278 lines
29 KiB
XML
278 lines
29 KiB
XML
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<mods:title id="5CCB1F3F37F08C05A18680E156B81C2F">The world’s tiniest land snails from Laos and Vietnam (Gastropoda, Pulmonata, Hypselostomatidae)</mods:title>
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<mods:namePart id="DE125E023EC0776D360E3E1975428B41">Páll-Gergely, Barna</mods:namePart>
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<mods:nameIdentifier id="E61F5B8F24DCC302F27B572D0757AB86" type="email">pall-gergely.barna@atk.hu & pallgergely2@gmail.com</mods:nameIdentifier>
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<mods:namePart id="BE85A0AA4A8797647F240B3F292E2076">Jochum, Adrienne</mods:namePart>
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<mods:nameIdentifier id="34BFD9AC6FD1101EB1C5DB17AF64DC8A" type="email">adrienne.jochum@senckenberg.de</mods:nameIdentifier>
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<mods:namePart id="E3ADBF55FA4D26FD91058A024FE22363">Anker, Katja</mods:namePart>
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<mods:namePart id="CF2826483B53EC1F5E0728D962010DD0">Hunyadi, András</mods:namePart>
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<mods:affiliation id="1C4DE48DA4E24B3DA59AECF9AACDBC6C">Adria sétány 10 G 2 / 5, 1148 Budapest, Hungary</mods:affiliation>
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<mods:namePart id="A44F948066D2A826AA1CB3BE450F12A2">Szabó, Ábel</mods:namePart>
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<mods:affiliation id="DDAAF5E0A361324B05C0A38A65297855">Lithosphere Fluid Research Laboratory, Institute of Geography and Earth Sciences, Eötvös University, Pázmány p. sétány 1 / C, 1117 Budapest, Hungary</mods:affiliation>
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<mods:namePart id="B506202B0EE824016146BDDEABF75C5C">Dányi, László</mods:namePart>
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<mods:affiliation id="09F797A91F02C802E2A32F8B14CD185D">Institute for Soil Sciences and Agricultural Chemistry, Centre for Agricultural Research, ELKH, Herman Ottó út 15, 1022 Budapest, Hungary</mods:affiliation>
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<mods:namePart id="6084D63C331C7A8A31E2B0817CAB84B7">Schilthuizen, Menno</mods:namePart>
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<mods:affiliation id="2B89C3D6C08A4D58F2A76D8BF08D4D8F">Naturalis Biodiversity Center, Darwinweg 2, 2333 CR Leiden, The Netherlands & Institute for Biology Leiden, Leiden University, Sylviusweg 72, 2333 BE Leiden, The Netherlands & Institute for Tropical Biology and Conservation, Universiti Malaysia Sabah, Jalan UMS, 88400 Kota Kinabalu, Sabah, Malaysia</mods:affiliation>
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<mods:title id="41432F694FB8BE31426368E71FBDB3EA">Contributions To Zoology</mods:title>
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Páll-Gergely, Anker & Vermeulen
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,
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, vegetation outside cave tall and woody, leg.
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<specimenCount id="AB99F5B2FFBBFF9752ACFCB2FCBAFCD6" box="[757,771,818,844]" pageId="7" pageNumber="69" type="generic">2</specimenCount>
|
||
<typeStatus id="62248099FFBBFF97534AFCB2FC3BFCD6" box="[787,898,818,844]" pageId="7" pageNumber="69">paratypes</typeStatus>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="0DF73466FFBBFF9753CEFCB2FC4FFCEA" ID-GBIF-Occurrence="3468252311" collectionCode="ZRC, MOL" country="Vietnam" pageId="7" pageNumber="69" specimenCount="411" stateProvince="Quang Ninh Province" typeStatus="paratype">
|
||
<collectionCode id="DB8EA6FEFFBBFF9753CEFCB2FC7AFCD6" box="[919,963,818,844]" collectionName="Singapore, National University of Singapore, Raffles Museum of Biodiversity Research, Zoological Reference Collection" pageId="7" pageNumber="69">ZRC</collectionCode>
|
||
.
|
||
<collectionCode id="DB8EA6FEFFBBFF975393FCB2FC47FCD6" box="[970,1022,818,844]" collectionName="MOL" pageId="7" pageNumber="69">MOL</collectionCode>
|
||
.23218/
|
||
<specimenCount id="AB99F5B2FFBBFF975415FCB2FBE1FCD6" box="[1100,1112,818,844]" pageId="7" pageNumber="69" type="generic">2</specimenCount>
|
||
<typeStatus id="62248099FFBBFF975430FCB2FD58FCEA" pageId="7" pageNumber="69">paratypes</typeStatus>
|
||
, jjV 17633/
|
||
<specimenCount id="AB99F5B2FFBBFF975300FCD5FC39FCF5" box="[857,896,853,879]" pageId="7" pageNumber="69" type="generic">409</specimenCount>
|
||
<typeStatus id="62248099FFBBFF9753DFFCD6FC4FFCEA" box="[902,1014,854,880]" pageId="7" pageNumber="69">paratypes</typeStatus>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="F5856DB0FFBBFF97529BFCFAFCFEFBB9" pageId="7" pageNumber="69" type="diagnosis">
|
||
<paragraph id="BD203E3BFFBBFF97529BFCFAFCFEFBB9" blockId="7.[676,1187,422,1742]" pageId="7" pageNumber="69">
|
||
<emphasis id="8FEBE229FFBBFF97529BFCFAFC8DFC09" box="[706,820,890,915]" italics="true" pageId="7" pageNumber="69">Diagnosis:</emphasis>
|
||
An
|
||
<taxonomicName id="7A9F45B8FFBBFF975332FCF9FBF4FC09" box="[875,1101,889,915]" pageId="7" pageNumber="69">
|
||
<emphasis id="8FEBE229FFBBFF975332FCF9FC49FC09" box="[875,1008,889,915]" italics="true" pageId="7" pageNumber="69">Angustopila</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
with a depressed-globular shell with dome-shaped spire, thick spiral striae, kidney-shaped aperture with single parietal denticle not reaching parietal callus.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="F5856DB0FFBBFF99529BFBAEFEF6FDB7" lastPageId="9" lastPageNumber="71" pageId="7" pageNumber="69" type="description">
|
||
<paragraph id="BD203E3BFFBBFF99529BFBAEFE85FE5B" blockId="7.[676,1187,422,1742]" lastBlockId="9.[129,640,171,1455]" lastPageId="9" lastPageNumber="71" pageId="7" pageNumber="69">
|
||
<emphasis id="8FEBE229FFBBFF97529BFBAEFCFCFBDD" box="[706,837,1070,1095]" italics="true" pageId="7" pageNumber="69">Description:</emphasis>
|
||
Shell tiny, off-white, depressed-globular form with domed spire; body whorl widest in standard apertural view; protoconch with 1.5 whorls and a minute and very uneven pattern of polygonal, rather sharp-crested ridges locally forming minute, conical peaks where ridges split (the structure also reminiscent of cross-sectional trabecular bone), no spiral striation discernible; teleoconch ornamented by some weak radial growth lines and much stronger, ropelike, equidistantly-arranged spiral striae (ca. 14–15 on body whorl from standard apertural view); the 3.25–3.5 whorls are separated by a deep suture; whorls slightly shouldered; aperture oblique to shell axis in lateral view; umbilicus relatively wide, less than one third of shell width; aperture kidney-shaped with strongly concave parietal side; sinulus wide, weakly separated due to weak parietal tooth; peristome expanded, not reflected; parietal callus strongly protruding, but in line with curvature of penultimate whorl and beyond aperture edge (profile) in lateral view (
|
||
<figureCitation id="25A422BEFFB5FF995275FEBBFDC9FECF" box="[556,624,315,341]" captionStart="FIGURE 3" captionStartId="8.[98,165,1712,1733]" captionTargetBox="[98,1156,164,1693]" captionTargetId="figure-3@8.[98,1156,164,1693]" captionText="FIGURE 3 Angustopila psammion n. sp. A:paratype, B–I: holotype." figureDoi="http://doi.org/10.5281/zenodo.6147320" httpUri="https://zenodo.org/record/6147320/files/figure.png" pageId="9" pageNumber="71">fig. 3C</figureCitation>
|
||
), detached from penultimate whorl; parietal tooth weak, low, short, rounded, does not reach peristome.
|
||
</paragraph>
|
||
<caption id="E9E06EB3FFB4FF98503BF930FD61F95C" ID-DOI="http://doi.org/10.5281/zenodo.6147320" ID-Zenodo-Dep="6147320" box="[98,728,1712,1734]" httpUri="https://zenodo.org/record/6147320/files/figure.png" pageId="8" pageNumber="70" startId="8.[98,165,1712,1733]" targetBox="[98,1156,164,1693]" targetPageId="8">
|
||
<paragraph id="BD203E3BFFB4FF98503BF930FD61F95C" blockId="8.[98,728,1712,1734]" box="[98,728,1712,1734]" pageId="8" pageNumber="70">
|
||
<pageTitle id="FD00E65CFFB4FF98503BF930FF0DF95F" box="[98,180,1712,1733]" pageId="8" pageNumber="70">FIGURE 3</pageTitle>
|
||
<taxonomicName id="7A9F45B8FFB4FF985082F930FE15F95F" authority="Páll-Gergely & Jochum & J.Vermeulen & Anker & Hunyadi & Örstan & Szabó, 2022" authorityName="Páll-Gergely & Jochum & J.Vermeulen & Anker & Hunyadi & Örstan & Szabó" authorityYear="2022" box="[219,428,1712,1733]" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="70" phylum="Mollusca" rank="species" species="psammion" status="n. sp.">
|
||
<emphasis id="8FEBE229FFB4FF985082F930FE15F95F" box="[219,428,1712,1733]" italics="true" pageId="8" pageNumber="70">Angustopila psammion</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="94D85F52FFB4FF9851E9F930FE59F95F" box="[432,480,1712,1733]" pageId="8" pageNumber="70" rank="species">n. sp.</taxonomicNameLabel>
|
||
A: paratype, B–I: holotype.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="BD203E3BFFB5FF9950C6FE4BFE12FE7F" blockId="9.[129,640,171,1455]" box="[159,427,459,485]" pageId="9" pageNumber="71">Soft anatomy unknown.</paragraph>
|
||
<paragraph id="BD203E3BFFB5FF9950C6FE6FFEF6FDB7" blockId="9.[129,640,171,1455]" pageId="9" pageNumber="71">
|
||
<emphasis id="8FEBE229FFB5FF9950C6FE6FFE1BFD93" box="[159,418,495,521]" italics="true" pageId="9" pageNumber="71">Measurements (in mm):</emphasis>
|
||
SW = 0.6–0.68, SH = 0.46–0.57 (n = 24).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="F5856DB0FFB5FF9950C6FDB6FE42FCEA" pageId="9" pageNumber="71" type="diagnosis">
|
||
<paragraph id="BD203E3BFFB5FF9950C6FDB6FE42FCEA" blockId="9.[129,640,171,1455]" pageId="9" pageNumber="71">
|
||
<emphasis id="8FEBE229FFB5FF9950C6FDB6FE22FDCA" box="[159,411,566,592]" italics="true" pageId="9" pageNumber="71">Differential diagnosis:</emphasis>
|
||
<taxonomicName id="7A9F45B8FFB5FF9951EDFDB6FECDFD02" authority="Dumrongrojwattana, Chuenit & Wongkamhaeng, 2021" authorityName="Dumrongrojwattana, Chuenit & Wongkamhaeng" authorityYear="2021" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="71" phylum="Mollusca" rank="species" species="pallgergelyi">
|
||
<emphasis id="8FEBE229FFB5FF9951EDFDB6FF6CFDEE" italics="true" pageId="9" pageNumber="71">Angustopila pallgergelyi</emphasis>
|
||
<bibRefCitation id="D90E43CAFFB5FF9950ABFDDAFECDFD02" author="Dumrongrojwattana, Chuenit & Wongkamhaeng" firstAuthor="Dumrongrojwattana" journalOrPublisher="Raffles Bulletin of Zoology" pageId="9" pageNumber="71" pagination="102 - 108" part="69" refId="ref7017" refString="Dumrongrojwattana, P., Chuenit, S. & Wongkamhaeng, K. (2021) A new species of the world's smallest cave snail of the genus Angustopila Jochum, Slapnik & Pall- Gergely in Jochum, et al., 2014 (Gastropoda: Hypselostomatidae) from eastern Thailand. Raffles Bulletin of Zoology, 69, 102 - 108." title="A new species of the world's smallest cave snail of the genus Angustopila Jochum, Slapnik & Pall- Gergely in Jochum, et al., 2014 (Gastropoda: Hypselostomatidae) from eastern Thailand" type="journal article" year="2021">Dumrongrojwattana, Chuenit & Wongkamhaeng, 2021</bibRefCitation>
|
||
</taxonomicName>
|
||
is similar in shell and aperture shape, but is larger, has a prominent palatal tooth (absent in
|
||
<taxonomicName id="7A9F45B8FFB5FF9951DFFD46FDC6FD7A" authority="Páll-Gergely & Jochum & J.Vermeulen & Anker & Hunyadi & Örstan & Szabó, 2022" authorityName="Páll-Gergely & Jochum & J.Vermeulen & Anker & Hunyadi & Örstan & Szabó" authorityYear="2022" box="[390,639,710,736]" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="71" phylum="Mollusca" rank="species" species="psammion" status="n. sp.">
|
||
<emphasis id="8FEBE229FFB5FF9951DFFD46FDC6FD7A" box="[390,639,710,736]" italics="true" pageId="9" pageNumber="71">Angustopila psammion</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="94D85F52FFB5FF9950D8FD6AFF01FC9E" box="[129,184,746,772]" pageId="9" pageNumber="71" rank="species">n. sp.</taxonomicNameLabel>
|
||
)and a stronger parietal tooth.
|
||
<taxonomicName id="7A9F45B8FFB5FF9951A4FD6AFF4DFCB2" authority="Páll-Gergely & Jochum & J.Vermeulen & Anker & Hunyadi & Örstan & Szabó, 2022" authorityName="Páll-Gergely & Jochum & J.Vermeulen & Anker & Hunyadi & Örstan & Szabó" authorityYear="2022" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="71" phylum="Mollusca" rank="species" species="coprologos" status="n. sp.">
|
||
<emphasis id="8FEBE229FFB5FF9951A4FD6AFF4DFCB2" italics="true" pageId="9" pageNumber="71">Angustopila coprologos</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="94D85F52FFB5FF9950A5FC8EFE8EFCB2" box="[252,311,782,808]" pageId="9" pageNumber="71" rank="species">n. sp.</taxonomicNameLabel>
|
||
is slightly larger, has a rough shell surface, and has an additional subcolumellar tooth and two palatal teeth.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="F5856DB0FFB5FF9950C6FCF9FEC9FC41" pageId="9" pageNumber="71" type="etymology">
|
||
<paragraph id="BD203E3BFFB5FF9950C6FCF9FEC9FC41" blockId="9.[129,640,171,1455]" pageId="9" pageNumber="71">
|
||
<emphasis id="8FEBE229FFB5FF9950C6FCF9FEA1FC09" box="[159,280,889,915]" italics="true" pageId="9" pageNumber="71">Etymology:</emphasis>
|
||
The specific epithet (ΨαμμϯΟν) means a grain of sand in Greek and is used as a noun in apposition.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="F5856DB0FFB5FF9950C6FC65FDEAFBDD" pageId="9" pageNumber="71" type="distribution">
|
||
<paragraph id="BD203E3BFFB5FF9950C6FC65FDEAFBDD" blockId="9.[129,640,171,1455]" pageId="9" pageNumber="71">
|
||
<emphasis id="8FEBE229FFB5FF9950C6FC65FE91FC65" box="[159,296,997,1023]" italics="true" pageId="9" pageNumber="71">Distribution:</emphasis>
|
||
This new species is known only from the
|
||
<typeStatus id="62248099FFB5FF99517EFB8AFEE1FBBE" box="[295,344,1034,1060]" pageId="9" pageNumber="71">type</typeStatus>
|
||
locality, Cap La Cave, Ha Long Bay, Quang Ninh Province, Vietnam.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="F5856DB0FFB5FF9950C6FBD1FD8BFA34" pageId="9" pageNumber="71" type="biology_ecology">
|
||
<paragraph id="BD203E3BFFB5FF9950C6FBD1FD8BFA34" blockId="9.[129,640,171,1455]" pageId="9" pageNumber="71">
|
||
<emphasis id="8FEBE229FFB5FF9950C6FBD1FDBDFBF1" box="[159,516,1105,1131]" italics="true" pageId="9" pageNumber="71">
|
||
Habitat:
|
||
<taxonomicName id="7A9F45B8FFB5FF99515BFBD1FDBDFBF1" authority="Páll-Gergely & Jochum & J.Vermeulen & Anker & Hunyadi & Örstan & Szabó, 2022" authorityName="Páll-Gergely & Jochum & J.Vermeulen & Anker & Hunyadi & Örstan & Szabó" authorityYear="2022" box="[258,516,1105,1131]" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="71" phylum="Mollusca" rank="species" species="psammion" status="n. sp.">Angustopila psammion</taxonomicName>
|
||
</emphasis>
|
||
<taxonomicNameLabel id="94D85F52FFB5FF995257FBD1FDF5FBF1" box="[526,588,1105,1131]" pageId="9" pageNumber="71" rank="species">n. sp.</taxonomicNameLabel>
|
||
was found in large numbers in small sediment deposits along the walls of a dry cave (Cap La Cave), in complete darkness. We assume that the sediment had fallen in through crevices in the rock, because it contains a species-rich assemblage of bleached, opaque shells of surface-dwelling terrestrial gastropods. Shells of
|
||
<taxonomicName id="7A9F45B8FFB5FF9950D8FAF0FE3DFA10" authority="Páll-Gergely & Jochum & J.Vermeulen & Anker & Hunyadi & Örstan & Szabó, 2022" authorityName="Páll-Gergely & Jochum & J.Vermeulen & Anker & Hunyadi & Örstan & Szabó" authorityYear="2022" box="[129,388,1392,1418]" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="71" phylum="Mollusca" rank="species" species="psammion" status="n. sp.">
|
||
<emphasis id="8FEBE229FFB5FF9950D8FAF0FE3DFA10" box="[129,388,1392,1418]" italics="true" pageId="9" pageNumber="71">Angustopila psammion</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="94D85F52FFB5FF9951C9FAF1FE69FA11" box="[400,464,1393,1419]" pageId="9" pageNumber="71" rank="species">n. sp.</taxonomicNameLabel>
|
||
, however, look fresh and are probably autochthonous.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |