treatments-xml/data/2C/04/87/2C0487A6FFCEFF9399EBFB7CFE04F9F1.xml

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<mods:title id="9F1FB5D78B42F7197B3F65398A17D69E">The Orbiniidae (Annelida: Polychaeta) of Pacific Costa Rica</mods:title>
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<heading id="FF5A81DCFFCEFF9E99EBFB7CFDFBFAF2" bold="true" box="[151,539,1265,1291]" fontSize="11" level="1" pageId="5" pageNumber="188" reason="1">
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<taxonomicName id="63AD4D33FFCEFF9E99EBFB7CFE64FAF2" ID-CoL="6P98K" box="[151,388,1265,1291]" class="Polychaeta" family="Orbiniidae" genus="Leodamas" kingdom="Animalia" pageId="5" pageNumber="188" phylum="Annelida" rank="species" species="hamatus" status="sp. nov.">
<emphasis id="96D9EAA2FFCEFF9E99EBFB7CFE64FAF2" bold="true" box="[151,388,1265,1291]" italics="true" pageId="5" pageNumber="188">Leodamas hamatus</emphasis>
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</emphasis>
</heading>
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<paragraph id="A41236B0FFCEFF9E99EBFA99FDC7FAD5" blockId="5.[151,551,1265,1325]" box="[151,551,1300,1325]" pageId="5" pageNumber="188">
<figureCitation id="3C962A35FFCEFF9E99EBFA99FEE2FAD5" box="[151,258,1300,1325]" captionStart="FIGURE 2" captionStartId="3.[151,250,1500,1522]" captionTargetBox="[174,1411,559,1474]" captionTargetId="figure@3.[173,1413,555,1478]" captionTargetPageId="3" captionText="FIGURE 2. Leitoscoloplos multipapillatus: A, Anterior end, lateral view. Leitoscoloplos panamensis: B, Station 29, Gulf of Nicoya specimen, anterior end, lateral view. Leitoscoloplos pugettensis: C, Station 22, Gulf of Nicoya specimen, anterior end, lateral view; D, Station 9, Bahia Culebra specimen, midbody abdominal setiger, anterior view; E, Station 9, Bahia Culebra specimen, posterior abdominal setiger, posterior view. Leodamas hamatus n. sp.: F, Holotype, posterior thoracic setiger, anterior view; G, Holotype, Posterior notopodial emergent hook. Scale bars: A, C = 1000 µm, B = 500 µm, D, E = 100 µm, F = 50 µm G = 10 µm." httpUri="https://zenodo.org/record/237220/files/figure.png" pageId="5" pageNumber="188">Figures 2</figureCitation>
(F–G), 3 (A–G), 4 (B–G)
</paragraph>
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<paragraph id="A41236B0FFCEFF9E99EBFAD1FEE0FA69" blockId="5.[151,1436,1372,1455]" pageId="5" pageNumber="188">
<treatmentCitationGroup id="84BD119EFFCEFF9E99EBFAD1FEE0FA69" pageId="5" pageNumber="188">
<taxonomicName id="63AD4D33FFCEFF9E99EBFAD1FD27FA8B" ID-CoL="4VSJZ" authority="Vargas et al. 1985: 336" authorityName="Vargas et al." authorityPageNumber="336" authorityYear="1985" box="[151,711,1372,1394]" class="Polychaeta" family="Orbiniidae" genus="Scoloplos" kingdom="Animalia" pageId="5" pageNumber="188" phylum="Annelida" rank="species" species="ohlini" subGenus="Leodamas">
<emphasis id="96D9EAA2FFCEFF9E99EBFAD1FF1AFA8B" box="[151,250,1372,1394]" italics="true" pageId="5" pageNumber="188">Scoloplos</emphasis>
(
<emphasis id="96D9EAA2FFCEFF9E9870FAD1FE94FA8B" box="[268,372,1372,1394]" italics="true" pageId="5" pageNumber="188">Leodamas</emphasis>
) 
<emphasis id="96D9EAA2FFCEFF9E98FBFAD1FE22FA8B" box="[391,450,1372,1394]" italics="true" pageId="5" pageNumber="188">ohlini</emphasis>
: 
<treatmentCitation id="250C10A1FFCEFF9E98AFFAD0FD27FA8B" author="Vargas" box="[467,711,1372,1394]" page="336" pageId="5" pageNumber="188" year="1985">
<bibRefCitation id="C03C4B41FFCEFF9E98AFFAD0FD74FA8B" author="Vargas" box="[467,660,1372,1394]" pageId="5" pageNumber="188" refString="Vargas, J. A., Dean, H. K., Maurer, D. &amp; Orellana, P. (1985) Lista preliminar de invertebrados asociadoa a los sedimentos del Golfo de Nicoya, Costa Rica. Brenesia, 24, 327 - 341." type="journal article" year="1985">
Vargas 
<emphasis id="96D9EAA2FFCEFF9E9B5DFAD0FDB6FA8B" box="[545,598,1372,1394]" italics="true" pageId="5" pageNumber="188">et al.</emphasis>
1985
</bibRefCitation>
: 336
</treatmentCitation>
</taxonomicName>
.— 
<treatmentCitation id="250C10A1FFCEFF9E9B98FAD1FC34FA8B" author="Maurer" box="[740,980,1372,1394]" page="47" pageId="5" pageNumber="188" year="1988">
<bibRefCitation id="C03C4B41FFCEFF9E9B98FAD1FC4DFA8B" author="Maurer" box="[740,941,1372,1394]" pageId="5" pageNumber="188" refString="Maurer, D., Vargas, J. &amp; Dean, H. (1988) Polychaetous Annelids from the Gulf of Nicoya, Costa Rica. Internationale Revue der Gesamten Hydrobiologie, 73 (1), 43 - 59. http: // dx. doi. org / 10.1002 / iroh. 19880730105" type="journal article" year="1988">
Maurer 
<emphasis id="96D9EAA2FFCEFF9E9A46FAD0FC8FFA8B" box="[826,879,1372,1394]" italics="true" pageId="5" pageNumber="188">et al.</emphasis>
1988
</bibRefCitation>
: 47
</treatmentCitation>
.— 
<treatmentCitation id="250C10A1FFCEFF9E9A8FFAD1FB41FA8B" author="Dean" box="[1011,1185,1372,1394]" page="74" pageId="5" pageNumber="188" year="1996">
<bibRefCitation id="C03C4B41FFCEFF9E9A8FFAD1FB99FA8B" author="Dean" box="[1011,1145,1372,1394]" pageId="5" pageNumber="188" refString="Dean, H. K. (1996 a) Subtidal benthic polychaetes (Annelida) of the Gulf of Nicoya, Costa Rica. Revista Biologia Tropical, 44 (Supplement 3), 69 - 80." type="journal article" year="1996" yearSuffix="a">Dean, 1996a</bibRefCitation>
: 74
</treatmentCitation>
; 2009: 146. Not 
<bibRefCitation id="C03C4B41FFCEFF9E9C29FAD1FF1DFA69" author="Ehlers" pageId="5" pageNumber="188" refString="Ehlers, E. (1901) Die Polychaeten des magellanischen und chilenischen Strandes. Ein faunistischer Versuch, Festschrift zur Feier des Underfunfzigjahrigen Bestehens der koniglichen Gesellschaft der Wissenschaft der Wissenschaften zu Gottingen, pp. 1 - 232, 25 plates." type="book chapter" year="1901">Ehlers, 1901</bibRefCitation>
.
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</paragraph>
<paragraph id="A41236B0FFCEFF9E99EBFA14FC39FA56" blockId="5.[151,1436,1372,1455]" box="[151,985,1433,1455]" pageId="5" pageNumber="188">
<treatmentCitationGroup id="84BD119EFFCEFF9E99EBFA14FC39FA56" box="[151,985,1433,1455]" pageId="5" pageNumber="188">
<taxonomicName id="63AD4D33FFCEFF9E99EBFA14FD43FA56" ID-CoL="856XS" authority="Dean, 1996a: 74" authorityName="Dean" authorityPageNumber="74" authorityYear="1996" box="[151,675,1433,1455]" class="Polychaeta" family="Orbiniidae" genus="Scoloplos" kingdom="Animalia" pageId="5" pageNumber="188" phylum="Annelida" rank="species" species="tribulosus" subGenus="Leodamas">
<emphasis id="96D9EAA2FFCEFF9E99EBFA14FE08FA56" box="[151,488,1433,1455]" italics="true" pageId="5" pageNumber="188">Scoloplos (Leodamas) tribulosus</emphasis>
: 
<treatmentCitation id="250C10A1FFCEFF9E988BFA17FD43FA56" author="Dean" box="[503,675,1433,1455]" page="74" pageId="5" pageNumber="188" year="1996">
<bibRefCitation id="C03C4B41FFCEFF9E988BFA17FD9CFA56" author="Dean" box="[503,636,1433,1455]" pageId="5" pageNumber="188" refString="Dean, H. K. (1996 a) Subtidal benthic polychaetes (Annelida) of the Gulf of Nicoya, Costa Rica. Revista Biologia Tropical, 44 (Supplement 3), 69 - 80." type="journal article" year="1996" yearSuffix="a">Dean, 1996a</bibRefCitation>
: 74
</treatmentCitation>
</taxonomicName>
; 2009: 146. Not 
<bibRefCitation id="C03C4B41FFCEFF9E9A2DFA14FC35FA56" author="Ehlers" box="[849,981,1433,1455]" pageId="5" pageNumber="188" refString="Ehlers, E. (1897) Polychaeten. Hamburger Magalhaensische Sammelreise, 1897, 1 - 148." type="journal article" year="1897">Ehlers, 1897</bibRefCitation>
.
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</paragraph>
</subSubSection>
<subSubSection id="ECB7653BFFCEFF9E99EBFA51FE62F831" pageId="5" pageNumber="188" type="materials_examined">
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<emphasis id="96D9EAA2FFCEFF9E99EBFA51FE9CFA0C" bold="true" box="[151,380,1500,1525]" pageId="5" pageNumber="188">Material examined</emphasis>
. Gulf of Nicoya: Punta Morales, South point, 10°3′30.2″N, 84°57′4.8″W, intertidal, muddy sand, 
<date id="D0131070FFCEFF9E99A7FA72FEDFF9E1" box="[219,319,1535,1560]" pageId="5" pageNumber="188" value="1995-07">Jul 1995</date>
, 
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<emphasis id="96D9EAA2FFCEFF9E9830FA72FE59F9E1" bold="true" box="[332,441,1535,1560]" pageId="5" pageNumber="188">Holotype</emphasis>
</typeStatus>
(
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50373); Sta. 2, 9º55′28″N, 84º52ˊ05″ W, 
<quantity id="63559B55FFCEFF9E9A8CF98DFBCDF9EE" box="[1008,1069,1536,1560]" metricMagnitude="1" metricUnit="m" metricValue="1.8" pageId="5" pageNumber="188" unit="m" value="18.0">18 m</quantity>
, muddy sand, 
<date id="D0131070FFCEFF9E9DAAF98DFA8AF9E1" box="[1238,1386,1535,1560]" pageId="5" pageNumber="188" value="1980-07-13">13 July 1980</date>
(1); Sta. 24, 9°49′25″N, 
<geoCoordinate id="C1995077FFCEFF9E9801F9A9FDE8F9C4" box="[381,520,1572,1597]" direction="west" orientation="longitude" pageId="5" pageNumber="188" precision="15" value="-84.68889">84°41′20&quot;W</geoCoordinate>
, 
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, sand, 
<date id="D0131070FFCEFF9E9BDAF9A9FCCDF9C5" box="[678,813,1572,1597]" pageId="5" pageNumber="188" value="1980-07-10">10 Jul 1980</date>
(1), 
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(5), 
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, 
<emphasis id="96D9EAA2FFCEFF9E9DC3F9A9FAAEF9C4" bold="true" box="[1215,1358,1572,1597]" pageId="5" pageNumber="188">
2 
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</emphasis>
(
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83993) (1), 
<date id="D0131070FFCEFF9E985CF9C5FE46F999" box="[288,422,1608,1632]" pageId="5" pageNumber="188" value="1982-04-04">4 Apr 1982</date>
, 1 
<typeStatus id="7B168812FFCEFF9E98B5F9CAFDD6F999" box="[457,566,1607,1632]" pageId="5" pageNumber="188" type="paratype">
<emphasis id="96D9EAA2FFCEFF9E98B5F9CAFDD6F999" bold="true" box="[457,566,1607,1632]" pageId="5" pageNumber="188">Paratype</emphasis>
</typeStatus>
(
<collectionCode id="C2BCAE75FFCEFF9E9B34F9C5FD67F999" LSID="urn:lsid:biocol.org:col:33791" box="[584,647,1608,1632]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="5" pageNumber="188">MCZ</collectionCode>
50374); Sta. 28, 9°52′16″N, 84°45′30″W, 
<quantity id="63559B55FFCEFF9E9D13F9CAFB4CF9A6" box="[1135,1196,1607,1632]" metricMagnitude="1" metricUnit="m" metricValue="2.6" pageId="5" pageNumber="188" unit="m" value="26.0">26 m</quantity>
, mud, 
<date id="D0131070FFCEFF9E9D87F9C5FA61F999" box="[1275,1409,1608,1632]" pageId="5" pageNumber="188" value="1982-04-01">1 Apr 1982</date>
, 2 (
<collectionCode id="C2BCAE75FFCEFF9E99E2F9E1FF3DF97D" LSID="urn:lsid:biocol.org:col:33791" box="[158,221,1644,1668]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="5" pageNumber="188">MCZ</collectionCode>
83996); Sta. 29, 9°54′55″N, 84°45′15″W, 
<quantity id="63559B55FFCEFF9E9BBCF9E1FD1CF97D" box="[704,764,1644,1668]" metricMagnitude="1" metricUnit="m" metricValue="1.8" pageId="5" pageNumber="188" unit="m" value="18.0">18 m</quantity>
, muddy sand, 
<date id="D0131070FFCEFF9E9ADDF9E1FBC2F97D" box="[929,1058,1644,1668]" pageId="5" pageNumber="188" value="1980-10-01">1 Oct 1980</date>
, 
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1 
<typeStatus id="7B168812FFCEFF9E9D38F9E1FB51F97C" box="[1092,1201,1644,1669]" pageId="5" pageNumber="188" type="paratype">Paratype</typeStatus>
</emphasis>
(
<collectionCode id="C2BCAE75FFCEFF9E9DBDF9E1FAC6F97D" box="[1217,1318,1644,1668]" pageId="5" pageNumber="188">MZUCR</collectionCode>
30-3) (6), 
<date id="D0131070FFCEFF9E99EBF91DFEC2F951" box="[151,290,1680,1704]" pageId="5" pageNumber="188" value="1981-01-27">27 Jan 1981</date>
, (
<emphasis id="96D9EAA2FFCEFF9E9844F902FEA6F951" bold="true" box="[312,326,1679,1704]" pageId="5" pageNumber="188">2</emphasis>
). 
<typeStatus id="7B168812FFCEFF9E9821F902FE35F951" box="[349,469,1679,1704]" pageId="5" pageNumber="188" type="paratype">
<emphasis id="96D9EAA2FFCEFF9E9821F902FE35F951" bold="true" box="[349,469,1679,1704]" pageId="5" pageNumber="188">Paratypes</emphasis>
</typeStatus>
(
<collectionCode id="C2BCAE75FFCEFF9E989AF91DFDACF951" box="[486,588,1680,1704]" pageId="5" pageNumber="188">MZUCR</collectionCode>
153-01 &amp; 
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31-02) (6), 
<date id="D0131070FFCEFF9E9AC4F91DFBD7F951" box="[952,1079,1680,1704]" pageId="5" pageNumber="188" value="1981-06-07">7 Jun 1981</date>
, 
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1 
<typeStatus id="7B168812FFCEFF9E9D25F902FB26F951" box="[1113,1222,1679,1704]" pageId="5" pageNumber="188" type="paratype">Paratype</typeStatus>
</emphasis>
(
<collectionCode id="C2BCAE75FFCEFF9E9DABF91DFAF6F951" LSID="urn:lsid:biocol.org:col:33791" box="[1239,1302,1680,1704]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="5" pageNumber="188">MCZ</collectionCode>
50375) (2), 
<date id="D0131070FFCEFF9E99EBF939FEFEF935" box="[151,286,1716,1740]" pageId="5" pageNumber="188" value="1982-04-01">1 Apr 1982</date>
, 
<emphasis id="96D9EAA2FFCEFF9E9850F939FE5AF934" bold="true" box="[300,442,1716,1741]" pageId="5" pageNumber="188">
2 
<typeStatus id="7B168812FFCEFF9E983EF939FE5AF934" box="[322,442,1716,1741]" pageId="5" pageNumber="188" type="paratype">Paratypes</typeStatus>
</emphasis>
(
<collectionCode id="C2BCAE75FFCEFF9E98B0F939FDEBF935" LSID="urn:lsid:biocol.org:col:33791" box="[460,523,1716,1740]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="5" pageNumber="188">MCZ</collectionCode>
50377 (SEM) (2); Sta. 30, 9°54′40″N, 84°45′50″W, 
<quantity id="63559B55FFCEFF9E9D13F939FB4CF935" box="[1135,1196,1716,1740]" metricMagnitude="1" metricUnit="m" metricValue="1.8" pageId="5" pageNumber="188" unit="m" value="18.0">18 m</quantity>
, muddy sand, 
<date id="D0131070FFCEFF9E9C2AF939FF32F909" pageId="5" pageNumber="188" value="1980-07-10">10 Jul 1980</date>
, 
<emphasis id="96D9EAA2FFCEFF9E999EF95AFE88F909" bold="true" box="[226,360,1751,1776]" pageId="5" pageNumber="188">
1 
<typeStatus id="7B168812FFCEFF9E9987F95AFE88F909" box="[251,360,1751,1776]" pageId="5" pageNumber="188" type="paratype">Paratype</typeStatus>
</emphasis>
(
<collectionCode id="C2BCAE75FFCEFF9E9800F955FE5BF909" LSID="urn:lsid:biocol.org:col:33791" box="[380,443,1752,1776]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="5" pageNumber="188">MCZ</collectionCode>
83992); Sta. 31, 9°44′00″N, 84°59′42″W, 
<quantity id="63559B55FFCEFF9E9ACBF955FC16F916" box="[951,1014,1752,1776]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="5" pageNumber="188" unit="m" value="20.0">20 m</quantity>
, mud/sand, 
<date id="D0131070FFCEFF9E9DFBF955FAE4F909" box="[1159,1284,1751,1776]" pageId="5" pageNumber="188" value="1980-07-09">9 Jul 1980</date>
(1); Sta. 35, 9°55′42″N, 84°47′40″W, 
<quantity id="63559B55FFCEFF9E98C0F971FE1AF8ED" box="[444,506,1788,1812]" metricMagnitude="1" metricUnit="m" metricValue="1.3" pageId="5" pageNumber="188" unit="m" value="13.0">13 m</quantity>
, sand, 
<date id="D0131070FFCEFF9E9B33F971FD38F8ED" box="[591,728,1788,1813]" pageId="5" pageNumber="188" value="1980-07-12">12 Jul 1980</date>
(1). Punta Morales: Playa Blanca, 
<geoCoordinate id="C1995077FFCEFF9E9D06F971FB3AF8EC" box="[1146,1242,1788,1813]" direction="north" orientation="latitude" pageId="5" pageNumber="188" precision="925" value="10.066667">10°04′N</geoCoordinate>
, 
<geoCoordinate id="C1995077FFCEFF9E9D94F971FAACF8EC" box="[1256,1356,1788,1813]" direction="west" orientation="longitude" pageId="5" pageNumber="188" precision="925" value="-84.96667">84°58′W</geoCoordinate>
, lower intertidal, muddy sand, 1986, 
<emphasis id="96D9EAA2FFCEFF9E9B79F892FDF3F8C1" bold="true" box="[517,531,1823,1848]" pageId="5" pageNumber="188">1</emphasis>
<typeStatus id="7B168812FFCEFF9E9B5CF892FD6DF8C1" box="[544,653,1823,1848]" pageId="5" pageNumber="188" type="paratype">Paratype</typeStatus>
(
<collectionCode id="C2BCAE75FFCEFF9E9BD9F8ADFCEAF8C1" box="[677,778,1824,1848]" pageId="5" pageNumber="188">MZUCR</collectionCode>
156-01), col: J. A. Vargas; South point, 10°3′30.2″N, 84°57′4.8″W, mid intertidal, muddy sand, 
<date id="D0131070FFCEFF9E9BFCF8C9FCF8F8A5" box="[640,792,1860,1884]" pageId="5" pageNumber="188" value="1988-08-29">29 Aug 1988</date>
, col: H. K. Dean, (2); east of South Point, 10°3′34.9″N, 84°57′6.0″W, intertidal fine sand, col: H. K. Dean, 
<date id="D0131070FFCEFF9E9B9CF8E5FC97F879" box="[736,887,1896,1920]" pageId="5" pageNumber="188" value="1988-08-29">29 Aug 1988</date>
, 
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2 
<typeStatus id="7B168812FFCEFF9E9AE6F8EAFBF2F879" box="[922,1042,1895,1920]" pageId="5" pageNumber="188" type="paratype">Paratypes</typeStatus>
</emphasis>
(
<collectionCode id="C2BCAE75FFCEFF9E9D5EF8E5FB81F879" LSID="urn:lsid:biocol.org:col:33791" box="[1058,1121,1896,1920]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="5" pageNumber="188">MCZ</collectionCode>
50378). Guanacaste, Bahia Culebra: Sta. 3, 
<geoCoordinate id="C1995077FFCEFF9E982FF801FE8EF85C" box="[339,366,1932,1957]" direction="north" orientation="latitude" pageId="5" pageNumber="188" precision="1" value="10.59525">10</geoCoordinate>
º35′42.9&quot;N, 85º39′16.6″W, 
<quantity id="63559B55FFCEFF9E9BD7F801FD08F85D" box="[683,744,1932,1956]" metricMagnitude="1" metricUnit="m" metricValue="1.3" pageId="5" pageNumber="188" unit="m" value="13.0">13 m</quantity>
, 
<date id="D0131070FFCEFF9E9B8AF801FC6EF85D" box="[758,910,1932,1957]" pageId="5" pageNumber="188" value="2011-05-19">19 May 2011</date>
(1); Sta. 12, 10º35′56.5″N, 85º40′34.6″W, 
<quantity id="63559B55FFCEFF9E9CFCF801FF4FF831" metricMagnitude="1" metricUnit="m" metricValue="1.2" pageId="5" pageNumber="188" unit="m" value="12.0">12 m</quantity>
, 
<date id="D0131070FFCEFF9E99C7F83DFEB1F831" box="[187,337,1968,1993]" pageId="5" pageNumber="188" value="2011-05-19">19 May 2011</date>
(1).
</paragraph>
</subSubSection>
<subSubSection id="ECB7653BFFCEFF9399BBF859FC35FE29" lastPageId="8" lastPageNumber="191" pageId="5" pageNumber="188" type="description">
<paragraph id="A41236B0FFCEFF9E99BBF859FA7DF815" blockId="5.[151,1437,1500,2029]" box="[199,1437,2004,2029]" pageId="5" pageNumber="188">
<emphasis id="96D9EAA2FFCEFF9E99BBF859FEB8F814" bold="true" box="[199,344,2004,2029]" pageId="5" pageNumber="188">Description.</emphasis>
An elongate species, all specimens incomplete, maximum thoracic width 0.30–1.23 mm.
</paragraph>
<paragraph id="A41236B0FFCDFF9D99EBFF1AFBDAFEE4" blockId="6.[151,1437,151,464]" pageId="6" pageNumber="189">
<typeStatus id="7B168812FFCDFF9D99EBFF1AFF1FFF49" box="[151,255,151,176]" pageId="6" pageNumber="189" type="holotype">Holotype</typeStatus>
incomplete, 
<quantity id="63559B55FFCDFF9D98DEFF15FE17FF56" box="[418,503,152,176]" metricMagnitude="-2" metricUnit="m" metricValue="3.2" pageId="6" pageNumber="189" unit="mm" value="32.0">32 mm</quantity>
long, 0.8 mm wide with 115 segments (
<figureCitation id="3C962A35FFCDFF9D9A92FF1AFBA1FF49" box="[1006,1089,151,176]" captionStart="FIGURE 3" captionStartId="6.[151,250,1831,1853]" captionTargetBox="[205,1384,507,1790]" captionTargetId="figure@6.[189,1397,491,1810]" captionTargetPageId="6" captionText="FIGURE 3. Leodamas hamatus n. sp.: A, holotype, anterior end, dorsal view; B, Holotype, posterior abdominal setiger, anterior view; C, Holotype, anterior thoracic mid-neuropodial spine; D &amp; E, Holotype, posterior thoracic mid-neuropodial spines; F, Holotype, abdominal notopodial furcate setae; G, Holotype, posterior neuropodial emergent hooked spine. Scale bars: A = 200 µm, B = 100 µm, C ‒ G = 10 µm." httpUri="https://zenodo.org/record/237221/files/figure.png" pageId="6" pageNumber="189">Figs. 3</figureCitation>
A, 4B). Prostomium conical; eyespots absent; paired dorsolateral nuchal organs present. Peristomium slightly shorter than prostomium, subequal to first setiger. Thorax with 13–21 setigers, 
<typeStatus id="7B168812FFCDFF9D9A68FF52FC96FF01" box="[788,886,223,248]" pageId="6" pageNumber="189" type="holotype">holotype</typeStatus>
with 18; transition to abdomen abrupt. Short, conical branchiae from setiger 
<quantity id="63559B55FFCDFF9D988BFE89FDC3FEE4" box="[503,547,260,285]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="6" pageNumber="189" unit="in" value="6.0">6 in</quantity>
all specimens, increasing in length posteriorly.
</paragraph>
<paragraph id="A41236B0FFCDFF9D99BBFEAAFC2DFE29" blockId="6.[151,1437,151,464]" pageId="6" pageNumber="189">
Thoracic notopodial presetal lobe a low ridge, postsetal lobe short, conical at first, increasing in length posteriorly; thoracic neuropodia with low presetal ridge, postsetal ridge somewhat higher (
<figureCitation id="3C962A35FFCDFF9D9DCCFEC1FB16FE9D" box="[1200,1270,332,357]" captionStart="FIGURE 2" captionStartId="3.[151,250,1500,1522]" captionTargetBox="[174,1411,559,1474]" captionTargetId="figure@3.[173,1413,555,1478]" captionTargetPageId="3" captionText="FIGURE 2. Leitoscoloplos multipapillatus: A, Anterior end, lateral view. Leitoscoloplos panamensis: B, Station 29, Gulf of Nicoya specimen, anterior end, lateral view. Leitoscoloplos pugettensis: C, Station 22, Gulf of Nicoya specimen, anterior end, lateral view; D, Station 9, Bahia Culebra specimen, midbody abdominal setiger, anterior view; E, Station 9, Bahia Culebra specimen, posterior abdominal setiger, posterior view. Leodamas hamatus n. sp.: F, Holotype, posterior thoracic setiger, anterior view; G, Holotype, Posterior notopodial emergent hook. Scale bars: A, C = 1000 µm, B = 500 µm, D, E = 100 µm, F = 50 µm G = 10 µm." httpUri="https://zenodo.org/record/237220/files/figure.png" pageId="6" pageNumber="189">Fig. 2</figureCitation>
F). Abdominal notopodial postsetal lobe conical, shorter than branchiae; abdominal neuropodia with presetal lobe a low ridge, postsetal lobe triangular; with a single well-developed, projecting acicula which enlarges with a hooked tip (
<figureCitation id="3C962A35FFCDFF9D9C13FE19FF45FE29" captionStart="FIGURE 3" captionStartId="6.[151,250,1831,1853]" captionTargetBox="[205,1384,507,1790]" captionTargetId="figure@6.[189,1397,491,1810]" captionTargetPageId="6" captionText="FIGURE 3. Leodamas hamatus n. sp.: A, holotype, anterior end, dorsal view; B, Holotype, posterior abdominal setiger, anterior view; C, Holotype, anterior thoracic mid-neuropodial spine; D &amp; E, Holotype, posterior thoracic mid-neuropodial spines; F, Holotype, abdominal notopodial furcate setae; G, Holotype, posterior neuropodial emergent hooked spine. Scale bars: A = 200 µm, B = 100 µm, C ‒ G = 10 µm." httpUri="https://zenodo.org/record/237221/files/figure.png" pageId="6" pageNumber="189">Fig. 3</figureCitation>
B, G) and becomes greatly emergent in far posterior setigers (
<figureCitation id="3C962A35FFCDFF9D9A18FE3AFC49FE29" box="[868,937,439,464]" captionStart="FIGURE 4" captionStartId="7.[151,250,1824,1846]" captionTargetBox="[154,1428,201,1801]" captionTargetId="figure@7.[150,1436,193,1803]" captionTargetPageId="7" captionText="FIGURE 4. Leitoscoloplos multipapillatus: A, Setigers 17 – 19, ventral view. Leodamas hamatus n. sp.: B, Paratype XXX Anterior region, lateral view; C, Anterior thoracic spines, setiger three, mid-neuropodial region; D, Posterior thoracic spines, setiger 15, mid-neuropodial region; E, Posterior thoracic spines, upper neuropodial region; F, Furcate notosetae, posterior abdominal region; G, Posterior emergent neuroaciculum, posterior abdominal region. Scale bars: A = 200 µm, B = 500 µm, C ‒ E, F, G = 10 µm. h = hoods, stp = stomach papillae." httpUri="https://zenodo.org/record/237222/files/figure.png" pageId="6" pageNumber="189">Fig. 4</figureCitation>
G).
</paragraph>
<caption id="F0D26638FFCDFF9D99EBF8AAFD97F863" httpUri="https://zenodo.org/record/237221/files/figure.png" pageId="6" pageNumber="189" targetBox="[205,1384,507,1790]" targetPageId="6">
<paragraph id="A41236B0FFCDFF9D99EBF8AAFD97F863" blockId="6.[151,1436,1831,1946]" pageId="6" pageNumber="189">
<emphasis id="96D9EAA2FFCDFF9D99EBF8AAFEF7F8C5" bold="true" box="[151,279,1831,1853]" pageId="6" pageNumber="189">FIGURE 3.</emphasis>
<taxonomicName id="63AD4D33FFCDFF9D985DF8AAFE0AF8C4" box="[289,490,1831,1853]" class="Polychaeta" family="Orbiniidae" genus="Leodamas" kingdom="Animalia" pageId="6" pageNumber="189" phylum="Annelida" rank="species" species="hamatus" status="sp. nov.">
<emphasis id="96D9EAA2FFCDFF9D985DF8AAFE0AF8C4" box="[289,490,1831,1853]" italics="true" pageId="6" pageNumber="189">Leodamas hamatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8DEA57D9FFCDFF9D988BF8A5FDD3F8C4" box="[503,563,1832,1853]" pageId="6" pageNumber="189" rank="species">
<emphasis id="96D9EAA2FFCDFF9D988BF8A5FDD3F8C4" bold="true" box="[503,563,1832,1853]" pageId="6" pageNumber="189">n. sp.</emphasis>
</taxonomicNameLabel>
: A, holotype, anterior end, dorsal view; B, Holotype, posterior abdominal setiger, anterior view; C, Holotype, anterior thoracic mid-neuropodial spine; D&amp;E, Holotype, posterior thoracic mid-neuropodial spines; F, Holotype, abdominal notopodial furcate setae; G, Holotype, posterior neuropodial emergent hooked spine. Scale bars: A = 200 µm, B = 100 µm, C‒G = 10 µm.
</paragraph>
</caption>
<caption id="F0D26638FFCCFF9C99EBF8ADFD28F848" httpUri="https://zenodo.org/record/237222/files/figure.png" pageId="7" pageNumber="190" targetBox="[154,1428,201,1801]" targetPageId="7">
<paragraph id="A41236B0FFCCFF9C99EBF8ADFD28F848" blockId="7.[151,1436,1824,1969]" pageId="7" pageNumber="190">
<emphasis id="96D9EAA2FFCCFF9C99EBF8ADFEF7F8CC" bold="true" box="[151,279,1824,1846]" pageId="7" pageNumber="190">FIGURE 4.</emphasis>
<taxonomicName id="63AD4D33FFCCFF9C9863F8ADFDB7F8CF" box="[287,599,1824,1846]" class="Polychaeta" family="Orbiniidae" genus="Leitoscoloplos" kingdom="Animalia" pageId="7" pageNumber="190" phylum="Annelida" rank="species" species="multipapillatus">
<emphasis id="96D9EAA2FFCCFF9C9863F8ADFDB7F8CF" box="[287,599,1824,1846]" italics="true" pageId="7" pageNumber="190">Leitoscoloplos multipapillatus</emphasis>
</taxonomicName>
: A, Setigers 17–19, ventral view. 
<taxonomicName id="63AD4D33FFCCFF9C9ABFF8ADFB6BF8CF" box="[963,1163,1824,1846]" class="Polychaeta" family="Orbiniidae" genus="Leodamas" kingdom="Animalia" pageId="7" pageNumber="190" phylum="Annelida" rank="species" species="hamatus" status="sp. nov.">
<emphasis id="96D9EAA2FFCCFF9C9ABFF8ADFB6BF8CF" box="[963,1163,1824,1846]" italics="true" pageId="7" pageNumber="190">Leodamas hamatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8DEA57D9FFCCFF9C9DEBF8ADFB32F8CC" box="[1175,1234,1824,1845]" pageId="7" pageNumber="190" rank="species">
<emphasis id="96D9EAA2FFCCFF9C9DEBF8ADFB32F8CC" bold="true" box="[1175,1234,1824,1845]" pageId="7" pageNumber="190">n. sp.</emphasis>
</taxonomicNameLabel>
: B, Paratype XXX Anterior region, lateral view; C, Anterior thoracic spines, setiger three, mid-neuropodial region; D, Posterior thoracic spines, setiger 15, mid-neuropodial region; E, Posterior thoracic spines, upper neuropodial region; F, Furcate notosetae, posterior abdominal region; G, Posterior emergent neuroaciculum, posterior abdominal region. Scale bars: A = 200 µm, B = 500 µm, C‒E, F, G = 10 µm. 
<emphasis id="96D9EAA2FFCCFF9C9814F816FE94F848" box="[360,372,1947,1969]" italics="true" pageId="7" pageNumber="190">h</emphasis>
= hoods, 
<emphasis id="96D9EAA2FFCCFF9C98A6F811FE17F848" box="[474,503,1948,1969]" italics="true" pageId="7" pageNumber="190">stp</emphasis>
= stomach papillae.
</paragraph>
</caption>
<paragraph id="A41236B0FFC3FF9399BBFF1AFC35FE29" blockId="8.[151,1437,151,1544]" pageId="8" pageNumber="191">
Thoracic notosetae camerated capillaries throughout; neurosetae include a small dorsal fascicle of 2‒7 thin capillaries and rows of heavy uncini arranged in 2‒4 complete longitudinal rows, with a partial posteriormost fifth row; uncini with 5‒8 transverse rows of small serrations on shaft (
<figureCitation id="3C962A35FFC3FF939A91FF52FBDFFF01" box="[1005,1087,223,248]" captionStart="FIGURE 3" captionStartId="6.[151,250,1831,1853]" captionTargetBox="[205,1384,507,1790]" captionTargetId="figure@6.[189,1397,491,1810]" captionTargetPageId="6" captionText="FIGURE 3. Leodamas hamatus n. sp.: A, holotype, anterior end, dorsal view; B, Holotype, posterior abdominal setiger, anterior view; C, Holotype, anterior thoracic mid-neuropodial spine; D &amp; E, Holotype, posterior thoracic mid-neuropodial spines; F, Holotype, abdominal notopodial furcate setae; G, Holotype, posterior neuropodial emergent hooked spine. Scale bars: A = 200 µm, B = 100 µm, C ‒ G = 10 µm." httpUri="https://zenodo.org/record/237221/files/figure.png" pageId="8" pageNumber="191">Figs. 3</figureCitation>
C–D, 4–D). First four or five thoracic setigers with narrow uncini with a blunt tip, and thin, appressed hood (sometimes eroded) (
<figureCitation id="3C962A35FFC3FF939D80FE89FAAAFEE5" box="[1276,1354,260,285]" captionStart="FIGURE 3" captionStartId="6.[151,250,1831,1853]" captionTargetBox="[205,1384,507,1790]" captionTargetId="figure@6.[189,1397,491,1810]" captionTargetPageId="6" captionText="FIGURE 3. Leodamas hamatus n. sp.: A, holotype, anterior end, dorsal view; B, Holotype, posterior abdominal setiger, anterior view; C, Holotype, anterior thoracic mid-neuropodial spine; D &amp; E, Holotype, posterior thoracic mid-neuropodial spines; F, Holotype, abdominal notopodial furcate setae; G, Holotype, posterior neuropodial emergent hooked spine. Scale bars: A = 200 µm, B = 100 µm, C ‒ G = 10 µm." httpUri="https://zenodo.org/record/237221/files/figure.png" pageId="8" pageNumber="191">Figs. 3</figureCitation>
C, 4C); subsequent thoracic uncini much more robust with thickened hood (
<figureCitation id="3C962A35FFC3FF939ADBFEAAFC15FEB9" box="[935,1013,295,320]" captionStart="FIGURE 3" captionStartId="6.[151,250,1831,1853]" captionTargetBox="[205,1384,507,1790]" captionTargetId="figure@6.[189,1397,491,1810]" captionTargetPageId="6" captionText="FIGURE 3. Leodamas hamatus n. sp.: A, holotype, anterior end, dorsal view; B, Holotype, posterior abdominal setiger, anterior view; C, Holotype, anterior thoracic mid-neuropodial spine; D &amp; E, Holotype, posterior thoracic mid-neuropodial spines; F, Holotype, abdominal notopodial furcate setae; G, Holotype, posterior neuropodial emergent hooked spine. Scale bars: A = 200 µm, B = 100 µm, C ‒ G = 10 µm." httpUri="https://zenodo.org/record/237221/files/figure.png" pageId="8" pageNumber="191">Figs. 3</figureCitation>
D, 4D) that together with curved tip, providing uncini with a bifurcate appearance; 5‒8 serrated ridges on shaft (
<figureCitation id="3C962A35FFC3FF939A8FFEC1FBA1FE9D" box="[1011,1089,332,357]" captionStart="FIGURE 3" captionStartId="6.[151,250,1831,1853]" captionTargetBox="[205,1384,507,1790]" captionTargetId="figure@6.[189,1397,491,1810]" captionTargetPageId="6" captionText="FIGURE 3. Leodamas hamatus n. sp.: A, holotype, anterior end, dorsal view; B, Holotype, posterior abdominal setiger, anterior view; C, Holotype, anterior thoracic mid-neuropodial spine; D &amp; E, Holotype, posterior thoracic mid-neuropodial spines; F, Holotype, abdominal notopodial furcate setae; G, Holotype, posterior neuropodial emergent hooked spine. Scale bars: A = 200 µm, B = 100 µm, C ‒ G = 10 µm." httpUri="https://zenodo.org/record/237221/files/figure.png" pageId="8" pageNumber="191">Figs. 3</figureCitation>
E, 4D). Uppermost few uncini short with wide rounded tip and a thick hood projecting slightly beyond the tip in all thoracic setigers (
<figureCitation id="3C962A35FFC3FF939C4BFEE2FA9CFE71" box="[1335,1404,367,392]" captionStart="FIGURE 4" captionStartId="7.[151,250,1824,1846]" captionTargetBox="[154,1428,201,1801]" captionTargetId="figure@7.[150,1436,193,1803]" captionTargetPageId="7" captionText="FIGURE 4. Leitoscoloplos multipapillatus: A, Setigers 17 – 19, ventral view. Leodamas hamatus n. sp.: B, Paratype XXX Anterior region, lateral view; C, Anterior thoracic spines, setiger three, mid-neuropodial region; D, Posterior thoracic spines, setiger 15, mid-neuropodial region; E, Posterior thoracic spines, upper neuropodial region; F, Furcate notosetae, posterior abdominal region; G, Posterior emergent neuroaciculum, posterior abdominal region. Scale bars: A = 200 µm, B = 500 µm, C ‒ E, F, G = 10 µm. h = hoods, stp = stomach papillae." httpUri="https://zenodo.org/record/237222/files/figure.png" pageId="8" pageNumber="191">Fig. 4</figureCitation>
E). Abdominal notosetae camerated capillaries accompanied by 1‒3 furcate setae posteriorly (
<figureCitation id="3C962A35FFC3FF939DEFFE19FB3FFE55" box="[1171,1247,404,429]" captionStart="FIGURE 3" captionStartId="6.[151,250,1831,1853]" captionTargetBox="[205,1384,507,1790]" captionTargetId="figure@6.[189,1397,491,1810]" captionTargetPageId="6" captionText="FIGURE 3. Leodamas hamatus n. sp.: A, holotype, anterior end, dorsal view; B, Holotype, posterior abdominal setiger, anterior view; C, Holotype, anterior thoracic mid-neuropodial spine; D &amp; E, Holotype, posterior thoracic mid-neuropodial spines; F, Holotype, abdominal notopodial furcate setae; G, Holotype, posterior neuropodial emergent hooked spine. Scale bars: A = 200 µm, B = 100 µm, C ‒ G = 10 µm." httpUri="https://zenodo.org/record/237221/files/figure.png" pageId="8" pageNumber="191">Figs. 3</figureCitation>
F, 4F), each with slightly unequal length tines and coarse spines along the median borders.
</paragraph>
</subSubSection>
<subSubSection id="ECB7653BFFC3FF9399BBFE51FB10FD09" pageId="8" pageNumber="191" type="discussion">
<paragraph id="A41236B0FFC3FF9399BBFE51FB10FD09" blockId="8.[151,1437,151,1544]" pageId="8" pageNumber="191">
<emphasis id="96D9EAA2FFC3FF9399BBFE51FED4FE0C" bold="true" box="[199,308,476,501]" pageId="8" pageNumber="191">Remarks</emphasis>
. 
<bibRefCitation id="C03C4B41FFC3FF939838FE51FE38FE0D" author="Blake" box="[324,472,476,501]" pageId="8" pageNumber="191" refString="Blake, J. A. (2000) A new genus and species of polychaete worm (Family Orbiniidae) from methane seeps in the Gulf of Mexico, with a review of the systematics and phylogenetic interrelationships of the genera of Orbiniidae. Cahiers de Biologie Marine, 41, 435 - 449." type="journal article" year="2000">Blake (2000)</bibRefCitation>
noted that 
<taxonomicName id="63AD4D33FFC3FF939B25FE51FD2CFE0D" box="[601,716,476,500]" class="Polychaeta" family="Orbiniidae" genus="Leodamas" kingdom="Animalia" pageId="8" pageNumber="191" phylum="Annelida" rank="genus">
<emphasis id="96D9EAA2FFC3FF939B25FE51FD2CFE0D" box="[601,716,476,500]" italics="true" pageId="8" pageNumber="191">Leodamas</emphasis>
</taxonomicName>
should be recognized as a genus separate from 
<taxonomicName id="63AD4D33FFC3FF939D90FE51FAB9FE0D" box="[1260,1369,476,500]" class="Polychaeta" family="Orbiniidae" genus="Scoloplos" kingdom="Animalia" pageId="8" pageNumber="191" phylum="Annelida" rank="genus">
<emphasis id="96D9EAA2FFC3FF939D90FE51FAB9FE0D" box="[1260,1369,476,500]" italics="true" pageId="8" pageNumber="191">Scoloplos</emphasis>
</taxonomicName>
as its thoracic neuropodial spines are thicker than the narrow, pointed spines of 
<taxonomicName id="63AD4D33FFC3FF939A98FD8DFBB1FDE1" box="[996,1105,512,536]" class="Polychaeta" family="Orbiniidae" genus="Scoloplos" kingdom="Animalia" pageId="8" pageNumber="191" phylum="Annelida" rank="genus">
<emphasis id="96D9EAA2FFC3FF939A98FD8DFBB1FDE1" box="[996,1105,512,536]" italics="true" pageId="8" pageNumber="191">Scoloplos</emphasis>
</taxonomicName>
and are not accompanied by numerous capillaries in the setal fascicle (see 
<taxonomicName id="63AD4D33FFC3FF939BD4FDA9FCB3FDC5" box="[680,851,548,572]" class="Polychaeta" family="Orbiniidae" genus="Scoloplos" kingdom="Animalia" pageId="8" pageNumber="191" phylum="Annelida" rank="species" species="cryptosetosa" status="sp. nov.">
<emphasis id="96D9EAA2FFC3FF939BD4FDA9FD56FDC2" box="[680,694,548,571]" italics="true" pageId="8" pageNumber="191">S</emphasis>
. 
<emphasis id="96D9EAA2FFC3FF939BBAFDA8FCB3FDC5" box="[710,851,549,572]" italics="true" pageId="8" pageNumber="191">cryptosetosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8DEA57D9FFC3FF939A20FDA9FC7CFDC5" box="[860,924,548,573]" pageId="8" pageNumber="191" rank="species">
<emphasis id="96D9EAA2FFC3FF939A20FDA9FC75FDC4" bold="true" box="[860,917,548,573]" pageId="8" pageNumber="191">n. sp</emphasis>
.
</taxonomicNameLabel>
below) as occurs in 
<taxonomicName id="63AD4D33FFC3FF939DF7FDA9FB18FDC5" box="[1163,1272,548,572]" class="Polychaeta" family="Orbiniidae" genus="Scoloplos" kingdom="Animalia" pageId="8" pageNumber="191" phylum="Annelida" rank="genus">
<emphasis id="96D9EAA2FFC3FF939DF7FDA9FB18FDC5" box="[1163,1272,548,572]" italics="true" pageId="8" pageNumber="191">Scoloplos</emphasis>
</taxonomicName>
. 
<bibRefCitation id="C03C4B41FFC3FF939C7BFDA9FA7CFDC4" author="Blake" box="[1287,1436,548,573]" pageId="8" pageNumber="191" refString="Blake, J. A. (1996) Family Orbiniidae Hartman, 1942. In: Blake, J. A., Hilbig, B. &amp; Scott, P. H. (Eds.), Taxonomic Atlas of the benthic fauna of the Santa Maria Basin and the Western Santa Barbara Channel, Volume 6. The Annelida, Part 3. Santa Barbara Museum of Natural History, Santa Barbara, California, pp. 1 - 26." type="book chapter" year="1996">Blake (1996)</bibRefCitation>
considered the subgenus 
<taxonomicName id="63AD4D33FFC3FF9398CEFDC5FD4DFD99" authority="(Leodamas)" baseAuthorityName="Leodamas" box="[434,685,584,608]" class="Polychaeta" family="Orbiniidae" genus="Scoloplos" kingdom="Animalia" pageId="8" pageNumber="191" phylum="Annelida" rank="genus">
<emphasis id="96D9EAA2FFC3FF9398CEFDC5FDFFFD99" box="[434,543,584,608]" italics="true" pageId="8" pageNumber="191">Scoloplos</emphasis>
(
<emphasis id="96D9EAA2FFC3FF939B4CFDC5FD43FD99" box="[560,675,584,608]" italics="true" pageId="8" pageNumber="191">Leodamas</emphasis>
)
</taxonomicName>
to differ from 
<taxonomicName id="63AD4D33FFC3FF939A2AFDC5FBABFD99" box="[854,1099,584,608]" class="Polychaeta" family="Orbiniidae" genus="Scoloplos" kingdom="Animalia" pageId="8" pageNumber="191" phylum="Annelida" rank="genus">
<emphasis id="96D9EAA2FFC3FF939A2AFDC5FC23FD99" box="[854,963,584,608]" italics="true" pageId="8" pageNumber="191">Scoloplos</emphasis>
(
<emphasis id="96D9EAA2FFC3FF939AA9FDC5FBA2FD99" box="[981,1090,584,608]" italics="true" pageId="8" pageNumber="191">Scoloplos</emphasis>
)
</taxonomicName>
in that the onset of branchiae occurred anterior to setiger 7 rather than posterior to setiger 8. Later, 
<bibRefCitation id="C03C4B41FFC3FF939AEBFDE1FBCAFD7D" author="Blake" box="[919,1066,620,645]" pageId="8" pageNumber="191" refString="Blake, J. A. (2000) A new genus and species of polychaete worm (Family Orbiniidae) from methane seeps in the Gulf of Mexico, with a review of the systematics and phylogenetic interrelationships of the genera of Orbiniidae. Cahiers de Biologie Marine, 41, 435 - 449." type="journal article" year="2000">Blake (2000)</bibRefCitation>
treated 
<taxonomicName id="63AD4D33FFC3FF939DFFFDE1FB16FD7D" box="[1155,1270,620,644]" class="Polychaeta" family="Orbiniidae" genus="Leodamas" kingdom="Animalia" pageId="8" pageNumber="191" phylum="Annelida" rank="genus">
<emphasis id="96D9EAA2FFC3FF939DFFFDE1FB16FD7D" box="[1155,1270,620,644]" italics="true" pageId="8" pageNumber="191">Leodamas</emphasis>
</taxonomicName>
as a full genus and noted that while most species in this genus have the first branchiae on the anterior part of the thorax at about setiger 6, there is one group of species represented by 
<taxonomicName id="63AD4D33FFC3FF939A74FD38FC78FD35" box="[776,920,692,716]" class="Polychaeta" family="Orbiniidae" genus="Leodamas" kingdom="Animalia" pageId="8" pageNumber="191" phylum="Annelida" rank="species" species="treadwelli">
<emphasis id="96D9EAA2FFC3FF939A74FD38FCF7FD35" box="[776,791,693,716]" italics="true" pageId="8" pageNumber="191">L</emphasis>
. 
<emphasis id="96D9EAA2FFC3FF939A54FD39FC78FD35" box="[808,920,692,716]" italics="true" pageId="8" pageNumber="191">treadwelli</emphasis>
</taxonomicName>
(see below) where the branchiae begin more posteriorly; the generic status of this group of species is currently being reviewed (Blake, in prep.).
</paragraph>
</subSubSection>
<subSubSection id="ECB7653BFFC3FF9399BBFD71FDE4FA81" pageId="8" pageNumber="191" type="reference_group">
<paragraph id="A41236B0FFC3FF9399BBFD71FD2EFB58" blockId="8.[151,1437,151,1544]" pageId="8" pageNumber="191">
<taxonomicName id="63AD4D33FFC3FF9399BBFD71FE42FCED" ID-CoL="6P98K" box="[199,418,764,788]" class="Polychaeta" family="Orbiniidae" genus="Leodamas" kingdom="Animalia" pageId="8" pageNumber="191" phylum="Annelida" rank="species" species="hamatus" status="sp. nov.">
<emphasis id="96D9EAA2FFC3FF9399BBFD71FE42FCED" box="[199,418,764,788]" italics="true" pageId="8" pageNumber="191">Leodamas hamatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8DEA57D9FFC3FF9398D7FD71FE0BFCEC" box="[427,491,764,789]" pageId="8" pageNumber="191" rank="species">
<emphasis id="96D9EAA2FFC3FF9398D7FD71FE0BFCEC" bold="true" box="[427,491,764,789]" pageId="8" pageNumber="191">n. sp.</emphasis>
</taxonomicNameLabel>
is unusual in the structural differences of the neuropodial uncini along the thorax. Blake (in prep.) has noted a similar transition in the neuropodial uncini along the thorax of 
<taxonomicName id="63AD4D33FFC3FF939DCBFCACFF2FFCA5" ID-CoL="67QXC" authority="Ehlers, 1900" authorityName="Ehlers" authorityYear="1900" class="Polychaeta" family="Orbiniidae" genus="Leodamas" kingdom="Animalia" pageId="8" pageNumber="191" phylum="Annelida" rank="species" species="cochleata">
<emphasis id="96D9EAA2FFC3FF939DCBFCACFB26FCC1" box="[1207,1222,801,824]" italics="true" pageId="8" pageNumber="191">L</emphasis>
. 
<emphasis id="96D9EAA2FFC3FF939DABFCADFAA4FCC1" box="[1239,1348,800,824]" italics="true" pageId="8" pageNumber="191">cochleata</emphasis>
<bibRefCitation id="C03C4B41FFC3FF939C31FC92FF2FFCA5" author="Ehlers" pageId="8" pageNumber="191" refString="Ehlers, E. (1900) Ehlers, E. 1900. Magellanische Anneliden gesammelt wahrend der schwedischen Expedition nach den Magellanslander, Nachrichten von der Gesellschaft der Wissenschaften zu Gottingen, Mathematisch-Physikalische Klasse, 206 - 223." type="journal article" year="1900">Ehlers, 1900</bibRefCitation>
</taxonomicName>
from Patagonia. The uncini of the first 2–3 setigers of 
<taxonomicName id="63AD4D33FFC3FF939A3BFCC8FC33FCA5" ID-CoL="67QXC" box="[839,979,836,860]" class="Polychaeta" family="Orbiniidae" genus="Leodamas" kingdom="Animalia" pageId="8" pageNumber="191" phylum="Annelida" rank="species" species="cochleata">
<emphasis id="96D9EAA2FFC3FF939A3BFCC8FCB6FCA5" box="[839,854,837,860]" italics="true" pageId="8" pageNumber="191">L</emphasis>
. 
<emphasis id="96D9EAA2FFC3FF939A1AFCC9FC33FCA5" box="[870,979,836,860]" italics="true" pageId="8" pageNumber="191">cochleata</emphasis>
</taxonomicName>
are straight shafts with a rounded apex, weak transverse ridges, and an elongate groove along one side. In the remaining thoracic setigers, however, the uncini have well-developed transverse ridges and a more expanded distal end with a round, cup-like concavity (see 
<bibRefCitation id="C03C4B41FFC3FF9399EBFC22FEC2FC31" author="Ehlers" box="[151,290,943,968]" pageId="8" pageNumber="191" refString="Ehlers, E. (1901) Die Polychaeten des magellanischen und chilenischen Strandes. Ein faunistischer Versuch, Festschrift zur Feier des Underfunfzigjahrigen Bestehens der koniglichen Gesellschaft der Wissenschaft der Wissenschaften zu Gottingen, pp. 1 - 232, 25 plates." type="book chapter" year="1901">Ehlers 1901</bibRefCitation>
: pl. 21, Figs. 20–21). The thoracic uncini of 
<taxonomicName id="63AD4D33FFC3FF939A66FC3CFC77FC31" ID-CoL="6P98K" box="[794,919,944,968]" class="Polychaeta" family="Orbiniidae" genus="Leodamas" kingdom="Animalia" pageId="8" pageNumber="191" phylum="Annelida" rank="species" species="hamatus" status="sp. nov.">
<emphasis id="96D9EAA2FFC3FF939A66FC3CFCC9FC31" box="[794,809,945,968]" italics="true" pageId="8" pageNumber="191">L</emphasis>
. 
<emphasis id="96D9EAA2FFC3FF939A44FC3DFC77FC31" box="[824,919,944,968]" italics="true" pageId="8" pageNumber="191">hamatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8DEA57D9FFC3FF939AE3FC22FC3EFC31" box="[927,990,943,968]" pageId="8" pageNumber="191" rank="species">
<emphasis id="96D9EAA2FFC3FF939AE3FC22FC3EFC31" bold="true" box="[927,990,943,968]" pageId="8" pageNumber="191">n. sp.</emphasis>
</taxonomicNameLabel>
also exhibit a transition in morphology of the uncini along the thorax but the uncini of anterior setigers are more pointed than those of 
<taxonomicName id="63AD4D33FFC3FF939DC0FC58FAA6FC15" ID-CoL="67QXC" box="[1212,1350,980,1004]" class="Polychaeta" family="Orbiniidae" genus="Leodamas" kingdom="Animalia" pageId="8" pageNumber="191" phylum="Annelida" rank="species" species="cochleata">
<emphasis id="96D9EAA2FFC3FF939DC0FC58FB2BFC15" box="[1212,1227,981,1004]" italics="true" pageId="8" pageNumber="191">L</emphasis>
. 
<emphasis id="96D9EAA2FFC3FF939DA5FC59FAA6FC15" box="[1241,1350,980,1004]" italics="true" pageId="8" pageNumber="191">cochleata</emphasis>
</taxonomicName>
, have a thin, distal hood, and more robust transverse ridges. These thoracic uncini are replaced posteriorly to setiger 5 by heavily ridged, thick uncini with a slightly curved apex and a thickened, oblique hood, sometimes giving the appearance of a biramus end (
<figureCitation id="3C962A35FFC3FF939898FBB2FDCFFBA1" box="[484,559,1087,1112]" captionStart="FIGURE 3" captionStartId="6.[151,250,1831,1853]" captionTargetBox="[205,1384,507,1790]" captionTargetId="figure@6.[189,1397,491,1810]" captionTargetPageId="6" captionText="FIGURE 3. Leodamas hamatus n. sp.: A, holotype, anterior end, dorsal view; B, Holotype, posterior abdominal setiger, anterior view; C, Holotype, anterior thoracic mid-neuropodial spine; D &amp; E, Holotype, posterior thoracic mid-neuropodial spines; F, Holotype, abdominal notopodial furcate setae; G, Holotype, posterior neuropodial emergent hooked spine. Scale bars: A = 200 µm, B = 100 µm, C ‒ G = 10 µm." httpUri="https://zenodo.org/record/237221/files/figure.png" pageId="8" pageNumber="191">Figs. 3</figureCitation>
E, 4D). Also, 
<taxonomicName id="63AD4D33FFC3FF939BB6FBCCFCA6FBA1" ID-CoL="6P98K" box="[714,838,1088,1112]" class="Polychaeta" family="Orbiniidae" genus="Leodamas" kingdom="Animalia" pageId="8" pageNumber="191" phylum="Annelida" rank="species" species="hamatus" status="sp. nov.">
<emphasis id="96D9EAA2FFC3FF939BB6FBCCFD39FBA1" box="[714,729,1089,1112]" italics="true" pageId="8" pageNumber="191">L</emphasis>
. 
<emphasis id="96D9EAA2FFC3FF939B9BFBCDFCA6FBA1" box="[743,838,1088,1112]" italics="true" pageId="8" pageNumber="191">hamatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8DEA57D9FFC3FF939A30FBB2FC6AFBA1" box="[844,906,1087,1112]" pageId="8" pageNumber="191" rank="species">
<emphasis id="96D9EAA2FFC3FF939A30FBB2FC6AFBA1" bold="true" box="[844,906,1087,1112]" pageId="8" pageNumber="191">n. sp.</emphasis>
</taxonomicNameLabel>
has a single, large, curved neuropodial acicular spine in posterior abdominal segments (
<figureCitation id="3C962A35FFC3FF939B1BFBE9FD4FFB85" box="[615,687,1124,1149]" captionStart="FIGURE 4" captionStartId="7.[151,250,1824,1846]" captionTargetBox="[154,1428,201,1801]" captionTargetId="figure@7.[150,1436,193,1803]" captionTargetPageId="7" captionText="FIGURE 4. Leitoscoloplos multipapillatus: A, Setigers 17 – 19, ventral view. Leodamas hamatus n. sp.: B, Paratype XXX Anterior region, lateral view; C, Anterior thoracic spines, setiger three, mid-neuropodial region; D, Posterior thoracic spines, setiger 15, mid-neuropodial region; E, Posterior thoracic spines, upper neuropodial region; F, Furcate notosetae, posterior abdominal region; G, Posterior emergent neuroaciculum, posterior abdominal region. Scale bars: A = 200 µm, B = 500 µm, C ‒ E, F, G = 10 µm. h = hoods, stp = stomach papillae." httpUri="https://zenodo.org/record/237222/files/figure.png" pageId="8" pageNumber="191">
Fig. 
<quantity id="63559B55FFC3FF939BE3FBE9FD4FFB85" box="[671,687,1124,1148]" metricMagnitude="-3" metricUnit="kg" metricValue="4.0" pageId="8" pageNumber="191" unit="g" value="4.0">4</quantity>
</figureCitation>
g). 
<taxonomicName id="63AD4D33FFC3FF939BAAFBE8FC84FB85" ID-CoL="67QXC" box="[726,868,1124,1148]" class="Polychaeta" family="Orbiniidae" genus="Leodamas" kingdom="Animalia" pageId="8" pageNumber="191" phylum="Annelida" rank="species" species="cochleata">
<emphasis id="96D9EAA2FFC3FF939BAAFBE8FD05FB85" box="[726,741,1125,1148]" italics="true" pageId="8" pageNumber="191">L</emphasis>
. 
<emphasis id="96D9EAA2FFC3FF939B8BFBE9FC84FB85" box="[759,868,1124,1148]" italics="true" pageId="8" pageNumber="191">cochleata</emphasis>
</taxonomicName>
possesses 1‒2 weakly curved spines in both the notopodia and neuropodia of abdominal segments.
</paragraph>
<paragraph id="A41236B0FFC3FF9399BBFB21FDE4FA81" blockId="8.[151,1437,151,1544]" pageId="8" pageNumber="191">
The unusual thickened hood in middle and posterior thoracic neuropodial uncini appears similar to the uncini of 
<taxonomicName id="63AD4D33FFC3FF9399C9FB5CFE10FB11" ID-CoL="85RHN" authority="Fauvel, 1902" authorityName="Fauvel" authorityYear="1902" box="[181,496,1231,1256]" class="Polychaeta" family="Orbiniidae" genus="Leodamas" kingdom="Animalia" pageId="8" pageNumber="191" phylum="Annelida" rank="species" species="chevalieri">
<emphasis id="96D9EAA2FFC3FF9399C9FB5CFF24FB11" box="[181,196,1233,1256]" italics="true" pageId="8" pageNumber="191">L</emphasis>
. 
<emphasis id="96D9EAA2FFC3FF9399AFFB5DFEA3FB11" box="[211,323,1232,1256]" italics="true" pageId="8" pageNumber="191">chevalieri</emphasis>
(
<bibRefCitation id="C03C4B41FFC3FF93982EFB42FE07FB11" author="Fauvel" box="[338,487,1231,1256]" pageId="8" pageNumber="191" refString="Fauvel, P. (1902) Annelides Polychetes de la Casamance rapportees par M. Aug Chevalier. Bulletin de la Societe Linneenne de Normandie, 5, 59 - 105." type="journal article" year="1902">Fauvel, 1902</bibRefCitation>
)
</taxonomicName>
(
<bibRefCitation id="C03C4B41FFC3FF939883FB42FD0BFB11" author="Fauvel" box="[511,747,1231,1256]" pageId="8" pageNumber="191" refString="Fauvel, P. (1902) Annelides Polychetes de la Casamance rapportees par M. Aug Chevalier. Bulletin de la Societe Linneenne de Normandie, 5, 59 - 105." type="journal article" year="1902">Fauvel 1902, Fig. 26</bibRefCitation>
) but there is no mention of variability of the uncini along the thorax of that species. 
<taxonomicName id="63AD4D33FFC3FF9398EDFB78FDFEFAF5" ID-CoL="85RHN" box="[401,542,1268,1292]" class="Polychaeta" family="Orbiniidae" genus="Leodamas" kingdom="Animalia" pageId="8" pageNumber="191" phylum="Annelida" rank="species" species="chevalieri">
<emphasis id="96D9EAA2FFC3FF9398EDFB78FE40FAF5" box="[401,416,1269,1292]" italics="true" pageId="8" pageNumber="191">L</emphasis>
. 
<emphasis id="96D9EAA2FFC3FF9398D2FB79FDFEFAF5" box="[430,542,1268,1292]" italics="true" pageId="8" pageNumber="191">chevalieri</emphasis>
</taxonomicName>
is similar to 
<taxonomicName id="63AD4D33FFC3FF939BD2FB78FCCAFAF5" ID-CoL="6P98K" box="[686,810,1268,1292]" class="Polychaeta" family="Orbiniidae" genus="Leodamas" kingdom="Animalia" pageId="8" pageNumber="191" phylum="Annelida" rank="species" species="hamatus" status="sp. nov.">
<emphasis id="96D9EAA2FFC3FF939BD2FB78FD5DFAF5" box="[686,701,1269,1292]" italics="true" pageId="8" pageNumber="191">L</emphasis>
. 
<emphasis id="96D9EAA2FFC3FF939BB7FB79FCCAFAF5" box="[715,810,1268,1292]" italics="true" pageId="8" pageNumber="191">hamatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8DEA57D9FFC3FF939A4CFB79FC8EFAF4" box="[816,878,1268,1293]" pageId="8" pageNumber="191" rank="species">
<emphasis id="96D9EAA2FFC3FF939A4CFB79FC8EFAF4" bold="true" box="[816,878,1268,1293]" pageId="8" pageNumber="191">n. sp.</emphasis>
</taxonomicNameLabel>
based on the length of the thorax, first occurrence of branchiae and morphology of the abdominal parapodia but lacks the large emergent, curved acicular spines in the posterior neuropodia (
<figureCitation id="3C962A35FFC3FF9398BEFAB1FDF2FAAD" box="[450,530,1340,1365]" captionStart="FIGURE 3" captionStartId="6.[151,250,1831,1853]" captionTargetBox="[205,1384,507,1790]" captionTargetId="figure@6.[189,1397,491,1810]" captionTargetPageId="6" captionText="FIGURE 3. Leodamas hamatus n. sp.: A, holotype, anterior end, dorsal view; B, Holotype, posterior abdominal setiger, anterior view; C, Holotype, anterior thoracic mid-neuropodial spine; D &amp; E, Holotype, posterior thoracic mid-neuropodial spines; F, Holotype, abdominal notopodial furcate setae; G, Holotype, posterior neuropodial emergent hooked spine. Scale bars: A = 200 µm, B = 100 µm, C ‒ G = 10 µm." httpUri="https://zenodo.org/record/237221/files/figure.png" pageId="8" pageNumber="191">Figs. 3</figureCitation>
B, G; 4E) as well as the furcate setae present in the posterior notopodia of 
<taxonomicName id="63AD4D33FFC3FF939CF9FAB0FF16FA81" ID-CoL="6P98K" class="Polychaeta" family="Orbiniidae" genus="Leodamas" kingdom="Animalia" pageId="8" pageNumber="191" phylum="Annelida" rank="species" species="hamatus" status="sp. nov.">
<emphasis id="96D9EAA2FFC3FF939CF9FAB0FA74FAAD" box="[1413,1428,1341,1364]" italics="true" pageId="8" pageNumber="191">L</emphasis>
. 
<emphasis id="96D9EAA2FFC3FF9399EBFAEDFF16FA81" box="[151,246,1376,1400]" italics="true" pageId="8" pageNumber="191">hamatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8DEA57D9FFC3FF939980FAD2FEDBFA81" box="[252,315,1375,1400]" pageId="8" pageNumber="191" rank="species">
<emphasis id="96D9EAA2FFC3FF939980FAD2FEDBFA81" bold="true" box="[252,315,1375,1400]" pageId="8" pageNumber="191">n. sp.</emphasis>
</taxonomicNameLabel>
(
<figureCitation id="3C962A35FFC3FF939837FAD2FE78FA81" box="[331,408,1375,1400]" captionStart="FIGURE 3" captionStartId="6.[151,250,1831,1853]" captionTargetBox="[205,1384,507,1790]" captionTargetId="figure@6.[189,1397,491,1810]" captionTargetPageId="6" captionText="FIGURE 3. Leodamas hamatus n. sp.: A, holotype, anterior end, dorsal view; B, Holotype, posterior abdominal setiger, anterior view; C, Holotype, anterior thoracic mid-neuropodial spine; D &amp; E, Holotype, posterior thoracic mid-neuropodial spines; F, Holotype, abdominal notopodial furcate setae; G, Holotype, posterior neuropodial emergent hooked spine. Scale bars: A = 200 µm, B = 100 µm, C ‒ G = 10 µm." httpUri="https://zenodo.org/record/237221/files/figure.png" pageId="8" pageNumber="191">Figs. 3</figureCitation>
F; 4E, F).
</paragraph>
</subSubSection>
<subSubSection id="ECB7653BFFC3FF9399BBFA09FF0FFA39" pageId="8" pageNumber="191" type="etymology">
<paragraph id="A41236B0FFC3FF9399BBFA09FF0FFA39" blockId="8.[151,1437,151,1544]" pageId="8" pageNumber="191">
<emphasis id="96D9EAA2FFC3FF9399BBFA09FEA6FA64" bold="true" box="[199,326,1412,1437]" pageId="8" pageNumber="191">Etymology</emphasis>
. 
<taxonomicName id="63AD4D33FFC3FF939825FA08FE5EFA65" box="[345,446,1413,1436]" class="Polychaeta" family="Orbiniidae" genus="Leodamas" kingdom="Animalia" pageId="8" pageNumber="191" phylum="Annelida" rank="species" species="hamatus" status="sp. nov.">
<emphasis id="96D9EAA2FFC3FF939825FA08FE5EFA65" box="[345,446,1413,1436]" italics="true" pageId="8" pageNumber="191">Hamatus</emphasis>
</taxonomicName>
is Latin for hooked, referring to the large curved neuropodial spines found on this species.
</paragraph>
</subSubSection>
<subSubSection id="ECB7653BFFC3FF9399BBFA41FE04F9F1" pageId="8" pageNumber="191" type="distribution">
<paragraph id="A41236B0FFC3FF9399BBFA41FE04F9F1" blockId="8.[151,1437,151,1544]" pageId="8" pageNumber="191">
<emphasis id="96D9EAA2FFC3FF9399BBFA41FE80FA1C" bold="true" box="[199,352,1484,1509]" pageId="8" pageNumber="191">Distribution.</emphasis>
Collected in muddy to sandy sediments from 
<quantity id="63559B55FFC3FF939A14FA41FC2BFA1D" box="[872,971,1484,1509]" metricMagnitude="1" metricUnit="m" metricValue="1.85" metricValueMax="2.6" metricValueMin="1.1" pageId="8" pageNumber="191" unit="m" value="18.5" valueMax="26.0" valueMin="11.0">11–26 m</quantity>
in the Gulf of Nicoya and from 
<quantity id="63559B55FFC3FF939C45FA41FA7CFA1D" box="[1337,1436,1484,1508]" metricMagnitude="1" metricUnit="m" metricValue="1.25" metricValueMax="1.3" metricValueMin="1.2" pageId="8" pageNumber="191" unit="m" value="12.5" valueMax="13.0" valueMin="12.0">12–13 m</quantity>
in Bahia Culebra, 
<collectingCountry id="DCBA7620FFC3FF93981FFA7DFE00F9F1" box="[355,480,1519,1544]" name="Costa Rica" pageId="8" pageNumber="191">Costa Rica</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>