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<document id="6C5E9EA67032004E80A2E2B3E198E495" ID-DOI="10.11646/zootaxa.1941.1.1" ID-ISSN="1175­5334" ID-Zenodo-Dep="5231015" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629573861311" checkinUser="felipe" docAuthor="Prieto, Carlos, Bálint, Zsolt, Boyer, Pierre &amp; Micó, Estefanía" docDate="2008" docId="18028E68FFB0AA426296BEBFFABDFB8B" docLanguage="en" docName="zt01941p024.pdf" docOrigin="Zootaxa 1941 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.1941.1.1" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Penaincisalia browni" docType="treatment" docVersion="4" lastPageNumber="5" masterDocId="E43BF610FFB2AA466201BB28FF9BFFC7" masterDocTitle="A review of the “ browni group ” of Penaincisalia with notes on their distribution and variability (Lepidoptera: Lycaenidae: Eumaeini)" masterLastPageNumber="24" masterPageNumber="1" pageNumber="3" updateTime="1699078074565" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="691ACB643B3431BEAD1ADEDE2175A3C2">A review of the “ browni group ” of Penaincisalia with notes on their distribution and variability (Lepidoptera: Lycaenidae: Eumaeini)</mods:title>
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<mods:namePart id="5AA315C0D55C4561C110D128C4121B68">Prieto, Carlos</mods:namePart>
<mods:affiliation id="6E11161EB1D8B0F0B359C8598442A675">Centro Iberoamericano de la Biodiversidad (CIBIO), Universidad de Alicante, Espa a. E-mail: cprieto 5000 @ yahoo. com. Hungarian Natural History Museum, Departament of Zoology, Budapest, Hungary. E-mail: balint @ nhmus. hu. Aix en Provence (France). E-mail: pierdom @ tiscali. fr. Grupo de Estudios en Geolog a, Ecolog a y Conservaci n (GECO), Universidad del Cauca, Colombia.</mods:affiliation>
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<mods:namePart id="78C4BE251F8A99F69ED6C9FC4536AFEB">Bálint, Zsolt</mods:namePart>
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<mods:namePart id="FFC7C90D817267ACFDACF0E65658BB4D">Boyer, Pierre</mods:namePart>
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<mods:namePart id="ECC46BE4754EA5A29BA70A155EA53B7C">Micó, Estefanía</mods:namePart>
<mods:affiliation id="38C99F492852935FFC787F8B79F94D47">Centro Iberoamericano de la Biodiversidad (CIBIO), Universidad de Alicante, Espa a. E-mail: cprieto 5000 @ yahoo. com. Hungarian Natural History Museum, Departament of Zoology, Budapest, Hungary. E-mail: balint @ nhmus. hu. Aix en Provence (France). E-mail: pierdom @ tiscali. fr. Grupo de Estudios en Geolog a, Ecolog a y Conservaci n (GECO), Universidad del Cauca, Colombia.</mods:affiliation>
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<mods:title id="6DC1A1EF4E28A45F1485859FA8743436">Zootaxa</mods:title>
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<mods:date id="0DDCA6576E8CBC0B2F29A27F887F605D">2008</mods:date>
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<mods:number id="FD42FE849992D7F9D6AA2973B7A30C0B">1941</mods:number>
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<treatment id="18028E68FFB0AA426296BEBFFABDFB8B" ID-DOI="http://doi.org/10.5281/zenodo.6315684" ID-Zenodo-Dep="6315684" LSID="urn:lsid:plazi:treatment:18028E68FFB0AA426296BEBFFABDFB8B" httpUri="http://treatment.plazi.org/id/18028E68FFB0AA426296BEBFFABDFB8B" lastPageId="4" lastPageNumber="5" pageId="2" pageNumber="3">
<subSubSection id="D8B16CF5FFB0AA446296BEBFFE08FA76" box="[151,403,1431,1457]" pageId="2" pageNumber="3" type="nomenclature">
<paragraph id="90143F7EFFB0AA446296BEBFFE08FA76" blockId="2.[151,403,1431,1457]" box="[151,403,1431,1457]" pageId="2" pageNumber="3">
<heading id="CB5C8812FFB0AA446296BEBFFE08FA76" bold="true" box="[151,403,1431,1457]" fontSize="11" level="1" pageId="2" pageNumber="3" reason="1">
<emphasis id="A2DFE36CFFB0AA446296BEBFFE08FA76" bold="true" box="[151,403,1431,1457]" pageId="2" pageNumber="3">
The “
<taxonomicName id="57AB44FDFFB0AA4462DCBEBFFEA9FA76" baseAuthorityName="Johnson" baseAuthorityYear="1992" box="[221,306,1431,1457]" class="Insecta" family="Lycaenidae" genus="Penaincisalia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="browni">
<emphasis id="A2DFE36CFFB0AA4462DCBEBFFEA9FA76" bold="true" box="[221,306,1431,1457]" italics="true" pageId="2" pageNumber="3">browni</emphasis>
</taxonomicName>
group”
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="D8B16CF5FFB0AA426296BEC0FABDFB8B" lastPageId="4" lastPageNumber="5" pageId="2" pageNumber="3" type="description">
<paragraph id="90143F7EFFB0AA446296BEC0FC07F965" blockId="2.[151,1437,1512,2018]" pageId="2" pageNumber="3">The wing patterns of all species of this group are very simple and consistent. The dorsal wing surface in males is structurally colored blue or purple, while the ventral wing surface is brown or reddish brown with simple and faint patterns of medial and submarginal bands. These subtle pattern elements proved to be among the most useful to distinguish species within the group. Secondary male wing characters, such as the shape and coloration of the androconial cluster, is also taxonomically useful.</paragraph>
<paragraph id="90143F7EFFB0AA4462C4BD98FCA6F885" blockId="2.[151,1437,1512,2018]" pageId="2" pageNumber="3">
The male valvae shapes, although also simple in structure, may differ significantly and consistently between species that are externally almost indistinguishable, like the species-pair
<taxonomicName id="57AB44FDFFB0AA446672BDF0FB7EF936" baseAuthorityName="Johnson" baseAuthorityYear="1992" box="[1139,1253,1752,1777]" class="Insecta" family="Lycaenidae" genus="Penaincisalia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="browni">
<emphasis id="A2DFE36CFFB0AA446672BDF0FB7EF936" box="[1139,1253,1752,1777]" italics="true" pageId="2" pageNumber="3">P. browni</emphasis>
</taxonomicName>
and
<emphasis id="A2DFE36CFFB0AA446722BDF0FA07F936" box="[1315,1436,1752,1777]" italics="true" pageId="2" pageNumber="3">
<taxonomicName id="57AB44FDFFB0AA446722BDF0FA03F936" authorityName="Robbins" authorityYear="2004" baseAuthorityName="Johnson" baseAuthorityYear="1992" box="[1315,1432,1752,1777]" class="Insecta" family="Lycaenidae" genus="Penaincisalia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="saraha">P. saraha</taxonomicName>
.
</emphasis>
Since, the entire view of the whole male genitalia doesnt provide sufficient information to discriminate species, we prefer to illustrate only the valvae in lateral view.
</paragraph>
<paragraph id="90143F7EFFB0AA4462C4BC78FBC7F855" blockId="2.[151,1437,1512,2018]" pageId="2" pageNumber="3">
The species of this presumably monophyletic group share the following combination of morphological characters which seem to be unique in
<taxonomicName id="57AB44FDFFB0AA44605CBC50FC2BF855" authority="sensu Robbins" authorityName="Robbins" box="[605,944,1912,1938]" class="Insecta" family="Lycaenidae" genus="Penaincisalia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="A2DFE36CFFB0AA44605CBC50FCDCF856" box="[605,839,1912,1937]" italics="true" pageId="2" pageNumber="3">Penaincisalia sensu</emphasis>
Robbins
</taxonomicName>
(See
<tableCitation id="DD290AC5FFB0AA4461F2BC50FBD2F855" box="[1011,1097,1912,1938]" captionStart="TABLE 1" captionStartId="4.[151,241,535,559]" captionText="TABLE 1. Distribution of characters (as numbered in the section “the browni species group), codes: present = “+”, absent = “-“,? = non available" pageId="2" pageNumber="3">Table 1</tableCitation>
):
</paragraph>
<paragraph id="90143F7EFFB0AA4462C4BC88FB1AF87D" blockId="2.[151,1437,1512,2018]" box="[197,1153,1952,1978]" pageId="2" pageNumber="3">
1-Hind wing tornus in males appears as a spatulate and bulbous lobe (
<figureCitation id="089023FBFFB0AA4461F9BC88FBEAF87D" box="[1016,1137,1952,1978]" captionStart="FIGURE 1" captionStartId="3.[151,255,1827,1851]" captionTargetBox="[303,1275,214,1791]" captionTargetId="figure-125@3.[283,1304,194,1803]" captionTargetPageId="3" captionText="FIGURE 1. General features in the external morphology of the “browni group”. (A): wing veins. (B): elements of “browni group” wing pattern indicating: DC, disc cell; mb, medial band; sm, submarginal band; m, marginal band; ds, discal spot; pb, postbasal pattern; BD, basal disc; dm, disc margin; AF, anal fold; SL, spatulate lobe. (C): wing shape of females indicating the different areas on the wing. ap, apical; b, basal; pb, postbasal; sm, submedial; m, medial; pm, postmedial; sma, submarginal; m, marginal; L, lateral lobe; ts, tail." figureDoi="http://doi.org/10.5281/zenodo.5231017" httpUri="https://zenodo.org/record/5231017/files/figure.png" pageId="2" pageNumber="3">Figure 1B</figureCitation>
).
</paragraph>
<paragraph id="90143F7EFFB0AA4262C4BCE0FCF8FF75" blockId="2.[151,1437,1512,2018]" lastBlockId="4.[151,1436,152,495]" lastPageId="4" lastPageNumber="5" pageId="2" pageNumber="3">
2-Sexual dimorphism in wing shape, where the females have a long tail from the CuA2 vein, accompanied by a small lateral lobe in the hindwing tornus (
<figureCitation id="089023FBFFB6AA4260B9BBB0FCCDFF75" box="[696,854,152,178]" captionStart="FIGURE 1" captionStartId="3.[151,255,1827,1851]" captionTargetBox="[303,1275,214,1791]" captionTargetId="figure-125@3.[283,1304,194,1803]" captionTargetPageId="3" captionText="FIGURE 1. General features in the external morphology of the “browni group”. (A): wing veins. (B): elements of “browni group” wing pattern indicating: DC, disc cell; mb, medial band; sm, submarginal band; m, marginal band; ds, discal spot; pb, postbasal pattern; BD, basal disc; dm, disc margin; AF, anal fold; SL, spatulate lobe. (C): wing shape of females indicating the different areas on the wing. ap, apical; b, basal; pb, postbasal; sm, submedial; m, medial; pm, postmedial; sma, submarginal; m, marginal; L, lateral lobe; ts, tail." figureDoi="http://doi.org/10.5281/zenodo.5231017" httpUri="https://zenodo.org/record/5231017/files/figure.png" pageId="4" pageNumber="5">Figure 1B, C</figureCitation>
).
</paragraph>
<caption id="C4D46FF6FFB1AA456296BC0BFCB5F802" ID-DOI="http://doi.org/10.5281/zenodo.5231017" ID-Zenodo-Dep="5231017" httpUri="https://zenodo.org/record/5231017/files/figure.png" pageId="3" pageNumber="4" startId="3.[151,255,1827,1851]" targetBox="[303,1275,214,1791]" targetPageId="3">
<paragraph id="90143F7EFFB1AA456296BC0BFCB5F802" blockId="3.[151,1436,1827,1989]" pageId="3" pageNumber="4">
<emphasis id="A2DFE36CFFB1AA456296BC0BFE8DF8FC" bold="true" box="[151,278,1827,1851]" pageId="3" pageNumber="4">FIGURE 1</emphasis>
. General features in the external morphology of the “
<taxonomicName id="57AB44FDFFB1AA456175BC0BFC24F8FD" baseAuthorityName="Johnson" baseAuthorityYear="1992" box="[884,959,1827,1850]" class="Insecta" family="Lycaenidae" genus="Penaincisalia" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="browni">
<emphasis id="A2DFE36CFFB1AA456175BC0BFC24F8FD" box="[884,959,1827,1850]" italics="true" pageId="3" pageNumber="4">browni</emphasis>
</taxonomicName>
group”.
<emphasis id="A2DFE36CFFB1AA456625BC0BFBC9F8FC" bold="true" box="[1060,1106,1827,1851]" pageId="3" pageNumber="4">(A):</emphasis>
wing veins.
<emphasis id="A2DFE36CFFB1AA4566E4BC0BFA89F8FC" bold="true" box="[1253,1298,1827,1851]" pageId="3" pageNumber="4">(B):</emphasis>
elements of “
<taxonomicName id="57AB44FDFFB1AA4562A2BC6EFF75F89A" baseAuthorityName="Johnson" baseAuthorityYear="1992" box="[163,238,1862,1885]" class="Insecta" family="Lycaenidae" genus="Penaincisalia" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="browni">
<emphasis id="A2DFE36CFFB1AA4562A2BC6EFF75F89A" box="[163,238,1862,1885]" italics="true" pageId="3" pageNumber="4">browni</emphasis>
</taxonomicName>
group” wing pattern indicating:
<emphasis id="A2DFE36CFFB1AA456052BC6EFDE3F89A" box="[595,632,1862,1885]" italics="true" pageId="3" pageNumber="4">DC</emphasis>
, disc cell;
<emphasis id="A2DFE36CFFB1AA4560F1BC6EFC8AF89A" box="[752,785,1862,1885]" italics="true" pageId="3" pageNumber="4">mb</emphasis>
, medial band;
<emphasis id="A2DFE36CFFB1AA4561B4BC6EFC49F89A" box="[949,978,1862,1885]" italics="true" pageId="3" pageNumber="4">sm</emphasis>
, submarginal band;
<emphasis id="A2DFE36CFFB1AA4566B0BC6EFB5FF89A" box="[1201,1220,1862,1885]" italics="true" pageId="3" pageNumber="4">m</emphasis>
, marginal band;
<emphasis id="A2DFE36CFFB1AA45677CBC6EFA0EF89A" box="[1405,1429,1862,1885]" italics="true" pageId="3" pageNumber="4">ds</emphasis>
, discal spot;
<emphasis id="A2DFE36CFFB1AA456319BC41FEA8F847" box="[280,307,1897,1920]" italics="true" pageId="3" pageNumber="4">pb</emphasis>
, postbasal pattern;
<emphasis id="A2DFE36CFFB1AA456005BC41FDBCF847" box="[516,551,1897,1920]" italics="true" pageId="3" pageNumber="4">BD</emphasis>
, basal disc;
<emphasis id="A2DFE36CFFB1AA4560AFBC41FD54F847" box="[686,719,1897,1920]" italics="true" pageId="3" pageNumber="4">dm</emphasis>
, disc margin;
<emphasis id="A2DFE36CFFB1AA456168BC41FC1CF847" box="[873,903,1897,1920]" italics="true" pageId="3" pageNumber="4">AF</emphasis>
, anal fold;
<emphasis id="A2DFE36CFFB1AA456602BC41FB84F847" box="[1027,1055,1897,1920]" italics="true" pageId="3" pageNumber="4">SL</emphasis>
, spatulate lobe.
<emphasis id="A2DFE36CFFB1AA4566CEBC40FB66F847" bold="true" box="[1231,1277,1896,1920]" pageId="3" pageNumber="4">(C):</emphasis>
wing shape of females indicating the different areas on the wing.
<emphasis id="A2DFE36CFFB1AA4560B1BCA3FD50F865" box="[688,715,1931,1954]" italics="true" pageId="3" pageNumber="4">ap</emphasis>
, apical;
<emphasis id="A2DFE36CFFB1AA456124BCA3FCA9F865" box="[805,818,1931,1954]" italics="true" pageId="3" pageNumber="4">b</emphasis>
, basal;
<emphasis id="A2DFE36CFFB1AA456185BCA3FC04F865" box="[900,927,1931,1954]" italics="true" pageId="3" pageNumber="4">pb</emphasis>
, postbasal;
<emphasis id="A2DFE36CFFB1AA45661CBCA3FBA1F865" box="[1053,1082,1931,1954]" italics="true" pageId="3" pageNumber="4">sm</emphasis>
, submedial;
<emphasis id="A2DFE36CFFB1AA4566C2BCA3FB4DF865" box="[1219,1238,1931,1954]" italics="true" pageId="3" pageNumber="4">m</emphasis>
, medial;
<emphasis id="A2DFE36CFFB1AA456738BCA3FAC1F865" box="[1337,1370,1931,1954]" italics="true" pageId="3" pageNumber="4">pm</emphasis>
, postmedial;
<emphasis id="A2DFE36CFFB1AA4562ECBC86FE83F802" box="[237,280,1966,1989]" italics="true" pageId="3" pageNumber="4">sma</emphasis>
, submarginal;
<emphasis id="A2DFE36CFFB1AA4563B6BC86FE51F802" box="[439,458,1966,1989]" italics="true" pageId="3" pageNumber="4">m</emphasis>
, marginal;
<emphasis id="A2DFE36CFFB1AA456045BC86FDC8F802" box="[580,595,1966,1989]" italics="true" pageId="3" pageNumber="4">L</emphasis>
, lateral lobe;
<emphasis id="A2DFE36CFFB1AA4560E4BC86FD6CF802" box="[741,759,1966,1989]" italics="true" pageId="3" pageNumber="4">ts</emphasis>
, tail.
</paragraph>
</caption>
<paragraph id="90143F7EFFB6AA4262C4BBE8FBE7FE95" blockId="4.[151,1436,152,495]" pageId="4" pageNumber="5">
3-Male androconial cluster comprised by a scent pad in the apical part of the discal cell and a minute scent patch situated distally at the erection of lower discocellular vein (
<figureCitation id="089023FBFFB6AA426197BBC0FB81FEC5" box="[918,1050,232,258]" captionStart="FIGURES 2633" captionStartId="7.[151,269,1907,1931]" captionTargetBox="[251,1334,1285,1877]" captionTargetId="figure-454@7.[248,1339,1277,1881]" captionTargetPageId="7" captionText="FIGURES 2633. Androconial clusters of the “browni group”. 26, P. browni; 27, P. saraha; 28, P. caeruleonota; 29, P. purpurea (Ecuador); 30, P. magnifica; 31, P.cuiva; 32, P.regala; 33. P. vittata." figureDoi="http://doi.org/10.5281/zenodo.5231021" httpUri="https://zenodo.org/record/5231021/files/figure.png" pageId="4" pageNumber="5">Figs 26-33</figureCitation>
). The scales in the scent pad and scent patch are arranged tightly and oriented almost perpendicularly to the wing membrane; they are identical in size (
<quantity id="5753929BFFB6AA4262F3BA10FEACFE96" box="[242,311,312,338]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="5" unit="m" value="100.0">100 µ</quantity>
in length and
<quantity id="5753929BFFB6AA4263DFBA10FD88FE96" box="[478,531,312,338]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="4" pageNumber="5" unit="m" value="30.0">30 µ</quantity>
in width) and nanostructures (Bálint
<emphasis id="A2DFE36CFFB6AA4261C9BA10FC67FE96" box="[968,1020,312,337]" italics="true" pageId="4" pageNumber="5">et al</emphasis>
., in prep.).
</paragraph>
<paragraph id="90143F7EFFB6AA4262C4BA48FF5EFE65" blockId="4.[151,1436,152,495]" pageId="4" pageNumber="5">
4-The male genital valva possesses a prominent, short triangular process at the ventral base (
<figureCitation id="089023FBFFB6AA426718BA48FF23FE65" captionStart="FIGURES 3443" captionStartId="9.[151,269,1939,1963]" captionTargetBox="[204,1383,680,1904]" captionTargetId="figure-258@9.[194,1392,674,1916]" captionTargetPageId="9" captionText="FIGURES 3443. Left valva in lateral view. 34, P. browni; 35, P. saraha; 36, P. caeruleonota (Loja, Ecuador) indicating the ventral keel; 37, P. caeruleonota (“Volcan Cabogana”, Ecuador); 38, P. vittata; 39, P.regala; 40, P. purpurea (Ecuador); 41, P. purpurea (Perú); 42, P.cuiva; 43, P. magnifica." figureDoi="http://doi.org/10.5281/zenodo.5231023" httpUri="https://zenodo.org/record/5231023/files/figure.png" pageId="4" pageNumber="5">Figure 34 43</figureCitation>
).
</paragraph>
<paragraph id="90143F7EFFB6AA4262C7BA98FCB1FE28" blockId="4.[151,1436,152,495]" pageId="4" pageNumber="5">
5-Female genital posterior portion of ductus bursae shorter than anterior portion of ductus bursae (
<figureCitation id="089023FBFFB6AA426743BA98FF78FE28" captionStart="FIGURES 4449" captionStartId="10.[151,269,1967,1991]" captionTargetBox="[260,1338,197,1934]" captionTargetId="figure-66@10.[251,1343,194,1940]" captionTargetPageId="10" captionText="FIGURES 4449. Female genital structures of the “browni group”. 44, P. browni (indicating the relation of posterior (p) and anterior (a) regions of the ductus bursae; 45, P. saraha; 46, P. purpurea; 47, P. magnifica; 48, P.cuiva; 49, P. vittata." figureDoi="http://doi.org/10.5281/zenodo.5231025" httpUri="https://zenodo.org/record/5231025/files/figure.png" pageId="4" pageNumber="5">Figures 4449</figureCitation>
; anterior and posterior portion defined in fig. 44).
</paragraph>
<caption id="C4D46FF6FFB6AA426296B93FFE47FD96" pageId="4" pageNumber="5" startId="4.[151,241,535,559]">
<paragraph id="90143F7EFFB6AA426296B93FFE47FD96" blockId="4.[151,1436,535,593]" pageId="4" pageNumber="5">
<emphasis id="A2DFE36CFFB6AA426296B93FFE96FDE8" bold="true" box="[151,269,535,559]" pageId="4" pageNumber="5">TABLE 1.</emphasis>
Distribution of characters (as numbered in the section “the
<taxonomicName id="57AB44FDFFB6AA4261ADB930FC6CFDE8" baseAuthorityName="Johnson" baseAuthorityYear="1992" box="[940,1015,536,559]" class="Insecta" family="Lycaenidae" genus="Penaincisalia" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="browni">
<emphasis id="A2DFE36CFFB6AA4261ADB930FC6CFDE8" box="[940,1015,536,559]" italics="true" pageId="4" pageNumber="5">browni</emphasis>
</taxonomicName>
species group), codes: present = “+”, absent = “-“,? = non available
</paragraph>
</caption>
<paragraph id="90143F7EFFB6AA42629EB95AFB4FFC46" pageId="4" pageNumber="5">
<table id="E2ABCDDEFFB655B9629EB95AFAD6FC46" box="[159,1357,626,897]" gridcols="7" gridrows="6" pageId="4" pageNumber="5">
<tr id="2E9B3D3CFFB655B9629EB95AFAD6FD6B" box="[159,1357,626,684]" gridrow="0" pageId="4" pageNumber="5">
<th id="6D4A5440FFB655B9629EB95AFEAEFD6B" box="[159,309,626,684]" gridcol="0" gridrow="0" pageId="4" pageNumber="5">
Species group (
<bibRefCitation id="F43A428FFFB6AA4262A7B9BDFEBEFD6B" author="Prieto, C." box="[166,293,661,684]" pageId="4" pageNumber="5" refId="ref12539" refString="Prieto, C. (2008) Taxonomia, Biogeografia y Relaciones Filogeneticas de Genero Altoandino PENAINCISALIA Johnson (Lepidoptera: Lycaenidae: Eumaeini). Unpublished Ph. D. dissertation, Universidad de Alicante, Espana [available in the HNHM library]." type="book" year="2008">Prieto 2008</bibRefCitation>
)
</th>
<th id="6D4A5440FFB655B9639CB95AFE73FD6B" box="[413,488,626,684]" gridcol="1" gridrow="0" pageId="4" pageNumber="5">
<taxonomicName id="57AB44FDFFB6AA42639CB95AFE73FD4E" baseAuthorityName="Johnson" baseAuthorityYear="1992" box="[413,488,626,649]" class="Insecta" family="Lycaenidae" genus="Penaincisalia" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="browni">
<emphasis id="A2DFE36CFFB6AA42639CB95AFE73FD4E" box="[413,488,626,649]" italics="true" pageId="4" pageNumber="5">browni</emphasis>
</taxonomicName>
</th>
<th id="6D4A5440FFB655B9602EB95AFDF7FD6B" box="[559,620,626,684]" gridcol="2" gridrow="0" pageId="4" pageNumber="5">
<taxonomicName id="57AB44FDFFB6AA42602EB95AFDF7FD4E" box="[559,620,626,649]" class="Insecta" family="Lycaenidae" genus="Penaincisalia" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="juliae">
<emphasis id="A2DFE36CFFB6AA42602EB95AFDF7FD4E" box="[559,620,626,649]" italics="true" pageId="4" pageNumber="5">juliae</emphasis>
</taxonomicName>
</th>
<th id="6D4A5440FFB655B960A8B95AFC98FD6B" box="[681,771,626,684]" gridcol="3" gridrow="0" pageId="4" pageNumber="5">
<emphasis id="A2DFE36CFFB6AA4260A8B95AFC98FD4E" box="[681,771,626,649]" italics="true" pageId="4" pageNumber="5">loxurina</emphasis>
</th>
<th id="6D4A5440FFB655B9614CB95AFC50FD6B" box="[845,971,626,684]" gridcol="4" gridrow="0" pageId="4" pageNumber="5">
<emphasis id="A2DFE36CFFB6AA42614CB95AFC50FD4E" box="[845,971,626,649]" italics="true" pageId="4" pageNumber="5">culminicola</emphasis>
</th>
<th id="6D4A5440FFB655B96627B95AFB19FD6B" box="[1062,1154,626,684]" gridcol="5" gridrow="0" pageId="4" pageNumber="5">
<emphasis id="A2DFE36CFFB6AA426627B95AFB19FD4E" box="[1062,1154,626,649]" italics="true" pageId="4" pageNumber="5">amatista</emphasis>
</th>
<th id="6D4A5440FFB655B966CAB95AFAD6FD6B" box="[1227,1357,626,684]" gridcol="6" gridrow="0" pageId="4" pageNumber="5">
<emphasis id="A2DFE36CFFB6AA4266CAB95AFAD6FD4E" box="[1227,1357,626,649]" italics="true" pageId="4" pageNumber="5">balzapamba</emphasis>
</th>
</tr>
<tr id="2E9B3D3CFFB655B9629EB9E8FAD6FD10" box="[159,1357,704,727]" gridrow="1" pageId="4" pageNumber="5">
<th id="6D4A5440FFB655B9629EB9E8FEAEFD10" box="[159,309,704,727]" gridcol="0" gridrow="1" pageId="4" pageNumber="5">Character 1</th>
<td id="6D4A5440FFB655B9639CB9E8FE73FD10" box="[413,488,704,727]" gridcol="1" gridrow="1" pageId="4" pageNumber="5">+</td>
<td id="6D4A5440FFB655B9602EB9E8FDF7FD10" box="[559,620,704,727]" gridcol="2" gridrow="1" pageId="4" pageNumber="5">+</td>
<td id="6D4A5440FFB655B960A8B9E8FC98FD10" box="[681,771,704,727]" gridcol="3" gridrow="1" pageId="4" pageNumber="5">-</td>
<td id="6D4A5440FFB655B9614CB9E8FC50FD10" box="[845,971,704,727]" gridcol="4" gridrow="1" pageId="4" pageNumber="5">-</td>
<td id="6D4A5440FFB655B96627B9E8FB19FD10" box="[1062,1154,704,727]" gridcol="5" gridrow="1" pageId="4" pageNumber="5">-</td>
<td id="6D4A5440FFB655B966CAB9E8FAD6FD10" box="[1227,1357,704,727]" gridcol="6" gridrow="1" pageId="4" pageNumber="5">-</td>
</tr>
<tr id="2E9B3D3CFFB655B9629EB9C2FAD6FCC6" box="[159,1357,746,769]" gridrow="2" pageId="4" pageNumber="5">
<th id="6D4A5440FFB655B9629EB9C2FEAEFCC6" box="[159,309,746,769]" gridcol="0" gridrow="2" pageId="4" pageNumber="5">Character 2</th>
<td id="6D4A5440FFB655B9639CB9C2FE73FCC6" box="[413,488,746,769]" gridcol="1" gridrow="2" pageId="4" pageNumber="5">+</td>
<td id="6D4A5440FFB655B9602EB9C2FDF7FCC6" box="[559,620,746,769]" gridcol="2" gridrow="2" pageId="4" pageNumber="5">?</td>
<td id="6D4A5440FFB655B960A8B9C2FC98FCC6" box="[681,771,746,769]" gridcol="3" gridrow="2" pageId="4" pageNumber="5">-</td>
<td id="6D4A5440FFB655B9614CB9C2FC50FCC6" box="[845,971,746,769]" gridcol="4" gridrow="2" pageId="4" pageNumber="5">-</td>
<td id="6D4A5440FFB655B96627B9C2FB19FCC6" box="[1062,1154,746,769]" gridcol="5" gridrow="2" pageId="4" pageNumber="5">-</td>
<td id="6D4A5440FFB655B966CAB9C2FAD6FCC6" box="[1227,1357,746,769]" gridcol="6" gridrow="2" pageId="4" pageNumber="5">-</td>
</tr>
<tr id="2E9B3D3CFFB655B9629EB83DFAD6FCEB" box="[159,1357,789,812]" gridrow="3" pageId="4" pageNumber="5">
<th id="6D4A5440FFB655B9629EB83DFEAEFCEB" box="[159,309,789,812]" gridcol="0" gridrow="3" pageId="4" pageNumber="5">Character 3</th>
<td id="6D4A5440FFB655B9639CB83DFE73FCEB" box="[413,488,789,812]" gridcol="1" gridrow="3" pageId="4" pageNumber="5">+</td>
<td id="6D4A5440FFB655B9602EB83DFDF7FCEB" box="[559,620,789,812]" gridcol="2" gridrow="3" pageId="4" pageNumber="5">+</td>
<td id="6D4A5440FFB655B960A8B83DFC98FCEB" box="[681,771,789,812]" gridcol="3" gridrow="3" pageId="4" pageNumber="5">+</td>
<td id="6D4A5440FFB655B9614CB83DFC50FCEB" box="[845,971,789,812]" gridcol="4" gridrow="3" pageId="4" pageNumber="5">+</td>
<td id="6D4A5440FFB655B96627B83DFB19FCEB" box="[1062,1154,789,812]" gridcol="5" gridrow="3" pageId="4" pageNumber="5">+</td>
<td id="6D4A5440FFB655B966CAB83DFAD6FCEB" box="[1227,1357,789,812]" gridcol="6" gridrow="3" pageId="4" pageNumber="5">-</td>
</tr>
<tr id="2E9B3D3CFFB655B9629EB868FAD6FC90" box="[159,1357,832,855]" gridrow="4" pageId="4" pageNumber="5">
<th id="6D4A5440FFB655B9629EB868FEAEFC90" box="[159,309,832,855]" gridcol="0" gridrow="4" pageId="4" pageNumber="5">Character 4</th>
<td id="6D4A5440FFB655B9639CB868FE73FC90" box="[413,488,832,855]" gridcol="1" gridrow="4" pageId="4" pageNumber="5">+</td>
<td id="6D4A5440FFB655B9602EB868FDF7FC90" box="[559,620,832,855]" gridcol="2" gridrow="4" pageId="4" pageNumber="5">-</td>
<td id="6D4A5440FFB655B960A8B868FC98FC90" box="[681,771,832,855]" gridcol="3" gridrow="4" pageId="4" pageNumber="5">-,+</td>
<td id="6D4A5440FFB655B9614CB868FC50FC90" box="[845,971,832,855]" gridcol="4" gridrow="4" pageId="4" pageNumber="5">-</td>
<td id="6D4A5440FFB655B96627B868FB19FC90" box="[1062,1154,832,855]" gridcol="5" gridrow="4" pageId="4" pageNumber="5">-</td>
<td id="6D4A5440FFB655B966CAB868FAD6FC90" box="[1227,1357,832,855]" gridcol="6" gridrow="4" pageId="4" pageNumber="5">-</td>
</tr>
<tr id="2E9B3D3CFFB655B9629EB842FAD6FC46" box="[159,1357,874,897]" gridrow="5" pageId="4" pageNumber="5">
<th id="6D4A5440FFB655B9629EB842FEAEFC46" box="[159,309,874,897]" gridcol="0" gridrow="5" pageId="4" pageNumber="5">Character 5</th>
<td id="6D4A5440FFB655B9639CB842FE73FC46" box="[413,488,874,897]" gridcol="1" gridrow="5" pageId="4" pageNumber="5">+</td>
<td id="6D4A5440FFB655B9602EB842FDF7FC46" box="[559,620,874,897]" gridcol="2" gridrow="5" pageId="4" pageNumber="5">-</td>
<td id="6D4A5440FFB655B960A8B842FC98FC46" box="[681,771,874,897]" gridcol="3" gridrow="5" pageId="4" pageNumber="5">-</td>
<td id="6D4A5440FFB655B9614CB842FC50FC46" box="[845,971,874,897]" gridcol="4" gridrow="5" pageId="4" pageNumber="5">-</td>
<td id="6D4A5440FFB655B96627B842FB19FC46" box="[1062,1154,874,897]" gridcol="5" gridrow="5" pageId="4" pageNumber="5">-</td>
<td id="6D4A5440FFB655B966CAB842FAD6FC46" box="[1227,1357,874,897]" gridcol="6" gridrow="5" pageId="4" pageNumber="5">-</td>
</tr>
</table>
</paragraph>
<paragraph id="90143F7EFFB6AA426296B882FABDFB8B" blockId="4.[151,1436,938,1100]" pageId="4" pageNumber="5">Characters: 1 = Hindwing tornus in males appears as a spatulate and bulbous lobe. 2 = Sexual dimorphism in wing shape, where the female has a long tail from the CuA2 vein, accompanied for a small lateral lobe in the hindwing anal corner. 3 = Male androconial cluster comprised by a scent pad in the apical part of the discal cell and a minute scent patch situated distally at the erection of lower discocellular vein. 4 = male genital valvae possesses a prominent, short triangular ventral process in the ventral edge. 5 = female genital ductus bursae posterior portion is shorter than the anterior one.</paragraph>
</subSubSection>
</treatment>
</document>