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<mods:title id="40507E18AEF85AFA4038594B92785F0C">The lineages of Stylocellidae (Arachnida: Opiliones: Cyphophthalmi)</mods:title>
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<treatment id="296DA64C707C9962DEAAFEDDA94BFCE2" ID-DOI="http://doi.org/10.5281/zenodo.5866108" ID-GBIF-Taxon="119557605" ID-Zenodo-Dep="5866108" LSID="urn:lsid:plazi:treatment:296DA64C707C9962DEAAFEDDA94BFCE2" httpUri="http://treatment.plazi.org/id/296DA64C707C9962DEAAFEDDA94BFCE2" lastPageNumber="28" pageId="27" pageNumber="28">
<subSubSection id="E9DE44D1707C9962DEAAFEDDAEFFFE3E" box="[151,723,367,393]" pageId="27" pageNumber="28" type="nomenclature">
<paragraph id="A17B175A707C9962DEAAFEDDAEFFFE3E" blockId="27.[151,723,367,393]" box="[151,723,367,393]" pageId="27" pageNumber="28">
<heading id="FA33A036707C9962DEAAFEDDAEFFFE3E" bold="true" box="[151,723,367,393]" fontSize="11" level="1" pageId="27" pageNumber="28" reason="1">
<emphasis id="93B0CB48707C9962DEAAFEDDAEFFFE3E" bold="true" box="[151,723,367,393]" pageId="27" pageNumber="28">
<taxonomicName id="66C46CD9707C9962DEAAFEDDAE6AFE3E" ID-CoL="7WXFB" authority="Shear, 1979" authorityName="Shear" authorityYear="1979" box="[151,582,367,393]" class="Arachnida" family="Stylocellidae" genus="Leptopsalis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="sedgwicki" status="comb. nov.">
<emphasis id="93B0CB48707C9962DEAAFEDDADB0FE3E" bold="true" box="[151,412,367,393]" italics="true" pageId="27" pageNumber="28">Leptopsalis sedgwicki</emphasis>
(
<bibRefCitation id="C5556AAB707C9962DF96FEDDAE12FE3E" author="Shear" box="[427,574,367,393]" firstAuthor="Shear" pageId="27" pageNumber="28" pagination="356" refId="ref17107" refString="Shear, W. A. (1979) Slylocellus sedgwicki n. sp., from Penang Island, Malaysia (Opiliones, Cyphophthalmida, Stylocellidae). Bulletin of the British Arachnological Society, 4, 356 - 360." type="journal article" year="1979">Shear, 1979</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="88837633707C9962DC73FEDDAEFFFE3E" box="[590,723,367,393]" pageId="27" pageNumber="28">new comb.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="E9DE44D1707C9962DEAAFE05AEE6FE7B" box="[151,714,438,461]" pageId="27" pageNumber="28" type="reference_group">
<paragraph id="A17B175A707C9962DEAAFE05AEE6FE7B" blockId="27.[151,714,438,461]" box="[151,714,438,461]" pageId="27" pageNumber="28">
<treatmentCitationGroup id="81D43074707C9962DEAAFE05AEE6FE7B" box="[151,714,438,461]" pageId="27" pageNumber="28">
<taxonomicName id="66C46CD9707C9962DEAAFE05AE78FE7B" ID-CoL="7Z54Y" authority="Shear, 1979: 359" authorityName="Shear" authorityPageNumber="359" authorityYear="1979" box="[151,596,438,461]" class="Arachnida" family="Stylocellidae" genus="Stylocellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="sedgwicki">
<emphasis id="93B0CB48707C9962DEAAFE05AD42FE7A" box="[151,366,439,461]" italics="true" pageId="27" pageNumber="28">Stylocellus sedgwicki</emphasis>
<bibRefCitation id="C5556AAB707C9962DF49FE04AE78FE7B" author="Shear" box="[372,596,438,460]" firstAuthor="Shear" page="359 - 360" pageId="27" pageNumber="28" pagination="356" refId="ref17107" refString="Shear, W. A. (1979) Slylocellus sedgwicki n. sp., from Penang Island, Malaysia (Opiliones, Cyphophthalmida, Stylocellidae). Bulletin of the British Arachnological Society, 4, 356 - 360." type="journal article" year="1979">
<treatmentCitation id="2065314B707C9962DF49FE04AE0CFE7B" author="Shear" box="[372,544,438,460]" page="359" pageId="27" pageNumber="28" year="1979">Shear, 1979: 359</treatmentCitation>
360
</bibRefCitation>
</taxonomicName>
, figs. 111.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="E9DE44D1707C9962DEAAFE4AAD14FC86" pageId="27" pageNumber="28" type="discussion">
<paragraph id="A17B175A707C9962DEAAFE4AAD14FC86" blockId="27.[151,1436,504,854]" pageId="27" pageNumber="28">
<emphasis id="93B0CB48707C9962DEAAFE4AAD0DFDA6" bold="true" box="[151,289,504,529]" pageId="27" pageNumber="28">Comments:</emphasis>
The lack of an anal gland pore or Ramblas organ, compressed ventral prosomal complex (including the lack of a sternum), and large first ventral cheliceral process place this species in Leptopsalinae
<emphasis id="93B0CB48707C9962DAC1FDAFA9B9FD82" bold="true" box="[1276,1429,541,566]" pageId="27" pageNumber="28">subfam. nov.</emphasis>
, and most likely in
<taxonomicName id="66C46CD9707C9962DF51FDF3ADC1FDEE" box="[364,493,577,601]" class="Arachnida" family="Stylocellidae" genus="Leptopsalis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="27" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis id="93B0CB48707C9962DF51FDF3ADC1FDEE" box="[364,493,577,601]" italics="true" pageId="27" pageNumber="28">Leptopsalis</emphasis>
</taxonomicName>
. The ozophores are long and thick, with a straight anterior edge; with the posterior slightly lowered they can appear perpendicular to the long body axis (as in
<bibRefCitation id="C5556AAB707C9962DDC8FDD7A8B7FDCA" author="Shear's" box="[1013,1179,613,637]" firstAuthor="Shear's" pageId="27" pageNumber="28" pagination="174" refId="ref17193" refString="Shear, W. A. (1993) New species in the opilionid genus Stylocellus from Malaysia, Indonesia and the Philippines (Opiliones, Cyphophthalmi, Stylocellidae). Bulletin of the British Arachnological Society, 9, 174 - 188." type="journal article" year="1993">Shears (1993)</bibRefCitation>
original drawing), but they are not entirely shaped like those of
<taxonomicName id="66C46CD9707C9962DC50FD3BAEC9FD16" box="[621,741,649,673]" class="Arachnida" family="Stylocellidae" genus="Stylocellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="27" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis id="93B0CB48707C9962DC50FD3BAEC9FD16" box="[621,741,649,673]" italics="true" pageId="27" pageNumber="28">Stylocellus</emphasis>
</taxonomicName>
nor as perpendicular as those of Fangensinae
<emphasis id="93B0CB48707C9962DACFFD3BA9A6FD15" bold="true" box="[1266,1418,649,674]" pageId="27" pageNumber="28">subfam. nov.</emphasis>
I have also analyzed a collection of juveniles by G. Cuccodoro and I. Löbl from the same area as the
<typeStatus id="7E7FA9F8707C9962DB32FD1CA913FD71" box="[1295,1343,686,710]" pageId="27" pageNumber="28">type</typeStatus>
locality (Penang Island [=
<collectingCountry id="D9D357CA707C9962DF5EFD63AD8FFD5E" box="[355,419,721,745]" name="Iceland" pageId="27" pageNumber="28">Pulau</collectingCountry>
Pinang]) that are consistent with this species in size (too large, it seems, to be
<taxonomicName id="66C46CD9707C9962DB3AFD60A9BFFD5E" box="[1287,1427,721,745]" class="Arachnida" family="Stylocellidae" genus="Miopsalis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="pulicaria">
<emphasis id="93B0CB48707C9962DB3AFD60A9BFFD5E" box="[1287,1427,721,745]" italics="true" pageId="27" pageNumber="28">M. pulicaria</emphasis>
</taxonomicName>
) and shape, and which are consistently recovered as
<taxonomicName id="66C46CD9707C9962DCE9FD47AF79FCBA" box="[724,853,757,781]" class="Arachnida" family="Stylocellidae" genus="Leptopsalis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="27" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis id="93B0CB48707C9962DCE9FD47AF79FCBA" box="[724,853,757,781]" italics="true" pageId="27" pageNumber="28">Leptopsalis</emphasis>
</taxonomicName>
in molecular phylogenies (Peninsula sp. 24,” MCZ DNA
<date id="D57A319A707C9962DEE8FCABAD05FC86" box="[213,297,793,817]" pageId="27" pageNumber="28" value="92-10-14">101492</date>
).
</paragraph>
</subSubSection>
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<paragraph id="A17B175A707C9962DEFAFC8FA94BFCE2" blockId="27.[151,1436,504,854]" box="[199,1383,829,854]" pageId="27" pageNumber="28">
<emphasis id="93B0CB48707C9962DEFAFC8FAD9EFCE1" bold="true" box="[199,434,829,854]" pageId="27" pageNumber="28">Material examined:</emphasis>
<materialsCitation id="11AC1D07707C9962DF84FC8FA94FFCE2" ID-GBIF-Occurrence="3443618310" box="[441,1379,829,854]" collectingDate="1976-06-07" collectionCode="AMNH" collectorName="W. Sedgwick" country="Malaysia" location="Penang Island" pageId="27" pageNumber="28" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="7E7FA9F8707C9962DF84FC8FAE0CFCE2" box="[441,544,829,853]" pageId="27" pageNumber="28" type="holotype">
<emphasis id="93B0CB48707C9962DF84FC8FAE0CFCE2" box="[441,544,829,853]" italics="true" pageId="27" pageNumber="28">Holotype</emphasis>
</typeStatus>
,
<specimenCount id="B7C2DCD3707C9962DC13FC8FAE4BFCE2" box="[558,615,829,853]" pageId="27" pageNumber="28" type="male">Male</specimenCount>
(
<collectionCode id="C7D58F9F707C9962DC4BFC8FAEFEFCE2" LSID="urn:lsid:biocol.org:col:34925" box="[630,722,829,853]" collectionName="USA, New York, New York, American Museum of Natural History" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="27" pageNumber="28">AMNH</collectionCode>
),
<collectingCountry id="D9D357CA707C9962DCD8FC8FAF62FCE2" box="[741,846,829,853]" name="Malaysia" pageId="27" pageNumber="28">Malaysia</collectingCountry>
,
<location id="A41B4181707C9962DD64FC8CAFD6FCE2" LSID="urn:lsid:plazi:treatment:296DA64C707C9962DEAAFEDDA94BFCE2:A41B4181707C9962DD64FC8CAFD6FCE2" box="[857,1018,829,854]" country="Malaysia" name="Penang Island" pageId="27" pageNumber="28">Penang Island</location>
,
<date id="D57A319A707C9962DA38FC8CA8A3FCE2" box="[1029,1167,829,853]" pageId="27" pageNumber="28" value="1976-06-07">
<collectingDate id="C53EC872707C9962DA38FC8CA8A3FCE2" box="[1029,1167,829,853]" pageId="27" pageNumber="28" value="1976-06-07">7 June 1976</collectingDate>
</date>
, col.
<collectorName id="0C31728C707C9962DAF7FC8CA94FFCE2" box="[1226,1379,829,853]" pageId="27" pageNumber="28">W. Sedgwick</collectorName>
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
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