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<document id="BC0B07D33F418FB393FDDEA77B0AF87A" ID-DOI="10.11646/zootaxa.4012.1.4" ID-GBIF-Dataset="d32dd252-5a36-4368-a636-6e688bc4ed13" ID-ISSN="1175-5326" ID-Zenodo-Dep="232879" ID-ZooBank="C3B3A06D-C18C-4B66-83E9-51B605E11862" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461226104448" checkinUser="plazi" docAuthor="Kobieluszova, Lucie &amp; Kundrata, Robin" docDate="2015" docId="C3140C0EFF8AFFF43DBFB7B75E5AC265" docLanguage="en" docName="zt04012p096.pdf" docOrigin="Zootaxa 4012 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Drilus obscuricornis Pic 1899" docType="treatment" docVersion="8" masterDocId="3F2D7476FF82FFFD3D28B31B5D1EC058" masterDocTitle="Taxonomic review of Drilus Olivier, 1790 (Elateridae: Agrypninae: Drilini) from Asia Minor, with descriptions of seven new species and comments on the female antennal morphology in Drilini" updateTime="1698604902572" updateUser="plazi">
<mods:mods id="4E4A1A646DDFA90C101C19F5071A2792" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="B6CDA18E7212FAB3C19B9A6C0C13C1CF">
<mods:title id="A2595030A054B301D4BCE1CCF1BB40DB">Taxonomic review of Drilus Olivier, 1790 (Elateridae: Agrypninae: Drilini) from Asia Minor, with descriptions of seven new species and comments on the female antennal morphology in Drilini</mods:title>
</mods:titleInfo>
<mods:name id="F0069A7AFD1023CF11F6239AF36EC78B" type="personal">
<mods:role id="7F1674E14103CFCF9356F3372937B28C">
<mods:roleTerm id="414CD21E8C0232727DDDEE11522B4642">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5C9E81D52435E1CE0E723BA063AC5E7D">Kobieluszova, Lucie</mods:namePart>
</mods:name>
<mods:name id="DA71E3D0A4F2662DF1257339FA6444C7" type="personal">
<mods:role id="3DCABDE36DBE274C28D8B606CB12DA6F">
<mods:roleTerm id="5D1A64E3F7271B5DC48C91841A8D1392">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="07995354F06D785C3EE98D76158F58BC">Kundrata, Robin</mods:namePart>
</mods:name>
<mods:typeOfResource id="514B3AABAE428456ABC31E1E9A1AC4F7">text</mods:typeOfResource>
<mods:relatedItem id="DC229644800AC59BC6A9122BA5AA5E4C" type="host">
<mods:titleInfo id="17EBC974F664400E8CBF79BC2470571C">
<mods:title id="3DE7DC5A02E921004B82724D4F481ECE">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="C03AFD4B1E6A3751F529398E1F3488B4">
<mods:date id="00A806F85014D9B71AB967599E829832">2015</mods:date>
<mods:detail id="65356DBF054469772314355C08EA917A" type="volume">
<mods:number id="4CA620BA270473ED3591492ABD4DC66E">4012</mods:number>
</mods:detail>
<mods:detail id="F7FB1D75BF57919AD5DC8BBDFCE4377A" type="issue">
<mods:number id="395BAF6BCDF385B55E3037A754F9887F">1</mods:number>
</mods:detail>
</mods:part>
</mods:relatedItem>
<mods:classification id="9C1696ACA0F8CD595C3B6D79598EFA12">journal volume</mods:classification>
<mods:identifier id="C4A39B5123C1C1E454264C1808800FB3" type="DOI">10.11646/zootaxa.4012.1.4</mods:identifier>
<mods:identifier id="FAC9659C1AF9AE149497C10FC6064BBE" type="GBIF-Dataset">d32dd252-5a36-4368-a636-6e688bc4ed13</mods:identifier>
<mods:identifier id="2B5ECCE53268B9F395B7031764A9F234" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="324131C0164B5DB3D47DEF59140813A7" type="Zenodo-Dep">232879</mods:identifier>
<mods:identifier id="E880E1EA8F0B1FEB3AD3502F630539DC" type="ZooBank">C3B3A06D-C18C-4B66-83E9-51B605E11862</mods:identifier>
</mods:mods>
<treatment id="C3140C0EFF8AFFF43DBFB7B75E5AC265" ID-DOI="http://doi.org/10.5281/zenodo.6112430" ID-GBIF-Taxon="119632739" ID-Zenodo-Dep="6112430" LSID="urn:lsid:plazi:treatment:C3140C0EFF8AFFF43DBFB7B75E5AC265" httpUri="http://treatment.plazi.org/id/C3140C0EFF8AFFF43DBFB7B75E5AC265" lastPageId="9" pageId="8">
<subSubSection id="03A7EE93FF8AFFF53DBFB7B75C72C4BF" pageId="8" type="nomenclature">
<paragraph id="4B02BD18FF8AFFF53DBFB7B75F19C49E" blockId="8.[151,519,1196,1255]" box="[151,519,1196,1222]" pageId="8">
<heading id="104A0A74FF8AFFF53DBFB7B75F19C49E" bold="true" box="[151,519,1196,1222]" fontSize="11" level="1" pageId="8" reason="1">
<taxonomicName id="8CBDC69BFF8AFFF53DBFB7B75F19C49E" authority="Pic, 1899" authorityName="Pic" authorityYear="1899" box="[151,519,1196,1222]" class="Insecta" family="Drilidae" genus="Drilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" phylum="Arthropoda" rank="species" species="obscuricornis">
<emphasis id="79C9610AFF8AFFF53DBFB7B75F19C49E" bold="true" box="[151,519,1196,1222]" pageId="8">
<emphasis id="79C9610AFF8AFFF53DBFB7B75C91C49E" bold="true" box="[151,399,1196,1222]" italics="true" pageId="8">Drilus obscuricornis</emphasis>
<bibRefCitation id="2F2CC0E9FF8AFFF53CBEB7B75F19C49E" author="Pic" box="[406,519,1196,1222]" pageId="8" refString="Pic, M. (1899) Diagnoses de coleopteres d`Orient. Bulletin de la Societe d`Histoire Naturelle d`Autun, 12, 204 - 210." type="journal article" year="1899">Pic, 1899</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="4B02BD18FF8AFFF53DBFB7D55C72C4BF" blockId="8.[151,519,1196,1255]" box="[151,364,1230,1255]" pageId="8">
(
<figureCitation id="D386A19DFF8AFFF53DB7B7D55DF9C4BF" box="[159,231,1230,1255]" captionStart="FIGURES 1 – 12" captionStartId="2.[151,264,1908,1931]" captionTargetBox="[181,1396,203,1850]" captionTargetId="figure@2.[172,1408,195,1856]" captionTargetPageId="2" captionText="FIGURES 1 – 12. Habitus images of Drilus species. 1, Drilus akbesianus (Fairmaire, 1895); 2, D. badius sp. nov.; 3, D. huijbregtsi sp. nov.; 4, D. latithorax Pic, 1902; 5, D. mertliki sp. nov.; 6, D. obscuricornis Pic, 1899; 7, D. rectus Schaufuss, 1867; 8, D. robustus sp. nov.; 9, D. sanliurfensis sp. nov.; 10, D. teunisseni sp. nov.; 11, D. turcicus sp. nov.; 12, D. iranicus Wittmer, 1967. Scale bars = 2.0 mm." httpUri="https://zenodo.org/record/232880/files/figure.png" pageId="8">Figs 6</figureCitation>
,
<figureCitation id="D386A19DFF8AFFF53DDBB7D45C0FC4BF" box="[243,273,1231,1255]" captionStart="FIGURES 13 – 21" captionStartId="3.[151,264,1423,1446]" captionTargetBox="[161,1430,204,1401]" captionTargetId="figure@3.[151,1436,193,1403]" captionTargetPageId="3" captionText="FIGURES 13 – 21. Pronota of Drilus species. 13, Drilus badius sp. nov.; 14, D. huijbregtsi sp. nov.; 15, D. latithorax Pic, 1902; 16, D. mertliki sp. nov.; 17, D. obscuricornis Pic, 1899; 18, D. robustus sp. nov.; 19, D. sanliurfensis sp. nov.; 20, D. teunisseni sp. nov.; 21, D. turcicus sp. nov. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/232881/files/figure.png" pageId="8">17</figureCitation>
,
<figureCitation id="D386A19DFF8AFFF53C35B7D55C25C4BF" box="[285,315,1230,1255]" captionStart="FIGURES 22 – 33" captionStartId="4.[151,264,1794,1817]" captionTargetBox="[155,1414,201,1764]" captionTargetId="figure@4.[151,1436,193,1773]" captionTargetPageId="4" captionText="FIGURES 22 – 33. Antennae of Drilus species. 22, Drilus badius sp. nov.; 23, D. huijbregtsi sp. nov.; 24, D. latithorax Pic, 1902; 25, D. mertliki sp. nov.; 26, D. obscuricornis Pic, 1899; 27, D. rectus Schaufuss, 1867; 28, D. robustus sp. nov.; 29, D. sanliurfensis sp. nov.; 30, D. teunisseni sp. nov.; 31, D. turcicus sp. nov. Drilus robustus sp. nov., male. 32, leg; 33, last abdominal segments. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/232882/files/figure.png" pageId="8">26</figureCitation>
,
<figureCitation id="D386A19DFF8AFFF53C6FB7D45C7AC4BF" box="[327,356,1231,1255]" captionStart="FIGURES 34 – 45" captionStartId="6.[151,264,1845,1868]" captionTargetBox="[163,1430,529,1821]" captionTargetId="figure@6.[151,1436,517,1825]" captionTargetPageId="6" captionText="FIGURES 34 – 45. Male genitalia of Drilus species. 34, Drilus badius sp. nov.; 35, D. huijbregtsi sp. nov.; 36, D. latithorax Pic, 1902; 37, D. mertliki sp. nov.; 38, D. obscuricornis Pic, 1899; 39, D. rectus Schaufuss, 1867 (Turkey, Adana Prov.); 40, D. robustus sp. nov.; 41, D. sanliurfensis sp. nov.; 42, D. teunisseni sp. nov.; 43, D. turcicus sp. nov.; 44, D. iranicus Wittmer, 1967; 45, Drilus sp. (Abkhazia). Scale bars = 0.25 mm." httpUri="https://zenodo.org/record/232883/files/figure.png" pageId="8">38</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="03A7EE93FF8AFFF53DBFB60C5F17C574" box="[151,521,1302,1324]" pageId="8" type="reference_group">
<paragraph id="4B02BD18FF8AFFF53DBFB60C5F17C574" blockId="8.[151,521,1302,1324]" box="[151,521,1302,1324]" pageId="8">
<treatmentCitationGroup id="6BAD9A36FF8AFFF53DBFB60C5F17C574" box="[151,521,1302,1324]" pageId="8">
<taxonomicName id="8CBDC69BFF8AFFF53DBFB60C5F1BC574" authority="Pic, 1899: 205" authorityName="Pic" authorityPageNumber="205" authorityYear="1899" box="[151,517,1302,1324]" class="Insecta" family="Drilidae" genus="Drilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" phylum="Arthropoda" rank="species" species="obscuricornis">
<emphasis id="79C9610AFF8AFFF53DBFB60C5C77C574" box="[151,361,1303,1324]" italics="true" pageId="8">Drilus obscuricornis</emphasis>
<treatmentCitation id="CA1C9B09FF8AFFF53C47B60D5F1BC574" author="Pic" box="[367,517,1302,1324]" page="205" pageId="8" year="1899">
<bibRefCitation id="2F2CC0E9FF8AFFF53C47B60D5CCCC574" author="Pic" box="[367,466,1302,1324]" pageId="8" refString="Pic, M. (1899) Diagnoses de coleopteres d`Orient. Bulletin de la Societe d`Histoire Naturelle d`Autun, 12, 204 - 210." type="journal article" year="1899">Pic, 1899</bibRefCitation>
: 205
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="03A7EE93FF8AFFF53DBFB6425E0DC5E1" pageId="8" type="materials_examined">
<paragraph id="4B02BD18FF8AFFF53DBFB6425FEFC5CE" blockId="8.[151,1437,1369,2006]" pageId="8">
<emphasis id="79C9610AFF8AFFF53DBFB6425C58C52A" bold="true" box="[151,326,1369,1394]" pageId="8">
<typeStatus id="940603BAFF8AFFF53DBFB6425DCCC52A" box="[151,210,1369,1394]" pageId="8">Type</typeStatus>
material.
</emphasis>
<typeStatus id="940603BAFF8AFFF53C65B6425CA6C52A" box="[333,440,1369,1394]" pageId="8" type="holotype">Holotype</typeStatus>
, male, &quot;Asie Min., Brousse,
<bibRefCitation id="2F2CC0E9FF8AFFF53E28B6425E7AC52A" author="Pic" box="[768,868,1369,1394]" pageId="8" refString="Pic, M. (1899) Diagnoses de coleopteres d`Orient. Bulletin de la Societe d`Histoire Naturelle d`Autun, 12, 204 - 210." type="journal article" year="1899">Pic 1899</bibRefCitation>
[printed label],
<typeStatus id="940603BAFF8AFFF53934B641594DC52A" box="[1052,1107,1370,1394]" pageId="8">Type</typeStatus>
[handwritten],
<typeStatus id="940603BAFF8AFFF5382BB6415824C52A" box="[1283,1338,1370,1394]" pageId="8">Type</typeStatus>
[printed red label],
<taxonomicName id="8CBDC69BFF8AFFF53C27B6665CCAC5CE" authority="Pic" authorityName="Pic" box="[271,468,1405,1430]" class="Insecta" family="Drilidae" genus="Drilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" phylum="Arthropoda" rank="species" species="obscuricornis">
<emphasis id="79C9610AFF8AFFF53C27B6665CB7C5CD" box="[271,425,1405,1429]" italics="true" pageId="8">obscuricornis</emphasis>
Pic
</taxonomicName>
[handwritten]&quot; (
<collectionCode id="2DAC25DDFF8AFFF53FA6B6655FFBC5CE" box="[654,741,1406,1430]" httpUri="http://grbio.org/cool/a5xg-x4h4" name="Museum d'Histoire Naturelle Perpignan" pageId="8">MHNP</collectionCode>
).
</paragraph>
<paragraph id="4B02BD18FF8AFFF53DEFB6BA5E0DC5E1" blockId="8.[151,1437,1369,2006]" box="[199,787,1441,1466]" pageId="8">
<emphasis id="79C9610AFF8AFFF53DEFB6BA5C7BC5E2" bold="true" box="[199,357,1441,1466]" pageId="8">
<typeStatus id="940603BAFF8AFFF53DEFB6BA5C1CC5E2" box="[199,258,1441,1466]" pageId="8">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="33AAFD88FF8AFFF53C44B6BA5CA3C5E2" box="[364,445,1441,1466]" name="Turkey" pageId="8">Turkey</collectingCountry>
, &quot;Anatolie, Brusse&quot; [= Bursa].
</paragraph>
</subSubSection>
<subSubSection id="03A7EE93FF8AFFF53DEFB6DE5FD5C636" pageId="8" type="diagnosis">
<paragraph id="4B02BD18FF8AFFF53DEFB6DE5FD5C636" blockId="8.[151,1437,1369,2006]" pageId="8">
<emphasis id="79C9610AFF8AFFF53DEFB6DE5C5FC586" bold="true" box="[199,321,1477,1502]" pageId="8">Diagnosis.</emphasis>
<taxonomicName id="8CBDC69BFF8AFFF53C62B6DE5F2AC585" box="[330,564,1477,1501]" class="Insecta" family="Drilidae" genus="Drilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" phylum="Arthropoda" rank="species" species="obscuricornis">
<emphasis id="79C9610AFF8AFFF53C62B6DE5F2AC585" box="[330,564,1477,1501]" italics="true" pageId="8">Drilus obscuricornis</emphasis>
</taxonomicName>
shares more or less uniform body coloration and weakly pectinate antennae with
<taxonomicName id="8CBDC69BFF8AFFF53DFAB6F15C5FC659" box="[210,321,1513,1537]" class="Insecta" family="Drilidae" genus="Drilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" phylum="Arthropoda" rank="species" species="badius" status="sp. nov.">
<emphasis id="79C9610AFF8AFFF53DFAB6F15DF8C659" box="[210,230,1514,1537]" italics="true" pageId="8">D</emphasis>
.
<emphasis id="79C9610AFF8AFFF53DDEB6F25C5FC659" box="[246,321,1513,1537]" italics="true" pageId="8">badius</emphasis>
</taxonomicName>
<emphasis id="79C9610AFF8AFFF53C62B6F25CB8C65A" bold="true" box="[330,422,1513,1538]" pageId="8">
<taxonomicNameLabel id="62FADC71FF8AFFF53C62B6F25CB8C65A" box="[330,422,1513,1538]" pageId="8" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="8CBDC69BFF8AFFF53C9EB6F15F4CC659" box="[438,594,1513,1537]" class="Insecta" family="Drilidae" genus="Drilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" phylum="Arthropoda" rank="species" species="huijbregtsi" status="sp. nov.">
<emphasis id="79C9610AFF8AFFF53C9EB6F15CD4C659" box="[438,458,1514,1537]" italics="true" pageId="8">D</emphasis>
.
<emphasis id="79C9610AFF8AFFF53CF2B6F25F4CC659" box="[474,594,1513,1537]" italics="true" pageId="8">huijbregtsi</emphasis>
</taxonomicName>
<emphasis id="79C9610AFF8AFFF53F74B6F25FA9C65A" bold="true" box="[604,695,1513,1538]" pageId="8">
<taxonomicNameLabel id="62FADC71FF8AFFF53F74B6F25FA9C65A" box="[604,695,1513,1538]" pageId="8" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, and
<taxonomicName id="8CBDC69BFF8AFFF53FD1B6F15E69C659" box="[761,887,1514,1537]" class="Insecta" family="Drilidae" genus="Drilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" phylum="Arthropoda" rank="species" species="turcicus" status="sp. nov.">
<emphasis id="79C9610AFF8AFFF53FD1B6F15E13C659" box="[761,781,1514,1537]" italics="true" pageId="8">D</emphasis>
.
<emphasis id="79C9610AFF8AFFF53E35B6F15E69C659" box="[797,887,1514,1537]" italics="true" pageId="8">turcicus</emphasis>
</taxonomicName>
<emphasis id="79C9610AFF8AFFF53E57B6F25EC5C65A" bold="true" box="[895,987,1513,1538]" pageId="8">
<taxonomicNameLabel id="62FADC71FF8AFFF53E57B6F25EC5C65A" box="[895,987,1513,1538]" pageId="8" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
However,
<taxonomicName id="8CBDC69BFF8AFFF53972B6F159D7C659" box="[1114,1225,1513,1537]" class="Insecta" family="Drilidae" genus="Drilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" phylum="Arthropoda" rank="species" species="badius" status="sp. nov.">
<emphasis id="79C9610AFF8AFFF53972B6F15970C659" box="[1114,1134,1514,1537]" italics="true" pageId="8">D</emphasis>
.
<emphasis id="79C9610AFF8AFFF53956B6F259D7C659" box="[1150,1225,1513,1537]" italics="true" pageId="8">badius</emphasis>
</taxonomicName>
<emphasis id="79C9610AFF8AFFF539FAB6F25830C65A" bold="true" box="[1234,1326,1513,1538]" pageId="8">
<taxonomicNameLabel id="62FADC71FF8AFFF539FAB6F25830C65A" box="[1234,1326,1513,1538]" pageId="8" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is lightly colored and has finer, less distinct pronotal punctation (
<figureCitation id="D386A19DFF8AFFF53E3EB5165E9EC67E" box="[790,896,1549,1574]" captionStart="FIGURES 1 – 12" captionStartId="2.[151,264,1908,1931]" captionTargetBox="[181,1396,203,1850]" captionTargetId="figure@2.[172,1408,195,1856]" captionTargetPageId="2" captionText="FIGURES 1 – 12. Habitus images of Drilus species. 1, Drilus akbesianus (Fairmaire, 1895); 2, D. badius sp. nov.; 3, D. huijbregtsi sp. nov.; 4, D. latithorax Pic, 1902; 5, D. mertliki sp. nov.; 6, D. obscuricornis Pic, 1899; 7, D. rectus Schaufuss, 1867; 8, D. robustus sp. nov.; 9, D. sanliurfensis sp. nov.; 10, D. teunisseni sp. nov.; 11, D. turcicus sp. nov.; 12, D. iranicus Wittmer, 1967. Scale bars = 2.0 mm." httpUri="https://zenodo.org/record/232880/files/figure.png" pageId="8">Figs 2, 6</figureCitation>
),
<taxonomicName id="8CBDC69BFF8AFFF53EBFB515592AC67D" box="[919,1076,1549,1573]" class="Insecta" family="Drilidae" genus="Drilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" phylum="Arthropoda" rank="species" species="huijbregtsi" status="sp. nov.">
<emphasis id="79C9610AFF8AFFF53EBFB5155EB5C67D" box="[919,939,1550,1573]" italics="true" pageId="8">D</emphasis>
.
<emphasis id="79C9610AFF8AFFF53E94B516592AC67D" box="[956,1076,1549,1573]" italics="true" pageId="8">huijbregtsi</emphasis>
</taxonomicName>
<emphasis id="79C9610AFF8AFFF53916B5165984C67E" bold="true" box="[1086,1178,1549,1574]" pageId="8">
<taxonomicNameLabel id="62FADC71FF8AFFF53916B5165984C67E" box="[1086,1178,1549,1574]" pageId="8" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
has finer, less distinct pronotal punctures and more slender male genitalia (
<figureCitation id="D386A19DFF8AFFF53FD2B52A5E9FC612" box="[762,897,1585,1610]" captionStart="FIGURES 13 – 21" captionStartId="3.[151,264,1423,1446]" captionTargetBox="[161,1430,204,1401]" captionTargetId="figure@3.[151,1436,193,1403]" captionTargetPageId="3" captionText="FIGURES 13 – 21. Pronota of Drilus species. 13, Drilus badius sp. nov.; 14, D. huijbregtsi sp. nov.; 15, D. latithorax Pic, 1902; 16, D. mertliki sp. nov.; 17, D. obscuricornis Pic, 1899; 18, D. robustus sp. nov.; 19, D. sanliurfensis sp. nov.; 20, D. teunisseni sp. nov.; 21, D. turcicus sp. nov. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/232881/files/figure.png" pageId="8">Figs 14, 17</figureCitation>
,
<figureCitation id="D386A19DFF8AFFF53EB8B52A5EC2C612" box="[912,988,1585,1610]" captionStart="FIGURES 34 – 45" captionStartId="6.[151,264,1845,1868]" captionTargetBox="[163,1430,529,1821]" captionTargetId="figure@6.[151,1436,517,1825]" captionTargetPageId="6" captionText="FIGURES 34 – 45. Male genitalia of Drilus species. 34, Drilus badius sp. nov.; 35, D. huijbregtsi sp. nov.; 36, D. latithorax Pic, 1902; 37, D. mertliki sp. nov.; 38, D. obscuricornis Pic, 1899; 39, D. rectus Schaufuss, 1867 (Turkey, Adana Prov.); 40, D. robustus sp. nov.; 41, D. sanliurfensis sp. nov.; 42, D. teunisseni sp. nov.; 43, D. turcicus sp. nov.; 44, D. iranicus Wittmer, 1967; 45, Drilus sp. (Abkhazia). Scale bars = 0.25 mm." httpUri="https://zenodo.org/record/232883/files/figure.png" pageId="8">35, 38</figureCitation>
), and
<taxonomicName id="8CBDC69BFF8AFFF5390EB52959BBC611" box="[1062,1189,1586,1609]" class="Insecta" family="Drilidae" genus="Drilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" phylum="Arthropoda" rank="species" species="turcicus" status="sp. nov.">
<emphasis id="79C9610AFF8AFFF5390EB5295924C611" box="[1062,1082,1586,1609]" italics="true" pageId="8">D</emphasis>
.
<emphasis id="79C9610AFF8AFFF53964B52959BBC611" box="[1100,1189,1586,1609]" italics="true" pageId="8">turcicus</emphasis>
</taxonomicName>
<emphasis id="79C9610AFF8AFFF53987B52A5812C612" bold="true" box="[1199,1292,1585,1610]" pageId="8">
<taxonomicNameLabel id="62FADC71FF8AFFF53987B52A5812C612" box="[1199,1292,1585,1610]" pageId="8" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
has a wider clypeus and short rounded paramera (
<figureCitation id="D386A19DFF8AFFF53F13B54E5FA0C636" box="[571,702,1621,1646]" captionStart="FIGURES 34 – 45" captionStartId="6.[151,264,1845,1868]" captionTargetBox="[163,1430,529,1821]" captionTargetId="figure@6.[151,1436,517,1825]" captionTargetPageId="6" captionText="FIGURES 34 – 45. Male genitalia of Drilus species. 34, Drilus badius sp. nov.; 35, D. huijbregtsi sp. nov.; 36, D. latithorax Pic, 1902; 37, D. mertliki sp. nov.; 38, D. obscuricornis Pic, 1899; 39, D. rectus Schaufuss, 1867 (Turkey, Adana Prov.); 40, D. robustus sp. nov.; 41, D. sanliurfensis sp. nov.; 42, D. teunisseni sp. nov.; 43, D. turcicus sp. nov.; 44, D. iranicus Wittmer, 1967; 45, Drilus sp. (Abkhazia). Scale bars = 0.25 mm." httpUri="https://zenodo.org/record/232883/files/figure.png" pageId="8">Figs 38, 43</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="03A7EE93FF8AFFF43DEFB5625C60C24F" lastPageId="9" pageId="8" type="description">
<paragraph id="4B02BD18FF8AFFF43DEFB5625C65C188" blockId="8.[151,1437,1369,2006]" lastBlockId="9.[151,1437,151,573]" lastPageId="9" pageId="8">
<emphasis id="79C9610AFF8AFFF53DEFB5625C47C6CA" bold="true" box="[199,345,1657,1682]" pageId="8">Description.</emphasis>
Male (
<typeStatus id="940603BAFF8AFFF53C80B5625F0FC6CA" box="[424,529,1657,1682]" pageId="8" type="holotype">holotype</typeStatus>
). Body 3.0 times longer than width at humeri, dark brown to black, covered with yellow pubescence (
<figureCitation id="D386A19DFF8AFFF53C54B5865CDFC6EE" box="[380,449,1693,1718]" captionStart="FIGURES 1 – 12" captionStartId="2.[151,264,1908,1931]" captionTargetBox="[181,1396,203,1850]" captionTargetId="figure@2.[172,1408,195,1856]" captionTargetPageId="2" captionText="FIGURES 1 – 12. Habitus images of Drilus species. 1, Drilus akbesianus (Fairmaire, 1895); 2, D. badius sp. nov.; 3, D. huijbregtsi sp. nov.; 4, D. latithorax Pic, 1902; 5, D. mertliki sp. nov.; 6, D. obscuricornis Pic, 1899; 7, D. rectus Schaufuss, 1867; 8, D. robustus sp. nov.; 9, D. sanliurfensis sp. nov.; 10, D. teunisseni sp. nov.; 11, D. turcicus sp. nov.; 12, D. iranicus Wittmer, 1967. Scale bars = 2.0 mm." httpUri="https://zenodo.org/record/232880/files/figure.png" pageId="8">Fig. 6</figureCitation>
). Head including eyes about as wide as anterior margin of pronotum (
<figureCitation id="D386A19DFF8AFFF539F1B5865832C6EE" box="[1241,1324,1693,1718]" captionStart="FIGURES 13 – 21" captionStartId="3.[151,264,1423,1446]" captionTargetBox="[161,1430,204,1401]" captionTargetId="figure@3.[151,1436,193,1403]" captionTargetPageId="3" captionText="FIGURES 13 – 21. Pronota of Drilus species. 13, Drilus badius sp. nov.; 14, D. huijbregtsi sp. nov.; 15, D. latithorax Pic, 1902; 16, D. mertliki sp. nov.; 17, D. obscuricornis Pic, 1899; 18, D. robustus sp. nov.; 19, D. sanliurfensis sp. nov.; 20, D. teunisseni sp. nov.; 21, D. turcicus sp. nov. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/232881/files/figure.png" pageId="8">Fig. 17</figureCitation>
), dorsally with irregularly shaped depression, rugose, lateral margins above antennal sockets elevated, with large shallow punctures, with sparse semierect pubescence; clypeal margin narrowed, almost straight. Eyes medium-sized, their frontal distance 1.8 times eye diameter. Mandibles curved, incisor margin with short tooth medially. Labrum transverse, partly hidden by clypeus; maxillary palpi slender, apical palpomere obliquely cut. Antennae weakly pectinate, reaching one third of elytral length, antennomeres 3–4 serrate, antennomeres 5–10 slightly pectinate, apical antennomere simple, longest (
<figureCitation id="D386A19DFF8AFFF53F14B46E5F8DC7D6" box="[572,659,1909,1934]" captionStart="FIGURES 22 – 33" captionStartId="4.[151,264,1794,1817]" captionTargetBox="[155,1414,201,1764]" captionTargetId="figure@4.[151,1436,193,1773]" captionTargetPageId="4" captionText="FIGURES 22 – 33. Antennae of Drilus species. 22, Drilus badius sp. nov.; 23, D. huijbregtsi sp. nov.; 24, D. latithorax Pic, 1902; 25, D. mertliki sp. nov.; 26, D. obscuricornis Pic, 1899; 27, D. rectus Schaufuss, 1867; 28, D. robustus sp. nov.; 29, D. sanliurfensis sp. nov.; 30, D. teunisseni sp. nov.; 31, D. turcicus sp. nov. Drilus robustus sp. nov., male. 32, leg; 33, last abdominal segments. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/232882/files/figure.png" pageId="8">Fig. 26</figureCitation>
). Pronotum slightly convex, transverse, widest at middle, 1.5 times wider posteriorly than length at midline. Anterior margin slightly concave, lateral margins convex, widened, slightly turned upward near posterior angles; posterior margin slightly emarginated at middle. Anterior angles inconspicuous, posterior angles distinct, rectangular (
<figureCitation id="D386A19DFF8BFFF43FD8B38C5E5BC0E8" box="[752,837,151,176]" captionStart="FIGURES 13 – 21" captionStartId="3.[151,264,1423,1446]" captionTargetBox="[161,1430,204,1401]" captionTargetId="figure@3.[151,1436,193,1403]" captionTargetPageId="3" captionText="FIGURES 13 – 21. Pronota of Drilus species. 13, Drilus badius sp. nov.; 14, D. huijbregtsi sp. nov.; 15, D. latithorax Pic, 1902; 16, D. mertliki sp. nov.; 17, D. obscuricornis Pic, 1899; 18, D. robustus sp. nov.; 19, D. sanliurfensis sp. nov.; 20, D. teunisseni sp. nov.; 21, D. turcicus sp. nov. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/232881/files/figure.png" pageId="9">Fig. 17</figureCitation>
); surface of disc sparsely covered with large shallow punctures, with sparse semierect setae, mainly at angles and lateral margins. Scutellum flat, gradually narrowed; apex rounded. Prosternum transverse, with deep transverse groove near frontal margin, frontal margin almost straight, prosternal process short, sharp, slightly elevated medially. Mesoventrite v-shaped, grooved medially. Elytra subparallel-sided, slightly rugose, shallowly punctured, covered with semierect pubescence. Legs slender, slightly compressed, with sparse, long, semierect setae, tarsomeres 1–4 gradually shortened, tarsomere 4 shortest, apical tarsomere longest, slender, slightly longer than combined lengths of tarsomeres 3 and 4. Aedeagus compact, with considerably curved phallus; paramera robust; phallobase wide, u-shaped (
<figureCitation id="D386A19DFF8BFFF4391BB28F5997C1F4" box="[1075,1161,404,429]" captionStart="FIGURES 34 – 45" captionStartId="6.[151,264,1845,1868]" captionTargetBox="[163,1430,529,1821]" captionTargetId="figure@6.[151,1436,517,1825]" captionTargetPageId="6" captionText="FIGURES 34 – 45. Male genitalia of Drilus species. 34, Drilus badius sp. nov.; 35, D. huijbregtsi sp. nov.; 36, D. latithorax Pic, 1902; 37, D. mertliki sp. nov.; 38, D. obscuricornis Pic, 1899; 39, D. rectus Schaufuss, 1867 (Turkey, Adana Prov.); 40, D. robustus sp. nov.; 41, D. sanliurfensis sp. nov.; 42, D. teunisseni sp. nov.; 43, D. turcicus sp. nov.; 44, D. iranicus Wittmer, 1967; 45, Drilus sp. (Abkhazia). Scale bars = 0.25 mm." httpUri="https://zenodo.org/record/232883/files/figure.png" pageId="9">Fig. 38</figureCitation>
). Female and immature stages are unknown.
</paragraph>
<paragraph id="4B02BD18FF8BFFF43DEFB2C75C60C24F" blockId="9.[151,1437,151,573]" pageId="9">
<emphasis id="79C9610AFF8BFFF43DEFB2C75C63C1AD" bold="true" box="[199,381,476,501]" pageId="9">Measurements.</emphasis>
BL
<quantity id="8C4510FDFF8BFFF43C99B2C75F14C1AC" box="[433,522,476,501]" metricMagnitude="-3" metricUnit="m" metricValue="5.4" pageId="9" unit="mm" value="5.4">5.4 mm</quantity>
, EL
<quantity id="8C4510FDFF8BFFF43F69B2C75F84C1AC" box="[577,666,476,500]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="9" unit="mm" value="4.3">4.3 mm</quantity>
, WHe
<quantity id="8C4510FDFF8BFFF43FC2B2C75E5DC1AC" box="[746,835,476,500]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="9" unit="mm" value="1.1">1.1 mm</quantity>
, WHum
<quantity id="8C4510FDFF8BFFF43E82B2C7591AC1AC" box="[938,1028,476,500]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="9" unit="mm" value="1.8">1.8 mm</quantity>
, PL 1.0 mm, PW
<quantity id="8C4510FDFF8BFFF439F9B2C75834C1AC" box="[1233,1322,476,501]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="9" unit="mm" value="1.5">1.5 mm</quantity>
, Edist
<quantity id="8C4510FDFF8BFFF43851B2C75DDBC24F" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="9" unit="mm" value="0.7">0.7 mm</quantity>
, Ediam
<quantity id="8C4510FDFF8BFFF43C08B11B5C64C24F" box="[288,378,512,536]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="9" unit="mm" value="0.4">0.4 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="03A7EE93FF8BFFF43DEFB13F5E5AC265" box="[199,836,548,573]" pageId="9" type="distribution">
<paragraph id="4B02BD18FF8BFFF43DEFB13F5E5AC265" blockId="9.[151,1437,151,573]" box="[199,836,548,573]" pageId="9">
<emphasis id="79C9610AFF8BFFF43DEFB13F5C7EC265" bold="true" box="[199,352,548,573]" pageId="9">Distribution.</emphasis>
Western
<collectingCountry id="33AAFD88FF8BFFF43CE1B13F5F07C265" box="[457,537,548,573]" name="Turkey" pageId="9">Turkey</collectingCountry>
(Bursa Province;
<figureCitation id="D386A19DFF8BFFF43FCCB13F5E28C265" box="[740,822,548,573]" captionStart="FIGURE 55" captionStartId="12.[151,250,1911,1933]" captionTargetBox="[160,1421,963,1875]" captionTargetId="figure@12.[151,1436,951,1889]" captionTargetPageId="12" captionText="FIGURE 55. The distribution of Drilus species in Asia Minor." httpUri="https://zenodo.org/record/232886/files/figure.png" pageId="9">Fig. 55</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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