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<document id="ECA73787CF07A872B862BBD8F13A9FDA" ID-CLB-Dataset="42344" ID-DOI="10.11646/zootaxa.3911.2.8" ID-GBIF-Dataset="1e357e76-0e29-4930-bd7e-29c66c8fffba" ID-ISSN="1175-5326" ID-Zenodo-Dep="237557" ID-ZooBank="FBF334D6-B43D-4EF6-BB5E-D89BDF528048" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460941719213" checkinUser="plazi" docAuthor="Jałoszyński, Paweł" docDate="2015" docId="201D64714330FFC8AEAF58A6FADEFC81" docLanguage="en" docName="zt03911p279.pdf" docOrigin="Zootaxa 3911 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cephennomicrus santubongensis Jałoszyński, 2015, sp. n." docType="treatment" docVersion="8" lastPageNumber="278" masterDocId="DC241C094333FFCDAE385E1EFFB0FFDC" masterDocTitle="Four new species of Cephennomicrus Reitter (Coleoptera, Staphylinidae, Scydmaeninae) from Malaysia" masterLastPageNumber="279" masterPageNumber="273" pageNumber="276" updateTime="1698584410400" updateUser="ExternalLinkService">
<mods:mods id="6E0C8685AF3D4B76966766271DF047E6" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="F7D3591F40F0B7297E37E3AF6B141998">
<mods:title id="F5D941B09D8E78069709FE7D8EF2BE24">Four new species of Cephennomicrus Reitter (Coleoptera, Staphylinidae, Scydmaeninae) from Malaysia</mods:title>
</mods:titleInfo>
<mods:name id="E382FE188CF29CE245D6E86AF8BF4336" type="personal">
<mods:role id="1252A515D3F90CA6BEEA8192EC18D046">
<mods:roleTerm id="600E2F083997652E21E5DF1A2030E29D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B417E9805B1206658882F5CA8D05745A">Jałoszyński, Paweł</mods:namePart>
</mods:name>
<mods:typeOfResource id="A6E42F2D0575612E6FAF7A3C4235EAFA">text</mods:typeOfResource>
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<mods:title id="E03306B1831C46B7DE83FDE32184732C">Zootaxa</mods:title>
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<mods:part id="B77F8891DE79759E6EF9CC1B93D5C2E5">
<mods:date id="6BEF52C5D40C9658B7C317358C9870EB">2015</mods:date>
<mods:detail id="A281C26AD330B2C0515FE370A4CC7F00" type="volume">
<mods:number id="C05386F3E2677489833C26869F92A849">3911</mods:number>
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<mods:number id="39CE012D15D6C55161D67AD8139D9D35">2</mods:number>
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<mods:start id="600BB43E1D86F133E10F660E9664430E">273</mods:start>
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<mods:classification id="8510FD2C584CF6332C95F2EC7CC679A8">journal article</mods:classification>
<mods:identifier id="280FA5365FA3D51F3498C45D80F7A127" type="CLB-Dataset">42344</mods:identifier>
<mods:identifier id="A64BC572D1E0EA492D225235C7DC1012" type="DOI">10.11646/zootaxa.3911.2.8</mods:identifier>
<mods:identifier id="9A07F9F6E906A9E2675ED15C0A7FAD11" type="GBIF-Dataset">1e357e76-0e29-4930-bd7e-29c66c8fffba</mods:identifier>
<mods:identifier id="F49BF325422B586B49880FF3B9534620" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="EF7F896FCB04BB208B94738C693921B0" type="Zenodo-Dep">237557</mods:identifier>
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</mods:mods>
<treatment id="201D64714330FFC8AEAF58A6FADEFC81" ID-DOI="http://doi.org/10.5281/zenodo.6097404" ID-GBIF-Taxon="119591477" ID-Zenodo-Dep="6097404" LSID="urn:lsid:plazi:treatment:201D64714330FFC8AEAF58A6FADEFC81" httpUri="http://treatment.plazi.org/id/201D64714330FFC8AEAF58A6FADEFC81" lastPageId="5" lastPageNumber="278" pageId="3" pageNumber="276">
<subSubSection id="E0AE86EC4330FFCEAEAF58A6FED5F92F" pageId="3" pageNumber="276" type="nomenclature">
<paragraph id="A80BD5674330FFCEAEAF58A6FDC2F90E" blockId="3.[151,626,1720,1780]" box="[151,626,1720,1746]" pageId="3" pageNumber="276">
<heading id="F343620B4330FFCEAEAF58A6FDC2F90E" bold="true" box="[151,626,1720,1746]" fontSize="11" level="1" pageId="3" pageNumber="276" reason="1">
<emphasis id="9AC009754330FFCEAEAF58A6FDC2F90E" bold="true" box="[151,626,1720,1746]" pageId="3" pageNumber="276">
<taxonomicName id="6FB4AEE44330FFCEAEAF58A6FD98F90E" ID-CoL="SFPR" box="[151,552,1720,1746]" class="Insecta" family="Staphylinidae" genus="Cephennomicrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="276" phylum="Arthropoda" rank="species" species="santubongensis" status="sp. nov.">
<emphasis id="9AC009754330FFCEAEAF58A6FD98F90E" bold="true" box="[151,552,1720,1746]" italics="true" pageId="3" pageNumber="276">Cephennomicrus santubongensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="81F3B40E4330FFCEAC1758A6FDC2F90E" box="[559,626,1720,1746]" pageId="3" pageNumber="276" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="A80BD5674330FFCEAEAF58C5FED5F92F" blockId="3.[151,626,1720,1780]" box="[151,357,1755,1780]" pageId="3" pageNumber="276">
(
<figureCitation id="308FC9E24330FFCEAEA758C5FF5DF92F" box="[159,237,1755,1780]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,264,1349,1372]" captionTargetBox="[162,1433,674,1320]" captionTargetId="figure@2.[151,1436,663,1328]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Dorsal habitus of holotype males. 1 ― Cephennomicrus curvidens sp. n.; 2 ― Cephennomicrus gentingensis sp. n.; 3 ― Cephennomicrus santubongensis sp. n.; 4 ― Cephennomicrus muluensis sp. n." httpUri="https://zenodo.org/record/237558/files/figure.png" pageId="3" pageNumber="276">Figs. 3</figureCitation>
,
<figureCitation id="308FC9E24330FFCEAEC258C2FEBAF92F" box="[250,266,1756,1779]" captionStart="FIGURES 5 – 8" captionStartId="3.[151,264,1591,1614]" captionTargetBox="[156,1428,997,1562]" captionTargetId="figure@3.[151,1436,988,1571]" captionTargetPageId="3" captionText="FIGURES 5 – 8. Body outline and distribution of macrosetae. 5 ― Cephennomicrus curvidens sp. n.; 6 ― Cephennomicrus gentingensis sp. n.; 7 ― Cephennomicrus santubongensis sp. n.; 8 ― Cephennomicrus muluensis sp. n." httpUri="https://zenodo.org/record/237559/files/figure.png" pageId="3" pageNumber="276">7</figureCitation>
,
<figureCitation id="308FC9E24330FFCEAF2E58C5FEEDF92F" box="[278,349,1755,1779]" captionStart="FIGURES 9 – 16" captionStartId="4.[151,264,1346,1369]" captionTargetBox="[159,1428,205,1314]" captionTargetId="figure@4.[151,1436,193,1325]" captionTargetPageId="4" captionText="FIGURES 9 – 16. Aedeagi of holotype males in ventral (9, 11, 13, 15) and lateral (10, 12, 14, 16) views. 9 – 10 ― Cephennomicrus curvidens sp. n.; 11 – 12 ― Cephennomicrus gentingensis sp. n.; 13 – 14 ― Cephennomicrus santubongensis sp. n.; 15 – 16 ― Cephennomicrus muluensis sp. n." httpUri="https://zenodo.org/record/237560/files/figure.png" pageId="3" pageNumber="276">13–14</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="E0AE86EC4330FFCEAEAF593DFA97F85F" pageId="3" pageNumber="276" type="materials_examined">
<paragraph id="A80BD5674330FFCEAEAF593DFA97F85F" blockId="3.[151,1436,1827,2031]" pageId="3" pageNumber="276">
<emphasis id="9AC009754330FFCEAEAF593DFC13F8E0" bold="true" box="[151,931,1827,1852]" pageId="3" pageNumber="276">
<typeStatus id="770F6BC54330FFCEAEAF593DFF62F8E0" box="[151,210,1827,1852]" pageId="3" pageNumber="276">Type</typeStatus>
material.
<typeStatus id="770F6BC54330FFCEAF74593DFE0DF8E0" box="[332,445,1827,1852]" pageId="3" pageNumber="276" type="holotype">Holotype</typeStatus>
:
<collectionCode id="CEA54DA24330FFCEAFF1593DFDA2F8E0" box="[457,530,1827,1852]" pageId="3" pageNumber="276">EAST</collectionCode>
<collectingCountry id="D0A395F74330FFCEAC21593DFD1FF8E0" box="[537,687,1827,1852]" name="Malaysia" pageId="3" pageNumber="276">MALAYSIA</collectingCountry>
(Borneo:
<collectingCountry id="D0A395F74330FFCEAD1F593DFC26F8E0" box="[807,918,1827,1852]" name="Malaysia" pageId="3" pageNumber="276">Sarawak</collectingCountry>
):
</emphasis>
♂, two labels: &quot;
<collectingCountry id="D0A395F74330FFCEAA6F593DFB5FF8E7" box="[1111,1263,1827,1851]" name="Malaysia" pageId="3" pageNumber="276">MALAYSIA</collectingCountry>
:
<collectingCountry id="D0A395F74330FFCEAAC2593DFAEDF8E0" box="[1274,1373,1827,1852]" name="Malaysia" pageId="3" pageNumber="276">Sarawak</collectingCountry>
, San- / tubong,
<quantity id="6F4C78824330FFCEAF2A5959FED1F883" box="[274,353,1862,1887]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="3" pageNumber="276" unit="km" value="20.0">20 km</quantity>
N Kuching / Camp Permai,
<quantity id="6F4C78824330FFCEACE95959FCA4F883" box="[721,788,1863,1887]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="276" unit="m" value="10.0">10 m</quantity>
,
<date id="DC0AF3A74330FFCEAD105958FBB1F883" box="[808,1025,1862,1887]" pageId="3" pageNumber="276" value="2003-08-05" valueMax="2003-08-10" valueMin="2003-08-05">5.-10. / VIII.2003</date>
, leg. A. Schulz&quot; [white, printed]; &quot;
<taxonomicName id="6FB4AEE44330FFCEAE9A5975FED0F85F" box="[162,352,1899,1923]" class="Insecta" family="Staphylinidae" genus="Cephennomicrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="276" phylum="Arthropoda" rank="genus">
<emphasis id="9AC009754330FFCEAE9A5975FED0F85F" box="[162,352,1899,1923]" italics="true" pageId="3" pageNumber="276">Cephennomicrus</emphasis>
</taxonomicName>
/
<taxonomicName id="6FB4AEE44330FFCEAF4E5975FD93F85F" box="[374,547,1899,1923]" class="Insecta" family="Staphylinidae" genus="Cephennomicrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="276" phylum="Arthropoda" rank="species" species="santubongensis">
<emphasis id="9AC009754330FFCEAF4E5975FD93F85F" box="[374,547,1899,1923]" italics="true" pageId="3" pageNumber="276">santubongensis</emphasis>
</taxonomicName>
m. / det. P. Jałoszyński, '14 /
<typeStatus id="770F6BC54330FFCEAD555975FBA7F85F" box="[877,1047,1899,1923]" pageId="3" pageNumber="276" type="holotype">HOLOTYPUS</typeStatus>
&quot; [red, printed] (
<collectionCode id="CEA54DA24330FFCEAAF05975FAAAF85F" LSID="urn:lsid:biocol.org:col:34840" box="[1224,1306,1899,1923]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="3" pageNumber="276">SMNS</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="E0AE86EC4330FFCEAEFF5990FC28F833" pageId="3" pageNumber="276" type="diagnosis">
<paragraph id="A80BD5674330FFCEAEFF5990FC28F833" blockId="3.[151,1436,1827,2031]" pageId="3" pageNumber="276">
<emphasis id="9AC009754330FFCEAEFF5990FEF1F87B" bold="true" box="[199,321,1934,1959]" pageId="3" pageNumber="276">Diagnosis.</emphasis>
BL about
<quantity id="6F4C78824330FFCEAFF75991FDABF87B" box="[463,539,1935,1959]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="276" unit="mm" value="1.0">1 mm</quantity>
; pronotum with two pairs of small antebasal pits; each elytron with two macrosetae; aedeagus in ventral view strongly elongate, with sharply delimited apical part longer than bulbous basal part and composed of dorsal and ventral plates of equal length.
</paragraph>
</subSubSection>
<caption id="FCCB85EF4337FFC9AEAF5B5CFC9FFA49" httpUri="https://zenodo.org/record/237560/files/figure.png" pageId="4" pageNumber="277" targetBox="[159,1428,205,1314]" targetPageId="4">
<paragraph id="A80BD5674337FFC9AEAF5B5CFC9FFA49" blockId="4.[151,1436,1346,1430]" pageId="4" pageNumber="277">
<emphasis id="9AC009754337FFC9AEAF5B5CFEE4FA84" bold="true" box="[151,340,1346,1369]" pageId="4" pageNumber="277">FIGURES 9–16.</emphasis>
Aedeagi of holotype males in ventral (9, 11, 13, 15) and lateral (10, 12, 14, 16) views. 9–10
<emphasis id="9AC009754337FFC9AEF15B7CFF52FAAB" bold="true" box="[201,226,1378,1399]" pageId="4" pageNumber="277">―</emphasis>
<taxonomicName id="6FB4AEE44337FFC9AEDB5B7CFDB7FAAB" box="[227,519,1378,1399]" class="Insecta" family="Staphylinidae" genus="Cephennomicrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="277" phylum="Arthropoda" rank="species" species="curvidens" status="sp. nov.">
<emphasis id="9AC009754337FFC9AEDB5B7CFDB7FAAB" box="[227,519,1378,1399]" italics="true" pageId="4" pageNumber="277">Cephennomicrus curvidens</emphasis>
</taxonomicName>
<emphasis id="9AC009754337FFC9AC255B7CFDD6FAAB" bold="true" box="[541,614,1378,1399]" pageId="4" pageNumber="277">
<taxonomicNameLabel id="81F3B40E4337FFC9AC255B7CFDD6FAAB" box="[541,614,1378,1399]" pageId="4" pageNumber="277" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
; 11–12
<emphasis id="9AC009754337FFC9ACFA5B7CFD6BFAAB" bold="true" box="[706,731,1378,1399]" pageId="4" pageNumber="277">―</emphasis>
<taxonomicName id="6FB4AEE44337FFC9ACE35B7CFBACFAAB" box="[731,1052,1378,1399]" class="Insecta" family="Staphylinidae" genus="Cephennomicrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="277" phylum="Arthropoda" rank="species" species="gentingensis" status="sp. nov.">
<emphasis id="9AC009754337FFC9ACE35B7CFBACFAAB" box="[731,1052,1378,1399]" italics="true" pageId="4" pageNumber="277">Cephennomicrus gentingensis</emphasis>
</taxonomicName>
<emphasis id="9AC009754337FFC9AA0A5B7CFBCBFAAB" bold="true" box="[1074,1147,1378,1399]" pageId="4" pageNumber="277">
<taxonomicNameLabel id="81F3B40E4337FFC9AA0A5B7CFBCBFAAB" box="[1074,1147,1378,1399]" pageId="4" pageNumber="277" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
; 13–14
<emphasis id="9AC009754337FFC9AAEF5B7CFB40FAAB" bold="true" box="[1239,1264,1378,1399]" pageId="4" pageNumber="277">―</emphasis>
<taxonomicName id="6FB4AEE44337FFC9AAC85B7CFE83FA49" class="Insecta" family="Staphylinidae" genus="Cephennomicrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="277" phylum="Arthropoda" rank="species" species="santubongensis" status="sp. nov.">
<emphasis id="9AC009754337FFC9AAC85B7CFE83FA49" italics="true" pageId="4" pageNumber="277">Cephennomicrus santubongensis</emphasis>
</taxonomicName>
<emphasis id="9AC009754337FFC9AF015B9EFEC2FA49" bold="true" box="[313,370,1408,1429]" pageId="4" pageNumber="277">
<taxonomicNameLabel id="81F3B40E4337FFC9AF015B9EFEC2FA49" box="[313,370,1408,1429]" pageId="4" pageNumber="277" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
; 15–16
<emphasis id="9AC009754337FFC9AF875B9EFE68FA49" bold="true" box="[447,472,1408,1429]" pageId="4" pageNumber="277">―</emphasis>
<taxonomicName id="6FB4AEE44337FFC9AFE05B9EFD5FFA49" box="[472,751,1408,1429]" class="Insecta" family="Staphylinidae" genus="Cephennomicrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="277" phylum="Arthropoda" rank="species" species="muluensis" status="sp. nov.">
<emphasis id="9AC009754337FFC9AFE05B9EFD5FFA49" box="[472,751,1408,1429]" italics="true" pageId="4" pageNumber="277">Cephennomicrus muluensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="81F3B40E4337FFC9ACCD5B9EFC9FFA49" box="[757,815,1408,1429]" pageId="4" pageNumber="277" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
</caption>
<subSubSection id="E0AE86EC4337FFC8AEFF5BDCFE2EFDE1" lastPageId="5" lastPageNumber="278" pageId="4" pageNumber="277" type="description">
<paragraph id="A80BD5674337FFC9AEFF5BDCFEADFA23" blockId="4.[151,1437,1474,2003]" pageId="4" pageNumber="277">
<emphasis id="9AC009754337FFC9AEFF5BDCFEE9FA07" bold="true" box="[199,345,1474,1499]" pageId="4" pageNumber="277">Description.</emphasis>
BL
<quantity id="6F4C78824337FFC9AFB15BDCFE40FA06" box="[393,496,1474,1499]" metricMagnitude="-3" metricUnit="m" metricValue="1.06" pageId="4" pageNumber="277" unit="mm" value="1.06">1.06 mm</quantity>
. Body of male (
<figureCitation id="308FC9E24337FFC9AC9A5BDCFD41FA07" box="[674,753,1474,1499]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,264,1349,1372]" captionTargetBox="[162,1433,674,1320]" captionTargetId="figure@2.[151,1436,663,1328]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Dorsal habitus of holotype males. 1 ― Cephennomicrus curvidens sp. n.; 2 ― Cephennomicrus gentingensis sp. n.; 3 ― Cephennomicrus santubongensis sp. n.; 4 ― Cephennomicrus muluensis sp. n." httpUri="https://zenodo.org/record/237558/files/figure.png" pageId="4" pageNumber="277">Figs. 3</figureCitation>
,
<figureCitation id="308FC9E24337FFC9ACC55BDDFCBCFA07" box="[765,780,1475,1499]" captionStart="FIGURES 5 – 8" captionStartId="3.[151,264,1591,1614]" captionTargetBox="[156,1428,997,1562]" captionTargetId="figure@3.[151,1436,988,1571]" captionTargetPageId="3" captionText="FIGURES 5 – 8. Body outline and distribution of macrosetae. 5 ― Cephennomicrus curvidens sp. n.; 6 ― Cephennomicrus gentingensis sp. n.; 7 ― Cephennomicrus santubongensis sp. n.; 8 ― Cephennomicrus muluensis sp. n." httpUri="https://zenodo.org/record/237559/files/figure.png" pageId="4" pageNumber="277">7</figureCitation>
) elongate and strongly convex, brown with slightly lighter macrosetae.
</paragraph>
<paragraph id="A80BD5674337FFC9AEFF5814FCB3F927" blockId="4.[151,1437,1474,2003]" pageId="4" pageNumber="277">
Head broadest at large, strongly convex and coarsely faceted eyes; HL
<quantity id="6F4C78824337FFC9ADC65814FBD7F9FE" box="[1022,1127,1546,1571]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" pageId="4" pageNumber="277" unit="mm" value="0.15">0.15 mm</quantity>
, HW
<quantity id="6F4C78824337FFC9AA955815FAA6F9FE" box="[1197,1302,1547,1571]" metricMagnitude="-4" metricUnit="m" metricValue="2.8" pageId="4" pageNumber="277" unit="mm" value="0.28">0.28 mm</quantity>
; vertex and frons confluent and convex; supraantennal tubercles feebly marked. Punctures on head dorsum shallow and distinct, slightly irregular in shape and unevenly distributed, separated by spaces 1–1.5x as wide as diameters of punctures; setae indiscernible under magnification
<date id="DC0AF3A74337FFC9ACDD5869FCA7F953" box="[741,791,1655,1679]" pageId="4" pageNumber="277" value="1980-10">80x.</date>
Antennae slender and with club composed of enlarged antennomeres X–XI, AnL
<quantity id="6F4C78824337FFC9AFE95884FD8DF96E" box="[465,573,1690,1715]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="277" unit="mm" value="0.5">0.50 mm</quantity>
, antennomeres I–II and IX elongate; III–VIII about as broad as long; X transverse; XI much longer than X and about 1.3x as long as broad; antennomeres IX–XI with sparse strongly erect and long setae among short and suberect basic setation.
</paragraph>
<paragraph id="A80BD5674337FFC8AEFF5916FEF7FF09" blockId="4.[151,1437,1474,2003]" lastBlockId="5.[151,1437,151,861]" lastPageId="5" lastPageNumber="278" pageId="4" pageNumber="277">
Pronotum (
<figureCitation id="308FC9E24337FFC9AF7D5918FE3AF8C3" box="[325,394,1798,1823]" captionStart="FIGURES 5 – 8" captionStartId="3.[151,264,1591,1614]" captionTargetBox="[156,1428,997,1562]" captionTargetId="figure@3.[151,1436,988,1571]" captionTargetPageId="3" captionText="FIGURES 5 – 8. Body outline and distribution of macrosetae. 5 ― Cephennomicrus curvidens sp. n.; 6 ― Cephennomicrus gentingensis sp. n.; 7 ― Cephennomicrus santubongensis sp. n.; 8 ― Cephennomicrus muluensis sp. n." httpUri="https://zenodo.org/record/237559/files/figure.png" pageId="4" pageNumber="277">Fig. 7</figureCitation>
) approximately subrectangular with convex anterior margin, broadest anterior to middle but indistinctly narrowing posteriorly; PL
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, PW
<quantity id="6F4C78824337FFC9ACD25935FCE1F89E" box="[746,849,1835,1859]" metricMagnitude="-4" metricUnit="m" metricValue="4.3" pageId="4" pageNumber="277" unit="mm" value="0.43">0.43 mm</quantity>
; anterior and lateral margins in anterior half broadly rounded, lateral margins in posterior half nearly straight; anterior corners visible only in anterodorsal view, distinct and sharp-angled but with rounded apices; posterior margin nearly straight with indistinct flattening in front of mesoscutellum; posterior corners slightly obtuse; pronotal base with two lateral pairs of small distinct pits. Punctures on pronotal disc smaller than those on frons and vertex but distinct, separated by spaces subequal to diameters of punctures; pronotal disc only with macrosetae: two lateral pairs and one posterior pair in front of mesoscutellum.
</paragraph>
<paragraph id="A80BD5674336FFC8AEFF5EC1FD1BFE9C" blockId="5.[151,1437,151,861]" pageId="5" pageNumber="278">
Elytra (
<figureCitation id="308FC9E24336FFC8AF265EC1FED4FF24" box="[286,356,223,248]" captionStart="FIGURES 5 – 8" captionStartId="3.[151,264,1591,1614]" captionTargetBox="[156,1428,997,1562]" captionTargetId="figure@3.[151,1436,988,1571]" captionTargetPageId="3" captionText="FIGURES 5 – 8. Body outline and distribution of macrosetae. 5 ― Cephennomicrus curvidens sp. n.; 6 ― Cephennomicrus gentingensis sp. n.; 7 ― Cephennomicrus santubongensis sp. n.; 8 ― Cephennomicrus muluensis sp. n." httpUri="https://zenodo.org/record/237559/files/figure.png" pageId="5" pageNumber="278">Fig. 7</figureCitation>
) oval, broadest anterior to middle; EL
<quantity id="6F4C78824336FFC8AD0A5EC1FC2BFF2B" box="[818,923,223,248]" metricMagnitude="-4" metricUnit="m" metricValue="5.8999999999999995" pageId="5" pageNumber="278" unit="mm" value="0.59">0.59 mm</quantity>
, EW
<quantity id="6F4C78824336FFC8ADD85EC1FBFAFF2B" box="[992,1098,223,248]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="278" unit="mm" value="0.5">0.50 mm</quantity>
, EI 1.18; humeral calli wellmarked, elongate. Punctures on elytra barely discernible under magnification
<date id="DC0AF3A74336FFC8AA1A5F1AFBFFFEC0" box="[1058,1103,260,284]" pageId="5" pageNumber="278" value="1980-10">80x</date>
; each elytron only with two lateral macrosetae.
<collectingCountry id="D0A395F74336FFC8AF485F39FE18FE9C" box="[368,424,295,320]" name="India" pageId="5" pageNumber="278">Hind</collectingCountry>
wings well developed.
</paragraph>
<paragraph id="A80BD5674336FFC8AEFF5F53FD68FEB9" blockId="5.[151,1437,151,861]" box="[199,728,332,357]" pageId="5" pageNumber="278">Legs moderately long and slender, unmodified.</paragraph>
<paragraph id="A80BD5674336FFC8AEFF5F71FDB3FDCB" blockId="5.[151,1437,151,861]" pageId="5" pageNumber="278">
Aedeagus (
<figureCitation id="308FC9E24336FFC8AF755F71FE6CFE54" box="[333,476,367,392]" captionStart="FIGURES 9 – 16" captionStartId="4.[151,264,1346,1369]" captionTargetBox="[159,1428,205,1314]" captionTargetId="figure@4.[151,1436,193,1325]" captionTargetPageId="4" captionText="FIGURES 9 – 16. Aedeagi of holotype males in ventral (9, 11, 13, 15) and lateral (10, 12, 14, 16) views. 9 – 10 ― Cephennomicrus curvidens sp. n.; 11 – 12 ― Cephennomicrus gentingensis sp. n.; 13 – 14 ― Cephennomicrus santubongensis sp. n.; 15 – 16 ― Cephennomicrus muluensis sp. n." httpUri="https://zenodo.org/record/237560/files/figure.png" pageId="5" pageNumber="278">Figs. 13–14</figureCitation>
) in ventral view strongly elongate; AeL
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; median lobe with sharply demarcated and very long apical part, apex subtrapezoidal and slightly concave; in lateral view apical projections composed of long ventral and dorsal plates; internal armature complicated and symmetrical, with a pair of very long and slender lateral rod-like projections curved mesally; parameres broad and not reaching apex of median lobe, each with two apical setae.
</paragraph>
<paragraph id="A80BD5674336FFC8AEFF5C3AFE2EFDE1" blockId="5.[151,1437,151,861]" box="[199,414,548,573]" pageId="5" pageNumber="278">Female. Unknown.</paragraph>
</subSubSection>
<subSubSection id="E0AE86EC4336FFC8AEFF5C59FDCBFDBC" box="[199,635,583,608]" pageId="5" pageNumber="278" type="distribution">
<paragraph id="A80BD5674336FFC8AEFF5C59FDCBFDBC" blockId="5.[151,1437,151,861]" box="[199,635,583,608]" pageId="5" pageNumber="278">
<emphasis id="9AC009754336FFC8AEFF5C59FED0FDBC" bold="true" box="[199,352,583,608]" pageId="5" pageNumber="278">Distribution.</emphasis>
East
<collectingCountry id="D0A395F74336FFC8AFA65C59FDB8FDBC" box="[414,520,583,608]" name="Malaysia" pageId="5" pageNumber="278">Malaysia</collectingCountry>
,
<collectingCountry id="D0A395F74336FFC8AC2B5C59FDC7FDBC" box="[531,631,583,608]" name="Malaysia" pageId="5" pageNumber="278">Sarawak</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="E0AE86EC4336FFC8AEFF5C72FD44FD59" box="[199,756,620,645]" pageId="5" pageNumber="278" type="etymology">
<paragraph id="A80BD5674336FFC8AEFF5C72FD44FD59" blockId="5.[151,1437,151,861]" box="[199,756,620,645]" pageId="5" pageNumber="278">
<emphasis id="9AC009754336FFC8AEFF5C72FEFEFD59" bold="true" box="[199,334,620,645]" pageId="5" pageNumber="278">Etymology.</emphasis>
Locotypical, after Mount Santubong.
</paragraph>
</subSubSection>
<subSubSection id="E0AE86EC4336FFC8AEFF5C91FADEFC81" pageId="5" pageNumber="278" type="discussion">
<paragraph id="A80BD5674336FFC8AEFF5C91FADEFC81" blockId="5.[151,1437,151,861]" pageId="5" pageNumber="278">
<emphasis id="9AC009754336FFC8AEFF5C91FE8BFD74" bold="true" box="[199,315,655,680]" pageId="5" pageNumber="278">Remarks.</emphasis>
The aedeagus of this otherwise unremarkable species is very unusual and not similar to the genitalia of any other
<taxonomicName id="6FB4AEE44336FFC8AF135CAAFE58FD10" box="[299,488,692,716]" class="Insecta" family="Staphylinidae" genus="Cephennomicrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="278" phylum="Arthropoda" rank="genus">
<emphasis id="9AC009754336FFC8AF135CAAFE58FD10" box="[299,488,692,716]" italics="true" pageId="5" pageNumber="278">Cephennomicrus</emphasis>
</taxonomicName>
. Strongly elongated aedeagi can also be found in the
<taxonomicName id="6FB4AEE44336FFC8AA695CAAFA2BFD10" box="[1105,1435,692,716]" class="Insecta" family="Staphylinidae" genus="Cephennomicrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="278" phylum="Arthropoda" rank="species" species="taiwanensis">
<emphasis id="9AC009754336FFC8AA695CAAFA2BFD10" box="[1105,1435,692,716]" italics="true" pageId="5" pageNumber="278">Cephennomicrus taiwanensis</emphasis>
</taxonomicName>
species group, but in that distinct Taiwanese lineage the apical projections are very short and the endophallus contains a distinct, strongly asymmetrical and usually looped flagellum (
<bibRefCitation id="CC25A8964336FFC8ADFC5CE2FB3AFCC8" author="Jaloszynski" box="[964,1162,764,789]" pageId="5" pageNumber="278" refString="Jaloszynski, P. (2009) Cephennomicrus Reitter (Coleoptera, Staphylinidae, Scydmaeninae) of Japan and Taiwan: taxonomic notes, ten new species and comparative morphology of nomurai and taiwanensis species groups. Zootaxa, 2145, 1 - 35." type="journal article" year="2009">Jałoszyński 2009</bibRefCitation>
). Another species with a strongly elongated median lobe is
<taxonomicName id="6FB4AEE44336FFC8AC0F5D3EFC9DFCE4" authority="Jaloszynski" authorityName="Jaloszynski" box="[567,813,799,824]" class="Insecta" family="Staphylinidae" genus="Cephennomicrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="278" phylum="Arthropoda" rank="species" species="lasios">
<emphasis id="9AC009754336FFC8AC0F5D3EFD2FFCE4" box="[567,671,800,824]" italics="true" pageId="5" pageNumber="278">C. lasios</emphasis>
Jałoszyński
</taxonomicName>
from the
<collectingCountry id="D0A395F74336FFC8AD955D01FB9EFCE4" box="[941,1070,799,824]" name="Philippines" pageId="5" pageNumber="278">Philippines</collectingCountry>
, but in that species the apical projections are short, and a long and looped flagellum can be seen inside the median lobe (
<bibRefCitation id="CC25A8964336FFC8AAB65D5AFAD2FC81" author="Jaloszynski" box="[1166,1378,836,861]" pageId="5" pageNumber="278" refString="Jaloszynski, P. (2010 b) Cephenniini of the Philippines. Part 4. Two new species of Cephennomicrus Reitter from Luzon (Coleoptera, Staphylinidae, Scydmaeninae). Genus, 21 (2), 197 - 204." type="journal article" year="2010" yearSuffix="b">Jałoszyński 2010b</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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