130 lines
14 KiB
XML
130 lines
14 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.130.1335" ID-GBIF-Dataset="b4081ab6-1cc8-46c2-9ccc-405ee4c43a22" ID-PMC="PMC3260778" ID-Pensoft-Pub="1313-2970-130-505" ID-PubMed="22259296" ModsDocAuthor="" ModsDocDate="2011" ModsDocID="1313-2970-130-505" ModsDocOrigin="ZooKeys 130" ModsDocTitle="Revision of fossil species of Dryinus belonging to lamellatus group, with description of a new species (Hymenoptera, Dryinidae)" checkinTime="1451249842480" checkinUser="pensoft" docAuthor="Olmi, Massimo & Guglielmino, Adalgisa" docDate="2011" docId="3F7F0BEABF530FF9C7C6D31AC0F17767" docLanguage="en" docName="ZooKeys 130: 505-514" docOrigin="ZooKeys 130" docSource="http://dx.doi.org/10.3897/zookeys.130.1335" docTitle="Dryinus grimaldii Olmi" docType="treatment" docVersion="3" lastPageNumber="509" masterDocId="FF8B262C425216709926FFA0FF86FF85" masterDocTitle="Revision of fossil species of Dryinus belonging to lamellatus group, with description of a new species (Hymenoptera, Dryinidae)" masterLastPageNumber="514" masterPageNumber="505" pageNumber="507" updateTime="1668152288687" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Revision of fossil species of Dryinus belonging to lamellatus group, with description of a new species (Hymenoptera, Dryinidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Olmi, Massimo</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Guglielmino, Adalgisa</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2011</mods:date>
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<mods:detail type="volume">
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<mods:number>130</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>505</mods:start>
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<mods:end>514</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.130.1335</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.130.1335</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-130-505</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152031048" LSID="urn:lsid:plazi:treatment:3F7F0BEABF530FF9C7C6D31AC0F17767" httpUri="http://treatment.plazi.org/id/3F7F0BEABF530FF9C7C6D31AC0F17767" lastPageId="4" lastPageNumber="509" pageId="2" pageNumber="507">
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<subSubSection pageId="2" pageNumber="507" type="nomenclature">
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<paragraph pageId="2" pageNumber="507">
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<taxonomicName LSID="http://species-id.net/wiki/Dryinus_grimaldii" authority="Olmi" class="Insecta" family="Ampulicidae" genus="Dryinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dryinus grimaldii" order="Hymenoptera" pageId="2" pageNumber="507" phylum="Arthropoda" rank="species" species="grimaldii">Dryinus grimaldii Olmi</taxonomicName>
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Figs 1-3
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</paragraph>
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</subSubSection>
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<subSubSection pageId="2" pageNumber="507" type="reference_group">
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<paragraph pageId="2" pageNumber="507">
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<taxonomicName class="Insecta" family="Ampulicidae" genus="Dryinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dryinus grimaldii" order="Hymenoptera" pageId="2" pageNumber="507" phylum="Arthropoda" rank="species" species="grimaldii">Dryinus grimaldii</taxonomicName>
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<bibRefCitation author="Olmi, M" journalOrPublisher="Redia" pageId="7" pageNumber="512" pagination="253 - 271" title="Dryinids and Embolemids in amber (Hymenoptera Dryinidae et Embolemidae)." volume="78" year="1995">Olmi 1995</bibRefCitation>
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: 254.
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</paragraph>
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<paragraph pageId="2" pageNumber="507">
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<taxonomicName class="Insecta" family="Ampulicidae" genus="Dryinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dryinus grimaldii" order="Hymenoptera" pageId="2" pageNumber="507" phylum="Arthropoda" rank="species" species="grimaldii">Dryinus grimaldii</taxonomicName>
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Olmi:
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<bibRefCitation pageId="2" pageNumber="507">Olmi 2000</bibRefCitation>
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: 65.
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</paragraph>
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<paragraph pageId="2" pageNumber="507">
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<taxonomicName class="Insecta" family="Ampulicidae" genus="Dryinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dryinus grimaldii" order="Hymenoptera" pageId="2" pageNumber="507" phylum="Arthropoda" rank="species" species="grimaldii">Dryinus grimaldii</taxonomicName>
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Olmi:
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<bibRefCitation author="Olmi, M" journalOrPublisher="Stuttgarter Beitraege zur Naturkunde, Serie B," pageId="7" pageNumber="512" pagination="1 - 58" title="New parasitic wasps from Baltic amber (Insecta: Hymenoptera: Dryinidae)." volume="306" year="2001">Olmi and Bechly 2001</bibRefCitation>
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: 45.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="2" pageNumber="507" type="type material">
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<paragraph pageId="2" pageNumber="507">Type material.</paragraph>
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<paragraph pageId="2" pageNumber="507">Holotype, female, Early Miocene amber from the Dominican Republic (16-19 Ma) (AMNH, No. DR-10-1426); same locality label, 1 female paratype (AMNH, No. DR-10-1423).</paragraph>
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</subSubSection>
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<subSubSection pageId="2" pageNumber="507" type="additional specimens examined">
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<paragraph pageId="2" pageNumber="507">Additional specimens examined.</paragraph>
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<paragraph pageId="2" pageNumber="507">same locality, three female specimens (GPJC).</paragraph>
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</subSubSection>
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<subSubSection pageId="2" pageNumber="507" type="diagnosis">
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<paragraph pageId="2" pageNumber="507">Diagnosis.</paragraph>
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<paragraph pageId="2" pageNumber="507">Female with enlarged claw not reduced and not spatulate (Figs 1, 2), longer than arolium; enlarged claw with two subapical teeth. Male unknown.</paragraph>
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</subSubSection>
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<subSubSection lastPageId="4" lastPageNumber="509" pageId="2" pageNumber="507" type="redescription">
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<paragraph pageId="2" pageNumber="507">Redescription.</paragraph>
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<paragraph lastPageId="4" lastPageNumber="509" pageId="2" pageNumber="507">
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Female: macropterous; length 4.3-6.3 mm. Colour difficult to discern, apparently testaceous, except two dark lateral spots on sides of pronotal disc, scutum, scutellum, propodeum and tegula dark; metasoma with dark transverse band. In paratype, legs with dark spots on coxae and clubs of femora; scutum apparently without dark lateral spots. In one specimen of GPJC labelled H-10-100, apparently scutum without lateral dark spots, scutellum not darkened, posterior surface of propodeum darkened. In two specimens of GPJC labelled H-10-23C, body totally testaceous, except petiole black and two brown spots on sides of scutum. Antenna 10-segmented, long and very slender, filiform, not thickened distally, covered with dense and short hairs; antennal segments of holotype in following proportions: 10:5:44:57:38:21:9:9:9:17; antenna more than nine times as long as head (length of head dorsally measured from occipital carina behind ocelli to distal apex of mandible): 219:22. Head weakly convex, apparently shiny, finely punctate, without apparent sculpture among punctures; clypeus and mandible not distinct; occipital carina apparently complete; occiput deeply excavated; eye normally bulging; POL = 2; OL = 1.5; OOL = 10; OPL = 1.5; TL = 5; greatest breadth of posterior ocellus longer than POL
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<pageBreakToken pageId="3" pageNumber="508" start="start">(</pageBreakToken>
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4:2); frontal line absent. Maxillary palpi not evident, apparently 6-segmented. Labial palpi not distinct. Pronotum apparently shiny, finely punctate, about as long as head, crossed by anterior strong transverse impression between anterior collar and disc; disc humped; posterior collar very short; pronotal tubercle reaching tegula. Scutum apparently shiny, finely punctate, slightly shorter than pronotum (19:22). Notauli complete, posteriorly separated; minimum distance between notauli about as long as greatest breadth of posterior ocelli. In one specimen of GPJC labelled H-10-100, notauli apparently almost complete, not reaching posterior margin of scutum. Scutellum apparently shorter than scutum (10:19), with sculpture not evident. Metanotum shorter than scutellum (6:10), with sculpture not evident. Propodeum longer than scutum (39:19), reticulate rugose, areolae very broad; posterior surface with two complete longitudinal keels; sculpture of median area of posterior surface not evident. In one specimen of GPJC labelled H-10-100, dorsal surface of propodeum with two median longitudinal and almost parallel keels. Shape of pronotum, scutum, scutellum, metanotum and propodeum usual for
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<taxonomicName lsidName="" pageId="3" pageNumber="508" rank="subFamily" subFamily="Dryininae">Dryininae</taxonomicName>
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. Forewing hyaline, without dark transverse bands, with usual venation of
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<taxonomicName lsidName="" pageId="3" pageNumber="508" rank="subFamily" subFamily="Dryininae">Dryininae</taxonomicName>
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; pterostigma narrow, much longer than broad (32:4); marginal cell apparently open; distal part of stigmal vein longer than proximal part (16:11); stigmal vein not S-shaped, forming angle between proximal and distal parts; forewing with usual three basal cells clearly enclosed by pigmented veins (costal, median and submedian cells). Hindwing hyaline, without dark transverse bands. Foreleg segments in following proportions: 55 (coxa): 53 (trochanter): 61 (femur): 60 (tibia): 18 (tarsal segment 1): 5 (tarsal segment 2): 8 (tarsal segment 3): 55 (tarsal
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<pageBreakToken pageId="4" pageNumber="509" start="start">segment</pageBreakToken>
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4): 78 (tarsal segment 5); foreleg chelate; enlarged claw much shorter than segment 5 of protarsus (42:78); protrochanter more than four times as long as broad (53:5)(greatest breadth measured on distal club), with long and slender proximal stalk, broadened after half-way; segments 2 and 3 of protarsus produced into hooks; rudimentary claw present; arolium much shorter than enlarged claw (8:42); enlarged claw with two strong subapical teeth and 1 row of 8 lamellae; subapical teeth of enlarged claw very strong, such as in
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<taxonomicName class="Insecta" family="Dryinidae" genus="Plesiodryinus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Plesiodryinus" order="Hymenoptera" pageId="4" pageNumber="509" phylum="Arthropoda" rank="genus">Plesiodryinus</taxonomicName>
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; distal apex of enlarged claw not spatulate. Segment 5 of protarsus with 2 rows of approximately 50 lamellae; distal apex with group of at least 20 lamellae (number of lamellae not evident). Midleg segments in following proportions: 22 (coxa): 7 (trochanter): 41 (femur): 60 (tibia): 17 (tarsal segment 1): 15 (tarsal segment 2): 15 (tarsal segment 3): 14 (tarsal segment 4); segment 5 of mesotarsus not distinct. Hindleg segments in following proportions: 27 (coxa): 11 (trochanter): 48 (femur): 80 (tibia); segments of metatarsus not distinct in holotype; segments of metatarsus of paratype in following proportions: 37 (tarsal segment 1): 19 (tarsal segment 2): 13 (tarsal segment 3): 10 (tarsal segment 4): 11 (tarsal segment 5). Metasoma without distinct and slender petiole. Shape and length of petiole usual for
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<taxonomicName lsidName="" pageId="4" pageNumber="509" rank="subFamily" subFamily="Dryininae">Dryininae</taxonomicName>
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. Shape, length and breadth of wings usual for
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<taxonomicName lsidName="" pageId="4" pageNumber="509" rank="subFamily" subFamily="Dryininae">Dryininae</taxonomicName>
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. Shape of body usual for
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<taxonomicName lsidName="" pageId="4" pageNumber="509" rank="subFamily" subFamily="Dryininae">Dryininae</taxonomicName>
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. Tibial spurs of holotype hardly visible, apparently 1/1/1; in one specimen of GPJC labelled H-10-100, tibial spurs distinctly 1/1/2.
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</paragraph>
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<paragraph pageId="4" pageNumber="509">Male: unknown.</paragraph>
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<caption pageId="4" pageNumber="509">
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<paragraph pageId="4" pageNumber="509">
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Figure 1.
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<taxonomicName class="Insecta" family="Ampulicidae" genus="Dryinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dryinus grimaldii" order="Hymenoptera" pageId="4" pageNumber="509" phylum="Arthropoda" rank="species" species="grimaldii">Dryinus grimaldii</taxonomicName>
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. Female holotype (from
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<bibRefCitation author="Olmi, M" journalOrPublisher="Redia" pageId="7" pageNumber="512" pagination="253 - 271" title="Dryinids and Embolemids in amber (Hymenoptera Dryinidae et Embolemidae)." volume="78" year="1995">Olmi 1995</bibRefCitation>
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). Length 4.3 mm.
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</paragraph>
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</caption>
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<caption pageId="4" pageNumber="509">
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<paragraph pageId="4" pageNumber="509">
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Figure 2.
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<taxonomicName class="Insecta" family="Ampulicidae" genus="Dryinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dryinus grimaldii" order="Hymenoptera" pageId="4" pageNumber="509" phylum="Arthropoda" rank="species" species="grimaldii">Dryinus grimaldii</taxonomicName>
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. Female specimen in lateral view (in GPJC, No. H-10-100). Length 6.3 mm.
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</paragraph>
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</caption>
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<caption pageId="4" pageNumber="509">
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<paragraph pageId="4" pageNumber="509">
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Figure 3.
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<taxonomicName class="Insecta" family="Ampulicidae" genus="Dryinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dryinus grimaldii" order="Hymenoptera" pageId="4" pageNumber="509" phylum="Arthropoda" rank="species" species="grimaldii">Dryinus grimaldii</taxonomicName>
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. Female specimen in dorsal view (in GPJC, No. H-10-100). Length 6.3 mm.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="4" pageNumber="509" type="hosts">
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<paragraph pageId="4" pageNumber="509">Hosts.</paragraph>
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<paragraph pageId="4" pageNumber="509">Unknown.</paragraph>
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</subSubSection>
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</treatment>
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</document> |