156 lines
17 KiB
XML
156 lines
17 KiB
XML
<document id="C7A7368D0631DDDDFA3982F594A4D8D4" ID-DOI="10.11646/zootaxa.3595.1.1" ID-GBIF-Dataset="6633d51b-c6e2-4664-a883-1972f94dce87" ID-ISSN="1175-5326" ID-Zenodo-Dep="246218" ID-ZooBank="0E34F9DE-B76C-4197-94D0-5A08A1F7C534" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460565569157" checkinUser="plazi" docAuthor="Clouse, Ronald M." docDate="2012" docId="296DA64C707A9967DEAAF89AAF26FD7B" docLanguage="en" docName="zt03595p034.pdf" docOrigin="Zootaxa 3595 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.3595.1.1" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Miopsalis globosa Schwendinger & Giribet 2004, new comb." docType="treatment" docVersion="14" lastPageNumber="31" masterDocId="D554DE3470679979DE3DFFB2AC2CFFB7" masterDocTitle="The lineages of Stylocellidae (Arachnida: Opiliones: Cyphophthalmi)" masterLastPageNumber="34" masterPageNumber="1" pageNumber="30" updateTime="1698320962157" updateUser="plazi">
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<mods:titleInfo id="990C76215A52061E43B0FD83D329C96B">
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<mods:title id="C0AD8B666130C48A99239027419C79E7">The lineages of Stylocellidae (Arachnida: Opiliones: Cyphophthalmi)</mods:title>
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<mods:namePart id="1CF089D21628748C52F34B0C24AE2541">Clouse, Ronald M.</mods:namePart>
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<mods:typeOfResource id="11732AD8CBD3BC15C529136F7C8BD9A3">text</mods:typeOfResource>
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<mods:title id="220C47A12F5941590F5AE97FBED5371D">Zootaxa</mods:title>
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<mods:part id="AAA4916B798CBCBC249B6F917EB88D1F">
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<mods:date id="09C78321E844C5963B47C0E741BB76D6">2012</mods:date>
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<mods:number id="1C24BDAB4F2ECD10F7AEA59FB4EE7F95">2012-12-21</mods:number>
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<mods:number id="F49BBB87DE388810C99C684405D3A18D">3595</mods:number>
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<mods:identifier id="D2E7654112F34044544466AA03FA3A9C" type="DOI">10.11646/zootaxa.3595.1.1</mods:identifier>
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<treatment id="296DA64C707A9967DEAAF89AAF26FD7B" ID-DOI="http://doi.org/10.5281/zenodo.5866114" ID-GBIF-Taxon="119557609" ID-Zenodo-Dep="5866114" LSID="urn:lsid:plazi:treatment:296DA64C707A9967DEAAF89AAF26FD7B" httpUri="http://treatment.plazi.org/id/296DA64C707A9967DEAAF89AAF26FD7B" lastPageId="30" lastPageNumber="31" pageId="29" pageNumber="30">
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<subSubSection id="E9DE44D1707A9964DEAAF89AAFB8F8F5" box="[151,916,1832,1858]" pageId="29" pageNumber="30" type="nomenclature">
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<paragraph id="A17B175A707A9964DEAAF89AAFB8F8F5" blockId="29.[151,916,1832,1858]" box="[151,916,1832,1858]" pageId="29" pageNumber="30">
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<heading id="FA33A036707A9964DEAAF89AAFB8F8F5" bold="true" box="[151,916,1832,1858]" fontSize="11" level="1" pageId="29" pageNumber="30" reason="1">
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<emphasis id="93B0CB48707A9964DEAAF89AAFB8F8F5" bold="true" box="[151,916,1832,1858]" pageId="29" pageNumber="30">
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<taxonomicName id="66C46CD9707A9964DEAAF89AAF24F8F5" ID-CoL="7XBX6" authority="Schwendinger & Giribet, 2004" authorityName="Schwendinger & Giribet" authorityYear="2004" box="[151,776,1832,1858]" class="Arachnida" family="Stylocellidae" genus="Miopsalis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="globosa" status="comb. nov.">
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<emphasis id="93B0CB48707A9964DEAAF89AAD5CF8F5" bold="true" box="[151,368,1832,1858]" italics="true" pageId="29" pageNumber="30">Miopsalis globosa</emphasis>
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(
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<bibRefCitation id="C5556AAB707A9964DF42F89AAED3F8F5" author="Schwendinger & Giribet" box="[383,767,1832,1858]" firstAuthor="Schwendinger" pageId="29" pageNumber="30" pagination="1421 - 1435" refId="ref17049" refString="Schwendinger, P. J., Giribet, G. & Steiner, H. (2004) A remarkable new cave-dwelling Stylocellus (Opiliones, Cyphophthalmi) from peninsular Malaysia, with a discussion on taxonomic characters in the family Stylocellidae. Journal of Natural History, 38 (11), 1421 - 1435." type="journal article" year="2004">Schwendinger & Giribet, 2004</bibRefCitation>
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)
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</taxonomicName>
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<taxonomicNameLabel id="88837633707A9964DD32F89AAFB8F8F5" box="[783,916,1832,1858]" pageId="29" pageNumber="30">new comb.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="E9DE44D1707A9964DEAAF8DDAF77F833" box="[151,859,1902,1925]" pageId="29" pageNumber="30" type="reference_group">
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<paragraph id="A17B175A707A9964DEAAF8DDAF77F833" blockId="29.[151,859,1902,1925]" box="[151,859,1902,1925]" pageId="29" pageNumber="30">
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<treatmentCitationGroup id="81D43074707A9964DEAAF8DDAF77F833" box="[151,859,1902,1925]" pageId="29" pageNumber="30">
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<taxonomicName id="66C46CD9707A9964DEAAF8DDAEC9F833" authority="Schwendinger & Giribet, 2004: 1423" authorityName="Schwendinger & Giribet" authorityPageNumber="1423" authorityYear="2004" box="[151,741,1902,1925]" class="Arachnida" family="Stylocellidae" genus="Stylocellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="globosus">
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<emphasis id="93B0CB48707A9964DEAAF8DDAD48F832" box="[151,356,1903,1925]" italics="true" pageId="29" pageNumber="30">Stylocellus globosus</emphasis>
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<treatmentCitation id="2065314B707A9964DF57F8DCAEC9F833" author="Schwendinger" box="[362,741,1902,1924]" page="1423" pageId="29" pageNumber="30" year="2004">
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<bibRefCitation id="C5556AAB707A9964DF57F8DCAEC9F833" author="Schwendinger & Giribet" box="[362,741,1902,1924]" firstAuthor="Schwendinger" page="1423" pageId="29" pageNumber="30" pagination="1421 - 1435" refId="ref17049" refString="Schwendinger, P. J., Giribet, G. & Steiner, H. (2004) A remarkable new cave-dwelling Stylocellus (Opiliones, Cyphophthalmi) from peninsular Malaysia, with a discussion on taxonomic characters in the family Stylocellidae. Journal of Natural History, 38 (11), 1421 - 1435." type="journal article" year="2004">Schwendinger & Giribet, 2004: 1423</bibRefCitation>
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</treatmentCitation>
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</taxonomicName>
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, figs. 1–29.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="E9DE44D1707A9967DEAAF803AD31FD33" lastPageId="30" lastPageNumber="31" pageId="29" pageNumber="30" type="discussion">
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<paragraph id="A17B175A707A9967DEAAF803AD31FD33" blockId="29.[151,1436,1969,2030]" lastBlockId="30.[151,1437,151,716]" lastPageId="30" lastPageNumber="31" pageId="29" pageNumber="30">
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<emphasis id="93B0CB48707A9964DEAAF803AD0DF87D" bold="true" box="[151,289,1969,1994]" pageId="29" pageNumber="30">Comments:</emphasis>
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This highly troglomorphic species has moved between Fangensinae
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<emphasis id="93B0CB48707A9964DA07F803A8FFF87D" bold="true" box="[1082,1235,1969,1994]" pageId="29" pageNumber="30">subfam. nov.</emphasis>
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and
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<taxonomicName id="66C46CD9707A9964DB33F803A951F87E" box="[1294,1405,1969,1993]" class="Arachnida" family="Stylocellidae" genus="Miopsalis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="29" pageNumber="30" phylum="Arthropoda" rank="genus">
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<emphasis id="93B0CB48707A9964DB33F803A951F87E" box="[1294,1405,1969,1993]" italics="true" pageId="29" pageNumber="30">Miopsalis</emphasis>
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</taxonomicName>
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in various morphometric phylogenetic analyses (
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<bibRefCitation id="C5556AAB707A9964DCA6F867AF42F85A" author="Clouse" box="[667,878,2005,2030]" etAl="et al." firstAuthor="Clouse" pageId="29" pageNumber="30" pagination="515" refId="ref16084" refString="Clouse, R. M., de Bivort, B. L. & Giribet, G. (2009) A phylogenetic analysis for the South-east Asian mite harvestman family Stylocellidae (Opiliones: Cyphophthalmi) - a combined analysis using morphometric and molecular data. Invertebrate Systematics, 23, 515 - 529." type="journal article" year="2009">
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Clouse
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<emphasis id="93B0CB48707A9964DCCFF865AF06F859" box="[754,810,2006,2030]" italics="true" pageId="29" pageNumber="30">et al.</emphasis>
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2009
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</bibRefCitation>
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). Although it has placed most commonly with the former, the case for its placement in
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<taxonomicName id="66C46CD970799967DC07FF2AAE85FF07" box="[570,681,152,176]" class="Arachnida" family="Stylocellidae" genus="Miopsalis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="30" pageNumber="31" phylum="Arthropoda" rank="genus">
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<emphasis id="93B0CB4870799967DC07FF2AAE85FF07" box="[570,681,152,176]" italics="true" pageId="30" pageNumber="31">Miopsalis</emphasis>
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</taxonomicName>
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appears stronger in light of the subfamily features identified here. The extremely rounded opisthosoma, with its lateral flange and shortened posterior, surely complicate morphometric studies and may push analyses to combine
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<taxonomicName id="66C46CD970799967DD23FF53AFB0FF4F" box="[798,924,224,248]" class="Arachnida" family="Stylocellidae" genus="Miopsalis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="globosa" status="comb. nov.">
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<emphasis id="93B0CB4870799967DD23FF53AFB0FF4F" box="[798,924,224,248]" italics="true" pageId="30" pageNumber="31">M. globosa</emphasis>
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</taxonomicName>
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<emphasis id="93B0CB4870799967DD99FF52A80FFF4F" bold="true" box="[932,1059,224,248]" pageId="30" pageNumber="31">
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<taxonomicNameLabel id="8883763370799967DD99FF52A80FFF4F" box="[932,1059,224,248]" pageId="30" pageNumber="31">new comb.</taxonomicNameLabel>
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</emphasis>
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with convergently troglomorphic species (which includes all of
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<taxonomicName id="66C46CD970799967DFCEFEB7AE4AFEAB" authority="(Schwendinger & Giribet 2005)" box="[499,614,261,284]" class="Arachnida" family="Stylocellidae" genus="Fangensis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="30" pageNumber="31" phylum="Arthropoda" rank="genus">
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<emphasis id="93B0CB4870799967DFCEFEB7AE4AFEAB" box="[499,614,261,284]" italics="true" pageId="30" pageNumber="31">Fangensis</emphasis>
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</taxonomicName>
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). It does not, however, present the most distinct feature of Fangensinae
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<emphasis id="93B0CB4870799967DEAAFE95AD02FEF7" bold="true" box="[151,302,295,320]" pageId="30" pageNumber="31">subfam. nov.</emphasis>
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, which is the enormous size of the fourth coxae. Three other discrepancies could be attributable to the opisthosoma of
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<taxonomicName id="66C46CD970799967DF6AFEFFADF0FED3" box="[343,476,332,356]" class="Arachnida" family="Stylocellidae" genus="Miopsalis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="globosa" status="comb. nov.">
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<emphasis id="93B0CB4870799967DF6AFEFFADF0FED3" box="[343,476,332,356]" italics="true" pageId="30" pageNumber="31">M. globosa</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="8883763370799967DFD7FEFEAE5CFED3" box="[490,624,332,356]" pageId="30" pageNumber="31">
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<emphasis id="93B0CB4870799967DFD7FEFEAE5CFED3" bold="true" box="[490,624,332,356]" pageId="30" pageNumber="31">new comb.</emphasis>
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</taxonomicNameLabel>
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being enlarged and widened, but they further weaken the case for placement in Fangensinae
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<emphasis id="93B0CB4870799967DFF8FEDDAE72FE3F" bold="true" box="[453,606,367,392]" pageId="30" pageNumber="31">subfam. nov.</emphasis>
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: the ozophores are not perpendicular to the long body axis, the posterior prosoma is not enlarged, and the fourth coxae do not have lateral edges parallel to the long body axis. Once we no longer consider Fangensinae
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<emphasis id="93B0CB4870799967DFD5FE05AEAFFE67" bold="true" box="[488,643,439,464]" pageId="30" pageNumber="31">subfam. nov.</emphasis>
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(nor, obviously,
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<emphasis id="93B0CB4870799967DD74FE0AAFEBFE67" box="[841,967,440,464]" italics="true" pageId="30" pageNumber="31">Meghalaya</emphasis>
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) as options for a generic placement, we have to consider two characters that might ally
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<taxonomicName id="66C46CD970799967DC84FE6FAF15FE43" box="[697,825,476,500]" class="Arachnida" family="Stylocellidae" genus="Miopsalis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="globosa">
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<emphasis id="93B0CB4870799967DC84FE6FAF15FE43" box="[697,825,476,500]" italics="true" pageId="30" pageNumber="31">M. globosa</emphasis>
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</taxonomicName>
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with
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<taxonomicName id="66C46CD970799967DD41FE6EAFD8FE43" box="[892,1012,476,500]" class="Arachnida" family="Stylocellidae" genus="Stylocellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="30" pageNumber="31" phylum="Arthropoda" rank="genus">
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<emphasis id="93B0CB4870799967DD41FE6EAFD8FE43" box="[892,1012,476,500]" italics="true" pageId="30" pageNumber="31">Stylocellus</emphasis>
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</taxonomicName>
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or
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<taxonomicName id="66C46CD970799967DA23FE6EA8B3FE43" box="[1054,1183,476,500]" class="Arachnida" family="Stylocellidae" genus="Leptopsalis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="30" pageNumber="31" phylum="Arthropoda" rank="genus">
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<emphasis id="93B0CB4870799967DA23FE6EA8B3FE43" box="[1054,1183,476,500]" italics="true" pageId="30" pageNumber="31">Leptopsalis</emphasis>
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</taxonomicName>
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: the strongly concave posterior gonostome and the rounded profile. But these are relatively variable characters across the family and contradicted by a combination of
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<taxonomicName id="66C46CD970799967DC73FD96AE90FD8B" box="[590,700,548,572]" class="Arachnida" family="Stylocellidae" genus="Miopsalis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="30" pageNumber="31" phylum="Arthropoda" rank="genus">
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<emphasis id="93B0CB4870799967DC73FD96AE90FD8B" box="[590,700,548,572]" italics="true" pageId="30" pageNumber="31">Miopsalis</emphasis>
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</taxonomicName>
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-like charactacter states: tapered ozophores, straight sternal opisthosomal sulci, large anal gland pore, large and flat ventral prosomal complex, and significant cheliceral sculpturing.
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</paragraph>
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</subSubSection>
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<subSubSection id="E9DE44D170799967DEFAFD3DAF26FD7B" pageId="30" pageNumber="31" type="materials_examined">
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<paragraph id="A17B175A70799967DEFAFD3DAF26FD7B" blockId="30.[151,1437,151,716]" pageId="30" pageNumber="31">
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<emphasis id="93B0CB4870799967DEFAFD3DAD9AFD1F" bold="true" box="[199,438,655,680]" pageId="30" pageNumber="31">Material examined:</emphasis>
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<materialsCitation id="11AC1D0770799967DFFDFD23AF2AFD7B" ID-GBIF-Occurrence="3443618301" pageId="30" pageNumber="31" typeStatus="paratype">
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<typeStatus id="7E7FA9F870799967DFFDFD23AE0BFD1F" box="[448,551,657,680]" pageId="30" pageNumber="31" type="paratype">
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<emphasis id="93B0CB4870799967DFFDFD23AE0BFD1F" box="[448,551,657,680]" italics="true" pageId="30" pageNumber="31">Paratype</emphasis>
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</typeStatus>
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, Male (
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<collectionCode id="C7D58F9F70799967DCBEFD3DAEEEFD10" LSID="urn:lsid:biocol.org:col:33791" box="[643,706,655,679]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="30" pageNumber="31">MCZ</collectionCode>
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44691),
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<collectingCountry id="D9D357CA70799967DD11FD3DAFB9FD10" box="[812,917,655,679]" name="Malaysia" pageId="30" pageNumber="31">Malaysia</collectingCountry>
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, Negeri Perak (Perak State), Gunung Lanno, Gua Monophyllaea,
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<date id="D57A319A70799967DF41FD06AE7AFD7B" box="[380,598,692,716]" pageId="30" pageNumber="31" value="2001-11-13">13 November 2001</date>
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, col. H. Steiner
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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</treatment>
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||
</document> |