142 lines
14 KiB
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142 lines
14 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1119.85400" ID-GBIF-Dataset="418bd521-e4b9-4ee6-981f-a6f471eab287" ID-Pensoft-Pub="1313-2970-1119-1" ID-Pensoft-UUID="F158A7C5261D52B69288A62F3C777CAF" ID-ZooBank="A3BE91C6B79344E1A886A803BF104D8B" ModsDocID="1313-2970-1119-1" checkinTime="1661532673136" checkinUser="pensoft" docAuthor="Jirapatrasilp, Parin, Sutcharit, Chirasak & Panha, Somsak" docDate="2022" docId="8900D54559A15D068D7C13B8E99736A2" docLanguage="en" docName="ZooKeys 1119: 1-115" docOrigin="ZooKeys 1119" docPubDate="2022-08-25" docSource="http://dx.doi.org/10.3897/zookeys.1119.85400" docTitle="Pupina excisa Moellendorff 1902" docType="treatment" docVersion="4" id="F158A7C5261D52B69288A62F3C777CAF" lastPageNumber="1" masterDocId="F158A7C5261D52B69288A62F3C777CAF" masterDocTitle="Annotated checklist of the operculated land snails from Thailand (Mollusca, Gastropoda, Caenogastropoda): the family Pupinidae, with descriptions of several new species and subspecies, and notes on classification of Pupina Vignard, 1829 and Pupinella Gray, 1850 from mainland Southeast Asia" masterLastPageNumber="115" masterPageNumber="1" pageNumber="1" updateTime="1661537468408" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Annotated checklist of the operculated land snails from Thailand (Mollusca, Gastropoda, Caenogastropoda): the family Pupinidae, with descriptions of several new species and subspecies, and notes on classification of Pupina Vignard, 1829 and Pupinella Gray, 1850 from mainland Southeast Asia</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Jirapatrasilp, Parin</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-5591-6724</mods:nameIdentifier>
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<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sutcharit, Chirasak</mods:namePart>
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<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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<mods:nameIdentifier type="email">jirasak4@yahoo.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Panha, Somsak</mods:namePart>
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<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand & Academy of Science, The Royal Society of Thailand, Bangkok 10300, Thailand</mods:affiliation>
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<mods:nameIdentifier type="email">somsak.pan@chula.ac.th</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2022</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2022-08-25</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>1119</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>115</mods:end>
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</mods:extent>
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</mods:part>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1119.85400</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1119.85400</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-1119-1</mods:identifier>
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<mods:identifier type="ZooBank">A3BE91C6B79344E1A886A803BF104D8B</mods:identifier>
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<mods:identifier type="Pensoft-UUID">F158A7C5261D52B69288A62F3C777CAF</mods:identifier>
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<treatment ID-GBIF-Taxon="199838968" LSID="urn:lsid:plazi:treatment:8900D54559A15D068D7C13B8E99736A2" httpUri="http://treatment.plazi.org/id/8900D54559A15D068D7C13B8E99736A2" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName LSID="8900D545-59A1-5D06-8D7C-13B8E99736A2" authority="Moellendorff, 1902" authorityName="Moellendorff" authorityYear="1902" class="Gastropoda" family="Pupinidae" genus="Pupina" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Pupina excisa" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="excisa">
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Pupina excisa
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<normalizedToken originalValue="Möllendorff">Moellendorff</normalizedToken>
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, 1902
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</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">
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<figureCitation captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Shells of Pupina arula species group from mainland Southeast Asia A, B Pupina arula A syntype UMZC I. 103025 and B specimen NHMUK 1888.12.4.109. C, D Pupina mouhoti, possible syntypes NHMUK ex. Cuming coll. E, F Pupina vescoi E syntype NHMUK 1893.2.4.767 and F specimen SMF 109956 / 1 G Pupina perakensis, lectotype SMF 109969 / 1 H Pupina excisa, lectotype SMF 110778 / 1 I-K Pupina exclamationis I, J syntypes MNHN-IM- 2000 - 35840 and K specimen NHMUK 1901.12.23.205 " forma minor ". Photo: J. Ablett, H. Taylor, NHM (A), A. Lardeur, P. Maestrati, MNHN (I, J)." figureDoi="10.3897/zookeys.1119.85400.figure29" httpUri="https://binary.pensoft.net/fig/735390" pageId="0" pageNumber="1">Figs 29H</figureCitation>
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<figureCitation captionStart="Figure 34" captionStartId="F34" captionText="Figure 34. A Pupina vescoi, specimen SMF 109956 / 1 from Cochin China B, C Pupina exclamationis B syntype MNHN-IM- 2000 - 35840 from Tonkin and C specimen NHMUK 1901.12.23.205 " forma minor " from Than-Moi, Tonkin D Pupina perakensis, lectotype SMF 109969 / 1 from Bukit Pondong, Perak E Pupina excisa, lectotype SMF 110778 / 1 from Kelantan F Pupina aureola, possible syntype NHMUK 1988.12.4.101 from Pinang. Photo: P. Maestrati, MNHN (B)." figureDoi="10.3897/zookeys.1119.85400.figure34" httpUri="https://binary.pensoft.net/fig/735395" pageId="0" pageNumber="1">, 34E</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="reference_group">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authorityName="Moellendorff" authorityYear="1902" class="Gastropoda" family="Pupinidae" genus="Pupina" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Pupina (Tylotoechus) excisa" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="excisa" subGenus="Tylotoechus">Pupina (Tylotoechus) excisa</taxonomicName>
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<normalizedToken originalValue="Möllendorff">Moellendorff</normalizedToken>
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, 1902a: 143. Type locality: Kelantan [Malaysia].
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<bibRefCitation author="Laidlaw, FF" journalOrPublisher="Journal of the Malaysian Branch of the Royal Asiatic Society" pageId="0" pageNumber="1" pagination="25 - 37" refId="B82" refString="Laidlaw, FF, 1928. A list of the land and fresh-water mollusca of the Malay Peninsula with notes. Journal of the Malaysian Branch of the Royal Asiatic Society 6: 25 - 37" title="A list of the land and fresh-water mollusca of the Malay Peninsula with notes." volume="6" year="1928">Laidlaw 1928</bibRefCitation>
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: 34.
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<bibRefCitation author="Zilch, A" journalOrPublisher="Archiv fuer Molluskenkunde" pageId="0" pageNumber="1" pagination="41 - 56" refId="B198" refString="Zilch, A, 1957. Die Typen und Typoide des Natur-Museums Senckenberg, 19: Mollusca, Cyclophoridae, Pupininae. Archiv fuer Molluskenkunde 86: 41 - 56" title="Die Typen und Typoide des Natur-Museums Senckenberg, 19: Mollusca, Cyclophoridae, Pupininae." volume="86" year="1957">Zilch 1957</bibRefCitation>
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: 45, pl. 2, fig. 18.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authorityName="Moellendorff" authorityYear="1902" class="Gastropoda" family="Pupinidae" genus="Pupina" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Pupina excisa" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="excisa">Pupina excisa</taxonomicName>
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-
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<bibRefCitation author="Chan, S-Y" journalOrPublisher="The Papustyla" pageId="0" pageNumber="1" pagination="4 - 5" refId="B17" refString="Chan, S-Y, 1998a. A brief collecting trip to Perak, West Malaysia, part one. The Papustyla 12: 4 - 5" title="A brief collecting trip to Perak, West Malaysia, part one." volume="12" year="1998 a">Chan 1998a</bibRefCitation>
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: 4, Ipoh, Perak.
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<bibRefCitation author="Chan, S-Y" journalOrPublisher="The Papustyla" pageId="0" pageNumber="1" pagination="1 - 2" refId="B18" refString="Chan, S-Y, 1998b. A brief collecting trip to Perak, West Malaysia, part two. The Papustyla 12: 1 - 2" title="A brief collecting trip to Perak, West Malaysia, part two." volume="12" year="1998 b">Chan 1998b</bibRefCitation>
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: 2.
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<bibRefCitation author="Maassen, WJM" journalOrPublisher="Proceedings of the United States National Museum" pageId="0" pageNumber="1" refId="B87" refString="Maassen, WJM, 2001. A preliminary checklist of the non-marine molluscs of West-Malaysia, A handlist. De Kreukel (Supplement): 1-155." title="A preliminary checklist of the non-marine molluscs of West-Malaysia, A handlist. De Kreukel (Supplement): 1 - 155." year="2001">Maassen 2001</bibRefCitation>
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: 41.
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<bibRefCitation author="Bavay, A" journalOrPublisher="Journal de Conchyliologie" pageId="0" pageNumber="1" refId="B8" refString="BEDO (2017) Land Snails: Checklist of Molluscan Biodiversity in Thailand.BEDO, Bangkok, 300 pp. [in Thai]" year="1904">BEDO 2017</bibRefCitation>
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: 90.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="type material">
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<paragraph pageId="0" pageNumber="1">Type material examined.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Lectotype</emphasis>
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SMF 110778/1 (Figs
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<figureCitation captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Shells of Pupina arula species group from mainland Southeast Asia A, B Pupina arula A syntype UMZC I. 103025 and B specimen NHMUK 1888.12.4.109. C, D Pupina mouhoti, possible syntypes NHMUK ex. Cuming coll. E, F Pupina vescoi E syntype NHMUK 1893.2.4.767 and F specimen SMF 109956 / 1 G Pupina perakensis, lectotype SMF 109969 / 1 H Pupina excisa, lectotype SMF 110778 / 1 I-K Pupina exclamationis I, J syntypes MNHN-IM- 2000 - 35840 and K specimen NHMUK 1901.12.23.205 " forma minor ". Photo: J. Ablett, H. Taylor, NHM (A), A. Lardeur, P. Maestrati, MNHN (I, J)." figureDoi="10.3897/zookeys.1119.85400.figure29" httpUri="https://binary.pensoft.net/fig/735390" pageId="0" pageNumber="1">29H</figureCitation>
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,
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<figureCitation captionStart="Figure 34" captionStartId="F34" captionText="Figure 34. A Pupina vescoi, specimen SMF 109956 / 1 from Cochin China B, C Pupina exclamationis B syntype MNHN-IM- 2000 - 35840 from Tonkin and C specimen NHMUK 1901.12.23.205 " forma minor " from Than-Moi, Tonkin D Pupina perakensis, lectotype SMF 109969 / 1 from Bukit Pondong, Perak E Pupina excisa, lectotype SMF 110778 / 1 from Kelantan F Pupina aureola, possible syntype NHMUK 1988.12.4.101 from Pinang. Photo: P. Maestrati, MNHN (B)." figureDoi="10.3897/zookeys.1119.85400.figure34" httpUri="https://binary.pensoft.net/fig/735395" pageId="0" pageNumber="1">34E</figureCitation>
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) from Kelantan.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="diagnosis">
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<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="1">Shell ovate with higher spire; last whorl ca. three quarters of shell height. Apertural lip somewhat thickened but not expanded; apertural lip when observed from lateral view angled. Columellar tooth curved downward like an earlobe.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="differential diagnosis">
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<paragraph pageId="0" pageNumber="1">Differential diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authorityName="Moellendorff" authorityYear="1902" class="Gastropoda" family="Pupinidae" genus="Pupina" kingdom="Animalia" lsidName="Pupina excisa" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="excisa">
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<emphasis italics="true" pageId="0" pageNumber="1">Pupina excisa</emphasis>
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</taxonomicName>
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can be distinguished from all other species in the
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<taxonomicName class="Gastropoda" family="Pupinidae" kingdom="Animalia" lsidName="P. arula" order="Architaenioglossa" pageId="0" pageNumber="1" rank="species" species="arula">
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<emphasis italics="true" pageId="0" pageNumber="1">P. arula</emphasis>
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</taxonomicName>
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species group from mainland Southeast Asia by an ovate shell shape with a higher spire, and an angled apertural lip when observed from lateral view.
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<taxonomicName authorityName="Moellendorff" authorityYear="1902" class="Gastropoda" family="Pupinidae" genus="Pupina" kingdom="Animalia" lsidName="Pupina excisa" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="excisa">
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<emphasis italics="true" pageId="0" pageNumber="1">Pupina excisa</emphasis>
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</taxonomicName>
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is different from
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<taxonomicName class="Gastropoda" family="Pupinidae" kingdom="Animalia" lsidName="P. mouhoti" order="Architaenioglossa" pageId="0" pageNumber="1" rank="species" species="mouhoti">
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<emphasis italics="true" pageId="0" pageNumber="1">P. mouhoti</emphasis>
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</taxonomicName>
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in having a thicker, more prominent parietal tooth.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="distribution">
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<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Kelantan and Perak States, Malaysia (
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<bibRefCitation author="Maassen, WJM" journalOrPublisher="Proceedings of the United States National Museum" pageId="0" pageNumber="1" refId="B87" refString="Maassen, WJM, 2001. A preliminary checklist of the non-marine molluscs of West-Malaysia, A handlist. De Kreukel (Supplement): 1-155." title="A preliminary checklist of the non-marine molluscs of West-Malaysia, A handlist. De Kreukel (Supplement): 1 - 155." year="2001">Maassen 2001</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |