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<mods:title id="183A88DF9245812253B6CA4D0A02B1F3">The Orbiniidae (Annelida: Polychaeta) of Pacific Costa Rica</mods:title>
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<paragraph id="A41236B0FFC3FF9399EBF9D1FDC6F98F" blockId="8.[151,550,1628,1687]" box="[151,550,1628,1654]" pageId="8" pageNumber="191">
<heading id="FF5A81DCFFC3FF9399EBF9D1FDC6F98F" bold="true" box="[151,550,1628,1654]" fontSize="11" level="1" pageId="8" pageNumber="191" reason="1">
<taxonomicName id="63AD4D33FFC3FF9399EBF9D1FDC6F98F" ID-CoL="3T6BP" authority="Eisig, 1914" authorityName="Eisig" authorityYear="1914" box="[151,550,1628,1654]" class="Polychaeta" family="Orbiniidae" genus="Leodamas" kingdom="Animalia" pageId="8" pageNumber="191" phylum="Annelida" rank="species" species="treadwelli">
<emphasis id="96D9EAA2FFC3FF9399EBF9D1FDC6F98F" bold="true" box="[151,550,1628,1654]" pageId="8" pageNumber="191">
<emphasis id="96D9EAA2FFC3FF9399EBF9D1FE73F98F" bold="true" box="[151,403,1628,1654]" italics="true" pageId="8" pageNumber="191">Leodamas treadwelli</emphasis>
<bibRefCitation id="C03C4B41FFC3FF9398E0F9D1FDC6F98F" author="Eisig" box="[412,550,1628,1654]" pageId="8" pageNumber="191" refString="Eisig, H. (1914) Zur Systematik, Anatomie und Morphologie der Ariciiden nebst Beitragen zur generellen Systematik. Mitteilungen aus der Zoologischen Station zu Neapel, 21 (6), 153 - 600." type="journal article" year="1914">Eisig, 1914</bibRefCitation>
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</paragraph>
<paragraph id="A41236B0FFC3FF9399EBF9F3FE7AF96E" blockId="8.[151,550,1628,1687]" box="[151,410,1662,1687]" pageId="8" pageNumber="191">
<figureCitation id="3C962A35FFC3FF9399EBF9F3FEE1F96E" box="[151,257,1662,1687]" captionStart="FIGURE 5" captionStartId="9.[151,250,1545,1567]" captionTargetBox="[173,1414,311,1516]" captionTargetId="figure@9.[160,1426,301,1524]" captionTargetPageId="9" captionText="FIGURE 5. Leodamas treadwelli: A, Station 28 Oct, Gulf of Nicoya specimen, anterior and midbody, lateral view; B, Station 28 Jun, Gulf of Nicoya specimen, notopodial furcate seta. Naineris setosa: C, Playa Blanca specimen, anterior end, dorsal view; D, Playa Blanca specimen, thoracic parapodia, posterior view; E, Playa Blanca specimen, anterior abdominal parapodia, posterior view; F, Playa Blanca specimen, posterior abdominal parapodia, posterior view; G, Playa Blanca specimen, notopodial furcate seta. Scoloplos cryptospinigerus n. sp.: H, Holotype, anterior end, dorsal view; I, Holotype, posterior thoracic neuropodial spines. Scale bars: A, D ‒ E, H = 500 µm, B, G, I = 10 µm, C, F = 1000 µm. s = neuropodial spines." httpUri="https://zenodo.org/record/237223/files/figure.png" pageId="8" pageNumber="191">Figures 5</figureCitation>
(A–B), 6 (C)
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<paragraph id="A41236B0FFC3FF9399EBF94AFC25F925" blockId="8.[151,1436,1734,1879]" box="[151,965,1734,1757]" pageId="8" pageNumber="191">
<treatmentCitationGroup id="84BD119EFFC3FF9399EBF94AFC25F925" box="[151,965,1734,1757]" pageId="8" pageNumber="191">
<taxonomicName id="63AD4D33FFC3FF9399EBF94AFDE9F925" ID-CoL="84X79" authority="Treadwell, 1901: 201" authorityName="Treadwell" authorityPageNumber="201" authorityYear="1901" box="[151,521,1734,1757]" class="Polychaeta" family="Orbiniidae" genus="Aricia" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="191" phylum="Annelida" rank="species" species="cirrata">
<emphasis id="96D9EAA2FFC3FF9399EBF94AFEC3F924" box="[151,291,1735,1757]" italics="true" pageId="8" pageNumber="191">Aricia cirrata</emphasis>
<treatmentCitation id="250C10A1FFC3FF939850F94BFDE9F925" author="Treadwell" box="[300,521,1734,1756]" page="201" pageId="8" pageNumber="191" year="1901">
<bibRefCitation id="C03C4B41FFC3FF939850F94BFE36F925" author="Treadwell" box="[300,470,1734,1756]" pageId="8" pageNumber="191" refString="Treadwell, A. L. (1901) The polychaetous annelids of Porto Rico. Bulletin of the United States Fisheries Commision, 20, 181 - 210." type="journal article" year="1901">Treadwell, 1901</bibRefCitation>
: 201
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, Figs. 54–57. Homonym, Not 
<bibRefCitation id="C03C4B41FFC3FF939A3FF94BFC25F925" author="Ehlers" box="[835,965,1734,1756]" pageId="8" pageNumber="191" refString="Ehlers, E. (1897) Polychaeten. Hamburger Magalhaensische Sammelreise, 1897, 1 - 148." type="journal article" year="1897">Ehlers, 1897</bibRefCitation>
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</paragraph>
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<treatmentCitationGroup id="84BD119EFFC3FF9399EBF968FDCEF8C1" pageId="8" pageNumber="191">
<taxonomicName id="63AD4D33FFC3FF9399EBF968FE18F902" ID-CoL="4VSM4" authority="Eisig, 1914" authorityName="Eisig" authorityYear="1914" box="[151,504,1765,1787]" class="Polychaeta" family="Orbiniidae" genus="Scoloplos" kingdom="Animalia" pageId="8" pageNumber="191" phylum="Annelida" rank="species" species="treadwelli">
<emphasis id="96D9EAA2FFC3FF9399EBF968FE8EF902" box="[151,366,1765,1787]" italics="true" pageId="8" pageNumber="191">Scoloplos treadwelli</emphasis>
<bibRefCitation id="C03C4B41FFC3FF939801F968FE18F902" author="Eisig" box="[381,504,1765,1787]" pageId="8" pageNumber="191" refString="Eisig, H. (1914) Zur Systematik, Anatomie und Morphologie der Ariciiden nebst Beitragen zur generellen Systematik. Mitteilungen aus der Zoologischen Station zu Neapel, 21 (6), 153 - 600." type="journal article" year="1914">Eisig, 1914</bibRefCitation>
</taxonomicName>
(new name for 
<taxonomicName id="63AD4D33FFC3FF939BC8F96BFC73F902" ID-CoL="84X79" authority="Treadwell" authorityName="Treadwell" box="[692,915,1765,1787]" class="Polychaeta" family="Orbiniidae" genus="Aricia" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="191" phylum="Annelida" rank="species" species="cirrata">
<emphasis id="96D9EAA2FFC3FF939BC8F96BFCFCF902" box="[692,796,1766,1787]" italics="true" pageId="8" pageNumber="191">A. cirrata</emphasis>
Treadwell
</taxonomicName>
).— 
<treatmentCitation id="250C10A1FFC3FF939ACBF968FB61F902" author="Augener" box="[951,1153,1765,1787]" page="69" pageId="8" pageNumber="191" year="1927">
<bibRefCitation id="C03C4B41FFC3FF939ACBF968FBB9F902" author="Augener" box="[951,1113,1765,1787]" pageId="8" pageNumber="191" refString="Augener, H. (1927) Polychaeten von Curacao. Bijdragen tot de Dierkunde, 25, 39 - 82." type="journal article" year="1927">Augener, 1927</bibRefCitation>
: 69
</treatmentCitation>
.— 
<treatmentCitation id="250C10A1FFC3FF939DE3F968FA9DF902" author="Hartman" box="[1183,1405,1765,1787]" page="283" pageId="8" pageNumber="191" year="1957">
<bibRefCitation id="C03C4B41FFC3FF939DE3F968FAA6F902" author="Hartman" box="[1183,1350,1765,1787]" pageId="8" pageNumber="191" refString="Hartman, O. (1957) Orbiniidae, Apistobranchidae, Paraonidae and Longosomidae. Allan Hancock Pacific Expeditions, 15, 211 - 393." type="journal article" year="1957">Hartman, 1957</bibRefCitation>
: 283
</treatmentCitation>
.— 
<treatmentCitation id="250C10A1FFC3FF9399BBF889FDFDF8E3" author="Maciolek" box="[199,541,1796,1818]" page="164" pageId="8" pageNumber="191" year="1978">
<bibRefCitation id="C03C4B41FFC3FF9399BBF889FE0AF8E3" author="Maciolek" box="[199,490,1796,1818]" pageId="8" pageNumber="191" refString="Maciolek, N. J. &amp; Holland, J. S. (1978) Scoloplos texana: a new orbiniid polychaete from South Texas, with notes on the related species Scoloplos treadwelli Eisig. Contributions in Marine Science, 21, 161 - 169." type="journal article" year="1978">
Maciolek &amp; 
<collectingCountry id="DCBA7620FFC3FF93982DF889FE45F8E3" box="[337,421,1796,1818]" name="Netherlands" pageId="8" pageNumber="191">Holland</collectingCountry>
, 1978
</bibRefCitation>
: 164
</treatmentCitation>
.— 
<treatmentCitation id="250C10A1FFC3FF939B47F889FCD9F8E3" author="Vargas" box="[571,825,1796,1818]" page="336" pageId="8" pageNumber="191" year="1985">
<bibRefCitation id="C03C4B41FFC3FF939B47F889FCE5F8E3" author="Vargas" box="[571,773,1796,1818]" pageId="8" pageNumber="191" refString="Vargas, J. A., Dean, H. K., Maurer, D. &amp; Orellana, P. (1985) Lista preliminar de invertebrados asociadoa a los sedimentos del Golfo de Nicoya, Costa Rica. Brenesia, 24, 327 - 341." type="journal article" year="1985">
Vargas 
<emphasis id="96D9EAA2FFC3FF939BF3F888FD26F8E3" box="[655,710,1796,1818]" italics="true" pageId="8" pageNumber="191">et al.</emphasis>
1985
</bibRefCitation>
: 336
</treatmentCitation>
.— 
<treatmentCitation id="250C10A1FFC3FF939A2AF889FBAEF8E3" author="Maurer" box="[854,1102,1796,1818]" page="47" pageId="8" pageNumber="191" year="1988">
<bibRefCitation id="C03C4B41FFC3FF939A2AF889FBC6F8E3" author="Maurer" box="[854,1062,1796,1818]" pageId="8" pageNumber="191" refString="Maurer, D., Vargas, J. &amp; Dean, H. (1988) Polychaetous Annelids from the Gulf of Nicoya, Costa Rica. Internationale Revue der Gesamten Hydrobiologie, 73 (1), 43 - 59. http: // dx. doi. org / 10.1002 / iroh. 19880730105" type="journal article" year="1988">
Maurer 
<emphasis id="96D9EAA2FFC3FF939AD2F888FC05F8E3" box="[942,997,1796,1818]" italics="true" pageId="8" pageNumber="191">et al.</emphasis>
1988
</bibRefCitation>
: 47
</treatmentCitation>
.— 
<treatmentCitation id="250C10A1FFC3FF939D10F889FAFEF8E3" author="Dean" box="[1132,1310,1796,1818]" page="74" pageId="8" pageNumber="191" year="1996">
<bibRefCitation id="C03C4B41FFC3FF939D10F889FB16F8E3" author="Dean" box="[1132,1270,1796,1818]" pageId="8" pageNumber="191" refString="Dean, H. K. (1996 a) Subtidal benthic polychaetes (Annelida) of the Gulf of Nicoya, Costa Rica. Revista Biologia Tropical, 44 (Supplement 3), 69 - 80." type="journal article" year="1996" yearSuffix="a">Dean, 1996a</bibRefCitation>
: 74
</treatmentCitation>
.—de León- 
<treatmentCitation id="250C10A1FFC3FF9399BBF8AFFDCAF8C1" author="Gonzalez" box="[199,554,1826,1848]" page="173" pageId="8" pageNumber="191" year="1996">
<bibRefCitation id="C03C4B41FFC3FF9399BBF8AFFE16F8C1" author="Gonzalez" box="[199,502,1826,1848]" pageId="8" pageNumber="191" refString="de Leon-Gonzalez, J. A. &amp; Rodriguez, J. A. (1996) Orbiniidae (Polychaeta) from soft bottom of the western coast of Baja California Peninsula, Mexico. Bulletin of Marine Science, 59, 169 - 174." type="journal article" year="1996">González &amp; Rodriguez, 1996</bibRefCitation>
: 173
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="A41236B0FFC3FF9399EBF8CCFCDBF8AE" blockId="8.[151,1436,1734,1879]" box="[151,827,1857,1879]" pageId="8" pageNumber="191">
<taxonomicName id="63AD4D33FFC3FF9399EBF8CCFDE2F8AE" ID-CoL="6Y9J7" authority="Dean, 1996: 74" authorityName="Dean" authorityPageNumber="74" authorityYear="1996" box="[151,514,1857,1879]" class="Polychaeta" family="Orbiniidae" genus="Scoloplos" kingdom="Animalia" pageId="8" pageNumber="191" phylum="Annelida" rank="species" species="armiger">
<emphasis id="96D9EAA2FFC3FF9399EBF8CCFEB3F8AE" box="[151,339,1857,1879]" italics="true" pageId="8" pageNumber="191">Scoloplos armiger</emphasis>
: Dean, 1996: 74
</taxonomicName>
; 2009: 146. Not 
<bibRefCitation id="C03C4B41FFC3FF939BD3F8CCFCD7F8AE" author="Muller" box="[687,823,1857,1879]" pageId="8" pageNumber="191" refString="Muller, O. F. (1776) Zoologiae Danicae Prodomus, seu Animalium Daniae et Norvegiae indigenarum. Characteres, nomina, et synonyma imprimis popularum. Havniae, typis Hallageriis, 274 pp." type="book" year="1776">Müller, 1776</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="ECB7653BFFC3FF9299EBF809FEF5FF01" lastPageId="9" lastPageNumber="192" pageId="8" pageNumber="191" type="materials_examined">
<paragraph id="A41236B0FFC3FF9299EBF809FEF5FF01" blockId="8.[151,1437,1924,2021]" lastBlockId="9.[151,1436,151,248]" lastPageId="9" lastPageNumber="192" pageId="8" pageNumber="191">
<emphasis id="96D9EAA2FFC3FF9399EBF809FE9CF864" bold="true" box="[151,380,1924,1949]" pageId="8" pageNumber="191">Material examined</emphasis>
. Gulf of Nicoya: Sta. 15, 9°57′40″N, 84°47′'00″W, 
<quantity id="63559B55FFC3FF939AACF809FBEDF865" box="[976,1037,1924,1949]" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="8" pageNumber="191" unit="m" value="15.0">15 m</quantity>
, sandy mud, 
<date id="D0131070FFC3FF939DD8F809FACAF865" box="[1188,1322,1924,1949]" pageId="8" pageNumber="191" value="1980-07-12">12 Jul 1980</date>
(1). STA. 24, 9°49′25″N, 84°41′20″W, 
<quantity id="63559B55FFC3FF939897F825FDC8F839" box="[491,552,1960,1984]" metricMagnitude="1" metricUnit="m" metricValue="1.1" pageId="8" pageNumber="191" unit="m" value="11.0">11 m</quantity>
, sand, 
<date id="D0131070FFC3FF939B01F825FD1FF839" box="[637,767,1960,1984]" pageId="8" pageNumber="191" value="1980-10-01">1 Oct 1980</date>
(1); Sta. 28, 9°52′16″N, 
<geoCoordinate id="C1995077FFC3FF939D54F825FB54F838" box="[1064,1204,1960,1985]" direction="west" orientation="longitude" pageId="8" pageNumber="191" precision="15" value="-84.75833">84°45′30&quot;W</geoCoordinate>
, 
<quantity id="63559B55FFC3FF939DB9F825FAE3F839" box="[1221,1283,1960,1985]" metricMagnitude="1" metricUnit="m" metricValue="2.6" pageId="8" pageNumber="191" unit="m" value="26.0">26 m</quantity>
, mud, 
<date id="D0131070FFC3FF939C29F825FF32F81D" pageId="8" pageNumber="191" value="1980-07-10">10 Jul 1980</date>
, 1 (
<collectionCode id="C2BCAE75FFC3FF939983F841FE89F81D" box="[255,361,1996,2020]" pageId="8" pageNumber="191">MZUCR</collectionCode>
), 
<date id="D0131070FFC3FF9398FCF841FDE2F81D" box="[384,514,1996,2020]" pageId="8" pageNumber="191" value="1980-10-01">1 Oct 1980</date>
(1), 
<date id="D0131070FFC3FF939B41F840FD2BF81D" box="[573,715,1996,2020]" pageId="8" pageNumber="191" value="1981-06-07">7 June 1981</date>
, 1 (
<collectionCode id="C2BCAE75FFC3FF939B85F841FC83F81D" box="[761,867,1996,2020]" pageId="8" pageNumber="191">MZUCR</collectionCode>
); Sta. 29, 9°54′55″N, 84°45′15″W, 
<quantity id="63559B55FFC3FF939C7EF841FAA0F81D" box="[1282,1344,1996,2020]" metricMagnitude="1" metricUnit="m" metricValue="1.8" pageId="8" pageNumber="191" unit="m" value="18.0">18 m</quantity>
, muddy sand, 
<date id="D0131070FFC2FF9299A5FF15FE81FF49" box="[217,353,152,176]" pageId="9" pageNumber="192" value="1981-01-29">29 Jan 1981</date>
(1); 
<date id="D0131070FFC2FF9298E4FF15FDFBFF49" box="[408,539,152,176]" pageId="9" pageNumber="192" value="1982-04-04">4 Apr 1982</date>
, 1 (
<collectionCode id="C2BCAE75FFC2FF929B38FF15FD4EFF49" box="[580,686,152,176]" pageId="9" pageNumber="192">MZUCR</collectionCode>
); Sta. 30, 9°54′40″N, 84°45′50″W, 
<quantity id="63559B55FFC2FF929D3CFF15FB9BFF56" box="[1088,1147,152,176]" metricMagnitude="1" metricUnit="m" metricValue="1.8" pageId="9" pageNumber="192" unit="m" value="18.0">18 m</quantity>
, muddy sand, 
<date id="D0131070FFC2FF929C5CFF15FA7CFF49" box="[1312,1436,152,176]" pageId="9" pageNumber="192" value="1980-10-01">1 Oct 1980</date>
(1). Punta Morales, Playa Blanca, lower intertidal, 10°3′47.2″ N, 84°57′9.2″ W, muddy sand, 
<date id="D0131070FFC2FF929D9AFF31FA9FFF2C" box="[1254,1407,188,213]" pageId="9" pageNumber="192" value="1986-12-20">20 Dec 1986</date>
, 3 (
<collectionCode id="C2BCAE75FFC2FF9299E2FF6DFEE9FF01" box="[158,265,224,248]" pageId="9" pageNumber="192">MZUCR</collectionCode>
).
</paragraph>
</subSubSection>
<caption id="F0D26638FFC2FF9299EBF984FF3DF92E" httpUri="https://zenodo.org/record/237223/files/figure.png" pageId="9" pageNumber="192" targetBox="[173,1414,311,1516]" targetPageId="9">
<paragraph id="A41236B0FFC2FF9299EBF984FF3DF92E" blockId="9.[151,1436,1545,1751]" pageId="9" pageNumber="192">
<emphasis id="96D9EAA2FFC2FF9299EBF984FEF4F9E6" bold="true" box="[151,276,1545,1567]" pageId="9" pageNumber="192">FIGURE 5.</emphasis>
<taxonomicName id="63AD4D33FFC2FF929866F984FE10F9E6" box="[282,496,1545,1567]" class="Polychaeta" family="Orbiniidae" genus="Leodamas" kingdom="Animalia" pageId="9" pageNumber="192" phylum="Annelida" rank="species" species="treadwelli">
<emphasis id="96D9EAA2FFC2FF929866F984FE10F9E6" box="[282,496,1545,1567]" italics="true" pageId="9" pageNumber="192">Leodamas treadwelli</emphasis>
</taxonomicName>
: A, Station 28 Oct, Gulf of Nicoya specimen, anterior and midbody, lateral view; B, Station 28 Jun, Gulf of Nicoya specimen, notopodial furcate seta. 
<taxonomicName id="63AD4D33FFC2FF929B82F9A4FC7EF9C7" box="[766,926,1577,1598]" class="Polychaeta" family="Orbiniidae" genus="Naineris" kingdom="Animalia" pageId="9" pageNumber="192" phylum="Annelida" rank="species" species="setosa">
<emphasis id="96D9EAA2FFC2FF929B82F9A4FC7EF9C7" box="[766,926,1577,1598]" italics="true" pageId="9" pageNumber="192">Naineris setosa</emphasis>
</taxonomicName>
: C, Playa Blanca specimen, anterior end, dorsal view; D, Playa Blanca specimen, thoracic parapodia, posterior view; E, Playa Blanca specimen, anterior abdominal parapodia, posterior view; F, Playa Blanca specimen, posterior abdominal parapodia, posterior view; G, Playa Blanca specimen, notopodial furcate seta. 
<taxonomicName id="63AD4D33FFC2FF929B7AF909FCFEF963" box="[518,798,1668,1690]" class="Polychaeta" family="Orbiniidae" genus="Scoloplos" kingdom="Animalia" pageId="9" pageNumber="192" phylum="Annelida" rank="species" species="cryptospinigerus" status="sp. nov.">
<emphasis id="96D9EAA2FFC2FF929B7AF909FCFEF963" box="[518,798,1668,1690]" italics="true" pageId="9" pageNumber="192">Scoloplos cryptospinigerus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8DEA57D9FFC2FF929A57F908FC86F963" box="[811,870,1669,1690]" pageId="9" pageNumber="192" rank="species">
<emphasis id="96D9EAA2FFC2FF929A57F908FC86F963" bold="true" box="[811,870,1669,1690]" pageId="9" pageNumber="192">n. sp.</emphasis>
</taxonomicNameLabel>
: H, Holotype, anterior end, dorsal view; I, Holotype, posterior thoracic neuropodial spines. Scale bars: A, D‒E, H = 500 µm, B, G, I = 10 µm, C, F= 1000 µm. 
<emphasis id="96D9EAA2FFC2FF929D86F929FAE4F940" box="[1274,1284,1700,1721]" italics="true" pageId="9" pageNumber="192">s</emphasis>
= neuropodial spines.
</paragraph>
</caption>
<subSubSection id="ECB7653BFFC2FF9199BBF973FE6EFE71" lastPageId="10" lastPageNumber="193" pageId="9" pageNumber="192" type="description">
<paragraph id="A41236B0FFC2FF9299BBF973FCBFF851" blockId="9.[151,1436,1790,2032]" pageId="9" pageNumber="192">
<emphasis id="96D9EAA2FFC2FF9299BBF973FEB9F8EE" bold="true" box="[199,345,1790,1815]" pageId="9" pageNumber="192">Description.</emphasis>
All specimens incomplete, maximum thoracic width 0.66–1.75 mm (
<figureCitation id="3C962A35FFC2FF929D20F973FB7EF8EE" box="[1116,1182,1790,1815]" captionStart="FIGURE 5" captionStartId="9.[151,250,1545,1567]" captionTargetBox="[173,1414,311,1516]" captionTargetId="figure@9.[160,1426,301,1524]" captionTargetPageId="9" captionText="FIGURE 5. Leodamas treadwelli: A, Station 28 Oct, Gulf of Nicoya specimen, anterior and midbody, lateral view; B, Station 28 Jun, Gulf of Nicoya specimen, notopodial furcate seta. Naineris setosa: C, Playa Blanca specimen, anterior end, dorsal view; D, Playa Blanca specimen, thoracic parapodia, posterior view; E, Playa Blanca specimen, anterior abdominal parapodia, posterior view; F, Playa Blanca specimen, posterior abdominal parapodia, posterior view; G, Playa Blanca specimen, notopodial furcate seta. Scoloplos cryptospinigerus n. sp.: H, Holotype, anterior end, dorsal view; I, Holotype, posterior thoracic neuropodial spines. Scale bars: A, D ‒ E, H = 500 µm, B, G, I = 10 µm, C, F = 1000 µm. s = neuropodial spines." httpUri="https://zenodo.org/record/237223/files/figure.png" pageId="9" pageNumber="192">Fig. 5</figureCitation>
A). Prostomium small, triangular, pointed; lacking eyespots; nuchal organs not apparent. Peristomium subequal to prostomium, approximately 1.25 times as long and equal in width to setiger 1. Thorax wide, dorso-ventrally flattened; 14–20 setigers long, transition to abdomen abrupt. Branchiae from first abdominal setiger in all specimens, small at first, more erect, longer than parapodial lobes, in posterior segments.
</paragraph>
<paragraph id="A41236B0FFC2FF9199BBF83EFE32FF2C" blockId="9.[151,1436,1790,2032]" lastBlockId="10.[151,1437,151,392]" lastPageId="10" lastPageNumber="193" pageId="9" pageNumber="192">
Thoracic notopodia with short digitate postsetal lobe, approaching setal length in posterior thoracic setigers; thoracic neuropodia low ridges. Abdominal notopodial lobe erect, postsetal lobe large, triangular, longer than branchiae in anterior setigers, shorter in subsequent setigers; neuropodial parapodial lobe erect, truncate; presetal lobe a weak ridge (
<figureCitation id="3C962A35FFC1FF919811FF31FE50FF2C" box="[365,432,188,213]" captionStart="FIGURE 6" captionStartId="10.[151,250,1716,1738]" captionTargetBox="[219,1361,452,1682]" captionTargetId="figure@10.[212,1374,445,1695]" captionTargetPageId="10" captionText="FIGURE 6. Leitoscoloplos pugettensis: A, Station 9, Bahia Culebra specimen, thoracic setiger, anterior view; B, Station 9, Bahia Culebra specimen, posterior notopodial flail setae. Leodamas treadwelli: C, Station 28 Jun, Gulf of Nicoya specimen, posterior abdominal setiger, anterior view; Scoloplos cryptospinigerus n. sp.: D, Holotype, anterior end, dorsal view; E, Holotype, thoracic parapodia, posterior view; F, Paratype, abdominal parapodia, anterior view; G, Paraytpe, thoracic neurosetal spines. Scale bars: A, F = 50 µm, B = 10 µm, C, E = 100 µm, D = 500 µm, G = 20 µm." httpUri="https://zenodo.org/record/237224/files/figure.png" pageId="10" pageNumber="193">Fig. 6</figureCitation>
C).
</paragraph>
<paragraph id="A41236B0FFC1FF9199BBFF52FE6EFE71" blockId="10.[151,1437,151,392]" pageId="10" pageNumber="193">
Thoracic notosetae camerated capillaries; thoracic neurosetae short, acicular, in two rows anteriorly, single row in posterior thoracic setigers (
<figureCitation id="3C962A35FFC1FF919B62FE89FD82FEE4" box="[542,610,260,285]" captionStart="FIGURE 5" captionStartId="9.[151,250,1545,1567]" captionTargetBox="[173,1414,311,1516]" captionTargetId="figure@9.[160,1426,301,1524]" captionTargetPageId="9" captionText="FIGURE 5. Leodamas treadwelli: A, Station 28 Oct, Gulf of Nicoya specimen, anterior and midbody, lateral view; B, Station 28 Jun, Gulf of Nicoya specimen, notopodial furcate seta. Naineris setosa: C, Playa Blanca specimen, anterior end, dorsal view; D, Playa Blanca specimen, thoracic parapodia, posterior view; E, Playa Blanca specimen, anterior abdominal parapodia, posterior view; F, Playa Blanca specimen, posterior abdominal parapodia, posterior view; G, Playa Blanca specimen, notopodial furcate seta. Scoloplos cryptospinigerus n. sp.: H, Holotype, anterior end, dorsal view; I, Holotype, posterior thoracic neuropodial spines. Scale bars: A, D ‒ E, H = 500 µm, B, G, I = 10 µm, C, F = 1000 µm. s = neuropodial spines." httpUri="https://zenodo.org/record/237223/files/figure.png" pageId="10" pageNumber="193">Fig. 5</figureCitation>
A); thoracic acicula accompanied by a few dorsal capillaries in posterior thoracic setigers. Abdominal notosetae camerated capillaries accompanied by a single furcate setae with unequal smooth tines with blunt tips; delicate thin teeth on inner faces of each tine (
<figureCitation id="3C962A35FFC1FF919D76FEC1FBB0FE9C" box="[1034,1104,332,357]" captionStart="FIGURE 5" captionStartId="9.[151,250,1545,1567]" captionTargetBox="[173,1414,311,1516]" captionTargetId="figure@9.[160,1426,301,1524]" captionTargetPageId="9" captionText="FIGURE 5. Leodamas treadwelli: A, Station 28 Oct, Gulf of Nicoya specimen, anterior and midbody, lateral view; B, Station 28 Jun, Gulf of Nicoya specimen, notopodial furcate seta. Naineris setosa: C, Playa Blanca specimen, anterior end, dorsal view; D, Playa Blanca specimen, thoracic parapodia, posterior view; E, Playa Blanca specimen, anterior abdominal parapodia, posterior view; F, Playa Blanca specimen, posterior abdominal parapodia, posterior view; G, Playa Blanca specimen, notopodial furcate seta. Scoloplos cryptospinigerus n. sp.: H, Holotype, anterior end, dorsal view; I, Holotype, posterior thoracic neuropodial spines. Scale bars: A, D ‒ E, H = 500 µm, B, G, I = 10 µm, C, F = 1000 µm. s = neuropodial spines." httpUri="https://zenodo.org/record/237223/files/figure.png" pageId="10" pageNumber="193">Fig. 5</figureCitation>
B); abdominal neurosetae all camerated capillaries.
</paragraph>
</subSubSection>
<caption id="F0D26638FFC1FF9199EBF939FB9CF8BC" httpUri="https://zenodo.org/record/237224/files/figure.png" pageId="10" pageNumber="193" targetBox="[219,1361,452,1682]" targetPageId="10">
<paragraph id="A41236B0FFC1FF9199EBF939FB9CF8BC" blockId="10.[151,1436,1716,1861]" pageId="10" pageNumber="193">
<emphasis id="96D9EAA2FFC1FF9199EBF939FEF5F930" bold="true" box="[151,277,1716,1738]" pageId="10" pageNumber="193">FIGURE 6.</emphasis>
<taxonomicName id="63AD4D33FFC1FF919861F939FDCEF933" box="[285,558,1716,1738]" class="Polychaeta" family="Orbiniidae" genus="Leitoscoloplos" kingdom="Animalia" pageId="10" pageNumber="193" phylum="Annelida" rank="species" species="pugettensis">
<emphasis id="96D9EAA2FFC1FF919861F939FDCEF933" box="[285,558,1716,1738]" italics="true" pageId="10" pageNumber="193">Leitoscoloplos pugettensis</emphasis>
</taxonomicName>
: A, Station 9, Bahia Culebra specimen, thoracic setiger, anterior view; B, Station 9, Bahia Culebra specimen, posterior notopodial flail setae. 
<taxonomicName id="63AD4D33FFC1FF919B96F95EFC21F910" box="[746,961,1747,1769]" class="Polychaeta" family="Orbiniidae" genus="Leodamas" kingdom="Animalia" pageId="10" pageNumber="193" phylum="Annelida" rank="species" species="treadwelli">
<emphasis id="96D9EAA2FFC1FF919B96F95EFC21F910" box="[746,961,1747,1769]" italics="true" pageId="10" pageNumber="193">Leodamas treadwelli</emphasis>
</taxonomicName>
: C, Station 28 Jun, Gulf of Nicoya specimen, posterior abdominal setiger, anterior view; 
<taxonomicName id="63AD4D33FFC1FF919B18F97FFC9CF8F1" box="[612,892,1778,1800]" class="Polychaeta" family="Orbiniidae" genus="Scoloplos" kingdom="Animalia" pageId="10" pageNumber="193" phylum="Annelida" rank="species" species="cryptospinigerus" status="sp. nov.">
<emphasis id="96D9EAA2FFC1FF919B18F97FFC9CF8F1" box="[612,892,1778,1800]" italics="true" pageId="10" pageNumber="193">Scoloplos cryptospinigerus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8DEA57D9FFC1FF919AF7F97FFC28F8FE" box="[907,968,1778,1799]" pageId="10" pageNumber="193" rank="species">
<emphasis id="96D9EAA2FFC1FF919AF7F97FFC28F8FE" bold="true" box="[907,968,1778,1799]" pageId="10" pageNumber="193">n. sp.</emphasis>
</taxonomicNameLabel>
: D, Holotype, anterior end, dorsal view; E, Holotype, thoracic parapodia, posterior view; F, Paratype, abdominal parapodia, anterior view; G, Paraytpe, thoracic neurosetal spines. Scale bars: A, F = 50 µm, B= 10 µm, C, E = 100 µm, D = 500 µm, G = 20 µm.
</paragraph>
</caption>
<subSubSection id="ECB7653BFFC1FF9099BBF8FCFE55FDE1" lastPageId="11" lastPageNumber="194" pageId="10" pageNumber="193" type="discussion">
<paragraph id="A41236B0FFC1FF9099BBF8FCFE55FDE1" blockId="10.[151,1436,1905,2003]" lastBlockId="11.[151,1436,151,752]" lastPageId="11" lastPageNumber="194" pageId="10" pageNumber="193">
<emphasis id="96D9EAA2FFC1FF9199BBF8FCFED4F873" bold="true" box="[199,308,1905,1930]" pageId="10" pageNumber="193">Remarks</emphasis>
. 
<taxonomicName id="63AD4D33FFC1FF91983FF8FFFD41F873" authority="Treadwell, 1901" authorityName="Treadwell" authorityYear="1901" box="[323,673,1905,1930]" class="Polychaeta" family="Orbiniidae" genus="Aricia" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="193" phylum="Annelida" rank="species" species="cirrata">
<emphasis id="96D9EAA2FFC1FF91983FF8FFFE3FF870" box="[323,479,1906,1929]" italics="true" pageId="10" pageNumber="193">Aricia cirrata</emphasis>
<bibRefCitation id="C03C4B41FFC1FF91989BF8FCFD41F873" author="Treadwell" box="[487,673,1905,1930]" pageId="10" pageNumber="193" refString="Treadwell, A. L. (1901) The polychaetous annelids of Porto Rico. Bulletin of the United States Fisheries Commision, 20, 181 - 210." type="journal article" year="1901">Treadwell, 1901</bibRefCitation>
</taxonomicName>
, originally described from 
<collectingCountry id="DCBA7620FFC1FF919AA9F8FFFBBEF873" box="[981,1118,1905,1930]" name="Puerto Rico" pageId="10" pageNumber="193">Puerto Rico</collectingCountry>
, is a homonym of 
<taxonomicName id="63AD4D33FFC1FF919C53F8FFFECAF857" authority="Ehlers, 1897" authorityName="Ehlers" authorityYear="1897" class="Polychaeta" family="Orbiniidae" genus="Aricia" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="193" phylum="Annelida" rank="species" species="cirrata">
<emphasis id="96D9EAA2FFC1FF919C53F8FFFA7CF870" box="[1327,1436,1906,1929]" italics="true" pageId="10" pageNumber="193">A. cirrata</emphasis>
<bibRefCitation id="C03C4B41FFC1FF9199EBF818FECAF857" author="Ehlers" box="[151,298,1941,1966]" pageId="10" pageNumber="193" refString="Ehlers, E. (1897) Polychaeten. Hamburger Magalhaensische Sammelreise, 1897, 1 - 148." type="journal article" year="1897">Ehlers, 1897</bibRefCitation>
</taxonomicName>
and was renamed 
<taxonomicName id="63AD4D33FFC1FF919B71F81BFD19F857" box="[525,761,1942,1966]" class="Polychaeta" family="Orbiniidae" genus="Scoloplos" kingdom="Animalia" pageId="10" pageNumber="193" phylum="Annelida" rank="species" species="treadwelli">
<emphasis id="96D9EAA2FFC1FF919B71F81BFD19F857" box="[525,761,1942,1966]" italics="true" pageId="10" pageNumber="193">Scoloplos treadwelli</emphasis>
</taxonomicName>
by 
<bibRefCitation id="C03C4B41FFC1FF919A50F818FC52F857" author="Eisig" box="[812,946,1941,1966]" pageId="10" pageNumber="193" refString="Eisig, H. (1914) Zur Systematik, Anatomie und Morphologie der Ariciiden nebst Beitragen zur generellen Systematik. Mitteilungen aus der Zoologischen Station zu Neapel, 21 (6), 153 - 600." type="journal article" year="1914">Eisig, 1914</bibRefCitation>
. This species has now been more widely reported from the Caribbean, and the Pacific coast of 
<collectingCountry id="DCBA7620FFC1FF919B8DF837FCAAF82A" box="[753,842,1978,2003]" name="Mexico" pageId="10" pageNumber="193">Mexico</collectingCountry>
; the species was redescribed by 
<bibRefCitation id="C03C4B41FFC1FF919DCBF836FA8CF82A" author="Hartman" box="[1207,1388,1978,2003]" pageId="10" pageNumber="193" refString="Hartman, O. (1957) Orbiniidae, Apistobranchidae, Paraonidae and Longosomidae. Allan Hancock Pacific Expeditions, 15, 211 - 393." type="journal article" year="1957">Hartman (1957)</bibRefCitation>
and 
<bibRefCitation id="C03C4B41FFC0FF9099EBFF1AFE00FF49" author="Maciolek" box="[151,480,151,176]" pageId="11" pageNumber="194" refString="Maciolek, N. J. &amp; Holland, J. S. (1978) Scoloplos texana: a new orbiniid polychaete from South Texas, with notes on the related species Scoloplos treadwelli Eisig. Contributions in Marine Science, 21, 161 - 169." type="journal article" year="1978">
Maciolek &amp; 
<collectingCountry id="DCBA7620FFC0FF909850FF1AFE66FF49" box="[300,390,151,176]" name="Netherlands" pageId="11" pageNumber="194">Holland</collectingCountry>
(1978)
</bibRefCitation>
. 
<bibRefCitation id="C03C4B41FFC0FF909892FF15FD36FF49" box="[494,726,152,176]" pageId="11" pageNumber="194" refString="Lopez, E., Cladera, P. &amp; San Martin, G. (2003) Two new species of the genus Leodamas (Orbiniidae: Scolecida: Polychaeta) from the Pacific coast of Panama. Journal of the Marine Biological Association of the United Kingdom, 83, 367 - 374. http: // dx. doi. org / 10.1017 / S 0025315403007215 h" type="journal article">
López, 
<emphasis id="96D9EAA2FFC0FF909B3BFF14FD61FF49" box="[583,641,152,176]" italics="true" pageId="11" pageNumber="194">et al.</emphasis>
(2003)
</bibRefCitation>
described 
<taxonomicName id="63AD4D33FFC0FF909A24FF14FB05FF49" authority="Lopez, Claders" authorityName="Lopez, Claders" box="[856,1253,151,176]" class="Polychaeta" family="Orbiniidae" genus="Leodamas" kingdom="Animalia" pageId="11" pageNumber="194" phylum="Annelida" rank="species" species="platythoracicus">
<emphasis id="96D9EAA2FFC0FF909A24FF14FC87FF49" box="[856,871,153,176]" italics="true" pageId="11" pageNumber="194">L</emphasis>
. 
<emphasis id="96D9EAA2FFC0FF909A05FF15FBC7FF49" box="[889,1063,152,176]" italics="true" pageId="11" pageNumber="194">platythoracicus</emphasis>
López, Claders
</taxonomicName>
, &amp; 
<bibRefCitation id="C03C4B41FFC0FF909C69FF15FF32FF2D" author="San" pageId="11" pageNumber="194" refString="Lopez, E., Cladera, P. &amp; San Martin, G. (2003) Two new species of the genus Leodamas (Orbiniidae: Scolecida: Polychaeta) from the Pacific coast of Panama. Journal of the Marine Biological Association of the United Kingdom, 83, 367 - 374. http: // dx. doi. org / 10.1017 / S 0025315403007215 h" type="journal article" year="2003">San Martín, 2003</bibRefCitation>
, from Pacific 
<collectingCountry id="DCBA7620FFC0FF909802FF30FE37FF2D" box="[382,471,189,212]" name="Panama" pageId="11" pageNumber="194">Panama</collectingCountry>
which is similar to 
<taxonomicName id="63AD4D33FFC0FF909BB7FF30FCBDFF2D" box="[715,861,188,212]" class="Polychaeta" family="Orbiniidae" genus="Scoloplos" kingdom="Animalia" pageId="11" pageNumber="194" phylum="Annelida" rank="species" species="treadwelli">
<emphasis id="96D9EAA2FFC0FF909BB7FF30FD3AFF2D" box="[715,730,189,212]" italics="true" pageId="11" pageNumber="194">L</emphasis>
. 
<emphasis id="96D9EAA2FFC0FF909B91FF31FCBDFF2D" box="[749,861,188,212]" italics="true" pageId="11" pageNumber="194">treadwelli</emphasis>
</taxonomicName>
in the thoracic length and setal morphology but differs in the nature of the abdominal neuropodia. The anterior abdominal neuropodia is erect and blunt-ended in both species but those of 
<taxonomicName id="63AD4D33FFC0FF9098B8FE88FD72FEE5" box="[452,658,260,284]" class="Polychaeta" family="Orbiniidae" genus="Leodamas" kingdom="Animalia" pageId="11" pageNumber="194" phylum="Annelida" rank="species" species="platythoracicus">
<emphasis id="96D9EAA2FFC0FF9098B8FE88FE33FEE5" box="[452,467,261,284]" italics="true" pageId="11" pageNumber="194">L</emphasis>
. 
<emphasis id="96D9EAA2FFC0FF909898FE89FD72FEE5" box="[484,658,260,284]" italics="true" pageId="11" pageNumber="194">platythoracicus</emphasis>
</taxonomicName>
have a notch on the lateral edge and a single, slightly protruding, aciculum. 
<taxonomicName id="63AD4D33FFC0FF909873FEA4FE40FEB9" box="[271,416,296,320]" class="Polychaeta" family="Orbiniidae" genus="Scoloplos" kingdom="Animalia" pageId="11" pageNumber="194" phylum="Annelida" rank="species" species="treadwelli">
<emphasis id="96D9EAA2FFC0FF909873FEA4FE40FEB9" box="[271,416,296,320]" italics="true" pageId="11" pageNumber="194">L. treadwelli</emphasis>
</taxonomicName>
lacks a notch in the anterior neuropodial lobe and the slightly projecting aciculum is accompanied by 2‒3 deeply embedded aciculae. 
<taxonomicName id="63AD4D33FFC0FF909BB9FEC0FCCFFE9D" box="[709,815,333,356]" class="Polychaeta" family="Orbiniidae" genus="Leodamas" kingdom="Animalia" pageId="11" pageNumber="194" phylum="Annelida" rank="species" species="texana">
<emphasis id="96D9EAA2FFC0FF909BB9FEC0FCCFFE9D" box="[709,815,333,356]" italics="true" pageId="11" pageNumber="194">L. texana</emphasis>
</taxonomicName>
(
<bibRefCitation id="C03C4B41FFC0FF909A41FEC1FB92FE9D" author="Maciolek" box="[829,1138,332,357]" pageId="11" pageNumber="194" refString="Maciolek, N. J. &amp; Holland, J. S. (1978) Scoloplos texana: a new orbiniid polychaete from South Texas, with notes on the related species Scoloplos treadwelli Eisig. Contributions in Marine Science, 21, 161 - 169." type="journal article" year="1978">
Maciolek &amp; 
<collectingCountry id="DCBA7620FFC0FF909AB3FEC1FBCCFE9C" box="[975,1068,332,357]" name="Netherlands" pageId="11" pageNumber="194">Holland</collectingCountry>
, 1978
</bibRefCitation>
) from the Gulf of 
<collectingCountry id="DCBA7620FFC0FF909C3AFEC1FA7BFE9C" box="[1350,1435,332,357]" name="Mexico" pageId="11" pageNumber="194">Mexico</collectingCountry>
is also similar to 
<taxonomicName id="63AD4D33FFC0FF909822FEFCFE0EFE71" box="[350,494,368,392]" class="Polychaeta" family="Orbiniidae" genus="Scoloplos" kingdom="Animalia" pageId="11" pageNumber="194" phylum="Annelida" rank="species" species="treadwelli">
<emphasis id="96D9EAA2FFC0FF909822FEFCFE0EFE71" box="[350,494,368,392]" italics="true" pageId="11" pageNumber="194">L. treadwelli</emphasis>
</taxonomicName>
but has only single rows of acicular spines in the thoracic neuropodia whereas 
<taxonomicName id="63AD4D33FFC0FF909CF9FEFCFEE8FE55" class="Polychaeta" family="Orbiniidae" genus="Scoloplos" kingdom="Animalia" pageId="11" pageNumber="194" phylum="Annelida" rank="species" species="treadwelli">
<emphasis id="96D9EAA2FFC0FF909CF9FEFCFA74FE71" box="[1413,1428,369,392]" italics="true" pageId="11" pageNumber="194">L</emphasis>
. 
<emphasis id="96D9EAA2FFC0FF9099EBFE19FEE8FE55" box="[151,264,404,428]" italics="true" pageId="11" pageNumber="194">treadwelli</emphasis>
</taxonomicName>
has several double rows of spines on anterior thoracic neuropodia and single rows in posterior thoracic segments. 
<taxonomicName id="63AD4D33FFC0FF909872FE34FCFDFE29" authority="Lopez, Cladera" authorityName="Lopez, Cladera" authorityYear="2003" box="[270,797,439,464]" class="Polychaeta" family="Orbiniidae" genus="Leodamas" kingdom="Animalia" pageId="11" pageNumber="194" phylum="Annelida" rank="species" species="minutis">
<emphasis id="96D9EAA2FFC0FF909872FE34FEFDFE29" box="[270,285,441,464]" italics="true" pageId="11" pageNumber="194">L</emphasis>
. 
<emphasis id="96D9EAA2FFC0FF909850FE35FE9FFE36" box="[300,383,440,463]" italics="true" pageId="11" pageNumber="194">minutis</emphasis>
<bibRefCitation id="C03C4B41FFC0FF9098FAFE35FCFDFE29" author="Lopez" box="[390,797,439,464]" pageId="11" pageNumber="194" refString="Lopez, E., Cladera, P. &amp; San Martin, G. (2003) Two new species of the genus Leodamas (Orbiniidae: Scolecida: Polychaeta) from the Pacific coast of Panama. Journal of the Marine Biological Association of the United Kingdom, 83, 367 - 374. http: // dx. doi. org / 10.1017 / S 0025315403007215 h" type="journal article" year="2003">López, Cladera &amp; San Martin, 2003</bibRefCitation>
</taxonomicName>
, also described from Pacific 
<collectingCountry id="DCBA7620FFC0FF909D1EFE35FB5EFE36" box="[1122,1214,440,463]" name="Panama" pageId="11" pageNumber="194">Panama</collectingCountry>
, is morphologically similar to 
<taxonomicName id="63AD4D33FFC0FF909872FE50FE7EFE0D" box="[270,414,476,500]" class="Polychaeta" family="Orbiniidae" genus="Scoloplos" kingdom="Animalia" pageId="11" pageNumber="194" phylum="Annelida" rank="species" species="treadwelli">
<emphasis id="96D9EAA2FFC0FF909872FE50FE7EFE0D" box="[270,414,476,500]" italics="true" pageId="11" pageNumber="194">L. treadwelli</emphasis>
</taxonomicName>
but is unusual in that the first three thoracic setigers are uniramous with notopodia and notosetae entirely absent.
</paragraph>
</subSubSection>
<subSubSection id="ECB7653BFFC0FF9099BBFDA9FA78FD09" pageId="11" pageNumber="194" type="distribution">
<paragraph id="A41236B0FFC0FF9099BBFDA9FA78FD09" blockId="11.[151,1436,151,752]" pageId="11" pageNumber="194">
<emphasis id="96D9EAA2FFC0FF9099BBFDA9FE80FDC4" bold="true" box="[199,352,548,573]" pageId="11" pageNumber="194">Distribution.</emphasis>
<taxonomicName id="63AD4D33FFC0FF909811FDA9FDBCFDC5" box="[365,604,548,572]" class="Polychaeta" family="Orbiniidae" genus="Leodamas" kingdom="Animalia" pageId="11" pageNumber="194" phylum="Annelida" rank="species" species="treadwelli">
<emphasis id="96D9EAA2FFC0FF909811FDA9FDBCFDC5" box="[365,604,548,572]" italics="true" pageId="11" pageNumber="194">Leodamas treadwelli</emphasis>
</taxonomicName>
was described by 
<bibRefCitation id="C03C4B41FFC0FF909A41FDA9FBE4FDC5" author="Treadwell" box="[829,1028,548,573]" pageId="11" pageNumber="194" refString="Treadwell, A. L. (1901) The polychaetous annelids of Porto Rico. Bulletin of the United States Fisheries Commision, 20, 181 - 210." type="journal article" year="1901">Treadwell (1901)</bibRefCitation>
as 
<taxonomicName id="63AD4D33FFC0FF909D4DFDA9FA77FDC5" authority="Treadwell, 1901" authorityName="Treadwell" authorityYear="1901" box="[1073,1431,548,573]" class="Polychaeta" family="Orbiniidae" genus="Aricia" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="194" phylum="Annelida" rank="species" species="cirrata">
<emphasis id="96D9EAA2FFC0FF909D4DFDA9FB2EFDC5" box="[1073,1230,548,572]" italics="true" pageId="11" pageNumber="194">Aricia cirrata</emphasis>
<bibRefCitation id="C03C4B41FFC0FF909DA5FDA9FA77FDC5" author="Treadwell" box="[1241,1431,548,573]" pageId="11" pageNumber="194" refString="Treadwell, A. L. (1901) The polychaetous annelids of Porto Rico. Bulletin of the United States Fisheries Commision, 20, 181 - 210." type="journal article" year="1901">Treadwell, 1901</bibRefCitation>
</taxonomicName>
, from 
<collectingCountry id="DCBA7620FFC0FF9099ABFDC5FEBFFD99" box="[215,351,583,608]" name="Puerto Rico" pageId="11" pageNumber="194">Puerto Rico</collectingCountry>
and has been subsequently been reported from 
<collectingCountry id="DCBA7620FFC0FF909AF6FDCAFC1BFD99" box="[906,1019,583,608]" name="Colombia" pageId="11" pageNumber="194">Colombia</collectingCountry>
, 
<collectingCountry id="DCBA7620FFC0FF909D76FDCAFBA8FD99" box="[1034,1096,583,608]" name="Cuba" pageId="11" pageNumber="194">Cuba</collectingCountry>
, 
<collectingCountry id="DCBA7620FFC0FF909D2AFDC5FB56FD99" box="[1110,1206,584,608]" name="Curaçao" pageId="11" pageNumber="194">Curaçao</collectingCountry>
, and 
<collectingCountry id="DCBA7620FFC0FF909D8BFDCAFAB1FD99" box="[1271,1361,583,608]" name="Jamaica" pageId="11" pageNumber="194">Jamaica</collectingCountry>
in the Caribbean (Dean, 2012). 
<bibRefCitation id="C03C4B41FFC0FF9098CAFDE0FD8BFD7C" author="Hartman" box="[438,619,620,645]" pageId="11" pageNumber="194" refString="Hartman, O. (1957) Orbiniidae, Apistobranchidae, Paraonidae and Longosomidae. Allan Hancock Pacific Expeditions, 15, 211 - 393." type="journal article" year="1957">Hartman (1957)</bibRefCitation>
reported this species from Acapulco on the Pacific coast of 
<collectingCountry id="DCBA7620FFC0FF909C6AFDE1FA8CFD7C" box="[1302,1388,620,645]" name="Mexico" pageId="11" pageNumber="194">Mexico</collectingCountry>
and 
<bibRefCitation id="C03C4B41FFC0FF9099EBFD02FE33FD51" box="[151,467,655,680]" pageId="11" pageNumber="194" refString="Maciolek, N. J. &amp; Holland, J. S. (1978) Scoloplos texana: a new orbiniid polychaete from South Texas, with notes on the related species Scoloplos treadwelli Eisig. Contributions in Marine Science, 21, 161 - 169." type="journal article">
Maciolek &amp; 
<collectingCountry id="DCBA7620FFC0FF90985AFD02FE60FD51" box="[294,384,655,680]" name="Netherlands" pageId="11" pageNumber="194">Holland</collectingCountry>
(1978)
</bibRefCitation>
later verified this identification. De 
<bibRefCitation id="C03C4B41FFC0FF909A0EFD02FAEDFD51" author="Leon-Gonzalez" box="[882,1293,655,680]" pageId="11" pageNumber="194" refString="de Leon-Gonzalez, J. A. &amp; Rodriguez, J. A. (1996) Orbiniidae (Polychaeta) from soft bottom of the western coast of Baja California Peninsula, Mexico. Bulletin of Marine Science, 59, 169 - 174." type="journal article" year="1996">León-González &amp; Rodriguez (1996)</bibRefCitation>
reported the species from Baja California, 
<collectingCountry id="DCBA7620FFC0FF909886FD39FDB3FD34" box="[506,595,692,717]" name="Mexico" pageId="11" pageNumber="194">Mexico</collectingCountry>
. The present records extend the distribution south to 
<collectingCountry id="DCBA7620FFC0FF909DB3FD39FAAEFD34" box="[1231,1358,692,717]" name="Costa Rica" pageId="11" pageNumber="194">Costa Rica</collectingCountry>
in the Central American eastern Pacific. Collected in muddy to sandy sediments from 
<quantity id="63559B55FFC0FF909D50FD5AFB73FD16" box="[1068,1171,727,752]" metricMagnitude="1" metricUnit="m" metricValue="1.85" metricValueMax="2.6" metricValueMin="1.1" pageId="11" pageNumber="194" unit="m" value="18.5" valueMax="26.0" valueMin="11.0">
11‒ 
<specimenCount id="B2ABFD39FFC0FF909D28FD5AFB73FD16" box="[1108,1171,727,752]" pageId="11" pageNumber="194" type="generic">26 m</specimenCount>
</quantity>
. in the Gulf of Nicoya.
</paragraph>
</subSubSection>
</treatment>
</document>