217 lines
23 KiB
XML
217 lines
23 KiB
XML
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<mods:title id="65550B683FE74F2DC57EEE313FBD4778">Systematics of the family Plectopylidae in Vietnam with additional information on Chinese taxa (Gastropoda, Pulmonata, Stylommatophora)</mods:title>
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<mods:namePart id="A8246CA48BF02B26759C45F8EEA8587E">Pall-Gergely, Barna</mods:namePart>
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<mods:namePart id="03139AD15DFC464B34F6E7E5A347C999">Hunyadi, Andras</mods:namePart>
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<mods:namePart id="9E7BD6B8D321FB87C0B5CA8D4AC05AF6">Ablett, Jonathan</mods:namePart>
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<mods:namePart id="BA338FA7E0A75C38E5C5B34B528ADF4B">Lương, Hao Văn</mods:namePart>
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<mods:namePart id="4D394E3BC805610A29A7C3AB1A94604E">Fred Naggs,</mods:namePart>
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<mods:namePart id="6669CC25B3EA8FCE443C03D320600831">Asami, Takahiro</mods:namePart>
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<mods:title id="CE2DB8FBFE389656EAA069F0632A501C">ZooKeys</mods:title>
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<mods:date id="1C2A44DC2A71E3F5E655D31256FBC499">2015</mods:date>
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<mods:number id="6932ABFB9F0B5A4E6AF8DE9B810BF3A8">473</mods:number>
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<mods:classification id="0BEB09F62FCC1B4946D7EB6A73FAC1E4">journal article</mods:classification>
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<mods:identifier id="57084D06D803A836AAEBDA45939404BD" type="DOI">http://dx.doi.org/10.3897/zookeys.473.8659</mods:identifier>
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<treatment id="D2A8FD5904B4166E88CC9D89A809CC4E" ID-GBIF-Taxon="152057797" LSID="urn:lsid:plazi:treatment:D2A8FD5904B4166E88CC9D89A809CC4E" httpUri="http://treatment.plazi.org/id/D2A8FD5904B4166E88CC9D89A809CC4E" lastPageId="23" lastPageNumber="23" pageId="21" pageNumber="22">
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<subSubSection id="33F67818179245365CCA08FC0379C783" pageId="21" pageNumber="22" type="multiple">
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<paragraph id="E88F2FC72E12102EEEF7C0429710072A" pageId="21" pageNumber="22">Taxon classification Animalia Stylommatophora Plectopylidae</paragraph>
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</subSubSection>
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<subSubSection id="772A4B2B52F6598BD8D171D28E236002" pageId="21" pageNumber="22" type="nomenclature">
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<paragraph id="8D39D8BCFE3B4D67F3CC76AFEFB39E3F" pageId="21" pageNumber="22">
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<taxonomicName id="9DFDE1FE28C47CDB5458FEDAF632C3F0" ID-CoL="3HK4G" authority="Gude, 1908" authorityName="Gude" authorityYear="1908" class="Gastropoda" family="Plectopylidae" genus="Gudeodiscus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gudeodiscus (Gudeodiscus) infralevis" order="Stylommatophora" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="infralevis" subGenus="Gudeodiscus">Gudeodiscus (Gudeodiscus?) infralevis (Gude, 1908)</taxonomicName>
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Figures 3
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<normalizedToken id="8173073F992D31E28DB723B880ED71FC" originalValue="D–E">D-E</normalizedToken>
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, 15
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<normalizedToken id="A9E5616C72B6C8D73F45A3C980B52E01" originalValue="A–D">A-D</normalizedToken>
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</paragraph>
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</subSubSection>
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<subSubSection id="25DB7D4F90529C8A82379770D3195955" pageId="22" pageNumber="23" type="reference_group">
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<paragraph id="1BB9E9F9BFC91CDC6691D4C179AB5B27" pageId="22" pageNumber="23">
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<taxonomicName id="5BCBB0199C463C42BDA2AE9DB1321610" class="Gastropoda" family="Plectopylidae" genus="Plectopylis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plectopylis infralevis" order="Stylommatophora" pageId="22" pageNumber="23" phylum="Mollusca" rank="species" species="infralevis">
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<pageBreakToken id="67FB4010CFE7B3DB759FFEA4FB63CDE0" pageId="22" pageNumber="23" start="start">Plectopylis</pageBreakToken>
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infralevis
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</taxonomicName>
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Gude 1908, Journal de Conchyliologie, 55: 345, 350, 352-353., Figs 3
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<normalizedToken id="606312D9FFF761A92E741A03E248054D" originalValue="a–e">a-e</normalizedToken>
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, Plate 7, Figs 4-6. ["Quang Huyen"].
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</paragraph>
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<paragraph id="113581B348786B4058B1EB4ED0093338" pageId="22" pageNumber="23">
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<taxonomicName id="8A64A28024C7F1DC89BEE9937C4E40AE" class="Gastropoda" family="Plectopylidae" genus="Plectopylis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plectopylis soror" order="Stylommatophora" pageId="22" pageNumber="23" phylum="Mollusca" rank="species" species="soror">Plectopylis soror</taxonomicName>
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Gude 1908, syn. n., Journal de Conchyliologie, 55: 355-357., Figs 5
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<normalizedToken id="60AAF63038B0791D54C91FFFCE21712D" originalValue="a–e">a-e</normalizedToken>
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, Plate 7, Figs 10-12. ["Quang Huyen"].
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</paragraph>
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<paragraph id="1A8B76B25DBB819478D59A5AFA5215C4" pageId="22" pageNumber="23">
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<taxonomicName id="6FDD76A8AD7947518D09BCD811263973" class="Gastropoda" family="Plectopylidae" genus="Gudeodiscus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gudeodiscus infralevis" order="Stylommatophora" pageId="22" pageNumber="23" phylum="Mollusca" rank="species" species="infralevis">Gudeodiscus infralevis</taxonomicName>
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, -
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<normalizedToken id="F134850D6488C95DFC54B582A1C302BC" originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
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& Hunyadi 2013, Archiv
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<normalizedToken id="C1965A727C5BA34E55A19843D1D2046B" originalValue="für">fuer</normalizedToken>
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Molluskenkunde, 142 (1): 8.
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</paragraph>
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<paragraph id="C68D50CC220C963148978122AC02A665" pageId="22" pageNumber="23">
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<taxonomicName id="2E9FB6EFA016BCA0BD441CF4176781FE" class="Gastropoda" family="Plectopylidae" genus="Gudeodiscus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gudeodiscus soror" order="Stylommatophora" pageId="22" pageNumber="23" phylum="Mollusca" rank="species" species="soror">Gudeodiscus soror</taxonomicName>
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, -
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<normalizedToken id="3CF6ABF970E9A27DC6C1EAEE88876B6E" originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
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& Hunyadi 2013, Archiv
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<normalizedToken id="C55DBEBB9D9555472DEDD0B59BCB0C0C" originalValue="für">fuer</normalizedToken>
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Molluskenkunde, 142 (1): 8.
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</paragraph>
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</subSubSection>
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<subSubSection id="3AD2587F0075622994FF03B864411E53" pageId="22" pageNumber="23" type="types examined">
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<paragraph id="A0E952C96CFC8AC47B4941297B56494B" pageId="22" pageNumber="23">Types examined.</paragraph>
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<paragraph id="AA02F02A9B2858D1A4023F27552E8806" pageId="22" pageNumber="23">
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Tonkin, Quang-Huyen, leg. Mansuy, MNHN 24604 (holotype of
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<taxonomicName id="BE6773710747FBB97832133FC71987F0" class="Insecta" family="Hybotidae" genus="Euthyneura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Euthyneura infralevis" order="Diptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="infralevis">infralevis</taxonomicName>
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, Figure 3D); Tonkin, Quang-Huyen, leg. Mansuy, MNHN 24585 (holotype of
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<taxonomicName id="75606081F0241DE6EA009179B501B1E8" class="Insecta" family="Hybotidae" genus="Euthyneura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Euthyneura soror" order="Diptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="soror">soror</taxonomicName>
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, Figure 3E).
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</paragraph>
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</subSubSection>
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<subSubSection id="3E7C544D134E07512D17887D526179A6" pageId="22" pageNumber="23" type="diagnosis">
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<paragraph id="3C34381D60686A816BE0F9915493670F" pageId="22" pageNumber="23">Diagnosis.</paragraph>
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<paragraph id="0852B87E9E62249AAFA469E75D74893C" pageId="22" pageNumber="23">Shell small, solid, discoid, with elevated spire, relatively deep umbilicus; relatively thin apertural lip and rather parallel, thick, straight palatal plicae. See also under remarks.</paragraph>
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</subSubSection>
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<subSubSection id="5A9C80A66E5D57BBCF542F26F5F83F02" pageId="22" pageNumber="23" type="measurements">
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<paragraph id="33B4846F37CBE2F46D265BDFAF7FB199" pageId="22" pageNumber="23">Measurements</paragraph>
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<paragraph id="99F3E835B87EFCD538265BB18A38DE25" pageId="22" pageNumber="23">
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(in mm): D = 13.9, D = 6.7 (
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<taxonomicName id="7CF4D080F2A7DF22F6FB1DB804013EC1" class="Insecta" family="Hybotidae" genus="Euthyneura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Euthyneura soror" order="Diptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="soror">soror</taxonomicName>
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holotype); D = 13.5, H = 6.6 (
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<taxonomicName id="52311E797FB4197189E641849CDCF107" class="Insecta" family="Hybotidae" genus="Euthyneura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Euthyneura infralevis" order="Diptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="infralevis">infralevis</taxonomicName>
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holotype).
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</paragraph>
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</subSubSection>
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<subSubSection id="16A1D6DE1B3CAEAEB09D3AB3437D1B47" pageId="22" pageNumber="23" type="differential diagnosis">
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<paragraph id="D6A72D589B2B0F0984AC79367A0F2E74" pageId="22" pageNumber="23">Differential diagnosis.</paragraph>
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<paragraph id="D2DA7B4AB7FA9626F34FC1AA27086621" pageId="22" pageNumber="23">
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Our knowledge on the intraspecific variety of the species is very limited (see Remarks). It seems that the thick, rather horizontal palatal plicae, the strong basal sculpture and the elevated spire distinguishes the species from the similar species (
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<taxonomicName id="2C1278E908AE5F2F1536FE4B38072C01" class="Gastropoda" family="Plectopylidae" genus="Gudeodiscus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gudeodiscus eroessi" order="Stylommatophora" pageId="22" pageNumber="23" phylum="Mollusca" rank="species" species="eroessi">Gudeodiscus eroessi</taxonomicName>
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,
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<taxonomicName id="AC9E2D38FCC8144159065BF09DF5C34F" class="Gastropoda" family="Plectopylidae" genus="Gudeodiscus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gudeodiscus multispira" order="Stylommatophora" pageId="22" pageNumber="23" phylum="Mollusca" rank="species" species="multispira">Gudeodiscus multispira</taxonomicName>
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,
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<taxonomicName id="4E3C716C115E7F9879EE6960207D50AB" class="Gastropoda" family="Plectopylidae" genus="Gudeodiscus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gudeodiscus soosi" order="Stylommatophora" pageId="22" pageNumber="23" phylum="Mollusca" rank="species" species="soosi">Gudeodiscus soosi</taxonomicName>
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,
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<taxonomicName id="980E5CD21666156FE291842A0ECDB738" class="Gastropoda" family="Plectopylidae" genus="Gudeodiscus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gudeodiscus yunnanensis" order="Stylommatophora" pageId="22" pageNumber="23" phylum="Mollusca" rank="species" species="yunnanensis">Gudeodiscus yunnanensis</taxonomicName>
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,
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<taxonomicName id="349888B609C0F27D719608C00BB6705A" class="Gastropoda" family="Plectopylidae" genus="Gudeodiscus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gudeodiscus cyrtochilus" order="Stylommatophora" pageId="22" pageNumber="23" phylum="Mollusca" rank="species" species="cyrtochilus">Gudeodiscus cyrtochilus</taxonomicName>
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and
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<taxonomicName id="CA86F3646AD133E3C60126B9BFBCEABD" class="Gastropoda" family="Plectopylidae" genus="Gudeodiscus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gudeodiscus fischeri" order="Stylommatophora" pageId="22" pageNumber="23" phylum="Mollusca" rank="species" species="fischeri">Gudeodiscus fischeri</taxonomicName>
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). The shell and aperture shape suggest that the closest relatives are
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<taxonomicName id="8315DCE600EEB00BD7A5B1470472F4D9" class="Gastropoda" family="Plectopylidae" genus="Gudeodiscus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gudeodiscus fischeri" order="Stylommatophora" pageId="22" pageNumber="23" phylum="Mollusca" rank="species" species="fischeri">Gudeodiscus fischeri</taxonomicName>
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and
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<taxonomicName id="A501DC6EEBA63F7E22A20AE7402741DF" class="Gastropoda" family="Plectopylidae" genus="Gudeodiscus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gudeodiscus suprafilaris" order="Stylommatophora" pageId="22" pageNumber="23" phylum="Mollusca" rank="species" species="suprafilaris">Gudeodiscus suprafilaris</taxonomicName>
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(see comparisons under those species).
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</paragraph>
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</subSubSection>
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<subSubSection id="EB0DAD7B13D104AAECFF8B3AE8EE5820" pageId="22" pageNumber="23" type="intraspecific diversity">
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<paragraph id="E367E5C86B1213B53B5B66955302489B" pageId="22" pageNumber="23">Intraspecific diversity.</paragraph>
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<paragraph id="A9CCCAE4EBDD8E8E7A3D2218191AB2C6" pageId="22" pageNumber="23">
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<taxonomicName id="7D2473AB548161A26D4FFE1E65F977ED" class="Gastropoda" family="Plectopylidae" genus="Plectopylis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plectopylis infralevis" order="Stylommatophora" pageId="22" pageNumber="23" phylum="Mollusca" rank="species" species="infralevis">Plectopylis infralevis</taxonomicName>
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and
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<taxonomicName id="B3027447F2C08ACCD9721AF4C64149DD" class="Gastropoda" family="Plectopylidae" genus="Plectopylis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plectopylis soror" order="Stylommatophora" pageId="22" pageNumber="23" phylum="Mollusca" rank="species" species="soror">Plectopylis soror</taxonomicName>
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are considered as conspecific (see Remarks). Only the holotypes of these taxa are known, therefore our knowledge on the intraspecific variability is limited.
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</paragraph>
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</subSubSection>
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<subSubSection id="077C847F3B1FE584A4681318038A6397" pageId="22" pageNumber="23" type="distribution">
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<paragraph id="90987EA6253DDFF93A414E9C4A34655B" pageId="22" pageNumber="23">Distribution.</paragraph>
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<paragraph id="D5A9B1623B3D00264D7D9F10B5954ACF" pageId="22" pageNumber="23">
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The type specimens of
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<taxonomicName id="E359AB5A5C21A1B972359696C8B7F205" class="Gastropoda" family="Plectopylidae" genus="Plectopylis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plectopylis infralevis" order="Stylommatophora" pageId="22" pageNumber="23" phylum="Mollusca" rank="species" species="infralevis">Plectopylis infralevis</taxonomicName>
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and
|
||
<taxonomicName id="26BAB661D4753D7688D9EE09F27517FD" class="Gastropoda" family="Plectopylidae" genus="Plectopylis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plectopylis soror" order="Stylommatophora" pageId="22" pageNumber="23" phylum="Mollusca" rank="species" species="soror">Plectopylis soror</taxonomicName>
|
||
(synonym of
|
||
<taxonomicName id="981DBFF10C04E163C56308B44135D062" class="Insecta" family="Hybotidae" genus="Euthyneura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Euthyneura infralevis" order="Diptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="infralevis">infralevis</taxonomicName>
|
||
) were collected in Quang Huyen (Quảng
|
||
<normalizedToken id="477FE98D6888AB00D99742FD5A2D02D7" originalValue="Uyên">Uyen</normalizedToken>
|
||
) (see Figure 39).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="BACC3583428F87E1212276561F0E1454" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23" type="remarks">
|
||
<paragraph id="9FF1E64FEED4C53A5C0F9DD0F37C62B7" pageId="22" pageNumber="23">Remarks.</paragraph>
|
||
<paragraph id="AE72D7D41316573B19FDAE430E4B60D1" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23">
|
||
Only the holotypes of
|
||
<taxonomicName id="7B8E4302620383240D9319157875993D" class="Gastropoda" family="Plectopylidae" genus="Plectopylis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plectopylis infralevis" order="Stylommatophora" pageId="23" pageNumber="24" phylum="Mollusca" rank="species" species="infralevis">
|
||
<pageBreakToken id="B592542DCE4E2295DF978E048A032DA4" pageId="23" pageNumber="24" start="start">Plectopylis</pageBreakToken>
|
||
infralevis
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="33D5ED36A105C14AA2FC8226E36326E8" class="Gastropoda" family="Plectopylidae" genus="Plectopylis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plectopylis soror" order="Stylommatophora" pageId="23" pageNumber="24" phylum="Mollusca" rank="species" species="soror">Plectopylis soror</taxonomicName>
|
||
are known. The notable differences between these two shells are the stronger sculpture, slightly shouldered body whorl and small apertural fold in
|
||
<taxonomicName id="4EBDE312F499F7BF97E68C0F41EE6E60" class="Insecta" family="Hybotidae" genus="Euthyneura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Euthyneura soror" order="Diptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="soror">soror</taxonomicName>
|
||
. Additionally, there are three lamellae in
|
||
<taxonomicName id="059F46F9DE3D0BD8F0E3E3B1550D94F6" class="Insecta" family="Hybotidae" genus="Euthyneura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Euthyneura infralevis" order="Diptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="infralevis">infralevis</taxonomicName>
|
||
versus only one in
|
||
<taxonomicName id="25258F1AC71657B4F1792A92BCB88E85" class="Insecta" family="Hybotidae" genus="Euthyneura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Euthyneura soror" order="Diptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="soror">soror</taxonomicName>
|
||
. The three vertical lamellae in the holotype of
|
||
<taxonomicName id="D08E8C546C5101CCFD6246D2792A56B9" class="Insecta" family="Hybotidae" genus="Euthyneura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Euthyneura infralevis" order="Diptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="infralevis">infralevis</taxonomicName>
|
||
is possibly the result of abnormal development. No other species of
|
||
<taxonomicName id="2BF57C0CF9B842B9A5AD925D62DCA802" family="Plectopylidae" lsidName="" pageId="23" pageNumber="24" rank="family">Plectopylidae</taxonomicName>
|
||
has three lamellae. Similar abnormal shells have been reported in
|
||
<taxonomicName id="531C250F632035ABEEC9F8FD8732A152" class="Gastropoda" family="Plectopylidae" genus="Gudeodiscus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gudeodiscus giardi" order="Stylommatophora" pageId="23" pageNumber="24" phylum="Mollusca" rank="species" species="giardi">Gudeodiscus giardi</taxonomicName>
|
||
(see
|
||
<bibRefCitation id="115C7A01CE73216C8CB8FFEFE3C4557E" author="Gude, GK" journalOrPublisher="Journal de Conchyliologie" pageId="82" pageNumber="83" pagination="345 - 357" title="Observation on a number of Plectopylis collected in Tonkin by M. Mansuy with description of four new species." volume="55" year="1908">Gude 1908</bibRefCitation>
|
||
). Consequently, we do not know what the characteristic type of parietal lamellae in this species is (=one or two). The differences between the two specimens suggest only intraspecific variance. Unfortunately we have no freshly-collected material of these two forms, but because of the high similarity between the two holotypes and same type locality we here synonymise
|
||
<taxonomicName id="CDA1CD3404EBD14A755167BFB4F523AC" class="Insecta" family="Hybotidae" genus="Euthyneura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Euthyneura soror" order="Diptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="soror">soror</taxonomicName>
|
||
with
|
||
<taxonomicName id="51D5728E07DD37A6A8D555D4B9BD3AA5" class="Insecta" family="Hybotidae" genus="Euthyneura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Euthyneura infralevis" order="Diptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="infralevis">infralevis</taxonomicName>
|
||
. These two names were published in the same paper (
|
||
<bibRefCitation id="AC090A5699BF4F5B10707A187436A3F7" author="Gude, GK" journalOrPublisher="Journal de Conchyliologie" pageId="82" pageNumber="83" pagination="345 - 357" title="Observation on a number of Plectopylis collected in Tonkin by M. Mansuy with description of four new species." volume="55" year="1908">Gude 1908</bibRefCitation>
|
||
), but
|
||
<taxonomicName id="D6648A1164CD6EF60AEB2768F910F54B" class="Insecta" family="Hybotidae" genus="Euthyneura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Euthyneura infralevis" order="Diptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="infralevis">infralevis</taxonomicName>
|
||
was described earlier in terms of page numbers.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |