treatments-xml/data/D2/7B/87/D27B87A3FFB2F134FEA9FC2692F5FC86.xml
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<mods:title id="44F8F3CEC6AD6D7575E9CDEB0B60F7FF">The Isoperla Of California (Plecoptera: Perlodidae); Larval Descriptions And A Key To 17 Western Nearctic Species</mods:title>
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<taxonomicName id="9DD24D36FFB2F132FEA9FC2691C4FBD8" ID-CoL="6MZYY" authority="Szczytko &amp; Stewart" authorityName="Szczytko &amp; Stewart" authorityYear="2004" box="[292,694,1011,1035]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="35" pageNumber="237" phylum="Arthropoda" rank="species" species="muir">
<emphasis id="68A6EAA7FFB2F132FEA9FC2692B6FBD8" bold="true" box="[292,452,1011,1035]" italics="true" pageId="35" pageNumber="237">Isoperla muir</emphasis>
Szczytko &amp; Stewart
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</heading>
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<subSubSection id="12C8653EFFB2F132FEF8FBC29114FBFC" box="[373,614,1047,1071]" pageId="35" pageNumber="237" type="description">
<paragraph id="5A6D36B5FFB2F132FEF8FBC29114FBFC" blockId="35.[292,694,1011,1071]" box="[373,614,1047,1071]" pageId="35" pageNumber="237">
(
<figureCitation id="C2E92A30FFB2F132FEF0FBC292A8FBFC" box="[381,474,1047,1071]" captionStart="Figs" captionStartId="9.[189,238,1823,1847]" captionTargetBox="[249,1383,235,1762]" captionTargetId="figure-57@9.[249,1383,235,1762]" captionTargetPageId="9" captionText="Figs. 2a-q. Isoperla lacinia habitus, 40x, Bar = 500µm. 2a. I. acula; 2b. I. adunca; 2c. I. baumanni; 2d. I. bifurcata; 2e. I. denningi; 2f. I. fulva; 2g. I. laucki; 2h. I. marmorata; 2i. I. miwok; 2j. I. mormona; 2k. I. muir; 2L. I. pinta; 2m. I. quinquepunctata; 2n. I. roguensis; 2o. I. sobria; 2p. I. sordida; 2q. I. tilasqua." figureDoi="http://doi.org/10.5281/zenodo.4760326" httpUri="https://zenodo.org/record/4760326/files/figure.png" pageId="35" pageNumber="237">Figs. 2k</figureCitation>
,
<figureCitation id="C2E92A30FFB2F132FE68FBC29172FBFC" box="[485,512,1047,1071]" captionStart="Figs" captionStartId="36.[189,238,1823,1847]" captionTargetBox="[317,1315,251,1773]" captionTargetId="figure-57@36.[317,1315,251,1773]" captionTargetPageId="36" captionText="Figs. 13a-h. Isoperla muir nymph and ventral maxilla, Muir Creek, Douglas County, OR. Submarginal setae groups A and B, Marginal setae C, Ventral surface setae D, and Dorsal surface setae DSS. White bars = 1 mm, 13e-f 100x, 13g 200x, and 13h 400x." figureDoi="http://doi.org/10.5281/zenodo.4760360" httpUri="https://zenodo.org/record/4760360/files/figure.png" pageId="35" pageNumber="237">13</figureCitation>
a-h,
<figureCitation id="C2E92A30FFB2F132FDBEFBC2913DFBFC" box="[563,591,1047,1071]" captionStart="Figs" captionStartId="54.[189,238,1610,1634]" captionTargetBox="[211,1421,251,1541]" captionTargetId="figure-57@54.[211,1421,251,1541]" captionTargetPageId="54" captionText="Figs. 20a-q. Isoperla front right leg habitus, Larvae, 3240x, Bar = 1mm. 20a. I. acula; 20b. I. adunca; 20c. I. baumanni; 20d. I. bifurcata; 20e. I. denningi; 20f. I. fulva; 20g. I. laucki; 20h. I. marmorata; 20i. I. miwok; 20j. I. mormona; 20k. I. muir; 20L. I. pinta; 20m. I. quinquepunctata; 20n. I. roguensis; 20o. I. sobria; 20p. I. sordida; 20q. I. tilasqua." figureDoi="http://doi.org/10.5281/zenodo.4760382" httpUri="https://zenodo.org/record/4760382/files/figure.png" pageId="35" pageNumber="237">20</figureCitation>
k)
</paragraph>
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<taxonomicName id="9DD24D36FFB2F132FF6EFBB49101FBA4" ID-CoL="6MZYY" authority="Szczytko &amp; Stewart 2004" authorityName="Szczytko &amp; Stewart" authorityYear="2004" box="[227,627,1121,1143]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="35" pageNumber="237" phylum="Arthropoda" rank="species" species="muir">
<emphasis id="68A6EAA7FFB2F132FF6EFBB49217FBA4" box="[227,357,1121,1143]" italics="true" pageId="35" pageNumber="237">Isoperla muir</emphasis>
<bibRefCitation id="3E434B44FFB2F132FEE1FBB49101FBA4" author="Szczytko &amp; Stewart" box="[364,627,1121,1143]" firstAuthor="Szczytko" pageId="35" pageNumber="237" pagination="233 - 243" refId="ref39226" refString="Szczytko, S. W. &amp; K. W. Stewart. 2004. Isoperla muir, a new species of western Nearctic Isoperla and a new larval description of Isoperla tilasqua Szczytko and Stewart, (Plecoptera: Perlodidae). Transactions of the American Entomological Society, 130 (2): 233 - 243." type="journal article" year="2004">Szczytko &amp; Stewart 2004</bibRefCitation>
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, 130:234-236.
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,
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, larva (reared), ovum.
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</paragraph>
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<emphasis id="68A6EAA7FFB2F132FF30FB1392D8FB0D" bold="true" box="[189,426,1222,1246]" pageId="35" pageNumber="237">Material examined.</emphasis>
<materialsCitation id="EABA3CE8FFB2F132FE3FFB12910BFA99" ID-GBIF-Occurrence="3111754308" collectingDate="27 / V / 2008, 31 / V / 2008" collectionCode="NMNH" collectorName="S. Szczytko &amp; K. Stewart" county="Douglas Co." location="Muir Creek" municipality="Hwy" pageId="35" pageNumber="237" specimenCount="1" specimenCount-male="1" stateProvince="OR" typeStatus="holotype">
TYPE:
<taxonomicName id="9DD24D36FFB2F132FD8EFB12913DFB0D" authorityName="Szczytko &amp; Stewart" authorityYear="2004" box="[515,591,1223,1246]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="35" pageNumber="237" phylum="Arthropoda" rank="species" species="muir">
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,
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:
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<collectingCounty id="B30C4E39FFB2F132FF30FB3F9224FAD1" box="[189,342,1258,1282]" pageId="35" pageNumber="237">Douglas Co.</collectingCounty>
</emphasis>
, Hwy 230, downstream from bridge,
<location id="5F0D606EFFB2F132FF30FADB9220FAF5" LSID="urn:lsid:plazi:treatment:D27B87A3FFB2F134FEA9FC2692F5FC86:5F0D606EFFB2F132FF30FADB9220FAF5" box="[189,338,1294,1318]" county="Douglas Co." municipality="Hwy" name="Muir Creek" pageId="35" pageNumber="237" stateProvince="OR">Muir Creek</location>
,
<collectingDate id="3E28E99DFFB2F132FEEAFADB92ACFAF5" box="[359,478,1294,1318]" pageId="35" pageNumber="237">
<date id="2E6C1075FFB2F132FEEAFADB92FBFAF5" box="[359,393,1294,1318]" pageId="35" pageNumber="237" value="2002-05-12">12/</date>
V/2002
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,
<collectorName id="F7275363FFB2F132FE7EFADB91F1FAF5" box="[499,643,1294,1318]" pageId="35" pageNumber="237">S. Szczytko</collectorName>
,
<collectorName id="F7275363FFB2F132FD15FADA906FFAF5" box="[664,797,1294,1318]" pageId="35" pageNumber="237">K. Stewart</collectorName>
(reared with exuviae) (
<collectionCode id="3CC3AE70FFB2F132FD80FAE7911EFA99" box="[525,620,1330,1354]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="35" pageNumber="237" type="Museum">NMNH</collectionCode>
);
</materialsCitation>
<materialsCitation id="EABA3CE8FFB2F132FD1AFAE49187FA66" ID-GBIF-Occurrence="3111754366" collectingDate="27 / V / 2008, 31 / V / 2008" county="Douglas Co." location="Muir Creek" municipality="Hwy" pageId="35" pageNumber="237" specimenCount="1" specimenCount-male="1" stateProvince="Oregon" typeStatus="holotype">
<emphasis id="68A6EAA7FFB2F132FD1AFAE4923AFABE" bold="true" pageId="35" pageNumber="237">
<typeStatus id="85698817FFB2F132FD1AFAE49230FABE" isOtherTypeStatus="true" pageId="35" pageNumber="237">Additional Specimens</typeStatus>
.
</emphasis>
<collectingRegion id="9816F857FFB2F132FED9FA8392BFFABD" box="[340,461,1366,1390]" country="United States of America" name="Oregon" pageId="35" pageNumber="237">OREGON</collectingRegion>
:
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<emphasis id="68A6EAA7FFB2F132FE51FA839102FABE" bold="true" box="[476,624,1365,1390]" pageId="35" pageNumber="237">Douglas Co</emphasis>
.
</collectingCounty>
,
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,
<location id="5F0D606EFFB2F132FF30FAAC923FFA42" LSID="urn:lsid:plazi:treatment:D27B87A3FFB2F134FEA9FC2692F5FC86:5F0D606EFFB2F132FF30FAAC923FFA42" box="[189,333,1401,1425]" county="Douglas Co." municipality="Muir Creek" name="Muir Camp" pageId="35" pageNumber="237" stateProvince="Oregon">Muir Camp</location>
,
<location id="5F0D606EFFB2F132FED6FAAF92E4FA41" LSID="urn:lsid:plazi:treatment:D27B87A3FFB2F134FEA9FC2692F5FC86:5F0D606EFFB2F132FED6FAAF92E4FA41" box="[347,406,1402,1426]" county="Douglas Co." municipality="Muir Creek" name="Hwy" pageId="35" pageNumber="237" stateProvince="Oregon">Hwy</location>
230 bridge,
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(
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)
<collectorName id="F7275363FFB2F132FC81FAAF938AFA66" pageId="35" pageNumber="237">E Hwy</collectorName>
138 intersection,
<collectingDate id="3E28E99DFFB2F132FE60FA489187FA66" box="[493,757,1437,1461]" isEnumeration="true" pageId="35" pageNumber="237">
<date id="2E6C1075FFB2F132FE60FA489117FA66" box="[493,613,1437,1461]" pageId="35" pageNumber="237" value="2008-05-27">27/V/2008</date>
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<date id="2E6C1075FFB2F132FDF0FA489187FA66" box="[637,757,1437,1461]" pageId="35" pageNumber="237" value="2008-05-31">31/V/2008</date>
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,
<materialsCitation id="EABA3CE8FFB2F132FC80FA489280FA0B" ID-GBIF-Occurrence="3111754342" location="Larvae" pageId="35" pageNumber="237" specimenCount="1" stateProvince="Oregon">
<collectorName id="F7275363FFB2F132FC80FA48925DFA0B" pageId="35" pageNumber="237">J. Sandberg</collectorName>
,
<location id="5F0D606EFFB2F132FEB7FA1492F9FA0B" LSID="urn:lsid:plazi:treatment:D27B87A3FFB2F134FEA9FC2692F5FC86:5F0D606EFFB2F132FEB7FA1492F9FA0B" box="[314,395,1473,1496]" name="Larvae" pageId="35" pageNumber="237" stateProvince="Oregon">Larvae</location>
(reared)
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.
</paragraph>
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<subSubSection id="12C8653EFFB2F134FF30FA319246FECF" lastPageId="37" lastPageNumber="239" pageId="35" pageNumber="237" type="description">
<paragraph id="5A6D36B5FFB2F134FF30FA319246FECF" blockId="35.[189,798,1221,1924]" lastBlockId="37.[189,797,225,854]" lastPageId="37" lastPageNumber="239" pageId="35" pageNumber="237">
<emphasis id="68A6EAA7FFB2F132FF30FA3192FCFA2F" bold="true" box="[189,398,1508,1532]" pageId="35" pageNumber="237">Male larva. Body</emphasis>
length of mature larva
<quantity id="9D2A9B50FFB2F132FD27FA319068FA28" box="[682,794,1508,1532]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" metricValueMax="1.3" metricValueMin="0.9" pageId="35" pageNumber="237" unit="mm" value="11.0" valueMax="13.0" valueMin="9.0">913 mm</quantity>
.
<emphasis id="68A6EAA7FFB2F132FF30F9DD92F4F9F3" bold="true" box="[189,390,1543,1568]" pageId="35" pageNumber="237">Dorsum of head</emphasis>
with contrasting pigment pattern and a mixture of fine light and dark clothing setae, anterior frontoclypeus margin unpigmented; light M shaped pattern anterior to median ocellus thin and well defined, not connected to light frontoclypeus area, median longitudinal light band absent, lateral thin arms directed posterolaterally, extending to antennal bases; posterior ocelli with completely enclosed small light areas along outer lateral margins; interocellar area entirely dark; occiput with irregular spinulae band extending from below eye to near median epicranial suture, enclosed completely by dark pigment (
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).
<emphasis id="68A6EAA7FFB2F132FB1DFED1979BFECF" bold="true" box="[1168,1257,260,284]" pageId="35" pageNumber="237">Lacinia</emphasis>
bidentate, total length
<quantity id="9D2A9B50FFB2F132FC10FEFD9740FE92" box="[925,1074,296,321]" metricMagnitude="-1" metricUnit="m" metricValue="7.805" metricValueMax="8.37" metricValueMin="7.24" pageId="35" pageNumber="237" unit="mm" value="780.5" valueMax="837.0" valueMin="724.0">724837 µm</quantity>
(
<figureCitation id="C2E92A30FFB2F132FBC5FEFD97DFFE93" box="[1096,1197,296,320]" captionStart="Figs" captionStartId="9.[189,238,1823,1847]" captionTargetBox="[249,1383,235,1762]" captionTargetId="figure-57@9.[249,1383,235,1762]" captionTargetPageId="9" captionText="Figs. 2a-q. Isoperla lacinia habitus, 40x, Bar = 500µm. 2a. I. acula; 2b. I. adunca; 2c. I. baumanni; 2d. I. bifurcata; 2e. I. denningi; 2f. I. fulva; 2g. I. laucki; 2h. I. marmorata; 2i. I. miwok; 2j. I. mormona; 2k. I. muir; 2L. I. pinta; 2m. I. quinquepunctata; 2n. I. roguensis; 2o. I. sobria; 2p. I. sordida; 2q. I. tilasqua." figureDoi="http://doi.org/10.5281/zenodo.4760326" httpUri="https://zenodo.org/record/4760326/files/figure.png" pageId="35" pageNumber="237">Figs. 2k</figureCitation>
,
<figureCitation id="C2E92A30FFB2F132FB4DFEFD9670FE93" box="[1216,1282,296,320]" captionStart="Figs" captionStartId="36.[189,238,1823,1847]" captionTargetBox="[317,1315,251,1773]" captionTargetId="figure-57@36.[317,1315,251,1773]" captionTargetPageId="36" captionText="Figs. 13a-h. Isoperla muir nymph and ventral maxilla, Muir Creek, Douglas County, OR. Submarginal setae groups A and B, Marginal setae C, Ventral surface setae D, and Dorsal surface setae DSS. White bars = 1 mm, 13e-f 100x, 13g 200x, and 13h 400x." figureDoi="http://doi.org/10.5281/zenodo.4760360" httpUri="https://zenodo.org/record/4760360/files/figure.png" pageId="35" pageNumber="237">13e- h</figureCitation>
,
<tableCitation id="1750030EFFB2F132FA98FEFD96E4FE93" box="[1301,1430,296,320]" captionStart-0="Table 2" captionStart-1="Table 3" captionStart-2="Table 4" captionStartId-0="4.[189,254,225,249]" captionStartId-1="4.[189,254,1147,1171]" captionStartId-2="5.[189,254,225,249]" captionTargetId-0="graphics-159@4.[288,1344,464,1066]" captionTargetId-1="graphics-2783@4.[344,1289,1315,1887]" captionTargetId-2="graphics-115@5.[423,1208,357,929]" captionTargetPageId-0="4" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="Table 2. Isoperla lacinia characters, seta counts from areas A-D and galea seta counts of left and right ventral surfaces (Fig. 1 &amp; Taxonomic Characters). Numbers within parentheses occurred only once. Characters: N = # of lacinia, C/I = Submarginal row Continuous or Interrupted, P/A = Present or Absent, SMR = submarginal row, DS = dorsal seta, DSS = dorsal surface setae, VSS = ventral surface setae, S = galea surface setae, A = galea apical setae." captionText-1="Table 3. Length and angle measurements of left and right lacinia ventral surfaces (Fig.1). TL = total length, ATL = apical tooth length, STL = subapical tooth length, PL = palm length, PWA = apical palm width, PA = palm angle, PWB = basal palm width." captionText-2="Table 4. Length measurements of left and right galea and maxillary palp ventral surfaces (Fig. 1). GL = galea length, PL 15 = palp segment length 15." httpUri-0="http://table.plazi.org/id/0EAD663DFF95F115FF30FF349239FE55" httpUri-1="http://table.plazi.org/id/0EAD663DFF95F115FF30FBAE9104FB09" httpUri-2="http://table.plazi.org/id/0EAD663DFF94F114FF30FF3491D4FECF" pageId="35" pageNumber="237" tableUuid-0="0EAD663DFF95F115FF30FF349239FE55" tableUuid-1="0EAD663DFF95F115FF30FBAE9104FB09" tableUuid-2="0EAD663DFF94F114FF30FF3491D4FECF">Tables 2-4</tableCitation>
); submarginal row (A+B) with 79 setae, groups A-B interrupted by gap below subapical tooth (SAT) inner margin (
<figureCitation id="C2E92A30FFB2F132FC60FE469739FE78" box="[1005,1099,403,427]" captionStart="Figs" captionStartId="36.[189,238,1823,1847]" captionTargetBox="[317,1315,251,1773]" captionTargetId="figure-57@36.[317,1315,251,1773]" captionTargetPageId="36" captionText="Figs. 13a-h. Isoperla muir nymph and ventral maxilla, Muir Creek, Douglas County, OR. Submarginal setae groups A and B, Marginal setae C, Ventral surface setae D, and Dorsal surface setae DSS. White bars = 1 mm, 13e-f 100x, 13g 200x, and 13h 400x." figureDoi="http://doi.org/10.5281/zenodo.4760360" httpUri="https://zenodo.org/record/4760360/files/figure.png" pageId="35" pageNumber="237">
Fig.
<quantity id="9D2A9B50FFB2F132FBADFE469739FE78" box="[1056,1099,403,427]" metricMagnitude="-2" metricUnit="kg" metricValue="1.3" pageId="35" pageNumber="237" unit="g" value="13.0">13g</quantity>
</figureCitation>
); 2 submarginal setae (A) the first inserted at base of apical tooth (AT) inner margin, the second located between AT and SAT inner margins, plus 1 thin marginal seta (TMS) adjacent to AT inner margin, sometimes obstructed from view by AT, submarginal seta (A), or broken, dorsal seta (DS) absent (
<figureCitation id="C2E92A30FFB2F132FBDAFDBD97DAFD53" box="[1111,1192,616,640]" captionStart="Figs" captionStartId="36.[189,238,1823,1847]" captionTargetBox="[317,1315,251,1773]" captionTargetId="figure-57@36.[317,1315,251,1773]" captionTargetPageId="36" captionText="Figs. 13a-h. Isoperla muir nymph and ventral maxilla, Muir Creek, Douglas County, OR. Submarginal setae groups A and B, Marginal setae C, Ventral surface setae D, and Dorsal surface setae DSS. White bars = 1 mm, 13e-f 100x, 13g 200x, and 13h 400x." figureDoi="http://doi.org/10.5281/zenodo.4760360" httpUri="https://zenodo.org/record/4760360/files/figure.png" pageId="35" pageNumber="237">Figs 13</figureCitation>
g-h); 57 submarginal setae (B) located past SAT inner margin (
<figureCitation id="C2E92A30FFB2F132FA94FD5E9607FD70" box="[1305,1397,651,675]" captionStart="Figs" captionStartId="36.[189,238,1823,1847]" captionTargetBox="[317,1315,251,1773]" captionTargetId="figure-57@36.[317,1315,251,1773]" captionTargetPageId="36" captionText="Figs. 13a-h. Isoperla muir nymph and ventral maxilla, Muir Creek, Douglas County, OR. Submarginal setae groups A and B, Marginal setae C, Ventral surface setae D, and Dorsal surface setae DSS. White bars = 1 mm, 13e-f 100x, 13g 200x, and 13h 400x." figureDoi="http://doi.org/10.5281/zenodo.4760360" httpUri="https://zenodo.org/record/4760360/files/figure.png" pageId="35" pageNumber="237">
Fig.
<quantity id="9D2A9B50FFB2F132FAC6FD5E9607FD70" box="[1355,1397,651,675]" metricMagnitude="-2" metricUnit="kg" metricValue="1.3" pageId="35" pageNumber="237" unit="g" value="13.0">13g</quantity>
</figureCitation>
); 9 11 marginal setae (C), initially long-stout and widely spaced, first marginal setae located under flap-like SAT inner margin (
<figureCitation id="C2E92A30FFB2F132FBACFD23970FFCDD" box="[1057,1149,758,782]" captionStart="Figs" captionStartId="36.[189,238,1823,1847]" captionTargetBox="[317,1315,251,1773]" captionTargetId="figure-57@36.[317,1315,251,1773]" captionTargetPageId="36" captionText="Figs. 13a-h. Isoperla muir nymph and ventral maxilla, Muir Creek, Douglas County, OR. Submarginal setae groups A and B, Marginal setae C, Ventral surface setae D, and Dorsal surface setae DSS. White bars = 1 mm, 13e-f 100x, 13g 200x, and 13h 400x." figureDoi="http://doi.org/10.5281/zenodo.4760360" httpUri="https://zenodo.org/record/4760360/files/figure.png" pageId="35" pageNumber="237">Fig. 13h</figureCitation>
arrow), last few shorter and widely spaced (
<figureCitation id="C2E92A30FFB2F132FBB9FCCC97E3FCE1" box="[1076,1169,793,818]" captionStart="Figs" captionStartId="36.[189,238,1823,1847]" captionTargetBox="[317,1315,251,1773]" captionTargetId="figure-57@36.[317,1315,251,1773]" captionTargetPageId="36" captionText="Figs. 13a-h. Isoperla muir nymph and ventral maxilla, Muir Creek, Douglas County, OR. Submarginal setae groups A and B, Marginal setae C, Ventral surface setae D, and Dorsal surface setae DSS. White bars = 1 mm, 13e-f 100x, 13g 200x, and 13h 400x." figureDoi="http://doi.org/10.5281/zenodo.4760360" httpUri="https://zenodo.org/record/4760360/files/figure.png" pageId="35" pageNumber="237">Fig. 13e</figureCitation>
); 3358 ventral surface setae (D) forming dense longitudinal band below submarginal and marginal setae, ending posteriorly at approximately ¾ the inner lacinia margin length, setae closest to inner margin protrude laterally past lacinia margin (
<figureCitation id="C2E92A30FFB2F132FC73FC1E972AFC30" box="[1022,1112,971,995]" captionStart="Figs" captionStartId="36.[189,238,1823,1847]" captionTargetBox="[317,1315,251,1773]" captionTargetId="figure-57@36.[317,1315,251,1773]" captionTargetPageId="36" captionText="Figs. 13a-h. Isoperla muir nymph and ventral maxilla, Muir Creek, Douglas County, OR. Submarginal setae groups A and B, Marginal setae C, Ventral surface setae D, and Dorsal surface setae DSS. White bars = 1 mm, 13e-f 100x, 13g 200x, and 13h 400x." figureDoi="http://doi.org/10.5281/zenodo.4760360" httpUri="https://zenodo.org/record/4760360/files/figure.png" pageId="35" pageNumber="237">Fig. 13f</figureCitation>
); dorsal surface setae (DSS) forming dense, laterally protruding, longitudinal band on and along inner-lateral margin, ending at approximately the posterior-most ventral surface setae (
<figureCitation id="C2E92A30FFB2F132FC1FFB8C909DFBA2" box="[914,1007,1113,1137]" captionStart="Figs" captionStartId="36.[189,238,1823,1847]" captionTargetBox="[317,1315,251,1773]" captionTargetId="figure-57@36.[317,1315,251,1773]" captionTargetPageId="36" captionText="Figs. 13a-h. Isoperla muir nymph and ventral maxilla, Muir Creek, Douglas County, OR. Submarginal setae groups A and B, Marginal setae C, Ventral surface setae D, and Dorsal surface setae DSS. White bars = 1 mm, 13e-f 100x, 13g 200x, and 13h 400x." figureDoi="http://doi.org/10.5281/zenodo.4760360" httpUri="https://zenodo.org/record/4760360/files/figure.png" pageId="35" pageNumber="237">Fig. 13f</figureCitation>
).
<emphasis id="68A6EAA7FFB2F132FB86FB8C9720FBA2" bold="true" box="[1035,1106,1113,1137]" pageId="35" pageNumber="237">Galea</emphasis>
with 2232 setae in sparse ventral row, apex with 56 setae.
<emphasis id="68A6EAA7FFB2F132FB67FBA896D1FB46" bold="true" box="[1258,1443,1149,1173]" pageId="35" pageNumber="237">Maxillary Palp</emphasis>
segments 23 with nearly straight, apically rounded setae.
<emphasis id="68A6EAA7FFB2F132FC02FB10977AFB0E" bold="true" box="[911,1032,1221,1245]" pageId="35" pageNumber="237">Pronotum</emphasis>
with thin indistinct median light line bordered by irregular dark longitudinal bands with adjacent light rugosites; discs each with brown comma shaped lateral areas, fine dark clothing setae and lateral margins without broad light bands (
<figureCitation id="C2E92A30FFB2F132FAFBFA86901FFA5D" captionStart="Figs" captionStartId="36.[189,238,1823,1847]" captionTargetBox="[317,1315,251,1773]" captionTargetId="figure-57@36.[317,1315,251,1773]" captionTargetPageId="36" captionText="Figs. 13a-h. Isoperla muir nymph and ventral maxilla, Muir Creek, Douglas County, OR. Submarginal setae groups A and B, Marginal setae C, Ventral surface setae D, and Dorsal surface setae DSS. White bars = 1 mm, 13e-f 100x, 13g 200x, and 13h 400x." figureDoi="http://doi.org/10.5281/zenodo.4760360" httpUri="https://zenodo.org/record/4760360/files/figure.png" pageId="35" pageNumber="237">Fig. 13b</figureCitation>
).
<emphasis id="68A6EAA7FFB2F132FC1EFAA397C9FA5D" bold="true" box="[915,1211,1398,1422]" pageId="35" pageNumber="237">Meso and metanotum</emphasis>
with contrasting pigment pattern and fine dark clothing setae (
<figureCitation id="C2E92A30FFB2F132FAFBFA4F9019FA05" captionStart="Figs" captionStartId="36.[189,238,1823,1847]" captionTargetBox="[317,1315,251,1773]" captionTargetId="figure-57@36.[317,1315,251,1773]" captionTargetPageId="36" captionText="Figs. 13a-h. Isoperla muir nymph and ventral maxilla, Muir Creek, Douglas County, OR. Submarginal setae groups A and B, Marginal setae C, Ventral surface setae D, and Dorsal surface setae DSS. White bars = 1 mm, 13e-f 100x, 13g 200x, and 13h 400x." figureDoi="http://doi.org/10.5281/zenodo.4760360" httpUri="https://zenodo.org/record/4760360/files/figure.png" pageId="35" pageNumber="237">Fig. 13c</figureCitation>
).
<emphasis id="68A6EAA7FFB2F132FC0EFA6B90C9FA05" bold="true" box="[899,955,1470,1494]" pageId="35" pageNumber="237">Legs</emphasis>
with numerous fine dark clothing setae and scattered erect spines on outer surface of femora, erect spines longest and concentrated on dorsal surfaces; fine silky setae absent on dorsal surface of femora, sparse on tibia (
<figureCitation id="C2E92A30FFB2F132FBDAF99997C6F9B7" box="[1111,1204,1612,1636]" captionStart="Figs" captionStartId="54.[189,238,1610,1634]" captionTargetBox="[211,1421,251,1541]" captionTargetId="figure-57@54.[211,1421,251,1541]" captionTargetPageId="54" captionText="Figs. 20a-q. Isoperla front right leg habitus, Larvae, 3240x, Bar = 1mm. 20a. I. acula; 20b. I. adunca; 20c. I. baumanni; 20d. I. bifurcata; 20e. I. denningi; 20f. I. fulva; 20g. I. laucki; 20h. I. marmorata; 20i. I. miwok; 20j. I. mormona; 20k. I. muir; 20L. I. pinta; 20m. I. quinquepunctata; 20n. I. roguensis; 20o. I. sobria; 20p. I. sordida; 20q. I. tilasqua." figureDoi="http://doi.org/10.5281/zenodo.4760382" httpUri="https://zenodo.org/record/4760382/files/figure.png" pageId="35" pageNumber="237">Fig. 20k</figureCitation>
); tibia with very faint transverse bands near proximal end.
<emphasis id="68A6EAA7FFB2F132FA95F9BA90F2F97F" bold="true" pageId="35" pageNumber="237">Abdominal terga</emphasis>
with two distinct longitudinal dark stripes; wide light median longitudinal band widest on middle segments, narrowing anteriorly and posteriorly; lateral pair of dark longitudinal stripes extending to lateral margins; numerous fine dark clothing setae and erect spines scattered dorsally; posterior margin with distinct and scattered long, and numerous short spines in a concentrated row (
<figureCitation id="C2E92A30FFB4F134FF48FED19254FECF" box="[197,294,260,284]" captionStart="Figs" captionStartId="36.[189,238,1823,1847]" captionTargetBox="[317,1315,251,1773]" captionTargetId="figure-57@36.[317,1315,251,1773]" captionTargetPageId="36" captionText="Figs. 13a-h. Isoperla muir nymph and ventral maxilla, Muir Creek, Douglas County, OR. Submarginal setae groups A and B, Marginal setae C, Ventral surface setae D, and Dorsal surface setae DSS. White bars = 1 mm, 13e-f 100x, 13g 200x, and 13h 400x." figureDoi="http://doi.org/10.5281/zenodo.4760360" httpUri="https://zenodo.org/record/4760360/files/figure.png" pageId="37" pageNumber="239">Fig. 13d</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="0EAD663DFFB5F135FF30F8CA9127F8AC" ID-DOI="http://doi.org/10.5281/zenodo.4760360" ID-Zenodo-Dep="4760360" httpUri="https://zenodo.org/record/4760360/files/figure.png" pageId="36" pageNumber="238" startId="36.[189,238,1823,1847]" targetBox="[317,1315,251,1773]" targetPageId="36">
<paragraph id="5A6D36B5FFB5F135FF30F8CA9127F8AC" blockId="36.[189,1441,1823,1919]" pageId="36" pageNumber="238">
Figs. 13a-h.
<taxonomicName id="9DD24D36FFB5F135FEC7F8CA92A8F8E4" authorityName="Szczytko &amp; Stewart" authorityYear="2004" box="[330,474,1823,1847]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="36" pageNumber="238" phylum="Arthropoda" rank="species" species="muir">
<emphasis id="68A6EAA7FFB5F135FEC7F8CA92A8F8E4" box="[330,474,1823,1847]" italics="true" pageId="36" pageNumber="238">Isoperla muir</emphasis>
</taxonomicName>
nymph and ventral maxilla, Muir Creek, Douglas County, OR. Submarginal setae groups A and B, Marginal setae C, Ventral surface setae D, and Dorsal surface setae DSS. White bars = 1 mm, 13e-f 100x, 13g 200x, and 13h 400x.
</paragraph>
</caption>
<subSubSection id="12C8653EFFB4F134FF30FEFD9225FE54" pageId="37" pageNumber="239" type="distribution">
<paragraph id="5A6D36B5FFB4F134FF30FEFD9225FE54" blockId="37.[189,797,225,854]" pageId="37" pageNumber="239">
<emphasis id="68A6EAA7FFB4F134FF30FEFD922EFE93" bold="true" box="[189,348,296,320]" pageId="37" pageNumber="239">Distribution.</emphasis>
<collectingRegion id="9816F857FFB4F134FEE4FEFD92B6FE93" box="[361,452,296,320]" country="United States of America" name="Oregon" pageId="37" pageNumber="239">Oregon</collectingRegion>
, known only from the
<typeStatus id="85698817FFB4F134FD64FEFD906FFE93" box="[745,797,296,320]" pageId="37" pageNumber="239">type</typeStatus>
location (
<bibRefCitation id="3E434B44FFB4F134FEA1FE999127FEB7" author="Szczytko &amp; Stewart" box="[300,597,332,356]" firstAuthor="Szczytko" pageId="37" pageNumber="239" pagination="233 - 243" refId="ref39226" refString="Szczytko, S. W. &amp; K. W. Stewart. 2004. Isoperla muir, a new species of western Nearctic Isoperla and a new larval description of Isoperla tilasqua Szczytko and Stewart, (Plecoptera: Perlodidae). Transactions of the American Entomological Society, 130 (2): 233 - 243." type="journal article" year="2004">Szczytko &amp; Stewart 2004</bibRefCitation>
) where it occurs with
<taxonomicName id="9DD24D36FFB4F134FF77FEA59222FE54" baseAuthorityName="Hagen" baseAuthorityYear="1874" box="[250,336,367,391]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="37" pageNumber="239" phylum="Arthropoda" rank="species" species="sobria">
<emphasis id="68A6EAA7FFB4F134FF77FEA59271FE54" box="[250,259,368,391]" italics="true" pageId="37" pageNumber="239">I</emphasis>
.
<emphasis id="68A6EAA7FFB4F134FE9DFEBA9222FE54" box="[272,336,367,391]" italics="true" pageId="37" pageNumber="239">sobria</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="12C8653EFFB4F134FF30FE4791BBFD14" pageId="37" pageNumber="239" type="diagnosis">
<paragraph id="5A6D36B5FFB4F134FF30FE4791BBFD14" blockId="37.[189,797,225,854]" pageId="37" pageNumber="239">
<emphasis id="68A6EAA7FFB4F134FF30FE47924DFE79" bold="true" box="[189,319,402,426]" pageId="37" pageNumber="239">Diagnosis.</emphasis>
The mature male larvae of
<taxonomicName id="9DD24D36FFB4F134FD13FE46919CFE79" authorityName="Szczytko &amp; Stewart" authorityYear="2004" box="[670,750,403,426]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="37" pageNumber="239" phylum="Arthropoda" rank="species" species="muir">
<emphasis id="68A6EAA7FFB4F134FD13FE46919CFE79" box="[670,750,403,426]" italics="true" pageId="37" pageNumber="239">I. muir</emphasis>
</taxonomicName>
are most similar to
<taxonomicName id="9DD24D36FFB4F134FEFFFE6292A6FE1D" box="[370,468,438,462]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="37" pageNumber="239" phylum="Arthropoda" rank="species" species="adunca">
<emphasis id="68A6EAA7FFB4F134FEFFFE6292A6FE1D" box="[370,468,438,462]" italics="true" pageId="37" pageNumber="239">I. adunca</emphasis>
</taxonomicName>
in that the submarginal row (A+B) setae form a continuous and regularly spaced row (
<figureCitation id="C2E92A30FFB4F134FF77FE2B9220FDC5" box="[250,338,510,534]" captionStart="Figs" captionStartId="36.[189,238,1823,1847]" captionTargetBox="[317,1315,251,1773]" captionTargetId="figure-57@36.[317,1315,251,1773]" captionTargetPageId="36" captionText="Figs. 13a-h. Isoperla muir nymph and ventral maxilla, Muir Creek, Douglas County, OR. Submarginal setae groups A and B, Marginal setae C, Ventral surface setae D, and Dorsal surface setae DSS. White bars = 1 mm, 13e-f 100x, 13g 200x, and 13h 400x." figureDoi="http://doi.org/10.5281/zenodo.4760360" httpUri="https://zenodo.org/record/4760360/files/figure.png" pageId="37" pageNumber="239">Figs. 13</figureCitation>
g-h).
<taxonomicName id="9DD24D36FFB4F134FE03FE2B916EFDC6" authorityName="Szczytko &amp; Stewart" authorityYear="2004" box="[398,540,510,534]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="37" pageNumber="239" phylum="Arthropoda" rank="species" species="muir">
<emphasis id="68A6EAA7FFB4F134FE03FE2B916EFDC6" box="[398,540,510,534]" italics="true" pageId="37" pageNumber="239">Isoperla muir</emphasis>
</taxonomicName>
can be separated from
<taxonomicName id="9DD24D36FFB4F134FF30FDF7925EFDEA" box="[189,300,545,569]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="37" pageNumber="239" phylum="Arthropoda" rank="species" species="adunca">
<emphasis id="68A6EAA7FFB4F134FF30FDF7925EFDEA" box="[189,300,545,569]" italics="true" pageId="37" pageNumber="239">I. adunca</emphasis>
</taxonomicName>
by the submarginal setae row (A+B) regularly spaced at about the width of each seta base at points of insertion (
<figureCitation id="C2E92A30FFB4F134FE37FDBD9153FD53" box="[442,545,616,640]" captionStart="Figs" captionStartId="36.[189,238,1823,1847]" captionTargetBox="[317,1315,251,1773]" captionTargetId="figure-57@36.[317,1315,251,1773]" captionTargetPageId="36" captionText="Figs. 13a-h. Isoperla muir nymph and ventral maxilla, Muir Creek, Douglas County, OR. Submarginal setae groups A and B, Marginal setae C, Ventral surface setae D, and Dorsal surface setae DSS. White bars = 1 mm, 13e-f 100x, 13g 200x, and 13h 400x." figureDoi="http://doi.org/10.5281/zenodo.4760360" httpUri="https://zenodo.org/record/4760360/files/figure.png" pageId="37" pageNumber="239">Figs. 13a</figureCitation>
, g-h), dorsal seta (DS) absent and first marginal seta (C) covered by flap-like subapical tooth inner margin base (
<figureCitation id="C2E92A30FFB4F134FDD6FD7A91CEFD14" box="[603,700,687,711]" captionStart="Figs" captionStartId="36.[189,238,1823,1847]" captionTargetBox="[317,1315,251,1773]" captionTargetId="figure-57@36.[317,1315,251,1773]" captionTargetPageId="36" captionText="Figs. 13a-h. Isoperla muir nymph and ventral maxilla, Muir Creek, Douglas County, OR. Submarginal setae groups A and B, Marginal setae C, Ventral surface setae D, and Dorsal surface setae DSS. White bars = 1 mm, 13e-f 100x, 13g 200x, and 13h 400x." figureDoi="http://doi.org/10.5281/zenodo.4760360" httpUri="https://zenodo.org/record/4760360/files/figure.png" pageId="37" pageNumber="239">Fig. 13h</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="12C8653EFFB4F134FF30FD0692F5FC86" pageId="37" pageNumber="239" type="discussion">
<paragraph id="5A6D36B5FFB4F134FF30FD0692F5FC86" blockId="37.[189,797,225,854]" pageId="37" pageNumber="239">
<emphasis id="68A6EAA7FFB4F134FF30FD069242FD38" bold="true" box="[189,304,723,747]" pageId="37" pageNumber="239">Remarks.</emphasis>
This distinctively pigmented species is rarely collected and is apparently restricted to the
<typeStatus id="85698817FFB4F134FF30FCCE9383FCE0" box="[189,241,795,819]" pageId="37" pageNumber="239">type</typeStatus>
locality. Limited rearing indicated emergence occurred in June.
</paragraph>
</subSubSection>
</treatment>
</document>