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<document id="40DBBD22C463A006FF95C975A37D096F" ID-DOI="10.5281/zenodo.197274" ID-GBIF-Dataset="c6a45517-5e2d-496e-a76c-091740a56475" ID-ISSN="1175-5326" ID-Zenodo-Dep="197274" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460263975273" checkinUser="plazi" docAuthor="Tarasov, Sergey I." docDate="2010" docId="200A8781FFD8F35500F091D09137204E" docLanguage="en" docName="zt02566p048.pdf" docOrigin="Zootaxa 2566" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Onthophagus gracilipes Boucomont 1914" docType="treatment" docVersion="8" lastPageNumber="46" masterDocId="DC33FFF9FFD9F35700679528925A2121" masterDocTitle="Seven new synonyms within the genus Onthophagus (Coleoptera: Scarabaeidae) from the Oriental Region" masterLastPageNumber="48" masterPageNumber="45" pageNumber="46" updateTime="1698245222659" updateUser="plazi">
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<mods:title id="79C607A05B76572022F19E89734996D0">Seven new synonyms within the genus Onthophagus (Coleoptera: Scarabaeidae) from the Oriental Region</mods:title>
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<mods:namePart id="E46E91990112C4ED2A83B1339AF87B16">Tarasov, Sergey I.</mods:namePart>
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<mods:date id="88F8E8F062F641D79602038D8CC22DA7">2010</mods:date>
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<treatment id="200A8781FFD8F35500F091D09137204E" ID-DOI="http://doi.org/10.5281/zenodo.6208722" ID-GBIF-Taxon="119393974" ID-Zenodo-Dep="6208722" LSID="urn:lsid:plazi:treatment:200A8781FFD8F35500F091D09137204E" httpUri="http://treatment.plazi.org/id/200A8781FFD8F35500F091D09137204E" lastPageId="2" lastPageNumber="46" pageId="1" pageNumber="46">
<subSubSection id="E0B9651CFFD8F35600F091D09033242E" box="[151,617,1271,1296]" pageId="1" pageNumber="46" type="nomenclature">
<paragraph id="A81C3697FFD8F35600F091D09033242E" blockId="1.[151,1325,1271,1359]" box="[151,617,1271,1296]" pageId="1" pageNumber="46">
<heading id="F35481FBFFD8F35600F091D09033242E" bold="true" box="[151,617,1271,1296]" fontSize="10" level="2" pageId="1" pageNumber="46" reason="2">
<taxonomicName id="6FA34D14FFD8F35600F091D09033242E" ID-CoL="49MKG" authority="Boucomont, 1914" authorityName="Boucomont" authorityYear="1914" box="[151,617,1271,1296]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="46" phylum="Arthropoda" rank="species" species="gracilipes">
<emphasis id="9AD7EA85FFD8F35600F091D09033242E" bold="true" box="[151,617,1271,1296]" pageId="1" pageNumber="46">
<emphasis id="9AD7EA85FFD8F35600F091D093C72431" bold="true" box="[151,413,1272,1296]" italics="true" pageId="1" pageNumber="46">Onthophagus gracilipes</emphasis>
<bibRefCitation id="CC324B66FFD8F35601C391DF9033242E" author="Boucomont" box="[420,617,1271,1295]" pageId="1" pageNumber="46" refString="Boucomont, A. (1914) Onthophagus asiatiques nouveaux ou peu connus. Annali del Museo Civico di Storia Naturale &quot; Giacomo Doria &quot;, 46, 210 - 243." type="journal article" year="1914">Boucomont, 1914</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="E0B9651CFFD8F35600F090309777246E" pageId="1" pageNumber="46" type="reference_group">
<paragraph id="A81C3697FFD8F35600F090309777246E" blockId="1.[151,1325,1271,1359]" pageId="1" pageNumber="46">
<treatmentCitationGroup id="88B311B9FFD8F35600F090309777246E" pageId="1" pageNumber="46">
<taxonomicName id="6FA34D14FFD8F35600F0903090C9240E" ID-CoL="49MKG" authority="Boucomont, 1914: 219" authorityName="Boucomont" authorityPageNumber="219" authorityYear="1914" box="[151,659,1304,1327]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="46" phylum="Arthropoda" rank="species" species="gracilipes">
<emphasis id="9AD7EA85FFD8F35600F0903093C2240E" box="[151,408,1304,1327]" italics="true" pageId="1" pageNumber="46">Onthophagus gracilipes</emphasis>
<treatmentCitation id="29021086FFD8F35601F9903090C9240E" author="Boucomont" box="[414,659,1304,1327]" page="219" pageId="1" pageNumber="46" year="1914">
<bibRefCitation id="CC324B66FFD8F35601F99030903B240E" author="Boucomont" box="[414,609,1304,1327]" pageId="1" pageNumber="46" refString="Boucomont, A. (1914) Onthophagus asiatiques nouveaux ou peu connus. Annali del Museo Civico di Storia Naturale &quot; Giacomo Doria &quot;, 46, 210 - 243." type="journal article" year="1914">Boucomont, 1914</bibRefCitation>
: 219
</treatmentCitation>
</taxonomicName>
(
<typeStatus id="77188835FFD8F35602C69030908A240E" box="[673,720,1304,1327]" pageId="1" pageNumber="46">type</typeStatus>
locality: Carin Cheba [
<collectingCountry id="D0B47607FFD8F35603AF9030966D240E" box="[968,1079,1304,1327]" name="Myanmar" pageId="1" pageNumber="46">Myanmar</collectingCountry>
])
<taxonomicName id="6FA34D14FFD8F35600F09010914D246E" ID-CoL="49MDS" authority="Frey, 1971: 95" authorityName="Frey" authorityPageNumber="95" authorityYear="1971" box="[151,791,1336,1359]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="46" phylum="Arthropoda" rank="species" species="laosensis" subGenus="Digithonthophagus">
<emphasis id="9AD7EA85FFD8F35600F09010902F246E" box="[151,629,1336,1359]" italics="true" pageId="1" pageNumber="46">Onthophagus (Digithonthophagus) laosensis</emphasis>
<treatmentCitation id="29021086FFD8F356021B9010914D246E" author="Frey" box="[636,791,1336,1359]" page="95" pageId="1" pageNumber="46" year="1971">
<bibRefCitation id="CC324B66FFD8F356021B901090AB246E" author="Frey" box="[636,753,1336,1359]" pageId="1" pageNumber="46" refString="Frey, G. (1971) Neue Afrikanischen und Asiatischen Onthophagus arten. Entomologische Arbeiten aus dem Museum G. Frey, 22, 93 - 98." type="journal article" year="1971">Frey, 1971</bibRefCitation>
: 95
</treatmentCitation>
</taxonomicName>
<taxonomicNameLabel id="81E457FEFFD8F356037A901F913A246E" box="[797,864,1335,1359]" pageId="1" pageNumber="46">
<emphasis id="9AD7EA85FFD8F356037A901F913A246E" bold="true" box="[797,864,1335,1359]" pageId="1" pageNumber="46">syn.n.</emphasis>
</taxonomicNameLabel>
(
<typeStatus id="77188835FFD8F356030A901091C7246E" box="[877,925,1336,1359]" pageId="1" pageNumber="46">type</typeStatus>
locality:
<collectingCountry id="D0B47607FFD8F35604669010966D246E" box="[1025,1079,1336,1359]" name="Laos" pageId="1" pageNumber="46">Laos</collectingCountry>
,
<collectingCountry id="D0B47607FFD8F35604259010962A246E" box="[1090,1136,1336,1359]" name="Iceland" pageId="1" pageNumber="46">Sala</collectingCountry>
Nam Chan
<collectingCountry id="D0B47607FFD8F35604979010977F246E" box="[1264,1317,1336,1359]" name="China" pageId="1" pageNumber="46">Chin</collectingCountry>
)
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="E0B9651CFFD8F35600F0905F961726EE" pageId="1" pageNumber="46" type="materials_examined">
<paragraph id="A81C3697FFD8F35600F0905F973024AE" blockId="1.[151,1436,1399,2031]" box="[151,1386,1399,1423]" pageId="1" pageNumber="46">
<emphasis id="9AD7EA85FFD8F35600F0905F93EB24AE" bold="true" box="[151,433,1399,1423]" pageId="1" pageNumber="46">
<typeStatus id="77188835FFD8F35600F0905F929524AE" box="[151,207,1399,1423]" pageId="1" pageNumber="46">Type</typeStatus>
material examined.
</emphasis>
<typeStatus id="77188835FFD8F35601D09050904524AE" box="[439,543,1400,1423]" pageId="1" pageNumber="46" type="lectotype">
<emphasis id="9AD7EA85FFD8F35601D09050904524AE" box="[439,543,1400,1423]" italics="true" pageId="1" pageNumber="46">Lectotype</emphasis>
</typeStatus>
of
<taxonomicName id="6FA34D14FFD8F35602259050909724AE" box="[578,717,1400,1423]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="46" phylum="Arthropoda" rank="species" species="gracilipes">
<emphasis id="9AD7EA85FFD8F35602259050909724AE" box="[578,717,1400,1423]" italics="true" pageId="1" pageNumber="46">
<collectingCountry id="D0B47607FFD8F35602259050900224AE" box="[578,600,1400,1423]" name="Iceland" pageId="1" pageNumber="46">O</collectingCountry>
. gracilipes
</emphasis>
</taxonomicName>
(
<collectionCode id="CEB2AE52FFD8F35602BC9050916A24AE" LSID="urn:lsid:biocol.org:col:34988" box="[731,816,1400,1423]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="1" pageNumber="46">MNHN</collectionCode>
)
<emphasis id="9AD7EA85FFD8F3560359905F91B524AE" bold="true" box="[830,1007,1399,1423]" pageId="1" pageNumber="46">here designated</emphasis>
, male bearing the following labels:
</paragraph>
<paragraph id="A81C3697FFD8F35600A290B09696248E" blockId="1.[151,1436,1399,2031]" box="[197,1228,1432,1455]" pageId="1" pageNumber="46">
1. White: Carin Cheba [printed]
<quantity id="6F5B9B72FFD8F356024790B090C5248E" box="[544,671,1432,1455]" metricMagnitude="2" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="1" pageNumber="46" unit="m" value="750.0" valueMax="1000.0" valueMin="500.0">5001000m</quantity>
[handwritten] L. Fea [printed]
<date id="DC1D1057FFD8F356038B90B0966F248E" box="[1004,1077,1432,1455]" pageId="1" pageNumber="46" value="1987-12">XII-87</date>
[handwritten]
</paragraph>
<paragraph id="A81C3697FFD8F35600A29090901024EE" blockId="1.[151,1436,1399,2031]" box="[197,586,1464,1487]" pageId="1" pageNumber="46">2. White, printed: Mus. Civ. Genova</paragraph>
<paragraph id="A81C3697FFD8F35600A290F091D724CE" blockId="1.[151,1436,1399,2031]" box="[197,909,1496,1519]" pageId="1" pageNumber="46">
3. White, printed: MUSÉUM
<collectionCode id="CEB2AE52FFD8F356026490F0901624CE" box="[515,588,1496,1519]" pageId="1" pageNumber="46">PARIS</collectionCode>
1936
<collectionCode id="CEB2AE52FFD8F35602E890F0908224CE" box="[655,728,1496,1519]" pageId="1" pageNumber="46">COLL</collectionCode>
. BOUCOMONT
</paragraph>
<paragraph id="A81C3697FFD8F35600A290D093E8272E" blockId="1.[151,1436,1399,2031]" box="[197,434,1528,1551]" pageId="1" pageNumber="46">
4. Red, printed:
<typeStatus id="77188835FFD8F356011690D093E8272E" box="[369,434,1528,1551]" pageId="1" pageNumber="46">Typus</typeStatus>
</paragraph>
<paragraph id="A81C3697FFD8F35600A2933091FE2708" blockId="1.[151,1436,1399,2031]" box="[197,932,1559,1583]" pageId="1" pageNumber="46">
5. White: Boucomont det. 191 [printed] 4 onthoph.
<taxonomicName id="6FA34D14FFD8F356028C93309109270E" box="[747,851,1560,1583]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="46" phylum="Arthropoda" rank="species" species="gracilipes" status="sp. nov.">gracilipes</taxonomicName>
<taxonomicNameLabel id="81E457FEFFD8F356033E933091D6270E" box="[857,908,1560,1583]" pageId="1" pageNumber="46" rank="species">n.sp.</taxonomicNameLabel>
</paragraph>
<paragraph id="A81C3697FFD8F35600A293109689276E" blockId="1.[151,1436,1399,2031]" box="[197,1235,1592,1615]" pageId="1" pageNumber="46">
6. Orange printed:
<typeStatus id="77188835FFD8F35601E99310905F276E" box="[398,517,1592,1615]" pageId="1" pageNumber="46" type="lectotype">Lectotypus</typeStatus>
<taxonomicName id="6FA34D14FFD8F356026C9310961E276E" authority="Boucomont, 1914 S. Tarasov" authorityName="Boucomont, 1914 S. Tarasov" box="[523,1092,1592,1615]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="46" phylum="Arthropoda" rank="species" species="gracilipes">
Onthophagus gracilipes
<bibRefCitation id="CC324B66FFD8F356036893109197276E" author="Boucomont" box="[783,973,1592,1615]" pageId="1" pageNumber="46" refString="Boucomont, A. (1914) Onthophagus asiatiques nouveaux ou peu connus. Annali del Museo Civico di Storia Naturale &quot; Giacomo Doria &quot;, 46, 210 - 243." type="journal article" year="1914">Boucomont, 1914</bibRefCitation>
S. Tarasov
</taxonomicName>
design. 2009
</paragraph>
<paragraph id="A81C3697FFD8F35600A293709132278E" blockId="1.[151,1436,1399,2031]" pageId="1" pageNumber="46">
Also, 1Ψ
<typeStatus id="77188835FFD8F356014D937093E7274E" box="[298,445,1624,1647]" pageId="1" pageNumber="46" type="paralectotype">
<emphasis id="9AD7EA85FFD8F356014D937093E7274E" box="[298,445,1624,1647]" italics="true" pageId="1" pageNumber="46">paralectotype</emphasis>
</typeStatus>
of
<taxonomicName id="6FA34D14FFD8F356018793709031274E" box="[480,619,1624,1647]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="46" phylum="Arthropoda" rank="species" species="gracilipes">
<emphasis id="9AD7EA85FFD8F356018793709031274E" box="[480,619,1624,1647]" italics="true" pageId="1" pageNumber="46">
<collectingCountry id="D0B47607FFD8F3560187937093AC274E" box="[480,502,1624,1647]" name="Iceland" pageId="1" pageNumber="46">O</collectingCountry>
. gracilipes
</emphasis>
</taxonomicName>
(
<collectionCode id="CEB2AE52FFD8F356021F93709097274E" LSID="urn:lsid:biocol.org:col:34988" box="[632,717,1624,1647]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="1" pageNumber="46">MNHN</collectionCode>
), bearing labels identical to
<typeStatus id="77188835FFD8F356039D93709606274E" box="[1018,1116,1624,1647]" pageId="1" pageNumber="46" type="lectotype">lectotype</typeStatus>
with the exception of the first label “Carin Ghecu
<quantity id="6F5B9B72FFD8F35601109350905227AE" box="[375,520,1656,1679]" metricMagnitude="3" metricUnit="m" metricValue="1.35" metricValueMax="1.4" metricValueMin="1.3" pageId="1" pageNumber="46" unit="m" value="1350.0" valueMax="1400.0" valueMin="1300.0">13001400m</quantity>
L. Fea
<date id="DC1D1057FFD8F35602079350909E27AE" box="[608,708,1656,1679]" pageId="1" pageNumber="46" value="1988-02" valueMax="1988-03-31" valueMin="1988-02-00">IIIII.88</date>
”. The
<typeStatus id="77188835FFD8F356036A9350912927AE" box="[781,883,1656,1679]" pageId="1" pageNumber="46" type="lectotype">lectotype</typeStatus>
is designated in order to preserve the stability of zoological nomenclature according to the article 74.7 (
<bibRefCitation id="CC324B66FFD8F35602BA93B09106278E" author="ICZN" box="[733,860,1688,1711]" pageId="1" pageNumber="46" refString="ICZN (1999) International Code of Zoological Nomenclature (4 th Edition). International Trust for Zoological Nomenclature, London, 306 pp." type="book" year="1999">
<collectionCode id="CEB2AE52FFD8F35602BA93B09147278E" box="[733,797,1688,1711]" pageId="1" pageNumber="46">ICZN</collectionCode>
1999
</bibRefCitation>
).
</paragraph>
<paragraph id="A81C3697FFD8F35600A2939091F627EE" blockId="1.[151,1436,1399,2031]" box="[197,940,1720,1743]" pageId="1" pageNumber="46">
<typeStatus id="77188835FFD8F35600A29390937D27EE" box="[197,295,1720,1743]" pageId="1" pageNumber="46" type="holotype">
<emphasis id="9AD7EA85FFD8F35600A29390937D27EE" box="[197,295,1720,1743]" italics="true" pageId="1" pageNumber="46">Holotype</emphasis>
</typeStatus>
of
<taxonomicName id="6FA34D14FFD8F356012D9390939727EE" box="[330,461,1720,1743]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="46" phylum="Arthropoda" rank="species" species="laosensis">
<emphasis id="9AD7EA85FFD8F356012D9390939727EE" box="[330,461,1720,1743]" italics="true" pageId="1" pageNumber="46">
<collectingCountry id="D0B47607FFD8F356012D9390933A27EE" box="[330,352,1720,1743]" name="Iceland" pageId="1" pageNumber="46">O</collectingCountry>
. laosensis
</emphasis>
</taxonomicName>
(NHB), female bearing the following labels:
</paragraph>
<paragraph id="A81C3697FFD8F35600A293F0908527CE" blockId="1.[151,1436,1399,2031]" box="[197,735,1752,1775]" pageId="1" pageNumber="46">
1. White, handwritten:
<collectingCountry id="D0B47607FFD8F35601DC93F093A927CE" box="[443,499,1752,1775]" name="Laos" pageId="1" pageNumber="46">Laos</collectingCountry>
:
<collectingCountry id="D0B47607FFD8F356019A93F0907127CE" box="[509,555,1752,1775]" name="Iceland" pageId="1" pageNumber="46">Sala</collectingCountry>
Nam Chan
<collectingCountry id="D0B47607FFD8F35602CC93F0908527CE" box="[683,735,1752,1775]" name="China" pageId="1" pageNumber="46">Chin</collectingCountry>
</paragraph>
<paragraph id="A81C3697FFD8F35600A293D0903F262E" blockId="1.[151,1436,1399,2031]" box="[197,613,1784,1807]" pageId="1" pageNumber="46">
2. White printed: VITALIS
<date id="DC1D1057FFD8F356018A93D0903F262E" box="[493,613,1784,1807]" pageId="1" pageNumber="46" value="1919" valueMax="1920">19191920</date>
</paragraph>
<paragraph id="A81C3697FFD8F35600A292309725260E" blockId="1.[151,1436,1399,2031]" box="[197,1407,1815,1839]" pageId="1" pageNumber="46">
3. White:
<taxonomicName id="6FA34D14FFD8F3560148923090CE260E" authority="(Digitonthophagus)" baseAuthorityName="Digitonthophagus" box="[303,660,1816,1839]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="46" phylum="Arthropoda" rank="genus">Onthophagus (Digitonthophagus)</taxonomicName>
n.sp. Ψ [handwritten] Dr. V. Balthasar det. [handwritten] 60. [printed]
</paragraph>
<paragraph id="A81C3697FFD8F35600A2921093EE266E" blockId="1.[151,1436,1399,2031]" box="[197,436,1848,1871]" pageId="1" pageNumber="46">
4. Red, printed:
<collectionCode id="CEB2AE52FFD8F3560116921093EE266E" box="[369,436,1848,1871]" pageId="1" pageNumber="46">
<typeStatus id="77188835FFD8F3560116921093EE266E" box="[369,436,1848,1871]" pageId="1" pageNumber="46">TYPE</typeStatus>
</collectionCode>
</paragraph>
<paragraph id="A81C3697FFD8F35600A29270937926AE" blockId="1.[151,1436,1399,2031]" pageId="1" pageNumber="46">
5. White:
<collectionCode id="CEB2AE52FFD8F356015792709328264E" box="[304,370,1880,1903]" pageId="1" pageNumber="46">
<typeStatus id="77188835FFD8F356015792709328264E" box="[304,370,1880,1903]" pageId="1" pageNumber="46">TYPE</typeStatus>
</collectionCode>
Ψ [handwritten]
<taxonomicName id="6FA34D14FFD8F356024B927090E0264E" box="[556,698,1880,1903]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="46" phylum="Arthropoda" rank="genus">Onthophagus</taxonomicName>
[printed]
<taxonomicName id="6FA34D14FFD8F356034092709614264E" box="[807,1102,1880,1903]" class="Insecta" family="Scarabaeidae" genus="Digitonthophagus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="46" phylum="Arthropoda" rank="species" species="laosensis" status="sp. nov.">Digitonthophagus laosensis</taxonomicName>
<taxonomicNameLabel id="81E457FEFFD8F3560432927096D2264E" box="[1109,1160,1880,1903]" pageId="1" pageNumber="46" rank="species">n.sp.</taxonomicNameLabel>
[handwritten] det.G.Frey, 196 [printed]
</paragraph>
<paragraph id="A81C3697FFD8F35600A292B09743268E" blockId="1.[151,1436,1399,2031]" box="[197,1305,1944,1967]" pageId="1" pageNumber="46">
6. Orange:
<typeStatus id="77188835FFD8F356015B92B09393268E" box="[316,457,1944,1967]" pageId="1" pageNumber="46" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="6FA34D14FFD8F35601A892B09166268E" authority="Frey, 1971" authorityName="Frey" authorityYear="1971" box="[463,828,1944,1967]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="46" phylum="Arthropoda" rank="species" species="laosensis">
Onthophagus laosensis
<bibRefCitation id="CC324B66FFD8F35602AC92B09166268E" author="Frey" box="[715,828,1944,1967]" pageId="1" pageNumber="46" refString="Frey, G. (1971) Neue Afrikanischen und Asiatischen Onthophagus arten. Entomologische Arbeiten aus dem Museum G. Frey, 22, 93 - 98." type="journal article" year="1971">Frey, 1971</bibRefCitation>
</taxonomicName>
[handwritten] S. Tarasov det. 2009 [printed]
</paragraph>
<paragraph id="A81C3697FFD8F35600A29290961726EE" blockId="1.[151,1436,1399,2031]" box="[197,1101,1976,1999]" pageId="1" pageNumber="46">
7. White:
<taxonomicName id="6FA34D14FFD8F35601489290902B26EE" authority="Bouc." authorityName="Bouc." box="[303,625,1976,1999]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="46" phylum="Arthropoda" rank="species" species="gracilipes">Onthophagus gracilipes Bouc.</taxonomicName>
[handwritten] S. Tarasov det. 2009 [printed]
</paragraph>
</subSubSection>
<subSubSection id="E0B9651CFFD8F35500A292FF9137204E" lastPageId="2" lastPageNumber="47" pageId="1" pageNumber="46" type="discussion">
<paragraph id="A81C3697FFD8F35500A292FF9137204E" blockId="1.[151,1436,1399,2031]" lastBlockId="2.[151,1437,152,367]" lastPageId="2" lastPageNumber="47" pageId="1" pageNumber="46">
<emphasis id="9AD7EA85FFD8F35600A292FF936926CE" bold="true" box="[197,307,2007,2031]" pageId="1" pageNumber="46">Remarks.</emphasis>
It seems that
<bibRefCitation id="CC324B66FFD8F35601AF92F0901D26CE" author="Frey" box="[456,583,2008,2031]" pageId="1" pageNumber="46" refString="Frey, G. (1971) Neue Afrikanischen und Asiatischen Onthophagus arten. Entomologische Arbeiten aus dem Museum G. Frey, 22, 93 - 98." type="journal article" year="1971">Frey (1971)</bibRefCitation>
did not thoroughly examine the description of
<taxonomicName id="6FA34D14FFD8F356042792F0969626CE" box="[1088,1228,2008,2031]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="46" phylum="Arthropoda" rank="species" species="gracilipes">
<emphasis id="9AD7EA85FFD8F356042792F0969626CE" box="[1088,1228,2008,2031]" italics="true" pageId="1" pageNumber="46">
<collectingCountry id="D0B47607FFD8F356042792F0960C26CE" box="[1088,1110,2008,2031]" name="Iceland" pageId="1" pageNumber="46">O</collectingCountry>
. gracilipes
</emphasis>
</taxonomicName>
and did not see the
<typeStatus id="77188835FFDBF35500F095B0929F218E" box="[151,197,152,175]" pageId="2" pageNumber="47">type</typeStatus>
series of the former. In the original description he compares
<taxonomicName id="6FA34D14FFDBF355032D95B09191218E" box="[842,971,152,175]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="species" species="laosensis">
<emphasis id="9AD7EA85FFDBF355032D95B09191218E" box="[842,971,152,175]" italics="true" pageId="2" pageNumber="47">
<collectingCountry id="D0B47607FFDBF355032D95B0913A218E" box="[842,864,152,175]" name="Iceland" pageId="2" pageNumber="47">O</collectingCountry>
. laosensis
</emphasis>
</taxonomicName>
with
<taxonomicName id="6FA34D14FFDBF355046095B0976E218E" authority="Balthasar, 1933" authorityName="Balthasar" authorityYear="1933" box="[1031,1332,152,175]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="species" species="troniceki">
<emphasis id="9AD7EA85FFDBF355046095B096DC218E" box="[1031,1158,152,175]" italics="true" pageId="2" pageNumber="47">
<collectingCountry id="D0B47607FFDBF355046095B09647218E" box="[1031,1053,152,175]" name="Iceland" pageId="2" pageNumber="47">O</collectingCountry>
. troniceki
</emphasis>
Balthasar, 1933
</taxonomicName>
. The
<typeStatus id="77188835FFDBF355050995B097C6218E" box="[1390,1436,152,175]" pageId="2" pageNumber="47">type</typeStatus>
series of Freys
<taxonomicName id="6FA34D14FFDBF35501589590939821EE" box="[319,450,184,207]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="species" species="laosensis">
<emphasis id="9AD7EA85FFDBF35501589590939821EE" box="[319,450,184,207]" italics="true" pageId="2" pageNumber="47">
<collectingCountry id="D0B47607FFDBF35501589590930F21EE" box="[319,341,184,207]" name="Iceland" pageId="2" pageNumber="47">O</collectingCountry>
. laosensis
</emphasis>
</taxonomicName>
includes only a female
<typeStatus id="77188835FFDBF35502DA9590914721EE" box="[701,797,184,207]" pageId="2" pageNumber="47" type="holotype">holotype</typeStatus>
, however that was not an obstacle for identification because the
<typeStatus id="77188835FFDBF35500D995F092B621CE" box="[190,236,216,239]" pageId="2" pageNumber="47">type</typeStatus>
series of
<taxonomicName id="6FA34D14FFDBF355013395F093BA21CE" box="[340,480,216,239]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="species" species="gracilipes">
<emphasis id="9AD7EA85FFDBF355013395F093BA21CE" box="[340,480,216,239]" italics="true" pageId="2" pageNumber="47">
<collectingCountry id="D0B47607FFDBF355013395F0933021CE" box="[340,362,216,239]" name="Iceland" pageId="2" pageNumber="47">O</collectingCountry>
. gracilipes
</emphasis>
</taxonomicName>
includes a female. I synonymize these two names because the
<typeStatus id="77188835FFDBF35504E495F096EB21CE" box="[1155,1201,216,239]" pageId="2" pageNumber="47">type</typeStatus>
series of
<taxonomicName id="6FA34D14FFDBF355057E95F097C621CE" box="[1305,1436,216,239]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="species" species="laosensis">
<emphasis id="9AD7EA85FFDBF355057E95F097C621CE" box="[1305,1436,216,239]" italics="true" pageId="2" pageNumber="47">
<collectingCountry id="D0B47607FFDBF355057E95F0977521CE" box="[1305,1327,216,239]" name="Iceland" pageId="2" pageNumber="47">O</collectingCountry>
. laosensis
</emphasis>
</taxonomicName>
and
<taxonomicName id="6FA34D14FFDBF35500A295D0930B202E" box="[197,337,248,271]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="species" species="parvidens">
<emphasis id="9AD7EA85FFDBF35500A295D0930B202E" box="[197,337,248,271]" italics="true" pageId="2" pageNumber="47">
<collectingCountry id="D0B47607FFDBF35500A295D09281202E" box="[197,219,248,271]" name="Iceland" pageId="2" pageNumber="47">O</collectingCountry>
. parvidens
</emphasis>
</taxonomicName>
are conspecific based on their same principle diagnostic character states of external morphology: the unique color pattern of the elytra and body, the unique groundplan of head, and the identical pattern of pronotum punctation. Additionally I have examined large material of
<taxonomicName id="6FA34D14FFDBF355036B941091CD206E" box="[780,919,312,335]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="species" species="gracilipes">
<emphasis id="9AD7EA85FFDBF355036B941091CD206E" box="[780,919,312,335]" italics="true" pageId="2" pageNumber="47">
<collectingCountry id="D0B47607FFDBF355036B94109178206E" box="[780,802,312,335]" name="Iceland" pageId="2" pageNumber="47">O</collectingCountry>
. gracilipes
</emphasis>
</taxonomicName>
from other localities in Indochina and these also match the two series of both names in the characters detailed above.
</paragraph>
</subSubSection>
</treatment>
</document>