234 lines
29 KiB
XML
234 lines
29 KiB
XML
<document id="961EEF765A50AF028A2C2A77BF72F3C4" ID-CLB-Dataset="30267" ID-DOI="10.11646/zootaxa.4407.4.4" ID-GBIF-Dataset="a850a99f-acfa-4f8e-b098-e7e783587f92" ID-ISSN="1175-5326" ID-Zenodo-Dep="1221080" ID-ZooBank="D112CD45-7907-4F42-9D32-0DC7DC3A39EB" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1524143490108" checkinUser="plazi" docAuthor="Shavrin, Alexey V." docDate="2018" docId="FD5D6D7EFFD4FF9A9D8066F0FBCB833E" docLanguage="en" docName="zootaxa.4407.4.4.pdf" docOrigin="Zootaxa 4407 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Mannerheimia taiwanica Shavrin 2018, sp.n." docType="treatment" docVersion="7" lastPageNumber="531" masterDocId="01641506FFD3FF909D176341FFB48743" masterDocTitle="Four new species of the genus Mannerheimia Mäklin, 1880 (Coleoptera: Staphylinidae: Omaliinae) from south-western China and Taiwan" masterLastPageNumber="532" masterPageNumber="521" pageNumber="528" updateTime="1698537035611" updateUser="ExternalLinkService">
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<mods:title id="2658896FE0DC95D43F79A412B9BFDBFB">Four new species of the genus Mannerheimia Mäklin, 1880 (Coleoptera: Staphylinidae: Omaliinae) from south-western China and Taiwan</mods:title>
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<mods:name id="1BEA4E9EBE2AA8E9F5442DDD7C716A51" type="personal">
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<mods:namePart id="8EB45C9C2FB30A607C452E66150BB273">Shavrin, Alexey V.</mods:namePart>
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<mods:title id="2FC95DEC756D87F261E64FDAA1BFA752">Zootaxa</mods:title>
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<mods:part id="7BDA8B6563162F98B750896C3E5CFBF4">
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<mods:date id="A81D4514B09BA1E2B823776822509B1E">2018</mods:date>
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<mods:number id="0D30D682B128D128437FCE55AECE9C7B">4407</mods:number>
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<treatment id="FD5D6D7EFFD4FF9A9D8066F0FBCB833E" ID-DOI="http://doi.org/10.5281/zenodo.5949835" ID-GBIF-Taxon="143605184" ID-Zenodo-Dep="5949835" LSID="urn:lsid:plazi:treatment:FD5D6D7EFFD4FF9A9D8066F0FBCB833E" httpUri="http://treatment.plazi.org/id/FD5D6D7EFFD4FF9A9D8066F0FBCB833E" lastPageId="10" lastPageNumber="531" pageId="7" pageNumber="528">
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<subSubSection id="3DEE8FE3FFD4FF979D8066F0FEC782AE" pageId="7" pageNumber="528" type="nomenclature">
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<paragraph id="754BDC68FFD4FF979D8066F0FDC8828F" blockId="7.[151,636,1457,1517]" box="[151,636,1457,1484]" pageId="7" pageNumber="528">
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<heading id="2E036B04FFD4FF979D8066F0FDC8828F" bold="true" box="[151,636,1457,1484]" fontSize="11" level="1" pageId="7" pageNumber="528" reason="1">
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<emphasis id="4780007AFFD4FF979D8066F0FDC8828F" bold="true" box="[151,636,1457,1484]" pageId="7" pageNumber="528">
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<taxonomicName id="B2F4A7EBFFD4FF979D8066F0FD80828F" ID-CoL="3XVMR" authority="Shavrin" authorityName="Shavrin" authorityYear="2018" box="[151,564,1457,1484]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="528" phylum="Arthropoda" rank="species" species="taiwanica" status="sp. nov.">
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<emphasis id="4780007AFFD4FF979D8066F0FE7C8288" bold="true" box="[151,456,1457,1483]" italics="true" pageId="7" pageNumber="528">Mannerheimia taiwanica</emphasis>
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Shavrin
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</taxonomicName>
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,
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<taxonomicNameLabel id="5CB3BD01FFD4FF979F5666F3FDC8828F" box="[577,636,1458,1484]" pageId="7" pageNumber="528" rank="species">sp.n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="754BDC68FFD4FF979D806695FEC782AE" blockId="7.[151,636,1457,1517]" box="[151,371,1492,1517]" pageId="7" pageNumber="528">
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(
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<figureCitation id="EDCFC0EDFFD4FF979D886695FF5A82AE" box="[159,238,1492,1517]" captionStart="FIGURES 1–4" captionStartId="3.[151,264,1963,1986]" captionTargetBox="[242,1406,245,1886]" captionTargetId="figure@3.[151,1436,235,1895]" captionTargetPageId="3" captionText="FIGURES 1–4. Habitus of Mannerheimia: 1—M. grandilobata (paratype), 2—M. emeishanensis (paratype), 3—M. maculata (paratype), 4—M. taiwanica (holotype).Scale bar:1.0 mm." httpUri="https://zenodo.org/record/1221082/files/figure.png" pageId="7" pageNumber="528">Figs. 4</figureCitation>
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,
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<figureCitation id="EDCFC0EDFFD4FF979DED6694FEAC82AE" box="[250,280,1493,1517]" captionStart="FIGURES 9–10" captionStartId="7.[151,264,1329,1352]" captionTargetBox="[201,1360,771,1291]" captionTargetId="figure@7.[187,1400,756,1330]" captionTargetPageId="7" captionText="FIGURES 9–10. Distribution of Mannerheimia in Yunnan and Sichuan, south-western China (9—M. grandilobata (black circles), M.emeishanensis (white circle), M.maculata (black triangle)) and Taiwan (10—M.taiwanica (black circles))." httpUri="https://zenodo.org/record/1221086/files/figure.png" pageId="7" pageNumber="528">10</figureCitation>
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,
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<figureCitation id="EDCFC0EDFFD4FF979C326695FED882AE" box="[293,364,1492,1517]" captionStart="FIGURES 12–15" captionStartId="9.[151,264,1980,2003]" captionTargetBox="[363,1366,219,1902]" captionTargetId="figure@9.[176,1411,193,1959]" captionTargetPageId="9" captionText="FIGURES 12–15. Aedeagus of Mannerheimia maculata (12–13: paratype) and M. taiwanica (14–15: paratype): 12, 14—parameral view, 13, 15—lateral view.Scale bar:0.1 mm." httpUri="https://zenodo.org/record/1221092/files/figure.png" pageId="7" pageNumber="528">14–15</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="3DEE8FE3FFD4FF979D80655DFC0A803A" pageId="7" pageNumber="528" type="materials_examined">
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<paragraph id="754BDC68FFD4FF979D80655DFD9B81E2" blockId="7.[151,1437,1564,2021]" pageId="7" pageNumber="528">
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<emphasis id="4780007AFFD4FF979D80655DFE668176" bold="true" box="[151,466,1564,1589]" pageId="7" pageNumber="528">
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<typeStatus id="AA4F62CAFFD4FF979D80655DFF668176" box="[151,210,1564,1589]" pageId="7" pageNumber="528">Type</typeStatus>
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material examined:
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</emphasis>
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<typeStatus id="AA4F62CAFFD4FF979CF6655DFDFD8176" box="[481,585,1564,1589]" pageId="7" pageNumber="528" type="holotype">Holotype</typeStatus>
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<specimenCount id="63F217E1FFD4FF979F4E655CFDC58177" box="[601,625,1565,1588]" pageId="7" pageNumber="528" type="male">♂</specimenCount>
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: ‘
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<collectingCountry id="0DE39CF8FFD4FF979F9E655CFD4F8176" box="[649,763,1565,1589]" name="Taiwan" pageId="7" pageNumber="528">TAIWAN</collectingCountry>
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<collectingRegion id="B730128AFFD4FF979E1D655DFCC68176" box="[778,882,1564,1589]" country="Taiwan" name="Taichung" pageId="7" pageNumber="528">Taichung</collectingRegion>
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| Hsien, Hsueshan | Chi-Ka
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<quantity id="B20C718DFFD4FF9799F6655DFA9B8176" box="[1249,1327,1564,1589]" metricMagnitude="3" metricUnit="m" metricValue="2.463" pageId="7" pageNumber="528" unit="m" value="2463.0">2463m</quantity>
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<date id="014AFAA8FFD4FF979828655DFA338176" box="[1343,1415,1564,1589]" pageId="7" pageNumber="528" value="1991-05-06">6.V.91</date>
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| A.Smetana [T67]’ <rectangular label, printed>, ‘?
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<taxonomicName id="B2F4A7EBFFD4FF979FE36501FC2B811A" authorityName="Shavrin" authorityYear="2018" box="[756,927,1600,1625]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="532" phylum="Arthropoda" rank="genus">Mannerheimia</taxonomicName>
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[handwritten] | M.K. Thayer det. 1998’ <rectangular label, printed>, ‘
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<typeStatus id="AA4F62CAFFD4FF979CE46524FD38813E" box="[499,652,1637,1661]" pageId="7" pageNumber="528" type="holotype">HOLOTYPE</typeStatus>
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<taxonomicName id="B2F4A7EBFFD4FF979FBB6525FCFB813E" authority="Shavrin, 2018" authorityName="Shavrin" authorityYear="2018" box="[684,847,1636,1661]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="532" phylum="Arthropoda" rank="genus" status="sp. nov.">Mannerheimia</taxonomicName>
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<taxonomicName id="B2F4A7EBFFD4FF979E786525FC6E813E" authority="Shavrin, 2018" authorityName="Shavrin" authorityYear="2018" box="[879,986,1636,1661]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="528" phylum="Arthropoda" rank="species" species="taiwanica" status="sp. nov.">taiwanica</taxonomicName>
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<taxonomicNameLabel id="5CB3BD01FFD4FF979EF06524FBA9813F" box="[999,1053,1637,1660]" pageId="7" pageNumber="528" rank="species">
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||
<emphasis id="4780007AFFD4FF979EF06524FBA9813F" bold="true" box="[999,1053,1637,1660]" pageId="7" pageNumber="528">sp.n.</emphasis>
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</taxonomicNameLabel>
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| Shavrin A.V. des. 2017’ <red rectangular label, printed> (
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<collectionCode id="13E544ADFFD4FF979CD965C8FD9781E2" box="[462,547,1673,1697]" country="Japan" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" pageId="7" pageNumber="528">NSMT</collectionCode>
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).
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</paragraph>
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<paragraph id="754BDC68FFD4FF979DD065ECFC0A803A" blockId="7.[151,1437,1564,2021]" pageId="7" pageNumber="528">
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Paratypes (4 specimens): 1 ♂ [specimen dissected]: ‘TAIWAN,Nantou | Hsien, Nenkaoshan | Tenchi Hut,
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<quantity id="B20C718DFFD4FF979D806591FF5081AA" box="[151,228,1744,1769]" metricMagnitude="3" metricUnit="m" metricValue="2.895" pageId="7" pageNumber="528" unit="m" value="2895.0">2895m</quantity>
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|
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<date id="014AFAA8FFD4FF979C156590FED381AA" box="[258,359,1745,1769]" pageId="7" pageNumber="528" value="1992-05-07">7.V.1992</date>
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| A. Smetana [T119]’ <rectangular label, printed> (NSMT); 1 ♂ [without aedeagus; apical abdominal segment glued on the same card under the specimen]: ‘TAIWAN Taichung | Hsien, Anmashan |
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<quantity id="B20C718DFFD4FF97985965B5FA28804E" box="[1358,1436,1780,1805]" metricMagnitude="3" metricUnit="m" metricValue="2.225" pageId="7" pageNumber="528" unit="m" value="2225.0">2225m</quantity>
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<date id="014AFAA8FFD4FF979D806458FF488072" box="[151,252,1817,1841]" pageId="7" pageNumber="528" value="1990-05-03">3.V.1990</date>
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| A. Smetana [T42]’ (CS); 2 ♀: ‘TAIWAN Taichung | Hsien, Anmashan, |
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<quantity id="B20C718DFFD4FF9799586459FB138073" box="[1103,1191,1816,1841]" metricMagnitude="3" metricUnit="m" metricValue="2.15" pageId="7" pageNumber="528" unit="m" value="2150.0">2150 m</quantity>
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,
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<date id="014AFAA8FFD4FF9799A36458FABF8072" box="[1204,1291,1817,1841]" pageId="7" pageNumber="528" value="1992-05-13">13.V.92</date>
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| A.Smetana [T129]’ (NSMT). All paratypes with additional red rectangular label: ‘PARATYPE |
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<taxonomicName id="B2F4A7EBFFD4FF979964647DFAA28016" authority="Shavrin, 2018" authorityName="Shavrin" authorityYear="2018" box="[1139,1302,1852,1877]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="532" phylum="Arthropoda" rank="genus" status="sp. nov.">Mannerheimia</taxonomicName>
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|
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<taxonomicName id="B2F4A7EBFFD4FF979826647DFA288016" authority="Shavrin, 2018" authorityName="Shavrin" authorityYear="2018" box="[1329,1436,1852,1877]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="528" phylum="Arthropoda" rank="species" species="taiwanica" status="sp. nov.">taiwanica</taxonomicName>
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<taxonomicNameLabel id="5CB3BD01FFD4FF979D806420FF78803B" box="[151,204,1889,1912]" pageId="7" pageNumber="528" rank="species">
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||
<emphasis id="4780007AFFD4FF979D806420FF78803B" bold="true" box="[151,204,1889,1912]" pageId="7" pageNumber="528">sp.n.</emphasis>
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||
</taxonomicNameLabel>
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| Shavrin A.V. des. 2017’ <red rectangular label, printed> (NSMT)
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="3DEE8FE3FFD4FF9A9DD064C5FCD98474" lastPageId="10" lastPageNumber="531" pageId="7" pageNumber="528" type="description">
|
||
<paragraph id="754BDC68FFD4FF979DD064C5FB7580A6" blockId="7.[151,1437,1564,2021]" pageId="7" pageNumber="528">
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||
<emphasis id="4780007AFFD4FF979DD064C5FEE580DE" bold="true" box="[199,337,1924,1949]" pageId="7" pageNumber="528">Description</emphasis>
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||
. Measurements (n=5): HW: 0.52–0.59; HL: 0.27–0.33; AL (
|
||
<typeStatus id="AA4F62CAFFD4FF97991664C5FBDE80DE" box="[1025,1130,1924,1949]" pageId="7" pageNumber="528" type="holotype">holotype</typeStatus>
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||
): 0.72; OL: 0.15–0.17; PL: 0.38–0.46; PW: 0.73–0.87; EL: 0.93–1.01; EW: 1.21–1.27; AW: 1.10–1.18; MTbL (
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<typeStatus id="AA4F62CAFFD4FF97999864E9FB4C8082" box="[1167,1272,1960,1985]" pageId="7" pageNumber="528" type="holotype">holotype</typeStatus>
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||
): 0.52; MTrL (
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<typeStatus id="AA4F62CAFFD4FF979D88648DFEBC80A6" box="[159,264,1996,2021]" pageId="7" pageNumber="528" type="holotype">holotype</typeStatus>
|
||
): 0.27 (MTrL 1–4: 0.17; MTrL 5: 0.10); AedL: 0.40; TL: 2.02–2.57 (
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<typeStatus id="AA4F62CAFFD4FF97991A648DFBC180A6" box="[1037,1141,1996,2021]" pageId="7" pageNumber="528" type="holotype">holotype</typeStatus>
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: 2.55).
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</paragraph>
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<paragraph id="754BDC68FFDBFF989DD063D6FBE2857F" blockId="8.[151,1437,151,573]" pageId="8" pageNumber="529">
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Body wide and convex. Head, elytra and abdomen reddish-brown (
|
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<typeStatus id="AA4F62CAFFDBFF989EAD63D6FBAA87F3" box="[954,1054,151,176]" pageId="8" pageNumber="529" type="holotype">holotype</typeStatus>
|
||
and one
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<typeStatus id="AA4F62CAFFDBFF98999263D9FB5187F3" box="[1157,1253,152,176]" pageId="8" pageNumber="529" type="paratype">paratype</typeStatus>
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||
with yellowishbrown abdomen); pronotum, laterobasal and sometimes apical portions of elytra, and paratergites yellow-brown; mouthparts, antennomeres and legs yellow. Forebody very shiny, without microsculpture; abdomen with fine isodiametric microsculpture. Head with irregular and sparse punctation, markedly denser on median portion between eyes and distinctly larger and deeper on basal portion of head and on neck; punctation of median portion of pronotum as that in median part of head but more regular and significantly denser, with interspaces between punctures as diameter of 1–3 puncture diameters, punctation of lateral portions of pronotum distinctly smaller, small oval medio-basal portion of pronotum impunctate; scutellum with several large punctures; punctation of elytra sparser than that of pronotum, denser and rugose on prescutellar portion, smaller on lateral and apical portions, median portions of each elytron with six to seven very tangled and vague longitudinal rows of punctures, remaining elytral punctures forming indistinct tangled more or less diagonal rows and becoming irregularly scattered laterally; abdominal tergites without visible punctation. Habitus as in
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<figureCitation id="EDCFC0EDFFDBFF9899196165FBE6857F" box="[1038,1106,548,573]" captionStart="FIGURES 1–4" captionStartId="3.[151,264,1963,1986]" captionTargetBox="[242,1406,245,1886]" captionTargetId="figure@3.[151,1436,235,1895]" captionTargetPageId="3" captionText="FIGURES 1–4. Habitus of Mannerheimia: 1—M. grandilobata (paratype), 2—M. emeishanensis (paratype), 3—M. maculata (paratype), 4—M. taiwanica (holotype).Scale bar:1.0 mm." httpUri="https://zenodo.org/record/1221082/files/figure.png" pageId="8" pageNumber="529">Fig. 4</figureCitation>
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.
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</paragraph>
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<caption id="218B8CE0FFDBFF989D80650BFF4C813C" httpUri="https://zenodo.org/record/1221088/files/figure.png" pageId="8" pageNumber="529" startId="8.[151,250,1610,1632]" targetBox="[151,1436,625,1589]" targetPageId="8">
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<paragraph id="754BDC68FFDBFF989D80650BFF4C813C" blockId="8.[151,1436,1610,1663]" pageId="8" pageNumber="529">
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||
<emphasis id="4780007AFFDBFF989D80650BFE96811C" bold="true" box="[151,290,1610,1632]" pageId="8" pageNumber="529">FIGURE 11.</emphasis>
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Type locality of
|
||
<taxonomicName id="B2F4A7EBFFDBFF989CC9650BFCB68123" authorityName="Shavrin" authorityYear="2018" box="[478,770,1610,1632]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="529" phylum="Arthropoda" rank="species" species="grandilobata">
|
||
<emphasis id="4780007AFFDBFF989CC9650BFCB68123" box="[478,770,1610,1632]" italics="true" pageId="8" pageNumber="529">Mannerheimia grandilobata</emphasis>
|
||
</taxonomicName>
|
||
in the Gaoligong Shan range, Yunnan, China (photograph: M. Schülke).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="754BDC68FFDBFF989DD065EAFABF803B" blockId="8.[151,1436,1707,1912]" pageId="8" pageNumber="529">
|
||
Head transverse, 1.7–1.9 times as wide as long, with distinctly elevated median portion, without occipital furrow between ocelli, with indistinct very short grooves in front of ocelli; anterior portion of head between antennal insertion and anterior margin of eye widely concave. Ocelli large and flattened; distance between ocelli about one and a half to two times as long as distance between ocellus and posterior margin of eye. Antennomere 4 as long as wide; length × width of antennomeres (
|
||
<typeStatus id="AA4F62CAFFDBFF989FD6647AFC9E8017" box="[705,810,1851,1876]" pageId="8" pageNumber="529" type="holotype">holotype</typeStatus>
|
||
): 1: 0.09 × 0.05; 2: 0.07 × 0.04; 3: 0.07 × 0.03; 4: 0.03 × 0.03; 5: 0.03 × 0.05; 6: 0.04 × 0.06; 7–8: 0.05 × 0.06; 9: 0.06 × 0.06; 10: 0.06 × 0.07; 11: 0.17 × 0.07.
|
||
</paragraph>
|
||
<caption id="218B8CE0FFDAFF999D8064FDFCBF80B1" httpUri="https://zenodo.org/record/1221092/files/figure.png" pageId="9" pageNumber="530" startId="9.[151,264,1980,2003]" targetBox="[363,1366,219,1902]" targetPageId="9">
|
||
<paragraph id="754BDC68FFDAFF999D8064FDFCBF80B1" blockId="9.[151,1435,1980,2034]" pageId="9" pageNumber="530">
|
||
<emphasis id="4780007AFFDAFF999D8064FDFEE88091" bold="true" box="[151,348,1980,2003]" pageId="9" pageNumber="530">FIGURES 12–15.</emphasis>
|
||
Aedeagus of
|
||
<taxonomicName id="B2F4A7EBFFDAFF999F1764FCFCB58090" authorityName="Shavrin" authorityYear="2018" box="[512,769,1981,2003]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="530" phylum="Arthropoda" rank="species" species="maculata">
|
||
<emphasis id="4780007AFFDAFF999F1764FCFCB58090" box="[512,769,1981,2003]" italics="true" pageId="9" pageNumber="530">Mannerheimia maculata</emphasis>
|
||
</taxonomicName>
|
||
(12
|
||
<emphasis id="4780007AFFDAFF999E2664FCFC8A8091" bold="true" box="[817,830,1981,2002]" pageId="9" pageNumber="530">–</emphasis>
|
||
13: paratype) and
|
||
<taxonomicName id="B2F4A7EBFFDAFF99991964FFFB288090" authorityName="Shavrin" authorityYear="2018" box="[1038,1180,1982,2003]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="530" phylum="Arthropoda" rank="species" species="taiwanica">
|
||
<emphasis id="4780007AFFDAFF99991964FFFB978090" box="[1038,1059,1982,2003]" italics="true" pageId="9" pageNumber="530">M</emphasis>
|
||
.
|
||
<emphasis id="4780007AFFDAFF99992F64FFFB288090" box="[1080,1180,1982,2003]" italics="true" pageId="9" pageNumber="530">taiwanica</emphasis>
|
||
</taxonomicName>
|
||
(14
|
||
<emphasis id="4780007AFFDAFF9999DB64FCFB6D8091" bold="true" box="[1228,1241,1981,2002]" pageId="9" pageNumber="530">–</emphasis>
|
||
15: paratype): 12, 14—parameral view, 13, 15—lateral view. Scale bar: 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="754BDC68FFD9FF9A9DD063D9FE9987BB" blockId="10.[151,1437,151,1149]" pageId="10" pageNumber="531">Pronotum transverse, 1.8–1.9 times as wide as long, 1.4 times as wide as head, widest about middle, gradually narrowed both anterad and posterad; anterior and posterior angles widely rounded; lateral portions narrow, without pit at middle.</paragraph>
|
||
<paragraph id="754BDC68FFD9FF9A9DD06245FCF78603" blockId="10.[151,1437,151,1149]" pageId="10" pageNumber="531">Elytra 1.2–1.3 times as wide as long, distinctly widened apicad, markedly more than twice as long as pronotum; latero-apical margin of elytra without crenulation.</paragraph>
|
||
<paragraph id="754BDC68FFD9FF9A9DD0620CFD288626" blockId="10.[151,1437,151,1149]" box="[199,668,332,357]" pageId="10" pageNumber="531">Metatarsus 1.9 times as long as metatibia.</paragraph>
|
||
<paragraph id="754BDC68FFD9FF9A9DD0622EFBAC86EE" blockId="10.[151,1437,151,1149]" pageId="10" pageNumber="531">Abdomen slightly narrower than elytra, with a pair of transverse moderately large tomentose spots on middle of tergite V; apical margin of abdominal tergite VII with narrow palisade fringe.</paragraph>
|
||
<paragraph id="754BDC68FFD9FF9A9DD062F6FE3F85C6" blockId="10.[151,1437,151,1149]" pageId="10" pageNumber="531">
|
||
Male. Protarsomeres 1–4 slightly widened. Apical margin of abdominal tergite VIII almost straight. Apical margin of abdominal sternite VIII slightly sinuate. Aedeagus (
|
||
<figureCitation id="EDCFC0EDFFD9FF9A9E79629DFC7786B7" box="[878,963,476,501]" captionStart="FIGURES 12–15" captionStartId="9.[151,264,1980,2003]" captionTargetBox="[363,1366,219,1902]" captionTargetId="figure@9.[176,1411,193,1959]" captionTargetPageId="9" captionText="FIGURES 12–15. Aedeagus of Mannerheimia maculata (12–13: paratype) and M. taiwanica (14–15: paratype): 12, 14—parameral view, 13, 15—lateral view.Scale bar:0.1 mm." httpUri="https://zenodo.org/record/1221092/files/figure.png" pageId="10" pageNumber="531">Fig. 14</figureCitation>
|
||
) with moderately wide basal part of the median lobe gradually narrowing toward narrow rounded apex; parameres wide, significantly distinctly apex of aedeagus, with very wide apical portions, gradually narrowing toward relatively obtuse apex; internal sac complicated, moderately narrow and short, with two fields of sclerites consisting of numerous thorns. Aedeagus laterally as in
|
||
<figureCitation id="EDCFC0EDFFD9FF9A9C22612DFE3385C6" box="[309,391,620,645]" captionStart="FIGURES 12–15" captionStartId="9.[151,264,1980,2003]" captionTargetBox="[363,1366,219,1902]" captionTargetId="figure@9.[176,1411,193,1959]" captionTargetPageId="9" captionText="FIGURES 12–15. Aedeagus of Mannerheimia maculata (12–13: paratype) and M. taiwanica (14–15: paratype): 12, 14—parameral view, 13, 15—lateral view.Scale bar:0.1 mm." httpUri="https://zenodo.org/record/1221092/files/figure.png" pageId="10" pageNumber="531">Fig. 15</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="754BDC68FFD9FF9A9DD061CEFA3A85EB" blockId="10.[151,1437,151,1149]" box="[199,1422,655,680]" pageId="10" pageNumber="531">Female. Protarsomeres 1–4 not widened. Apical margins of abdominal tergite VIII and sternite VIII rounded.</paragraph>
|
||
<paragraph id="754BDC68FFD9FF9A9DD061F5FCD98474" blockId="10.[151,1437,151,1149]" pageId="10" pageNumber="531">
|
||
<emphasis id="4780007AFFD9FF9A9DD061F5FE06858E" bold="true" box="[199,434,692,717]" pageId="10" pageNumber="531">Comparative notes.</emphasis>
|
||
Based on the body size, pale coloration of the body and general shape of the aedeagus, the new species is most similar to the Japanese
|
||
<taxonomicName id="B2F4A7EBFFD9FF9A9FD16198FC5B85B3" authority="(Sharp, 1889)" authorityName="Sharp" authorityYear="1889" baseAuthorityName="Sharp" baseAuthorityYear="1889" box="[710,1007,727,752]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="531" phylum="Arthropoda" rank="species" species="curtella">
|
||
<emphasis id="4780007AFFD9FF9A9FD16198FCF285B3" box="[710,838,728,752]" italics="true" pageId="10" pageNumber="531">M. curtella</emphasis>
|
||
(
|
||
<bibRefCitation id="1165A199FFD9FF9A9E4F6196FC5C85B3" box="[856,1000,727,752]" pageId="10" pageNumber="531" refId="ref8145" refString="Sharp, D. (1889) The Staphylinidae of Japan. The Annals and Magazine of Natural History, Series 6, 3, 463 - 476. https: // doi. org / 10.1080 / 00222938909460371" type="journal article">Sharp, 1889</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, known from Honshu, from which it differs by the shape of the pronotum (widest at about middle), shorter elytra, as well as by the narrower apical portion of the median lobe of the aedeagus and wider parameres.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="3DEE8FE3FFD9FF9A9DD06005FE21831B" pageId="10" pageNumber="531" type="distribution">
|
||
<paragraph id="754BDC68FFD9FF9A9DD06005FF5B84C3" blockId="10.[151,1437,151,1149]" pageId="10" pageNumber="531">
|
||
<emphasis id="4780007AFFD9FF9A9DD06005FED4841E" bold="true" box="[199,352,836,861]" pageId="10" pageNumber="531">Distribution.</emphasis>
|
||
The new species is known from several locations (
|
||
<figureCitation id="EDCFC0EDFFD9FF9A9E8E6005FC5E841F" box="[921,1002,836,861]" captionStart="FIGURES 9–10" captionStartId="7.[151,264,1329,1352]" captionTargetBox="[201,1360,771,1291]" captionTargetId="figure@7.[187,1400,756,1330]" captionTargetPageId="7" captionText="FIGURES 9–10. Distribution of Mannerheimia in Yunnan and Sichuan, south-western China (9—M. grandilobata (black circles), M.emeishanensis (white circle), M.maculata (black triangle)) and Taiwan (10—M.taiwanica (black circles))." httpUri="https://zenodo.org/record/1221086/files/figure.png" pageId="10" pageNumber="531">Fig. 10</figureCitation>
|
||
) in
|
||
<collectingRegion id="B730128AFFD9FF9A99026005FBC9841E" box="[1045,1149,836,861]" country="Taiwan" name="Taichung" pageId="10" pageNumber="531">Taichung</collectingRegion>
|
||
Hsien and
|
||
<collectingRegion id="B730128AFFD9FF9A99EC6004FA2C841E" box="[1275,1432,836,861]" country="Taiwan" name="Nantou" pageId="10" pageNumber="531">Nantou Hsien</collectingRegion>
|
||
,
|
||
<collectingCountry id="0DE39CF8FFD9FF9A9D806026FF5F84C3" box="[151,235,871,896]" name="Taiwan" pageId="10" pageNumber="531">Taiwan</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="754BDC68FFD9FF9A9DD060CDFE21831B" blockId="10.[151,1437,151,1149]" pageId="10" pageNumber="531">
|
||
<emphasis id="4780007AFFD9FF9A9DD060CDFEFE84E6" bold="true" box="[199,330,908,933]" pageId="10" pageNumber="531">Bionomics.</emphasis>
|
||
Specimens were collected at elevations from
|
||
<quantity id="B20C718DFFD9FF9A9E5960CDFBB484E7" box="[846,1024,908,933]" metricMagnitude="3" metricUnit="m" metricValue="2.56" metricValueMax="2.895" metricValueMin="2.225" pageId="10" pageNumber="531" unit="m" value="2560.0" valueMax="2895.0" valueMin="2225.0">
|
||
<date id="014AFAA8FFD9FF9A9E5960CDFC5684E6" box="[846,994,908,933]" pageId="10" pageNumber="531" value="2225" valueMax="2895">2225 to 2895</date>
|
||
m
|
||
</quantity>
|
||
a.s.l. and were taken by sifting from moldy rotten wood, humus and other debris on and along fallen large trees in original forest (locality: T67), from leaf litter and other debris along fallen large trees in original forest (locality: T42), from wet debris (leaf litter twigs, moss) along vertical walls along the trail in open
|
||
<taxonomicName id="B2F4A7EBFFD9FF9A9E0560B9FCE58353" box="[786,849,1016,1040]" class="Pinopsida" family="Pinaceae" genus="Abies" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="10" pageNumber="531" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="4780007AFFD9FF9A9E0560B9FCE58353" box="[786,849,1016,1040]" italics="true" pageId="10" pageNumber="531">Abies</emphasis>
|
||
</taxonomicName>
|
||
forest with lush undergrowth (locality: T119), from fallen leaves and fermenting parts of buds of deciduous trees, accumulated along the road through mixed deciduous forest (locality: T129).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="3DEE8FE3FFD9FF9A9DD06725FBCB833E" box="[199,1151,1124,1149]" pageId="10" pageNumber="531" type="etymology">
|
||
<paragraph id="754BDC68FFD9FF9A9DD06725FBCB833E" blockId="10.[151,1437,151,1149]" box="[199,1151,1124,1149]" pageId="10" pageNumber="531">
|
||
<emphasis id="4780007AFFD9FF9A9DD06725FEFA833E" bold="true" box="[199,334,1124,1149]" pageId="10" pageNumber="531">Etymology.</emphasis>
|
||
The new species is named after
|
||
<collectingCountry id="0DE39CF8FFD9FF9A9FAD6725FCB8833E" box="[698,780,1124,1149]" name="Taiwan" pageId="10" pageNumber="531">Taiwan</collectingCountry>
|
||
where the species was collected.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |