis: this species is very similar both in colour pattern and wing venation to many extant species of the genus, but has a more extended light mark at the base of the pterostigma. It is much larger than the two fossil species from Messel, even than
<taxonomicName id="B3B3A2FBFFAFFFF78E834626FEB30A09" authorityName="Khalaim" authorityYear="2008" box="[186,295,1870,1894]" class="Insecta" family="Ichneumonidae" genus="Xanthopimpla" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="659" phylum="Arthropoda" rank="species" species="biamosa">
<emphasis id="46C7056AFFAFFFF78E834626FEB30A09" box="[186,295,1870,1894]" italics="true" pageId="16" pageNumber="659">X. biamosa</emphasis>
</taxonomicName>
(forewing 8.0 mm), from which it can be further distinguished by the lateral dark marks on T
<quantity id="B34B749DFFAFFFF78C434600FD0B0AEE" box="[634,671,1897,1921]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="16" pageNumber="659" unit="in" value="1.0">1 in</quantity>
the latter. From
<taxonomicName id="B3B3A2FBFFAFFFF78EFD46EAFE310AF4" authorityName="Klopfstein" authorityYear="2022" baseAuthorityName="Henriksen" baseAuthorityYear="1922" box="[196,421,1923,1947]" class="Insecta" family="Ichneumonidae" genus="Epitheronia" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="659" phylum="Arthropoda" rank="species" species="stigmatica" status="comb. nov.">
<emphasis id="46C7056AFFAFFFF78EFD46EAFE310AF4" box="[196,421,1923,1947]" italics="true" pageId="16" pageNumber="659">Epitheronia stigmatica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5DF4B811FFAFFFF78F9446EDFD9F0AF4" box="[429,523,1923,1947]" pageId="16" pageNumber="659" rank="species">n. comb.</taxonomicNameLabel>
, it can be distinguished by the larger size, light coloured mesoscutum, more sinuous 4Rs in the front wing, and even shorter 1-Cu compared to cu-a in the hind wing.
</paragraph>
</subSubSection>
<subSubSection id="3CA98AF3FFAFFFF68D14421CFE480901" lastPageId="17" lastPageNumber="660" pageId="16" pageNumber="659" type="description">
<paragraph id="740CD978FFAFFFF78D14421CFC290EE0" blockId="16.[811,1457,885,1102]" box="[813,957,885,911]" pageId="16" pageNumber="659">DESCRIPTION</paragraph>
<paragraph id="740CD978FFAFFFF78D1442FFFC3F0EC0" blockId="16.[811,1457,885,1102]" box="[813,939,918,943]" pageId="16" pageNumber="659">
<emphasis id="46C7056AFFAFFFF78D1442FFFC3F0EC0" box="[813,939,918,943]" italics="true" pageId="16" pageNumber="659">Preservation</emphasis>
</paragraph>
<paragraph id="740CD978FFAFFFF78D1442DCFB190921" blockId="16.[811,1457,885,1102]" pageId="16" pageNumber="659">
<emphasis id="46C7056AFFAFFFF78D1442DCFC2E0EA1" bold="true" box="[813,954,948,975]" pageId="16" pageNumber="659">Dorsal view.</emphasis>
Head partially preserved, including extreme base of antennae. Mesosoma rather well preserved, including propodeal carination; all four wings nearly complete, partial hind legs visible.Metasoma almost complete; ovipositor partly, sheaths nearly entirely preserved.
</paragraph>
<paragraph id="740CD978FFAFFFF78D14451DFA1F0862" blockId="16.[813,1456,1140,1294]" pageId="16" pageNumber="659">
<emphasis id="46C7056AFFAFFFF78D14451DFC4E09E1" bold="true" box="[813,986,1140,1166]" pageId="16" pageNumber="659">
Body
<quantity id="B34B749DFFAFFFF78D48451DFC4E09E1" box="[881,986,1140,1166]" metricMagnitude="-2" metricUnit="m" metricValue="1.13" pageId="16" pageNumber="659" unit="mm" value="11.3">11.3 mm</quantity>
</emphasis>
. Yellow to light orange, with dark brown compound eyes, wing base, wing veins except basal
<emphasis id="46C7056AFFAFFFF78B4145FCFA1D09C1" box="[1400,1417,1173,1198]" italics="true" pageId="16" pageNumber="659">c.</emphasis>
3
<superScript id="83C67430FFAFFFF78BA645FCFA3B09C9" attach="none" box="[1439,1455,1173,1190]" fontSize="7" pageId="16" pageNumber="659">rd</superScript>
of pterostigma; dark paired spots evident on T3, seemingly absent from the other tergites (see also discussion of colour preservation); ovipositor light orange, sheaths brownish.
</paragraph>
<paragraph id="740CD978FFAFFFF78D14445AFC2508E2" blockId="16.[813,1457,1331,1421]" pageId="16" pageNumber="659">
<emphasis id="46C7056AFFAFFFF78D14445AFCE40822" bold="true" box="[813,880,1331,1357]" pageId="16" pageNumber="659">Head.</emphasis>
Distorted, with compound eyes overlapping; scape, pedicel and part of first flagellomeres preserved, of normal dimensions.
</paragraph>
<paragraph id="740CD978FFAFFFF78D1444DAFA3A0AE3" blockId="16.[810,1457,1459,1932]" pageId="16" pageNumber="659">
<emphasis id="46C7056AFFAFFFF78D1444DAFC3108A2" bold="true" box="[813,933,1459,1485]" pageId="16" pageNumber="659">Mesosoma.</emphasis>
Rather short and stout; deep notauli converging on basal ¼, then parallel; dark patches at fore wing base probably corresponding to axial sclerites; scutellum rather short, with transverse carinulae in prescutellar groove; propodeum with distinct carination, corresponding to nearly complete basal and probably also apical transverse, lateromedian and maybe lateral longitudinal and pleural carinae. Fore wing
<quantity id="B34B749DFFAFFFF78B68471BFA3F0BE2" box="[1361,1451,1650,1677]" metricMagnitude="-3" metricUnit="m" metricValue="9.2" pageId="16" pageNumber="659" unit="mm" value="9.2">9.2 mm</quantity>
; areolet closed, slightly petiolate above, quadrate with uneven sides (4-M very short, 2r-m a bit shorter than 3r-m); 2m-cu with two bullae, with a weak zig-zag that anteriorly is bent outwards, then inwards in front of the posterior bulla; pterostigma 3.5 × longer than wide; 1cu-a meeting M + Cu opposite 1-M; 3-Cu a bit longer than 2cu-a; radial cell 3.5 × longer than wide. Hind wing with 1-Rs about 3 × longer than 1rs-m, 1-Cu 0.3 × as long as cu-a. Hind leg not well preserved, but rather stout.
</paragraph>
<paragraph id="740CD978FFAFFFF78D1446D8FA250A83" blockId="16.[813,1457,1969,2028]" pageId="16" pageNumber="659">
<emphasis id="46C7056AFFAFFFF78D1446D8FC3E0AA4" bold="true" box="[813,938,1969,1995]" pageId="16" pageNumber="659">Metasoma.</emphasis>
With T1 about 0.7 × longer than wide, with strong median longitudinal carina converging at least on
</paragraph>
<caption id="20CC89F0FFAEFFF68EBD425EFD1A0E30" ID-DOI="http://doi.org/10.5281/zenodo.6819436" ID-Zenodo-Dep="6819436" httpUri="https://zenodo.org/record/6819436/files/figure.png" pageId="17" pageNumber="660" startId="17.[132,143,823,840]" targetBox="[132,1439,215,780]" targetPageId="17">
<paragraph id="740CD978FFAEFFF68EBD425EFD1A0E30" blockId="17.[132,1455,823,863]" pageId="17" pageNumber="660">
FIG. 11. — Holotype of
<taxonomicName id="B3B3A2FBFFAEFFF68F71425EFDB50E27" authority="Klopfstein, 2022" authorityName="Klopfstein" authorityYear="2022" box="[328,545,823,840]" class="Insecta" family="Ichneumonidae" genus="Xanthopimpla" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="660" phylum="Arthropoda" rank="species" species="crescendae" status="sp. nov.">
<emphasis id="46C7056AFFAEFFF68F71425EFDB50E27" box="[328,545,823,840]" italics="true" pageId="17" pageNumber="660">Xanthopimpla crescendae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5DF4B811FFAEFFF68C11425EFDCD0E27" box="[552,601,823,840]" pageId="17" pageNumber="660" rank="species">n. sp.</taxonomicNameLabel>
(FUR-11214), photograph (
<emphasis id="46C7056AFFAEFFF68D07425EFCDF0E27" bold="true" box="[830,843,823,840]" pageId="17" pageNumber="660">A</emphasis>
) with details and interpretative drawing (
<emphasis id="46C7056AFFAEFFF68A98425EFB3A0E27" bold="true" box="[1185,1198,823,840]" pageId="17" pageNumber="660">B</emphasis>
), where dotted lines represent uncertain and/or interpolated interpretations. Scale bar: 1 mm.
</paragraph>
</caption>
<paragraph id="740CD978FFAEFFF68EBD42FCFE480901" blockId="17.[131,776,917,1135]" pageId="17" pageNumber="660">
basal two-thirds; T2 0.4 × as long as wide, with basal oblique grooves indicated and maybe a preapical transverse groove present. T3 with paired dark patches on basal half and preapical transverse groove indicated. T4 with weak shadows of paired marks, unclear on remainder of tergites. Ovipositor sheaths
<emphasis id="46C7056AFFAEFFF68EE0455CFF7E0921" box="[217,234,1077,1102]" italics="true" pageId="17" pageNumber="660">c.</emphasis>
0.35 × as long as fore wing; rather broad, covered with very dense and short hairs.
</paragraph>
</subSubSection>
</treatment>
</document>