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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.502.9049" ID-PMC="PMC4443589" ID-Pensoft-Pub="1313-2970-502-39" ID-Pensoft-UUID="6270FFBD2E505746FFB3AD432267D82E" ID-PubMed="26019663" ID-Zenodo-Dep="578839" ID-ZooBank="57086377EE59465484398B309B7374A0" ModsDocID="1313-2970-502-39" checkinTime="1451244355627" checkinUser="pensoft" docAuthor="Vives, Eduard" docDate="2015" docId="0AC327FA5130C5722991C055D9B763CD" docLanguage="en" docName="ZooKeys 502: 39-60" docOrigin="ZooKeys 502" docPubDate="2015-05-04" docSource="http://dx.doi.org/10.3897/zookeys.502.9049" docTitle="Trypogeus coarctatus Holzschuh 2006" docType="treatment" docVersion="5" id="6270FFBD2E505746FFB3AD432267D82E" lastPageNumber="49" masterDocId="6270FFBD2E505746FFB3AD432267D82E" masterDocTitle="Revision of the genus Trypogeus Lacordaire, 1869 (Cerambycidae, Dorcasominae)" masterLastPageNumber="60" masterPageNumber="39" pageNumber="48" updateTime="1668160313917" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Revision of the genus Trypogeus Lacordaire, 1869 (Cerambycidae, Dorcasominae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Vives, Eduard</mods:namePart>
<mods:affiliation>Facultat de Biologia, Departament de Biologia Animal, Avda. Diagonal, 643, 08028 Barcelona, Spain</mods:affiliation>
<mods:nameIdentifier type="email">eduard_vives@hotmail.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2015</mods:date>
<mods:detail type="pubDate">
<mods:number>2015-05-04</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>502</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>39</mods:start>
<mods:end>60</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.502.9049</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.502.9049</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-502-39</mods:identifier>
<mods:identifier type="ZooBank">57086377EE59465484398B309B7374A0</mods:identifier>
<mods:identifier type="Pensoft-UUID">6270FFBD2E505746FFB3AD432267D82E</mods:identifier>
<mods:identifier type="Zenodo-Dep">578839</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="152059963" LSID="urn:lsid:plazi:treatment:0AC327FA5130C5722991C055D9B763CD" httpUri="http://treatment.plazi.org/id/0AC327FA5130C5722991C055D9B763CD" lastPageId="10" lastPageNumber="49" pageId="9" pageNumber="48">
<subSubSection pageId="9" pageNumber="48" type="nomenclature">
<paragraph pageId="9" pageNumber="48">
<taxonomicName LSID="0AC327FA-5130-C572-2991-C055D9B763CD" authority="Holzschuh, 2006" authorityName="Holzschuh" authorityYear="2006" class="Insecta" family="Cerambycidae" genus="Trypogeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trypogeus coarctatus" order="Coleoptera" pageId="9" pageNumber="48" phylum="Arthropoda" rank="species" species="coarctatus">Trypogeus coarctatus Holzschuh, 2006</taxonomicName>
<figureCitation captionStart="Figures 1520" captionStartId="F4" captionText="Figures 15 - 20. 15 Trypogeus albicornis Lacordaire, holotype female, Malaysia; 16 Trypogeus aureopubens Pic, holotype male, Yunnan 17 Trypogeus barclayi Vives, holotype male, Brunei 18 Trypogeus cabigasi Vives, holotype female, Philippines 19 Trypogeus coarctatus Holzschuh, holotype male, Sumatra, (Photo: L. Dembicky) 20 Trypogeus javanicus Aurivillius, syntype male, Java." pageId="9" pageNumber="48">Fig. 19</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="10" pageNumber="49" type="reference_group">
<paragraph pageId="10" pageNumber="49">
<taxonomicName authorityName="Nonfried" authorityYear="1894" class="Insecta" family="Cerambycidae" genus="Trypogeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trypogeus fuscus" order="Coleoptera" pageId="10" pageNumber="49" phylum="Arthropoda" rank="species" species="fuscus">
<pageBreakToken pageId="10" pageNumber="49" start="start">Trypogeus</pageBreakToken>
fuscus
</taxonomicName>
: auct. nec
<taxonomicName class="Insecta" family="Cerambycidae" genus="Toxotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Toxotus fuscus" order="Coleoptera" pageId="10" pageNumber="49" phylum="Arthropoda" rank="species" species="fuscus">
<emphasis italics="true" pageId="10" pageNumber="49">Toxotus fuscus</emphasis>
</taxonomicName>
Nonfried, 1894: 209.
</paragraph>
<paragraph pageId="10" pageNumber="49">
<taxonomicName authorityName="Holzschuh" authorityYear="2006" class="Insecta" family="Cerambycidae" genus="Trypogeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trypogeus coarctatus" order="Coleoptera" pageId="10" pageNumber="49" phylum="Arthropoda" rank="species" species="coarctatus">Trypogeus coarctatus</taxonomicName>
Holzschuh, 2006: 207.
</paragraph>
</subSubSection>
<subSubSection pageId="10" pageNumber="49" type="material studied">
<paragraph pageId="10" pageNumber="49">Material studied.</paragraph>
<paragraph pageId="10" pageNumber="49">
1 male from Indonesia, Bandar Baru, Sumatra Utara, 3-V-1999, S. Tsuyuki leg. (NOC,); 1 male from Indonesia, N Sumatra, Brastagi, 27.IV-4.V-1988, A. Saito leg. (
<emphasis italics="true" pageId="10" pageNumber="49">in</emphasis>
NOC); 1 female from Indonesia, SW Sumatra, Marang, W. Doherty leg, 1890 (ex coll. Oberthur) (MNHN); 1 female from Indonesia, W Sumatra, Merapi, IV.1991, S. Ymada leg. (KMC).
</paragraph>
</subSubSection>
<subSubSection pageId="10" pageNumber="49" type="redescription">
<paragraph pageId="10" pageNumber="49">Redescription.</paragraph>
<paragraph pageId="10" pageNumber="49">
Size of the male: length 10-12 mm; width 3.6 mm. Size of the female: length 14-17 mm; width 4.2 mm. The general colour of the integument is testaceous yellow, males are darker and browner. Head brown except for the upper part which is yellowish. The antennae of males are completely brown, except for the last antennomere which is yellow; in females the scapus is testaceous, the antennomeres 3-8 are brown and the remaining 9-11 are yellow. The legs of the males are brown except for the femora which are mostly yellow. The legs of the females are completely yellowish. Pronotum brown in males with the discal area yellowish, in females it is almost completely testaceous. Entirely brown elytra, except for the basal area which is yellower, slightly lighter in the females. Elytra wider than pronotum with very obtuse humeri, rounded and projecting upwards. The suture is bordered and protruding. Apex of elytra rounded and dehiscent. The elytra are narrowed after the middle, widening slightly at apical fifth. Underside of the body and head brown except gular area which is yellow. Brown epimerae and abdominal sternites yellowish in both sexes. Large head with very short broad neck. The mandibles are long and almost entirely covered in golden tomentum. Translucid epistome, rectangular labrum bearing long golden setae on the free margin. Maxillary palpi longer than mandibles, with the last segment fusiform. Labial palpi shorter with the last segment cup-shaped. Base of the antennae with a granulous crest that extends to base of mandibles. Large prominent eyes, medium granulation. Posterior part of the head with dense golden tomentum. Antennae of males slightly longer than elytra, in females only reaching apical third. Long slender antennomeres, slightly flattened and angulose at the external distal part from antennomeres 4-10, the last is fusiform. Cylindrical pronotum, almost as wide as long in males, transverse in females, weakly arched sides with a barely protruding gibbosity before the middle. Discal area with four rather indistinct protuberances, the two anterior ones are very close to each other, the posterior pair slightly transverse in the males. Anterior border of pronotum finely margined, the posterior border is sinuate with double margination. Narrow prosternum, prosternal process in the shape of a fine lamina, hardly dilated behind. Widely opened procoxal cavities. Conical prominent coxae contiguous. Mesocoxae slightly separated in males and much more so in females, the metacoxae, contiguous in males, are separated in the females. Triangular scutellum rounded at apex. Elytral apex round and dehiscent. Elytra covered by golden tomentum which becomes reddish or black at the apex and margin of the apical third. Legs short and robust, covered by long silky golden pubescence. Tibiae slightly dilated apically. The male aedeagus (Fig.
<figureCitation captionStart="Figures 1114" captionStartId="F3" captionText="Figures 11 - 14. Tegmen, dorsal and lateral view of aedeagus 11 Trypogeus cabigasi Vives 12 Trypogeus albicornis Lacordaire 13 Trypogeus barclayi Vives 14 Trypogeus coarctatus Holzschuh, dorsal and lateral view of aedeagus and endophallus (drawings by Dr. N. Ohbayashi)." pageId="10" pageNumber="49">14</figureCitation>
) is long and slightly arched, acuminated at the apex, the lower lamina is distinctlylonger than the upper. Very simple endophalus lacking interior sclerites.
</paragraph>
</subSubSection>
<subSubSection pageId="10" pageNumber="49" type="remarks">
<paragraph pageId="10" pageNumber="49">Remarks.</paragraph>
<paragraph pageId="10" pageNumber="49">
Two syntype specimens described as
<taxonomicName class="Insecta" family="Cerambycidae" genus="Toxotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Toxotus fuscus" order="Coleoptera" pageId="10" pageNumber="49" phylum="Arthropoda" rank="species" species="fuscus">
<emphasis italics="true" pageId="10" pageNumber="49">Toxotus fuscus</emphasis>
</taxonomicName>
Nonfried, 1894, were examined that correspond in reality to
<taxonomicName authorityName="Pascoe" authorityYear="1886" class="Insecta" family="Cerambycidae" genus="Philus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Philus ophthalmicus" order="Coleoptera" pageId="10" pageNumber="49" phylum="Arthropoda" rank="species" species="ophthalmicus">
<emphasis italics="true" pageId="10" pageNumber="49">Philus ophthalmicus</emphasis>
</taxonomicName>
Pascoe, 1886 (Fig.
<figureCitation captionStart="Figures 18" captionStartId="F1" captionText="Figures 1 - 8. 1 Trypogeus barclayi Vives, left wing 2 Trypogeus sericeus female, tarsi from posterior leg 3 Trypogeus gressitti Miroshnikov, dorsal view of aedeagus 4 tegmen 5 pygidium and spiculum gastrale 6 Philus ophthalmicus Pascoe, (= Lectotype of Toxotus fuscus Nonfried) 7 - 8 Trypogeus fuscus labels of lectotype (DEI)." pageId="10" pageNumber="49">6</figureCitation>
). It is hereafter established that the species assigned as
<emphasis italics="true" pageId="10" pageNumber="49">
<taxonomicName authorityName="Lacordaire" authorityYear="1869" class="Insecta" family="Cerambycidae" genus="Trypogeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trypogeus" order="Coleoptera" pageId="10" pageNumber="49" phylum="Arthropoda" rank="genus">Trypogeus</taxonomicName>
fuscus auct.
</emphasis>
belong to
<taxonomicName authorityName="Holzschuh" authorityYear="2006" class="Insecta" family="Cerambycidae" genus="Trypogeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trypogeus coarctatus" order="Coleoptera" pageId="10" pageNumber="49" phylum="Arthropoda" rank="species" species="coarctatus">
<emphasis italics="true" pageId="10" pageNumber="49">Trypogeus coarctatus</emphasis>
</taxonomicName>
Holzschuh. A lectotype is designated between the two syntypes studied by Nonfried, in order to establish the exact taxonomic status of the species. Lectotype, 1 male from NW Sumatra, Tebing Tinggi, Dr. Schultheiss; 12.V.1884, (DEI) (Figs
<figureCitation captionStart="Figures 18" captionStartId="F1" captionText="Figures 1 - 8. 1 Trypogeus barclayi Vives, left wing 2 Trypogeus sericeus female, tarsi from posterior leg 3 Trypogeus gressitti Miroshnikov, dorsal view of aedeagus 4 tegmen 5 pygidium and spiculum gastrale 6 Philus ophthalmicus Pascoe, (= Lectotype of Toxotus fuscus Nonfried) 7 - 8 Trypogeus fuscus labels of lectotype (DEI)." pageId="10" pageNumber="49">7-8</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="10" pageNumber="49" type="distribution">
<paragraph pageId="10" pageNumber="49">Distribution.</paragraph>
<paragraph pageId="10" pageNumber="49">Indonesia (Sumatra).</paragraph>
</subSubSection>
</treatment>
</document>