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<mods:title id="7E916D447CDCFC244CE5619501B26725">New anaxyelids (Hymenoptera: Anaxyelidae) from the Cretaceous of Asia</mods:title>
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<paragraph id="163FA7510520B06716F9F90706B77AA3" blockId="2.[151,645,1747,1773]" box="[151,645,1747,1773]" pageId="2" pageNumber="343">
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<emphasis id="24F47B430520B06716F9F90706B77AA3" bold="true" box="[151,645,1747,1773]" pageId="2" pageNumber="343">
Subfamily
<taxonomicName id="D180DCD20520B0671770F90706B77AA3" authority="Martynov, 1925" authorityName="Martynov" authorityYear="1925" box="[286,645,1747,1773]" class="Insecta" family="Anaxyelidae" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="343" phylum="Arthropoda" rank="subFamily" subFamily="Anaxyelinae">
Anaxyelinae
<bibRefCitation id="7211DAA00520B06717AEF90706B77AA3" author="Martynov, A. V." box="[448,645,1747,1773]" pageId="2" pageNumber="343" pagination="753 - 762" refId="ref8602" refString="Martynov, A. V. (1925) To the knowledge of fossil insects from Jurassic beds in Turkestan. Bulletin de l'Academie des Sciences de l'URSS, 19, 753 - 762." type="journal article" year="1925">Martynov, 1925</bibRefCitation>
</taxonomicName>
</emphasis>
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</paragraph>
<paragraph id="163FA7510520B06716F9F8CF068F7B7B" blockId="2.[151,701,1819,1845]" box="[151,701,1819,1845]" pageId="2" pageNumber="343">
<heading id="4D77103D0520B06716F9F8CF068F7B7B" bold="true" box="[151,701,1819,1845]" fontSize="11" level="1" pageId="2" pageNumber="343" reason="1">
<emphasis id="24F47B430520B06716F9F8CF068F7B7B" bold="true" box="[151,701,1819,1845]" pageId="2" pageNumber="343">
Identification key for genera of
<taxonomicName id="D180DCD20520B067144CF8CF068F7B7B" authorityName="Martynov" authorityYear="1925" box="[546,701,1819,1845]" class="Insecta" family="Anaxyelidae" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="343" phylum="Arthropoda" rank="subFamily" subFamily="Anaxyelinae">Anaxyelinae</taxonomicName>
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<key id="6211F0600520B06616F9F88B01AE7E41" lastPageId="3" lastPageNumber="344" pageId="2" pageNumber="343">
<keyStep id="AD74BFF40520B06716F9F88B01AE7BDC" pageId="2" pageNumber="343">
<paragraph id="163FA7510520B06716F9F88B01AE7B3A" blockId="2.[151,1436,1887,2027]" box="[151,1436,1887,1909]" pageId="2" pageNumber="343">
<keyLead id="AD7104640520B06716F9F88B01AE7B3A" box="[151,1436,1887,1909]" pageId="2" pageNumber="343">1. Pterostigma widely desclerotized......................................................................... 2</keyLead>
</paragraph>
<paragraph id="163FA7510520B06716F9F8A901AE7BDC" blockId="2.[151,1436,1887,2027]" box="[151,1436,1917,1939]" pageId="2" pageNumber="343">
<keyLead id="AD7104640520B06716F9F8A901AE7BDC" box="[151,1436,1917,1939]" pageId="2" pageNumber="343">- Pterostigma sclerotized, rarely with tiny desclerotized spot.................................................... 3</keyLead>
</paragraph>
</keyStep>
<keyStep id="AD74BFF40520B06716F9F84E01AE7BA5" pageId="2" pageNumber="343">
<paragraph id="163FA7510520B06716F9F84E01AE7BFE" blockId="2.[151,1436,1887,2027]" box="[151,1436,1946,1968]" pageId="2" pageNumber="343">
<keyLead id="AD7104640520B06716F9F84E01AE7BFE" box="[151,1436,1946,1968]" pageId="2" pageNumber="343">
2. Apical section of ovipositor sheath at least 1.5 × as long as basal section, ovipositor cylindric.....
<taxonomicName id="D180DCD20520B06712CAF84E01AE7BFE" authority="Martynov, 1925" authorityName="Martynov" authorityYear="1925" box="[1188,1436,1946,1968]" class="Insecta" family="Anaxyelidae" genus="Anaxyela" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="343" phylum="Arthropoda" rank="genus">
<emphasis id="24F47B430520B06712CAF84E00CF7BE1" box="[1188,1277,1946,1967]" italics="true" pageId="2" pageNumber="343">Anaxyela</emphasis>
<bibRefCitation id="7211DAA00520B067136DF84E01AE7BFE" author="Martynov, A. V." box="[1283,1436,1946,1968]" pageId="2" pageNumber="343" pagination="753 - 762" refId="ref8602" refString="Martynov, A. V. (1925) To the knowledge of fossil insects from Jurassic beds in Turkestan. Bulletin de l'Academie des Sciences de l'URSS, 19, 753 - 762." type="journal article" year="1925">Martynov, 1925</bibRefCitation>
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</paragraph>
<paragraph id="163FA7510520B06716F9F86C01AE7BA5" blockId="2.[151,1436,1887,2027]" pageId="2" pageNumber="343">
<keyLead id="AD7104640520B06716F9F86C01AE7BA5" pageId="2" pageNumber="343">
- Apical section of ovipositor sheath as long as or shorter than basal section, ovipositor tapered apically..............................................................................................
<taxonomicName id="D180DCD20520B0671207F80101AE7BA5" authority="Rasnitsyn, 1969" authorityName="Rasnitsyn" authorityYear="1969" box="[1129,1436,2005,2027]" class="Insecta" family="Anaxyelidae" genus="Sphenosyntexis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="343" phylum="Arthropoda" rank="genus">
<emphasis id="24F47B430520B0671207F80100CE7BA4" box="[1129,1276,2005,2026]" italics="true" pageId="2" pageNumber="343">Sphenosyntexis</emphasis>
<bibRefCitation id="7211DAA00520B067136CF80101AE7BA5" author="Rasnitsyn, A. P." box="[1282,1436,2005,2027]" pageId="2" pageNumber="343" refId="ref8807" refString="Rasnitsyn, A. P. (1969) Pрoiсkozdiнii i evoЛЮtiЯ нiZSik piрipoнcatoKрыЛыk [Origin and Evolution of the Lower Hyme- noptera]. Nauka, Moscow, 196 pp. [in Russian, Proceedings of the Paleontological Institute, Academy of Sciences USSR, 123]" type="book" year="1969">Rasnitsyn, 1969</bibRefCitation>
</taxonomicName>
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</paragraph>
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<keyStep id="AD74BFF40521B06616F9FF4E01AE7D49" pageId="3" pageNumber="344">
<paragraph id="163FA7510521B06616F9FF4E01AE7C83" blockId="3.[151,1437,153,527]" pageId="3" pageNumber="344">
<keyLead id="AD7104640521B06616F9FF4E01AE7C83" pageId="3" pageNumber="344">
3. Antenna serrate; hind wing with r-m joining M well before m-cu, M between r-m and m-cu 2 × as long as r-m.........................................................................................
<taxonomicName id="D180DCD20521B06612E7FF6301AE7C83" authority="Rasnitsyn, 1968" authorityName="Rasnitsyn" authorityYear="1968" box="[1161,1436,183,205]" class="Insecta" family="Anaxyelidae" genus="Anasyntexis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="344" phylum="Arthropoda" rank="genus">
<emphasis id="24F47B430521B06612E7FF6300CE7C82" box="[1161,1276,183,204]" italics="true" pageId="3" pageNumber="344">Anasyntexis</emphasis>
<bibRefCitation id="7211DAA00521B066136CFF6301AE7C83" author="Rasnitsyn, A. P." box="[1282,1436,183,205]" pageId="3" pageNumber="344" pagination="190 - 236" refId="ref8747" refString="Rasnitsyn, A. P. (1968) Нovыi мiZoZoйсkii piлiлЬЩiki [New Mesozoic sawflies (Hymenoptera, Symphyta)]. In: Rohdendorf, B. B. (Ed.), ЮрсKii нaсiKoмыi Kaрatay [Jurassic insects of Karatau]. Nauka, Moscow, pp. 190 - 236. [in Russian]" type="book chapter" year="1968">Rasnitsyn, 1968</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="163FA7510521B06616F9FF0101AE7D49" blockId="3.[151,1437,153,527]" pageId="3" pageNumber="344">
<keyLead id="AD7104640521B06616F9FF0101AE7D49" pageId="3" pageNumber="344">- Antenna smooth, rarely moniliform, but never serrate; hind wing with r-m joining M before or after m-cu, but never displaced basad from m-cu for more than 1 × r-m length.............................................................. 4</keyLead>
</paragraph>
</keyStep>
<keyStep id="AD74BFF40521B06616F9FEDB01AE7D11" pageId="3" pageNumber="344">
<paragraph id="163FA7510521B06616F9FEDB01AE7D0C" blockId="3.[151,1437,153,527]" pageId="3" pageNumber="344">
<keyLead id="AD7104640521B06616F9FEDB01AE7D0C" pageId="3" pageNumber="344">
4. Apical section of ovipositor sheath at least 3 × as long as basal section; ovipositor as long as or longer than body.......................................................................................
<taxonomicName id="D180DCD20521B06612E6FEF801AE7D0C" authority="Rasnitsyn, 1968" authorityName="Rasnitsyn" authorityYear="1968" box="[1160,1436,300,322]" class="Insecta" family="Anaxyelidae" genus="Kulbastavia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="344" phylum="Arthropoda" rank="genus">
<emphasis id="24F47B430521B06612E6FEF800CE7D0F" box="[1160,1276,300,321]" italics="true" pageId="3" pageNumber="344">Kulbastavia</emphasis>
<bibRefCitation id="7211DAA00521B066136CFEF801AE7D0C" author="Rasnitsyn, A. P." box="[1282,1436,300,322]" pageId="3" pageNumber="344" pagination="190 - 236" refId="ref8747" refString="Rasnitsyn, A. P. (1968) Нovыi мiZoZoйсkii piлiлЬЩiki [New Mesozoic sawflies (Hymenoptera, Symphyta)]. In: Rohdendorf, B. B. (Ed.), ЮрсKii нaсiKoмыi Kaрatay [Jurassic insects of Karatau]. Nauka, Moscow, pp. 190 - 236. [in Russian]" type="book chapter" year="1968">Rasnitsyn, 1968</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="163FA7510521B06616F9FE9E01AE7D11" blockId="3.[151,1437,153,527]" box="[151,1436,329,351]" pageId="3" pageNumber="344">
<keyLead id="AD7104640521B06616F9FE9E01AE7D11" box="[151,1436,329,351]" pageId="3" pageNumber="344">- Apical section of ovipositor sheath less than 1.5 × as long as basal section; ovipositor significantly shorter than body...... 5</keyLead>
</paragraph>
</keyStep>
<keyStep id="AD74BFF40521B06616F9FEB301AE7DD4" pageId="3" pageNumber="344">
<paragraph id="163FA7510521B06616F9FEB301AE7D33" blockId="3.[151,1437,153,527]" box="[151,1436,359,381]" pageId="3" pageNumber="344">
<keyLead id="AD7104640521B06616F9FEB301AE7D33" box="[151,1436,359,381]" pageId="3" pageNumber="344">
5. Ovipositor very short, its apical section less than 0.5 × as long as basal section.............
<taxonomicName id="D180DCD20521B0661204FEB301AE7D33" authority="Rasnitsyn, 1969" authorityName="Rasnitsyn" authorityYear="1969" box="[1130,1436,359,381]" class="Insecta" family="Anaxyelidae" genus="Brachysyntexis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="344" phylum="Arthropoda" rank="genus">
<emphasis id="24F47B430521B0661204FEB300C97D32" box="[1130,1275,359,380]" italics="true" pageId="3" pageNumber="344">Brachysyntexis</emphasis>
<bibRefCitation id="7211DAA00521B066136CFEB301AE7D33" author="Rasnitsyn, A. P." box="[1282,1436,359,381]" pageId="3" pageNumber="344" refId="ref8807" refString="Rasnitsyn, A. P. (1969) Pрoiсkozdiнii i evoЛЮtiЯ нiZSik piрipoнcatoKрыЛыk [Origin and Evolution of the Lower Hyme- noptera]. Nauka, Moscow, 196 pp. [in Russian, Proceedings of the Paleontological Institute, Academy of Sciences USSR, 123]" type="book" year="1969">Rasnitsyn, 1969</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="163FA7510521B06616F9FE5101AE7DD4" blockId="3.[151,1437,153,527]" box="[151,1436,388,410]" pageId="3" pageNumber="344">
<keyLead id="AD7104640521B06616F9FE5101AE7DD4" box="[151,1436,388,410]" pageId="3" pageNumber="344">- Ovipositor rather long, its apical section as long as or longer than basal section.................................... 6</keyLead>
</paragraph>
</keyStep>
<keyStep id="AD74BFF40521B06616F9FE7501AE7E41" pageId="3" pageNumber="344">
<paragraph id="163FA7510521B06616F9FE7501AE7D9B" blockId="3.[151,1437,153,527]" pageId="3" pageNumber="344">
<keyLead id="AD7104640521B06616F9FE7501AE7D9B" pageId="3" pageNumber="344">
6. Basal part of flagellum, consisting of one or a few flagellomeres, very robust, much thicker than its apical part; fore wing with 5-M as long as or slightly shorter than 2m-cu..........................................
<taxonomicName id="D180DCD20521B06612FBFE6B01AE7D9B" authority="Rasnitsyn, 1968" authorityName="Rasnitsyn" authorityYear="1968" box="[1173,1436,447,469]" class="Insecta" family="Anaxyelidae" genus="Syntexyela" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="344" phylum="Arthropoda" rank="genus">
<emphasis id="24F47B430521B06612FBFE6B00CE7D9A" box="[1173,1276,447,468]" italics="true" pageId="3" pageNumber="344">Syntexyela</emphasis>
<bibRefCitation id="7211DAA00521B066136CFE6B01AE7D9B" author="Rasnitsyn, A. P." box="[1282,1436,447,469]" pageId="3" pageNumber="344" pagination="190 - 236" refId="ref8747" refString="Rasnitsyn, A. P. (1968) Нovыi мiZoZoйсkii piлiлЬЩiki [New Mesozoic sawflies (Hymenoptera, Symphyta)]. In: Rohdendorf, B. B. (Ed.), ЮрсKii нaсiKoмыi Kaрatay [Jurassic insects of Karatau]. Nauka, Moscow, pp. 190 - 236. [in Russian]" type="book chapter" year="1968">Rasnitsyn, 1968</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="163FA7510521B06616F9FE0901AE7E41" blockId="3.[151,1437,153,527]" pageId="3" pageNumber="344">
<keyLead id="AD7104640521B06616F9FE0901AE7E41" pageId="3" pageNumber="344">
- Flagellum setiform, gradually tapering apically; fore wing with 5-M less than 0.5 × as long as 2m-cu.................................................................................................
<taxonomicName id="D180DCD20521B06612E4FE2E01AE7E41" authority="Rasnitsyn, 1969" authorityName=": Rasnitsyn" authorityYear="1969" box="[1162,1436,505,527]" class="Insecta" family="Anaxyelidae" genus="Urosyntexis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="344" phylum="Arthropoda" rank="genus">
<emphasis id="24F47B430521B06612E4FE2E00CE7E41" box="[1162,1276,506,527]" italics="true" pageId="3" pageNumber="344">Urosyntexis</emphasis>
<bibRefCitation id="7211DAA00521B066136CFE2D01AE7E41" author="Rasnitsyn, A. P." box="[1282,1436,505,527]" pageId="3" pageNumber="344" refId="ref8807" refString="Rasnitsyn, A. P. (1969) Pрoiсkozdiнii i evoЛЮtiЯ нiZSik piрipoнcatoKрыЛыk [Origin and Evolution of the Lower Hyme- noptera]. Nauka, Moscow, 196 pp. [in Russian, Proceedings of the Paleontological Institute, Academy of Sciences USSR, 123]" type="book" year="1969">Rasnitsyn, 1969</bibRefCitation>
</taxonomicName>
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</subSubSection>
<subSubSection id="5E9AF4DA0521B06616A9FDEA053D7F36" pageId="3" pageNumber="344" type="discussion">
<paragraph id="163FA7510521B06616A9FDEA053D7F36" blockId="3.[151,1437,574,888]" pageId="3" pageNumber="344">
<emphasis id="24F47B430521B06616A9FDEA05097E16" bold="true" box="[199,315,574,600]" pageId="3" pageNumber="344">Remarks.</emphasis>
The genus
<taxonomicName id="D180DCD20521B06617D1FDEB06767E16" authorityName=": Rasnitsyn" authorityYear="1969" box="[447,580,575,600]" class="Insecta" family="Anaxyelidae" genus="Urosyntexis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="344" phylum="Arthropoda" rank="genus">
<emphasis id="24F47B430521B06617D1FDEB06767E16" box="[447,580,575,600]" italics="true" pageId="3" pageNumber="344">Urosyntexis</emphasis>
</taxonomicName>
was separated from
<taxonomicName id="D180DCD20521B0661541FDEA07957E16" authorityName="Rasnitsyn" authorityYear="1968" box="[815,935,574,600]" class="Insecta" family="Anaxyelidae" genus="Syntexyela" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="344" phylum="Arthropoda" rank="genus">
<emphasis id="24F47B430521B0661541FDEA07957E16" box="[815,935,574,600]" italics="true" pageId="3" pageNumber="344">Syntexyela</emphasis>
</taxonomicName>
basing on a few findings from Karatau,
<collectingCountry id="6E97E7C10521B066131CFDEB04CB7E32" name="Kazakhstan" pageId="3" pageNumber="344">Kazakhstan</collectingCountry>
(
<bibRefCitation id="7211DAA00521B0661765FDB605F77E33" author="Rasnitsyn, A. P." box="[267,453,610,637]" pageId="3" pageNumber="344" refId="ref8807" refString="Rasnitsyn, A. P. (1969) Pрoiсkozdiнii i evoЛЮtiЯ нiZSik piрipoнcatoKрыЛыk [Origin and Evolution of the Lower Hyme- noptera]. Nauka, Moscow, 196 pp. [in Russian, Proceedings of the Paleontological Institute, Academy of Sciences USSR, 123]" type="book" year="1969">Rasnitsyn, 1969</bibRefCitation>
). Subsequent finding of Cretaceous
<taxonomicName id="D180DCD20521B066150DFDB601AF7E33" authority="(Zhang, Rasnitsyn, 2006)" authorityName="Zhang, Rasnitsyn" authorityYear="2006" baseAuthorityName="Zhang, Rasnitsyn" baseAuthorityYear="2006" box="[867,1437,610,637]" class="Insecta" family="Anaxyelidae" genus="Syntexyela" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="344" phylum="Arthropoda" rank="species" species="continentalis">
<emphasis id="24F47B430521B066150DFDB600467E32" box="[867,1140,610,636]" italics="true" pageId="3" pageNumber="344">Syntexyela continentalis</emphasis>
(
<bibRefCitation id="7211DAA00521B06612EAFDB601A77E33" author="Zhang, H. &amp; Rasnitsyn, A. P." box="[1156,1429,610,637]" pageId="3" pageNumber="344" pagination="279 - 284" refId="ref9633" refString="Zhang, H. &amp; Rasnitsyn, A. P. (2006) Two new anaxyelid sawflies (Insecta, Hymenoptera, Siricoidea) from the Yixian Formation of western Liaoning, China. Cretaceous Research, 27, 279 - 284. https: // doi. org / 10.1016 / j. cretres. 2005.11.001" type="journal article" year="2006">Zhang, Rasnitsyn, 2006</bibRefCitation>
)
</taxonomicName>
have filled the morphological gap between these genera and made its distinguishing rather problematic. The not described representatives of
<taxonomicName id="D180DCD20521B066178EFD7F06577E8A" authorityName=": Rasnitsyn" authorityYear="1969" box="[480,613,683,708]" class="Insecta" family="Anaxyelidae" genus="Urosyntexis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="344" phylum="Arthropoda" rank="genus">
<emphasis id="24F47B430521B066178EFD7F06577E8A" box="[480,613,683,708]" italics="true" pageId="3" pageNumber="344">Urosyntexis</emphasis>
</taxonomicName>
and/or
<taxonomicName id="D180DCD20521B06614AEFD7E070A7E8A" authorityName="Rasnitsyn" authorityYear="1968" box="[704,824,682,708]" class="Insecta" family="Anaxyelidae" genus="Syntexyela" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="344" phylum="Arthropoda" rank="genus">
<emphasis id="24F47B430521B06614AEFD7E070A7E8A" box="[704,824,682,708]" italics="true" pageId="3" pageNumber="344">Syntexyela</emphasis>
</taxonomicName>
were also found in the Jurassic of Daohugou,
<collectingCountry id="6E97E7C10521B0661337FD7E01AE7E8A" box="[1369,1436,682,708]" name="China" pageId="3" pageNumber="344">China</collectingCountry>
(personal data), and the identification of these specimens basing on the characters originally proposed by
<bibRefCitation id="7211DAA00521B0661340FD1A04E87F42" author="Rasnitsyn, A. P." pageId="3" pageNumber="344" pagination="190 - 236" refId="ref8747" refString="Rasnitsyn, A. P. (1968) Нovыi мiZoZoйсkii piлiлЬЩiki [New Mesozoic sawflies (Hymenoptera, Symphyta)]. In: Rohdendorf, B. B. (Ed.), ЮрсKii нaсiKoмыi Kaрatay [Jurassic insects of Karatau]. Nauka, Moscow, pp. 190 - 236. [in Russian]" type="book chapter" year="1968">Rasnitsyn (1968</bibRefCitation>
,
<bibRefCitation id="7211DAA00521B0661689FD2605127F43" author="Rasnitsyn, A. P." box="[231,288,754,781]" pageId="3" pageNumber="344" refId="ref8807" refString="Rasnitsyn, A. P. (1969) Pрoiсkozdiнii i evoЛЮtiЯ нiZSik piрipoнcatoKрыЛыk [Origin and Evolution of the Lower Hyme- noptera]. Nauka, Moscow, 196 pp. [in Russian, Proceedings of the Paleontological Institute, Academy of Sciences USSR, 123]" type="book" year="1969">1969</bibRefCitation>
) is impossible. The revision and, probably, the synonymization of these two genera is required. However, this revision without the inclusion of the Chinese collections will not be reliable. Here I preliminary assign the new species to
<taxonomicName id="D180DCD20521B066172FFCEF05F47F1A" authorityName=": Rasnitsyn" authorityYear="1969" box="[321,454,827,852]" class="Insecta" family="Anaxyelidae" genus="Urosyntexis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="344" phylum="Arthropoda" rank="genus">
<emphasis id="24F47B430521B066172FFCEF05F47F1A" box="[321,454,827,852]" italics="true" pageId="3" pageNumber="344">Urosyntexis</emphasis>
</taxonomicName>
basing on its antenna without thickened basal flagellomeres, short 5-M and rather short ovipositor.
</paragraph>
</subSubSection>
</treatment>
</document>