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<document id="837E8D36F61D3204C6C1B04A06BD7831" ID-CLB-Dataset="31258" ID-DOI="10.11646/zootaxa.4361.1.1" ID-GBIF-Dataset="b15f5f11-6a00-4158-8ec8-3d0004605c28" ID-ISSN="1175-5326" ID-Zenodo-Dep="1069337" ID-ZooBank="53489D29-C68F-44FD-9EA2-CFCA7B949630" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1512125394505" checkinUser="plazi" docAuthor="Ellenrieder, Natalia Von &amp; Garrison, Rosser W." docDate="2017" docId="D2332A59FFBB4E41FF5DFA0FFC286CD3" docLanguage="en" docName="zootaxa.4361.1.1.pdf.imf" docOrigin="Zootaxa 4361 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Protoneura corculum Calvert 1907" docType="treatment" docVersion="10" lastPageNumber="23" masterDocId="2E0A5221FFAE4E57FFCAFF8BFFB86B47" masterDocTitle="A synopsis of the Neotropical genus Protoneura (Odonata: Coenagrionidae)" masterLastPageNumber="76" masterPageNumber="1" pageNumber="22" updateTime="1698506080776" updateUser="ExternalLinkService">
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<mods:title id="3A8A9B556340F26C00E6456B8445DF34">A synopsis of the Neotropical genus Protoneura (Odonata: Coenagrionidae)</mods:title>
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<mods:namePart id="5EFEE486D629AAEDF7CD03CAF0847408">Ellenrieder, Natalia Von</mods:namePart>
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<mods:namePart id="8D49BE7AE4CDAEC71A29BD63A11D7A04">Garrison, Rosser W.</mods:namePart>
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<treatment id="D2332A59FFBB4E41FF5DFA0FFC286CD3" ID-DOI="http://doi.org/10.5281/zenodo.6030141" ID-GBIF-Taxon="137582229" ID-Zenodo-Dep="6030141" LSID="urn:lsid:plazi:treatment:D2332A59FFBB4E41FF5DFA0FFC286CD3" httpUri="http://treatment.plazi.org/id/D2332A59FFBB4E41FF5DFA0FFC286CD3" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22">
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<heading id="016D2C23FFBB4E42FF5DFA0FFDF36ED9" bold="true" box="[151,587,1412,1438]" fontSize="11" level="1" pageId="21" pageNumber="22" reason="1">
<taxonomicName id="9D9AE0CCFFBB4E42FF5DFA0FFDF36ED9" ID-CoL="4N6FY" authority="Calvert, 1907" authorityName="Calvert" authorityYear="1907" box="[151,587,1412,1438]" class="Insecta" family="Protoneuridae" genus="Protoneura" kingdom="Animalia" order="Odonata" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="corculum">
<emphasis id="68EE475DFFBB4E42FF5DFA0FFDF36ED9" bold="true" box="[151,587,1412,1438]" pageId="21" pageNumber="22">
<emphasis id="68EE475DFFBB4E42FF5DFA0FFE246ED9" bold="true" box="[151,412,1412,1438]" italics="true" pageId="21" pageNumber="22">Protoneura corculum</emphasis>
<bibRefCitation id="3E0BE6BEFFBB4E42FE6EFA0FFDF36ED9" author="Calvert" box="[420,587,1412,1438]" pageId="21" pageNumber="22" refString="Calvert, P. P. (1907) Odonata. In: Godman, F. d. C. &amp; Salvin, O. (Eds.), Biologia Centrali Americana: Insecta Neuroptera. R. H. Porter &amp; Dulau Co., London, pp. 309 - 404." type="book chapter" year="1907">Calvert, 1907</bibRefCitation>
</emphasis>
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</paragraph>
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<figureCitation id="C2A187CAFFBB4E42FF5DFA2DFF5B6EF8" box="[151,227,1446,1471]" captionStart="FIGURES 58" captionStartId="17.[151,264,1532,1555]" captionTargetBox="[162,1429,524,1500]" captionTargetId="figure@17.[151,1436,519,1510]" captionTargetPageId="17" captionText="FIGURES 58. Male body scan; head in dorsal view, thorax and abdomen in lateral view (not to scale). (5) P. calverti; (6) P. capillaris; (7) P.cara; (8) P.corculum." httpUri="https://zenodo.org/record/1069341/files/figure.png" pageId="21" pageNumber="22">Figs. 8</figureCitation>
(
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habitus), 32 (
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habitus), 54 (
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mes. plate), 78 (gen. lig.), 104 (
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app.), 121 (map)
</paragraph>
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<taxonomicName id="9D9AE0CCFFBB4E42FF5DFA7BFD816D43" ID-CoL="4N6FY" authority="Calvert, 1907: 395" authorityName="Calvert" authorityPageNumber="395" authorityYear="1907" box="[151,569,1518,1541]" class="Insecta" family="Protoneuridae" genus="Protoneura" kingdom="Animalia" order="Odonata" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="corculum">
<emphasis id="68EE475DFFBB4E42FF5DFA7BFEC96D43" box="[151,369,1518,1541]" italics="true" pageId="21" pageNumber="22">Protoneura corculum</emphasis>
<treatmentCitation id="DB3BBD5EFFBB4E42FEB2FA65FD816D43" author="Calvert" box="[376,569,1518,1540]" page="395" pageId="21" pageNumber="22" year="1907">
<bibRefCitation id="3E0BE6BEFFBB4E42FEB2FA65FDBF6D43" author="Calvert" box="[376,519,1518,1540]" pageId="21" pageNumber="22" refString="Calvert, P. P. (1907) Odonata. In: Godman, F. d. C. &amp; Salvin, O. (Eds.), Biologia Centrali Americana: Insecta Neuroptera. R. H. Porter &amp; Dulau Co., London, pp. 309 - 404." type="book chapter" year="1907">Calvert, 1907</bibRefCitation>
: 395
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397, pl. 10,
<figureCitation id="C2A187CAFFBB4E42FD08FA65FC836D43" box="[706,827,1518,1540]" captionStart="FIGURES 4144" captionStartId="48.[151,264,1190,1213]" captionTargetBox="[158,1421,203,1156]" captionTargetId="figure@48.[151,1436,193,1169]" captionTargetPageId="48" captionText="FIGURES 4144. Female body scan; head in dorsal view, thorax and abdomen in lateral view (not to scale). (41) P. sanguinipes; (42) P. scintilla; (43) P. sulfurata; (44) P. tenuis. Insert in Fig. 42: posterior lobe of pronotum in dorsal view; in Fig.43: pterothorax in dorsal view." httpUri="https://zenodo.org/record/1069359/files/figure.png" pageId="21" pageNumber="22">Figs. 4144</figureCitation>
(in key, description of
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, illustrations of
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S10, base of wing, and
<specimenCount id="4C9C50C6FFBB4E42FEFAF985FEFB6D64" box="[304,323,1550,1571]" pageId="21" pageNumber="22" type="female"></specimenCount>
pronotum and mesostigmal plates);—
<bibRefCitation id="3E0BE6BEFFBB4E42FD08F986FD756D05" author="Williamson" pageId="21" pageNumber="22" refString="Williamson, E. (1915) Notes on neotropical dragonflies or Odonata. Proceedings of the U. S. Natural History Museum, 48, 601 - 638. https: // doi. org / 10.5479 / si. 00963801.48 - 2089.601" type="journal article" year="1915">
Williamson (1915: 619, 623,
<figureCitation id="C2A187CAFFBB4E42FC27F986FB8B6D64" box="[1005,1075,1549,1571]" captionStart="FIGURES 14" captionStartId="8.[151,264,1958,1981]" captionTargetBox="[167,1430,957,1933]" captionTargetId="figure@8.[151,1436,951,1934]" captionTargetPageId="8" captionText="FIGURES 14. Male body scan; head in dorsal view, thorax and abdomen in lateral view (not to scale). (1) P. ailsa; (2) P. amatoria; (3) P.aurantiaca; (4) P.caligata." httpUri="https://zenodo.org/record/1069339/files/figure.png" pageId="21" pageNumber="22">Figs. 2</figureCitation>
,
<figureCitation id="C2A187CAFFBB4E42FBF4F986FBC76D64" box="[1086,1151,1549,1571]" captionStart="FIGURES 1316" captionStartId="30.[151,264,1592,1615]" captionTargetBox="[161,1426,597,1565]" captionTargetId="figure@30.[151,1436,591,1570]" captionTargetPageId="30" captionText="FIGURES 1316. Male body scan; head in dorsal view, thorax and abdomen in lateral view (not to scale).(13) P. paucinervis; (14) P.peramans; (15) P.rojiza; (16) P. romanae." httpUri="https://zenodo.org/record/1069345/files/figure.png" pageId="21" pageNumber="22">13, 14</figureCitation>
; discussion of relationships, in key, illustration of wings and thoracic pattern)
</bibRefCitation>
;—
<bibRefCitation id="3E0BE6BEFFBB4E42FD21F9A7FCC46D05" author="Alayo" box="[747,892,1580,1602]" pageId="21" pageNumber="22" refString="Alayo, D. P. (1968 a) Las libelulas de Cuba (Insecta-Odonata). Torreia, Nueva Series No. 2 (Parte I: Texto) 102 pp." type="book" year="1968" yearSuffix="a">Alayo (1968a</bibRefCitation>
,
<treatmentCitation id="DB3BBD5EFFBB4E42FC43F9A7FC7A6D05" author="Alayo" box="[905,962,1580,1602]" page="74" pageId="21" pageNumber="22" year="1968">
<bibRefCitation id="3E0BE6BEFFBB4E42FC43F9A7FC216D05" author="Alayo" box="[905,921,1580,1602]" pageId="21" pageNumber="22" refString="Alayo, D. P. (1968 b) Las libelulas de Cuba (Insecta-Odonata). Torreia, Nueva Serie No. 3 (Parte II: Laminas) 54 pp." type="book" year="1968" yearSuffix="b">b</bibRefCitation>
: 74
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, 75,
<figureCitation id="C2A187CAFFBB4E42FC33F9A7FBD96D05" box="[1017,1121,1580,1602]" captionStart="FIGURES 2124" captionStartId="42.[151,264,1594,1617]" captionTargetBox="[159,1430,593,1567]" captionTargetId="figure@42.[151,1436,591,1572]" captionTargetPageId="42" captionText="FIGURES 2124. Male body scan; head in dorsal view, thorax and abdomen in lateral view (not to scale). (21) P. viridis; (22) P. woytkowskii form a; (23)P.woytkowskii form b; (24) P.woytkowskii form c." httpUri="https://zenodo.org/record/1069349/files/figure.png" pageId="21" pageNumber="22">Figs. 23B</figureCitation>
,
<figureCitation id="C2A187CAFFBB4E42FBA5F9A7FB236D05" box="[1135,1179,1580,1602]" captionStart="FIGURES 3740" captionStartId="47.[151,264,1190,1213]" captionTargetBox="[166,1416,198,1156]" captionTargetId="figure@47.[151,1436,193,1169]" captionTargetPageId="47" captionText="FIGURES 3740. Female body scan; head in dorsal view, thorax and abdomen in lateral view (not to scale). (37) P. paucinervis; (38) P.peramans; (39) P.rojiza; (40) P.romanae. Insert in Fig. 40: pronotum in dorsal view." httpUri="https://zenodo.org/record/1069357/files/figure.png" pageId="21" pageNumber="22">39B</figureCitation>
,
<figureCitation id="C2A187CAFFBB4E42FB63F9A7FAB86D05" box="[1193,1280,1580,1602]" captionStart="FIGURES 4144" captionStartId="48.[151,264,1190,1213]" captionTargetBox="[158,1421,203,1156]" captionTargetId="figure@48.[151,1436,193,1169]" captionTargetPageId="48" captionText="FIGURES 4144. Female body scan; head in dorsal view, thorax and abdomen in lateral view (not to scale). (41) P. sanguinipes; (42) P. scintilla; (43) P. sulfurata; (44) P. tenuis. Insert in Fig. 42: posterior lobe of pronotum in dorsal view; in Fig.43: pterothorax in dorsal view." httpUri="https://zenodo.org/record/1069359/files/figure.png" pageId="21" pageNumber="22">
<date id="2E24BD8FFFBB4E42FB63F9A7FB746D05" box="[1193,1228,1580,1602]" pageId="21" pageNumber="22" value="1942-01">42I</date>
, 42J
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; key to Cuban species, illustrations of
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wings, head, thorax, and S10);—
<bibRefCitation id="3E0BE6BEFFBB4E42FCDEF9C1FA956D27" author="Paulson" box="[788,1325,1610,1632]" pageId="21" pageNumber="22" refString="Paulson, D. R. (1982) Odonata, pp. 249 - 277. In: Hurlbert, S. H. &amp; Villalobos-Figueroa, A. (Eds.), Aquatic biota of Mexico, Central America and the West Indies. San Diego State University, San Diego, California." type="book chapter" year="1982">
Paulson (1982: 252, 260;
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<collectingRegion id="985E55ADFFBB4E42FBD6F9C1FBD46D27" box="[1052,1132,1610,1632]" country="Mexico" name="Mexico" pageId="21" pageNumber="22">Mexico</collectingRegion>
</collectingCountry>
,
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<collectingRegion id="985E55ADFFBB4E42FBBFF9C1FB5D6D27" box="[1141,1253,1610,1632]" country="Guatemala" name="Guatemala" pageId="21" pageNumber="22">Guatemala</collectingRegion>
</collectingCountry>
,
<collectingCountry id="228DDBDFFFBB4E42FB25F9C1FA9E6D27" box="[1263,1318,1610,1632]" name="Cuba" pageId="21" pageNumber="22">Cuba</collectingCountry>
)
</bibRefCitation>
;—
<bibRefCitation id="3E0BE6BEFFBB4E42FA81F9C1FE336D38" author="Measey" pageId="21" pageNumber="22" refString="Measey, G. J. (1994) Some Odonata from Belize, Central America. Notulae odonatologicae, 4 (3), 40 - 46." type="journal article" year="1994">
Measey (1994: 45;
<collectingCountry id="228DDBDFFFBB4E42FE89F9E2FE3C6D38" box="[323,388,1641,1663]" name="Belize" pageId="21" pageNumber="22">
<collectingRegion id="985E55ADFFBB4E42FE89F9E2FE3C6D38" box="[323,388,1641,1663]" country="Belize" name="Belize" pageId="21" pageNumber="22">Belize</collectingRegion>
</collectingCountry>
)
</bibRefCitation>
;—
<bibRefCitation id="3E0BE6BEFFBB4E42FE63F9E2FCE36D38" author="Boomsma" box="[425,859,1641,1663]" pageId="21" pageNumber="22" refString="Boomsma, T. &amp; Dunkle, S. W. (1996) Odonata of Belize. Odonatologica, 25 (1), 17 - 29." type="journal article" year="1996">
Boomsma &amp; Dunkle (1996: 25;
<collectingCountry id="228DDBDFFFBB4E42FCD9F9E2FCEC6D38" box="[787,852,1641,1663]" name="Belize" pageId="21" pageNumber="22">
<collectingRegion id="985E55ADFFBB4E42FCD9F9E2FCEC6D38" box="[787,852,1641,1663]" country="Belize" name="Belize" pageId="21" pageNumber="22">Belize</collectingRegion>
</collectingCountry>
)
</bibRefCitation>
;—
<bibRefCitation id="3E0BE6BEFFBB4E42FCB3F9E2FEA36DD9" author="Gonzalez-Soriano" pageId="21" pageNumber="22" refString="Gonzalez-Soriano, E. &amp; Novelo-Gutierrez, R. (1996) 14. Odonata, pp. 147 - 167. In: Llorente- Bousquets, J., Garcia-Aldrete, A. N. &amp; Gonzalez-Soriano, E. (Eds.), Biodiversidad, taxonomia y biogeografia de artropodos de Mexico: Hacia una sintesis de su conocimiento. Universidad Autonoma de Mexico, Mexico." type="book chapter" year="1996">
González-Soriano &amp; Novelo-Gutiérrez (1996: 162;
<collectingCountry id="228DDBDFFFBB4E42FF0DF903FEAC6DD9" box="[199,276,1672,1694]" name="Mexico" pageId="21" pageNumber="22">
<collectingRegion id="985E55ADFFBB4E42FF0DF903FEAC6DD9" box="[199,276,1672,1694]" country="Mexico" name="Mexico" pageId="21" pageNumber="22">Mexico</collectingRegion>
</collectingCountry>
)
</bibRefCitation>
;—
<bibRefCitation id="3E0BE6BEFFBB4E42FEF3F903FCD16DD9" author="Trapero-Quintana" box="[313,873,1672,1694]" pageId="21" pageNumber="22" refString="Trapero-Quintana, A. D. &amp; Naranjo-Lopez, J. C. (2003) Revision of the order Odonata in Cuba. Bulletin of American Odonatology, 7 (2), 23 - 40." type="journal article" year="2003">
Trapero-Quintana &amp; Naranjo-López (2003: 27;
<collectingCountry id="228DDBDFFFBB4E42FCE7F903FCDA6DD9" box="[813,866,1672,1694]" name="Cuba" pageId="21" pageNumber="22">Cuba</collectingCountry>
)
</bibRefCitation>
;—Trapero-Quintana &amp; Naranjo-López (2004: 180; in key for
<collectingCountry id="228DDBDFFFBB4E42FEDFF92DFEF36DFB" box="[277,331,1702,1724]" name="Cuba" pageId="21" pageNumber="22">Cuba</collectingCountry>
);—
<bibRefCitation id="3E0BE6BEFFBB4E42FEA5F92DFB516D9C" author="Westfall" pageId="21" pageNumber="22" refString="Westfall, Jr., M. J. &amp; May, M. L. (2006) Damselflies of North America. Revised edition. Scientific Publishers, Inc., Gainesville, FL, xii + 502 pp." type="book" year="2006">
Westfall &amp; May (2006: 431433, 438, 439, Figs. 223A, 224C, 226C; in key to northern representatives of genus, characterization of adults, illustrations of
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thorax and S10,
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pronotum and mesostigmal plates)
</bibRefCitation>
;—
<treatmentCitation id="DB3BBD5EFFBB4E42FACDF94EFE886DBD" author="Garrison" page="20" pageId="21" pageNumber="22" year="2003">
<bibRefCitation id="3E0BE6BEFFBB4E42FACDF94EFEBE6DBD" author="Garrison" pageId="21" pageNumber="22" refString="Garrison, R. W., von Ellenrieder, N. &amp; O'Brien, M. F. (2003) An annotated list of the name bearing types of species-group names in Odonata preserved in the University of Michigan, Museum of Zoology. Occasional Papers of the Museum of Zoology, University of Michigan, 736, 1 - 73." type="journal article" year="2003">
Garrison
<emphasis id="68EE475DFFBB4E42FAA3F94DFA256D9C" box="[1385,1437,1733,1755]" italics="true" pageId="21" pageNumber="22">et al.</emphasis>
(2003
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: 20
</treatmentCitation>
;
<typeStatus id="852125EDFFBB4E42FE8BF96EFED56DBD" box="[321,365,1765,1786]" pageId="21" pageNumber="22">type</typeStatus>
information);—
<bibRefCitation id="3E0BE6BEFFBB4E42FDDDF96FFC476DBD" author="Pessacq" box="[535,1023,1764,1786]" pageId="21" pageNumber="22" refString="Pessacq, P. (2008) Phylogeny of Neotropical Protoneuridae (Odonata: Zygoptera) and a preliminary study of their relationship with related families. Systematic Entomology, 33, 511 - 528. https: // doi. org / 10.1111 / j. 1365 - 3113.2007.00414. x" type="journal article" year="2008">Pessacq (2008: 527; in phylogenetic analysis)</bibRefCitation>
;—
<bibRefCitation id="3E0BE6BEFFBB4E42FBD6F96FFEC16C5F" author="Trapero-Quintana" pageId="21" pageNumber="22" refString="Trapero-Quintana, A. D. &amp; Torres-Cambas, Y. (2008) Actualizacion sistematica de la odonatofauna cubana (Insecta: Odonata). Cocuyo, 17, 25 - 28." type="journal article" year="2008">
Trapero-Quintana &amp; Torres-Cambas (2008: 26;
<collectingCountry id="228DDBDFFFBB4E42FEF6F889FECB6C5F" box="[316,371,1794,1816]" name="Cuba" pageId="21" pageNumber="22">Cuba</collectingCountry>
)
</bibRefCitation>
;—
<treatmentCitation id="DB3BBD5EFFBB4E42FE5DF889FD166C5F" author="Garrison" box="[407,686,1794,1816]" page="379" pageId="21" pageNumber="22" year="2010">
<bibRefCitation id="3E0BE6BEFFBB4E42FE5DF889FDC16C5F" author="Garrison" box="[407,633,1794,1816]" pageId="21" pageNumber="22" refString="Garrison, R. W., von Ellenrieder, N. &amp; Louton, J. A. (2010) Damselfly genera of the New World. An Illustrated and Annotated Key to the Zygoptera. The Johns Hopkins University Press, Baltimore, xiv + 490 pp, + 24 pls." type="book" year="2010">
Garrison
<emphasis id="68EE475DFFBB4E42FE31F888FD896C5F" box="[507,561,1794,1816]" italics="true" pageId="21" pageNumber="22">et al.</emphasis>
(2010
</bibRefCitation>
: 379
</treatmentCitation>
, 381, Figs. 2510, 2511, 2514; illustrations
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genital ligula and S10);— González-Soriano
<emphasis id="68EE475DFFBB4E42FE42F8A9FE786C70" box="[392,448,1825,1847]" italics="true" pageId="21" pageNumber="22">et al.</emphasis>
(2011: 184;
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<collectingRegion id="985E55ADFFBB4E42FD86F8AAFD236C70" box="[588,667,1825,1847]" country="Mexico" name="Mexico" pageId="21" pageNumber="22">Mexico</collectingRegion>
</collectingCountry>
);—
<bibRefCitation id="3E0BE6BEFFBB4E42FD0AF8AAFEA56C11" author="Gonzalez-Soriano" pageId="21" pageNumber="22" refString="Gonzalez-Soriano, E. &amp; Novelo-Gutierrez, R. (2011) Libelulas (Insecta: Odonata), pp. 295 - 306. In: Cruz Angon, A. (Ed.) La Biodiversidad en Veracruz, estudio de estado. Comision Nacional para el Conocimiento y Uso de la Biodiversidad (CONABIO), Mexico." type="book chapter" year="2011">
González-Soriano &amp; Novelo-Gutiérrez (2011: 2, Appendix VIII.21.2;
<collectingCountry id="228DDBDFFFBB4E42FF0DF8CBFEAE6C11" box="[199,278,1856,1878]" name="Mexico" pageId="21" pageNumber="22">
<collectingRegion id="985E55ADFFBB4E42FF0DF8CBFEAE6C11" box="[199,278,1856,1878]" country="Mexico" name="Mexico" pageId="21" pageNumber="22">Mexico</collectingRegion>
</collectingCountry>
)
</bibRefCitation>
;—
<bibRefCitation id="3E0BE6BEFFBB4E42FEF0F8CBFD756C11" author="Meurgey" box="[314,717,1856,1878]" pageId="21" pageNumber="22" refString="Meurgey, F. (2013) A catalogue of the West Indian dragonflies (Insecta: Odonata). Annales de la Societe entomologique de France (N. S.): International Journal of Entomology, 49 (3), 298 - 334. https: // doi. org / 10.1080 / 00379271.2013.848066" type="journal article" year="2013">Meurgey (2013: 300, 305; distribution)</bibRefCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="5A259B4FFFBB4E42FF5DF809FB626CDD" blockId="21.[151,1436,1922,2019]" box="[151,1242,1922,1947]" pageId="21" pageNumber="22">
<materialsCitation id="EAF29112FFBB4E42FF5DF809FB626CDD" ID-GBIF-Occurrence="1703062704" box="[151,1242,1922,1947]" collectingDate="1905-02-18" collectionCode="UMMZ" collectorName="E. B. Williamson" country="Guatemala" location="Livingston" pageId="21" pageNumber="22" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="68EE475DFFBB4E42FF5DF809FEF06CDC" bold="true" box="[151,328,1922,1947]" pageId="21" pageNumber="22">
Primary
<typeStatus id="852125EDFFBB4E42FEC9F809FEFB6CDC" box="[259,323,1922,1947]" pageId="21" pageNumber="22">types</typeStatus>
.
</emphasis>
<typeStatus id="852125EDFFBB4E42FE85F809FE0F6CDC" box="[335,439,1922,1947]" pageId="21" pageNumber="22" type="holotype">Holotype</typeStatus>
<specimenCount id="4C9C50C6FFBB4E42FE74F808FE6E6CDD" box="[446,470,1923,1946]" pageId="21" pageNumber="22" type="male"></specimenCount>
.
<collectingCountry id="228DDBDFFFBB4E42FE2BF809FDE76CDC" box="[481,607,1922,1947]" name="Guatemala" pageId="21" pageNumber="22">Guatemala</collectingCountry>
:
<location id="5F45CD94FFBB4E42FDA3F809FD5D6CDC" LSID="urn:lsid:plazi:treatment:D2332A59FFBB4E41FF5DFA0FFC286CD3:5F45CD94FFBB4E42FDA3F809FD5D6CDC" box="[617,741,1922,1947]" country="Guatemala" name="Livingston" pageId="21" pageNumber="22">Livingston</location>
,
<date id="2E24BD8FFFBB4E42FD3BF808FCDD6CDC" box="[753,869,1922,1947]" pageId="21" pageNumber="22" value="1905-02-18">
<collectingDate id="3E604467FFBB4E42FD3BF808FCDD6CDC" box="[753,869,1922,1947]" pageId="21" pageNumber="22" value="1905-02-18">18 ii 1905</collectingDate>
</date>
,
<collectorName id="F76FFE99FFBB4E42FCBBF808FB8A6CDC" box="[881,1074,1922,1947]" pageId="21" pageNumber="22">E. B. Williamson</collectorName>
leg. [
<collectionCode id="3C8B038AFFBB4E42FBBAF808FB686CDD" box="[1136,1232,1923,1946]" country="USA" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="21" pageNumber="22">UMMZ</collectionCode>
].
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="1280C8C4FFBB4E41FF0DF82DFE056E12" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22" type="description">
<paragraph id="5A259B4FFFBB4E41FF0DF82DFD7469C3" blockId="21.[151,1436,1922,2019]" lastBlockId="22.[151,1437,151,1941]" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22">
<materialsCitation id="EAF29112FFBB4E42FF0DF82DFCD66CF8" ID-GBIF-Occurrence="1703062749" box="[199,878,1958,1983]" country="Mexico" location="MEXICO" pageId="21" pageNumber="22" specimenCount="36" specimenCount-female="19" specimenCount-male="17" stateProvince="MEXICO">
<emphasis id="68EE475DFFBB4E42FF0DF82DFE7F6CF8" bold="true" box="[199,455,1958,1983]" pageId="21" pageNumber="22">Specimens examined.</emphasis>
<specimenCount id="4C9C50C6FFBB4E42FE07F82CFDBD6CF9" box="[461,517,1959,1983]" pageId="21" pageNumber="22" type="male">17 ♂</specimenCount>
<specimenCount id="4C9C50C6FFBB4E42FDC6F82CFDFF6CF9" box="[524,583,1959,1983]" pageId="21" pageNumber="22" type="female">19 ♀</specimenCount>
:
<collectingCountry id="228DDBDFFFBB4E42FD9EF82CFD7D6CF8" box="[596,709,1959,1983]" name="Mexico" pageId="21" pageNumber="22">
<collectingRegion id="985E55ADFFBB4E42FD9EF82CFD7D6CF8" box="[596,709,1959,1983]" country="Mexico" name="Mexico" pageId="21" pageNumber="22">MEXICO</collectingRegion>
</collectingCountry>
,
<collectingRegion id="985E55ADFFBB4E42FD1AF82CFC926CF9" box="[720,810,1959,1982]" country="Mexico" name="Yucatan" pageId="21" pageNumber="22">Yucatan</collectingRegion>
State
</materialsCitation>
:
<materialsCitation id="EAF29112FFBB4E42FCB2F82CFD266CA5" ID-GBIF-Occurrence="1703062682" collectingDate="1932-07-22" collectionCode="UMMZ" collectorName="Xlacah Cenote &amp; E. P. Creaser" country="Mexico" elevation="6" latitude="21.083334" location="Merida" longLatPrecision="1265" longitude="-89.583336" pageId="21" pageNumber="22" specimenCount="1" specimenCount-male="1" stateProvince="Merida">
<specimenCount id="4C9C50C6FFBB4E42FCB2F82CFC1C6CF9" box="[888,932,1959,1983]" pageId="21" pageNumber="22" type="male">1 ♂</specimenCount>
,
<collectorName id="F76FFE99FFBB4E42FC7AF82DFBE16CF8" box="[944,1113,1958,1983]" pageId="21" pageNumber="22">Xlacah Cenote</collectorName>
,
<collectingRegion id="985E55ADFFBB4E42FBAFF82DFB0E6CF8" box="[1125,1206,1958,1983]" country="Venezuela" name="Merida" pageId="21" pageNumber="22">Merida</collectingRegion>
{
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,
<geoCoordinate id="3FAEFD88FFBB4E42FAE4F82DFA206CF9" box="[1326,1432,1958,1983]" direction="west" orientation="longitude" pageId="21" pageNumber="22" precision="925" value="-89.583336">89°35' W</geoCoordinate>
,
<elevation id="D1B77C7CFFBB4E42FF5DF841FF706CA4" box="[151,200,1994,2019]" metricMagnitude="0" metricUnit="m" metricValue="6.0" pageId="21" pageNumber="22" unit="m" value="6.0">
<quantity id="9D6236AAFFBB4E42FF5DF841FF706CA4" box="[151,200,1994,2019]" metricMagnitude="0" metricUnit="m" metricValue="6.0" pageId="21" pageNumber="22" unit="m" value="6.0">6 m</quantity>
</elevation>
},
<date id="2E24BD8FFFBB4E42FF15F840FEDB6CA4" box="[223,355,1994,2019]" pageId="21" pageNumber="22" value="1932-07-22">
<collectingDate id="3E604467FFBB4E42FF15F840FEDB6CA4" box="[223,355,1994,2019]" pageId="21" pageNumber="22" value="1932-07-22">22 vii 1932</collectingDate>
</date>
,
<collectorName id="F76FFE99FFBB4E42FEA4F840FE4F6CA4" box="[366,503,1995,2019]" pageId="21" pageNumber="22">E.P. Creaser</collectorName>
leg. [
<collectionCode id="3C8B038AFFBB4E42FDFDF840FD2F6CA5" box="[567,663,1995,2018]" country="USA" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="21" pageNumber="22">UMMZ</collectionCode>
]
</materialsCitation>
;
<materialsCitation id="EAF29112FFBB4E41FD60F840FE3F6B93" ID-GBIF-Occurrence="1703062720" collectingDate="1976-08-18" collectorName="R. W. Garrison &amp; J. A. Garrison" country="Mexico" elevation="349" lastPageId="22" lastPageNumber="23" latitude="18.426111" location="Caldera Lake" longLatPrecision="21" longitude="-95.09473" pageId="21" pageNumber="22" specimenCount="19" specimenCount-female="11" specimenCount-male="8" stateProvince="Veracruz">
<collectingRegion id="985E55ADFFBB4E42FD60F840FCB66CA5" box="[682,782,1995,2018]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="21" pageNumber="22">Veracruz</collectingRegion>
State:
<specimenCount id="4C9C50C6FFBB4E42FC96F840FC3F6CA5" box="[860,903,1995,2019]" pageId="21" pageNumber="22" type="male">8 ♂</specimenCount>
<specimenCount id="4C9C50C6FFBB4E42FC45F840FC7E6CA5" box="[911,966,1995,2019]" pageId="21" pageNumber="22" type="female">11 ♀</specimenCount>
(six pairs in tandem),
<location id="5F45CD94FFBB4E42FB0CF841FAD86CA4" LSID="urn:lsid:plazi:treatment:D2332A59FFBB4E41FF5DFA0FFC286CD3:5F45CD94FFBB4E42FB0CF841FAD86CA4" box="[1222,1376,1994,2019]" country="Mexico" latitude="18.426111" longLatPrecision="21" longitude="-95.09473" name="Caldera Lake" pageId="21" pageNumber="22" stateProvince="Veracruz">Caldera Lake</location>
, near Motel Playa Azul and Laguna Catemaco (
<geoCoordinate id="3FAEFD88FFB84E41FDBFFF1CFCBB6BE8" box="[629,771,151,176]" direction="north" orientation="latitude" pageId="22" pageNumber="23" precision="15" value="18.426111">18°25'34'' N</geoCoordinate>
,
<geoCoordinate id="3FAEFD88FFB84E41FCC5FF1CFC2B6BE8" box="[783,915,151,176]" direction="west" orientation="longitude" pageId="22" pageNumber="23" precision="15" value="-95.09473">95°5'41'' W</geoCoordinate>
,
<quantity id="9D6236AAFFB84E41FC55FF13FC526BF7" box="[927,1002,152,176]" metricMagnitude="2" metricUnit="m" metricValue="3.4899999999999998" pageId="22" pageNumber="23" unit="m" value="349.0">
<elevation id="D1B77C7CFFB84E41FC55FF13FC526BF7" box="[927,1002,152,176]" metricMagnitude="2" metricUnit="m" metricValue="3.4899999999999998" pageId="22" pageNumber="23" unit="m" value="349.0">349 m</elevation>
</quantity>
),
<date id="2E24BD8FFFB84E41FC35FF13FB226BF7" box="[1023,1178,151,176]" pageId="22" pageNumber="23" value="1976-08-18">
<collectingDate id="3E604467FFB84E41FC35FF13FB226BF7" box="[1023,1178,151,176]" pageId="22" pageNumber="23" value="1976-08-18">18 viii 1976</collectingDate>
</date>
,
<collectorName id="F76FFE99FFB84E41FB6CFF13FAF16BF7" box="[1190,1353,151,176]" pageId="22" pageNumber="23">R.W. Garrison</collectorName>
&amp;
<collectorName id="F76FFE99FFB84E41FAA5FF13FF416B92" pageId="22" pageNumber="23">J.A. Garrison</collectorName>
leg. [RWG]
</materialsCitation>
;
<materialsCitation id="EAF29112FFB84E41FE58FF37FC3E6B93" ID-GBIF-Occurrence="1703062709" box="[402,902,188,213]" collectingDate="1976-08-18" collectionCode="CSCA" collectorName="R. W. Garrison &amp; J. A. Garrison" country="Mexico" elevation="349" latitude="18.426111" location="Caldera Lake" longLatPrecision="21" longitude="-95.09473" pageId="22" pageNumber="23" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Veracruz">
<specimenCount id="4C9C50C6FFB84E41FE58FF37FE046B93" box="[402,444,188,212]" pageId="22" pageNumber="23" type="male">1 ♂</specimenCount>
<specimenCount id="4C9C50C6FFB84E41FE09FF37FE566B93" box="[451,494,188,212]" pageId="22" pageNumber="23" type="female">1 ♀</specimenCount>
(in tandem), same data but [
<collectionCode id="3C8B038AFFB84E41FCE4FF37FCC76B93" box="[814,895,188,212]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33156" name="California State Collection of Arthropods" pageId="22" pageNumber="23">CSCA</collectionCode>
]
</materialsCitation>
;
<materialsCitation id="EAF29112FFB84E41FC5BFF37FA3B6B93" ID-GBIF-Occurrence="1703062714" box="[913,1411,188,213]" collectingDate="1976-08-18" collectionCode="FSCA" collectorName="R. W. Garrison &amp; J. A. Garrison" country="Mexico" elevation="349" latitude="18.426111" location="Caldera Lake" longLatPrecision="21" longitude="-95.09473" pageId="22" pageNumber="23" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Veracruz">
<specimenCount id="4C9C50C6FFB84E41FC5BFF37FC046B93" box="[913,956,188,212]" pageId="22" pageNumber="23" type="male">1 ♂</specimenCount>
<specimenCount id="4C9C50C6FFB84E41FC09FF37FC556B93" box="[963,1005,188,212]" pageId="22" pageNumber="23" type="female">1 ♀</specimenCount>
(in tandem), same data but [
<collectionCode id="3C8B038AFFB84E41FAE7FF37FAC46B93" box="[1325,1404,188,212]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="22" pageNumber="23">FSCA</collectionCode>
]
</materialsCitation>
;
<materialsCitation id="EAF29112FFB84E41FA44FF37FD2F6BB0" ID-GBIF-Occurrence="1703062686" collectingDate="1976-08-18" collectionCode="UMMZ" collectorName="R. W. Garrison &amp; J. A. Garrison" country="Mexico" elevation="349" latitude="18.426111" location="Caldera Lake" longLatPrecision="21" longitude="-95.09473" pageId="22" pageNumber="23" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Veracruz">
<specimenCount id="4C9C50C6FFB84E41FA44FF37FF146BB0" pageId="22" pageNumber="23" type="male">1 ♂</specimenCount>
<specimenCount id="4C9C50C6FFB84E41FF7FFF6BFF5A6BB0" box="[181,226,224,248]" pageId="22" pageNumber="23" type="female">1 ♀</specimenCount>
(in tandem), same data but [
<collectionCode id="3C8B038AFFB84E41FDFAFF6BFD286BB0" box="[560,656,224,247]" country="USA" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="22" pageNumber="23">UMMZ</collectionCode>
]
</materialsCitation>
;
<materialsCitation id="EAF29112FFB84E41FD6EFF6BFF556A07" ID-GBIF-Occurrence="1703062689" collectingDate="1982-08-07" collectingDateMax="1982-08-14" collectingDateMin="1982-08-07" collectorName="R. W. Garrison" country="Mexico" county="Lagunas Azul" elevation="600" latitude="18.42" location="Los Tuxtlas Biological Station" longLatPrecision="21" longitude="-95.11" municipality="Emilia" pageId="22" pageNumber="23" specimenCount="1" specimenCount-male="1" stateProvince="Veracruz">
<specimenCount id="4C9C50C6FFB84E41FD6EFF6BFD6B6BB0" box="[676,723,224,248]" pageId="22" pageNumber="23" type="male">1 ♂</specimenCount>
, vicinity of
<collectingCounty id="B344E3C3FFB84E41FCABFF6BFBB86BBF" box="[865,1024,223,248]" pageId="22" pageNumber="23">Lagunas Azul</collectingCounty>
and
<collectingMunicipality id="BA410135FFB84E41FBF6FF54FB316BBF" box="[1084,1161,223,248]" pageId="22" pageNumber="23">Emilia</collectingMunicipality>
,
<location id="5F45CD94FFB84E41FB5DFF6BFF516A5A" LSID="urn:lsid:plazi:treatment:D2332A59FFBB4E41FF5DFA0FFC286CD3:5F45CD94FFB84E41FB5DFF6BFF516A5A" country="Mexico" county="Lagunas Azul" latitude="18.42" longLatPrecision="21" longitude="-95.11" municipality="Emilia" name="Los Tuxtlas Biological Station" pageId="22" pageNumber="23" stateProvince="Veracruz">Los Tuxtlas Biological Station</location>
, about
<locationDeviation id="4B97826DFFB84E41FE8AFE8FFDE06A5B" box="[320,600,260,285]" pageId="22" pageNumber="23">
<quantity id="9D6236AAFFB84E41FE8AFE8FFE336A5A" box="[320,395,260,285]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="22" pageNumber="23" unit="km" value="30.0">30 km</quantity>
NE of Catemaco
</locationDeviation>
(
<geoCoordinate id="3FAEFD88FFB84E41FDA6FE8FFD456A5B" box="[620,765,260,285]" direction="north" orientation="latitude" pageId="22" pageNumber="23" precision="15" value="18.42">18°25'12'' N</geoCoordinate>
,
<geoCoordinate id="3FAEFD88FFB84E41FCC1FE8FFC2A6A5B" box="[779,914,260,285]" direction="west" orientation="longitude" pageId="22" pageNumber="23" precision="15" value="-95.11">95°6'36'' W</geoCoordinate>
,
<quantity id="9D6236AAFFB84E41FC6AFE8FFC566A5B" box="[928,1006,260,285]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="22" pageNumber="23" unit="m" value="600.0">
<elevation id="D1B77C7CFFB84E41FC6AFE8FFC566A5B" box="[928,1006,260,285]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="22" pageNumber="23" unit="m" value="600.0">600 m</elevation>
</quantity>
),
<date id="2E24BD8FFFB84E41FBCCFE8FFB0C6A5B" box="[1030,1204,260,285]" pageId="22" pageNumber="23" value="1982-08-07" valueMax="1982-08-14" valueMin="1982-08-07">
<collectingDate id="3E604467FFB84E41FBCCFE8FFB0C6A5B" box="[1030,1204,260,285]" pageId="22" pageNumber="23" value="1982-08-07" valueMax="1982-08-14" valueMin="1982-08-07">714 viii 1982</collectingDate>
</date>
,
<collectorName id="F76FFE99FFB84E41FB09FE8EFAD06A5A" box="[1219,1384,260,285]" pageId="22" pageNumber="23">R.W. Garrison</collectorName>
leg. [RWG]
</materialsCitation>
;
<materialsCitation id="EAF29112FFB84E41FF37FEA3FD2B6A22" ID-GBIF-Occurrence="1703062716" collectingDate="1978-08-03" collectionCode="FSCA" collectorName="Laguna &amp; E. Gonzalez" country="Belize" elevation="508" latitude="18.411388" location="Cayo Dist." longLatPrecision="21" longitude="-95.13084" pageId="22" pageNumber="23" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="BELIZE">
<specimenCount id="4C9C50C6FFB84E41FF37FEA3FE936A78" box="[253,299,296,320]" pageId="22" pageNumber="23" type="male">2 ♂</specimenCount>
<specimenCount id="4C9C50C6FFB84E41FEFDFEA3FEDE6A78" box="[311,358,296,320]" pageId="22" pageNumber="23" type="female">1 ♀</specimenCount>
,
<collectorName id="F76FFE99FFB84E41FEB3FEA3FE756A07" box="[377,461,296,320]" pageId="22" pageNumber="23">Laguna</collectorName>
del Cacahuateno, Los Tuxtlas {
<geoCoordinate id="3FAEFD88FFB84E41FC8DFEACFC646A78" box="[839,988,295,320]" direction="north" orientation="latitude" pageId="22" pageNumber="23" precision="15" value="18.411388">18°24'41'' N</geoCoordinate>
,
<geoCoordinate id="3FAEFD88FFB84E41FC26FEACFBCB6A78" box="[1004,1139,295,320]" direction="west" orientation="longitude" pageId="22" pageNumber="23" precision="15" value="-95.13084">95°7'51'' W</geoCoordinate>
,
<quantity id="9D6236AAFFB84E41FB49FEACFB6D6A07" box="[1155,1237,295,320]" metricMagnitude="2" metricUnit="m" metricValue="5.08" pageId="22" pageNumber="23" unit="m" value="508.0">
<elevation id="D1B77C7CFFB84E41FB49FEACFB6D6A07" box="[1155,1237,295,320]" metricMagnitude="2" metricUnit="m" metricValue="5.08" pageId="22" pageNumber="23" unit="m" value="508.0">508 m</elevation>
</quantity>
},
<date id="2E24BD8FFFB84E41FB25FEA3FACC6A07" box="[1263,1396,295,320]" pageId="22" pageNumber="23" value="1978-08-03">
<collectingDate id="3E604467FFB84E41FB25FEA3FACC6A07" box="[1263,1396,295,320]" pageId="22" pageNumber="23" value="1978-08-03">3 viii 1978</collectingDate>
</date>
,
<collectorName id="F76FFE99FFB84E41FA4EFEA3FEB96A22" pageId="22" pageNumber="23">E. González</collectorName>
leg. [
<collectionCode id="3C8B038AFFB84E41FE88FEC7FE296A23" box="[322,401,332,356]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="22" pageNumber="23">FSCA</collectionCode>
].
<collectingRegion id="985E55ADFFB84E41FE6FFEC6FDB06A23" box="[421,520,333,356]" country="Belize" name="Belize" pageId="22" pageNumber="23">
<collectingCountry id="228DDBDFFFB84E41FE6FFEC6FDB06A23" box="[421,520,333,356]" name="Belize" pageId="22" pageNumber="23">BELIZE</collectingCountry>
</collectingRegion>
,
<location id="5F45CD94FFB84E41FDDFFEC7FD2B6A22" LSID="urn:lsid:plazi:treatment:D2332A59FFBB4E41FF5DFA0FFC286CD3:5F45CD94FFB84E41FDDFFEC7FD2B6A22" box="[533,659,332,357]" country="Belize" latitude="18.411388" longLatPrecision="21" longitude="-95.13084" name="Cayo Dist." pageId="22" pageNumber="23" stateProvince="BELIZE">Cayo Dist.</location>
</materialsCitation>
: 1 ♂, stream on W highway at Unitedville School,
<date id="2E24BD8FFFB84E41FB29FEC7FAE26A23" box="[1251,1370,332,356]" pageId="22" pageNumber="23" value="1993-05-30">30 v 1993</date>
, S.W. Dunkle leg. [FSCA];
<materialsCitation id="EAF29112FFB84E41FE42FEE4FEA26AEA" ID-GBIF-Occurrence="1703062698" collectorName="San Antonio" country="Belize" elevation="5" location="Southern" pageId="22" pageNumber="23" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Toledo">
<collectingRegion id="985E55ADFFB84E41FE42FEE4FE6D6ACF" box="[392,469,367,392]" country="Belize" name="Toledo" pageId="22" pageNumber="23">Toledo</collectingRegion>
Dist.:
<specimenCount id="4C9C50C6FFB84E41FDEBFEFBFDF36AC0" box="[545,587,368,392]" pageId="22" pageNumber="23" type="male">1 ♂</specimenCount>
<specimenCount id="4C9C50C6FFB84E41FD98FEFBFDC56AC0" box="[594,637,368,392]" pageId="22" pageNumber="23" type="female">1 ♀</specimenCount>
(in tandem),
<collectorName id="F76FFE99FFB84E41FCDEFEFBFC1C6ACF" box="[788,932,367,392]" pageId="22" pageNumber="23">San Antonio</collectorName>
, about
<quantity id="9D6236AAFFB84E41FC3EFEE4FBA76AC0" box="[1012,1055,367,392]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="22" pageNumber="23" unit="m" value="5.0">
<elevation id="D1B77C7CFFB84E41FC3EFEE4FBA76AC0" box="[1012,1055,367,392]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="22" pageNumber="23" unit="m" value="5.0">5 m</elevation>
</quantity>
W of
<location id="5F45CD94FFB84E41FBACFEE4FB736ACF" LSID="urn:lsid:plazi:treatment:D2332A59FFBB4E41FF5DFA0FFC286CD3:5F45CD94FFB84E41FBACFEE4FB736ACF" box="[1126,1227,367,392]" country="Belize" name="Southern" pageId="22" pageNumber="23" stateProvince="Toledo">Southern</location>
highway junction, borrow pits
</materialsCitation>
,
<materialsCitation id="EAF29112FFB84E41FEECFE1FFAA86AEB" ID-GBIF-Occurrence="1703062692" box="[294,1296,404,429]" collectingDate="1993-06-08" collectorName="S. W. Dunkle" country="Belize" elevation="124" latitude="16.233334" location="Bladen River" longLatPrecision="1282" longitude="-89.01667" pageId="22" pageNumber="23" specimenCount="1" stateProvince="Toledo">
<quantity id="9D6236AAFFB84E41FEECFE1FFEE16AEA" box="[294,345,404,429]" metricMagnitude="3" metricUnit="m" metricValue="3.218688" pageId="22" pageNumber="23" unit="mi" value="2.0">2 mi</quantity>
N of
<location id="5F45CD94FFB84E41FE51FE1FFD976AEA" LSID="urn:lsid:plazi:treatment:D2332A59FFBB4E41FF5DFA0FFC286CD3:5F45CD94FFB84E41FE51FE1FFD976AEA" box="[411,559,404,429]" country="Belize" latitude="16.233334" longLatPrecision="1282" longitude="-89.01667" name="Bladen River" pageId="22" pageNumber="23" stateProvince="Toledo">Bladen River</location>
{
<geoCoordinate id="3FAEFD88FFB84E41FD8AFE1FFD136AEB" box="[576,683,404,429]" direction="north" orientation="latitude" pageId="22" pageNumber="23" precision="925" value="16.233334">16°14' N</geoCoordinate>
,
<geoCoordinate id="3FAEFD88FFB84E41FD7DFE1FFCB26AEB" box="[695,778,404,428]" direction="west" orientation="longitude" pageId="22" pageNumber="23" precision="925" value="-89.01667">89°1'W</geoCoordinate>
,
<quantity id="9D6236AAFFB84E41FCDDFE1FFCDC6AEA" box="[791,868,404,429]" metricMagnitude="2" metricUnit="m" metricValue="1.24" pageId="22" pageNumber="23" unit="m" value="124.0">
<elevation id="D1B77C7CFFB84E41FCDDFE1FFCDC6AEA" box="[791,868,404,429]" metricMagnitude="2" metricUnit="m" metricValue="1.24" pageId="22" pageNumber="23" unit="m" value="124.0">124 m</elevation>
</quantity>
},
<date id="2E24BD8FFFB84E41FCB0FE1FFC506AEB" box="[890,1000,404,429]" pageId="22" pageNumber="23" value="1993-06-08">
<collectingDate id="3E604467FFB84E41FCB0FE1FFC506AEB" box="[890,1000,404,429]" pageId="22" pageNumber="23" value="1993-06-08">8 vi 1993</collectingDate>
</date>
,
<collectorName id="F76FFE99FFB84E41FC39FE1FFB3A6AEA" box="[1011,1154,404,429]" pageId="22" pageNumber="23">S.W. Dunkle</collectorName>
leg. [RWG]
</materialsCitation>
;
<materialsCitation id="EAF29112FFB84E41FAD6FE1FFD416AB3" ID-GBIF-Occurrence="1703062680" collectingDate="1993-06-07" collectionCode="FSCA" collectorName="Piedra Lagoon &amp; Punta Gorda &amp; S. W. Dunkle &amp; El Peten Dept." country="Guatemala" latitude="16.1" location="Southern Highway" longLatPrecision="1283" longitude="-88.78333" pageId="22" pageNumber="23" specimenCount="1" specimenCount-female="1" stateProvince="GUATEMALA">
<specimenCount id="4C9C50C6FFB84E41FAD6FE1FFAFE6AEB" box="[1308,1350,404,428]" pageId="22" pageNumber="23" type="female">1 ♀</specimenCount>
,
<collectorName id="F76FFE99FFB84E41FA9FFE1FFF576A97" pageId="22" pageNumber="23">Piedra Lagoon</collectorName>
,
<quantity id="9D6236AAFFB84E41FF34FE33FEF16A97" box="[254,329,439,464]" metricMagnitude="3" metricUnit="m" metricValue="1.4484096" pageId="22" pageNumber="23" unit="mi" value="0.9">0.9 mi</quantity>
S of junction
<location id="5F45CD94FFB84E41FE3DFE3CFD766A97" LSID="urn:lsid:plazi:treatment:D2332A59FFBB4E41FF5DFA0FFC286CD3:5F45CD94FFB84E41FE3DFE3CFD766A97" box="[503,718,439,464]" country="Guatemala" latitude="16.1" longLatPrecision="1283" longitude="-88.78333" name="Southern Highway" pageId="22" pageNumber="23" stateProvince="GUATEMALA">Southern Highway</location>
and
<collectorName id="F76FFE99FFB84E41FCC1FE33FC246A97" box="[779,924,439,464]" pageId="22" pageNumber="23">Punta Gorda</collectorName>
road {
<geoCoordinate id="3FAEFD88FFB84E41FC26FE3CFBF46A88" box="[1004,1100,439,464]" direction="north" orientation="latitude" pageId="22" pageNumber="23" precision="925" value="16.1">16°6' N</geoCoordinate>
,
<geoCoordinate id="3FAEFD88FFB84E41FB91FE33FB756A88" box="[1115,1229,440,464]" direction="west" orientation="longitude" pageId="22" pageNumber="23" precision="925" value="-88.78333">88°47' W</geoCoordinate>
],
<date id="2E24BD8FFFB84E41FB2EFE33FAE06A97" box="[1252,1368,439,464]" pageId="22" pageNumber="23" value="1993-06-07">
<collectingDate id="3E604467FFB84E41FB2EFE33FAE06A97" box="[1252,1368,439,464]" pageId="22" pageNumber="23" value="1993-06-07">7 vi 1993</collectingDate>
</date>
,
<collectorName id="F76FFE99FFB84E41FAADFE33FF526AB2" pageId="22" pageNumber="23">S.W. Dunkle</collectorName>
leg. [
<collectionCode id="3C8B038AFFB84E41FEE6FE57FEC36AB3" box="[300,379,476,500]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="22" pageNumber="23">FSCA</collectionCode>
].
<collectingCountry id="228DDBDFFFB84E41FE44FE57FD876AB3" box="[398,575,476,500]" name="Guatemala" pageId="22" pageNumber="23">
<collectingRegion id="985E55ADFFB84E41FE44FE57FD876AB3" box="[398,575,476,500]" country="Guatemala" name="Guatemala" pageId="22" pageNumber="23">GUATEMALA</collectingRegion>
</collectingCountry>
,
<collectorName id="F76FFE99FFB84E41FD87FE57FD416AB3" box="[589,761,476,501]" pageId="22" pageNumber="23">
<collectingRegion id="985E55ADFFB84E41FD87FE57FD156AB3" box="[589,685,476,501]" country="Guatemala" name="Peten" pageId="22" pageNumber="23">El Petén</collectingRegion>
Dept.
</collectorName>
</materialsCitation>
:
<materialsCitation id="EAF29112FFB84E41FCCFFE57FD0B695F" ID-GBIF-Occurrence="1703062683" collectingDate="1963-07-29" collectionCode="FSCA" collectorName="Tikal &amp; T. W. Donnelly" country="Guatemala" elevation="194" latitude="17.216667" location="Aguada" longLatPrecision="1279" longitude="-89.6" pageId="22" pageNumber="23" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Aguada">
<specimenCount id="4C9C50C6FFB84E41FCCFFE57FC896AB3" box="[773,817,476,500]" pageId="22" pageNumber="23" type="male">1 ♂</specimenCount>
<specimenCount id="4C9C50C6FFB84E41FCF1FE57FCDF6AB3" box="[827,871,476,500]" pageId="22" pageNumber="23" type="female">1 ♀</specimenCount>
(in copula),
<collectingRegion id="985E55ADFFB84E41FC37FE57FBEF6AB2" box="[1021,1111,476,501]" country="Puerto Rico" name="Aguada" pageId="22" pageNumber="23">Aguada</collectingRegion>
,
<collectorName id="F76FFE99FFB84E41FBAFFE57FB276AB2" box="[1125,1183,476,501]" pageId="22" pageNumber="23">Tikal</collectorName>
{
<geoCoordinate id="3FAEFD88FFB84E41FB78FE57FAA66AB3" box="[1202,1310,476,501]" direction="north" orientation="latitude" pageId="22" pageNumber="23" precision="925" value="17.216667">17°13' N</geoCoordinate>
,
<geoCoordinate id="3FAEFD88FFB84E41FAE6FE57FA206AB3" box="[1324,1432,476,501]" direction="west" orientation="longitude" pageId="22" pageNumber="23" precision="925" value="-89.6">89°36' W</geoCoordinate>
,
<quantity id="9D6236AAFFB84E41FF5DFE74FF65695F" box="[151,221,511,536]" metricMagnitude="2" metricUnit="m" metricValue="1.94" pageId="22" pageNumber="23" unit="m" value="194.0">
<elevation id="D1B77C7CFFB84E41FF5DFE74FF65695F" box="[151,221,511,536]" metricMagnitude="2" metricUnit="m" metricValue="1.94" pageId="22" pageNumber="23" unit="m" value="194.0">194m</elevation>
</quantity>
},
<date id="2E24BD8FFFB84E41FF3BFD8BFECB695F" box="[241,371,511,536]" pageId="22" pageNumber="23" value="1963-07-29">
<collectingDate id="3E604467FFB84E41FF3BFD8BFECB695F" box="[241,371,511,536]" pageId="22" pageNumber="23" value="1963-07-29">29 vii 1963</collectingDate>
</date>
,
<collectorName id="F76FFE99FFB84E41FEB7FD8BFD98695F" box="[381,544,511,536]" pageId="22" pageNumber="23">T.W. Donnelly</collectorName>
leg. [
<collectionCode id="3C8B038AFFB84E41FD94FD8BFD14695F" box="[606,684,512,536]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="22" pageNumber="23">FSCA</collectionCode>
]
</materialsCitation>
; 1 ♀, E of Laguna Perdida {17°3' N, 90°13' W,
<quantity id="9D6236AAFFB84E41FB0EFD8BFABA695F" box="[1220,1282,511,536]" metricMagnitude="1" metricUnit="m" metricValue="7.4" pageId="22" pageNumber="23" unit="m" value="74.0">74 m</quantity>
},
<date id="2E24BD8FFFB84E41FADDFD8BFA20695F" box="[1303,1432,511,536]" pageId="22" pageNumber="23" value="1970-07-19">19 vii 1970</date>
, J.T. MacLean leg. [FSCA]; 1 ♂, Flores,
<date id="2E24BD8FFFB84E41FDA3FDAFFCAA697B" box="[617,786,548,573]" pageId="22" pageNumber="23" value="1980-06-24" valueMax="1980-06-30" valueMin="1980-06-24">2430 vi 1980</date>
, W. Brehme leg. [FSCA];
<materialsCitation id="EAF29112FFB84E41FB8CFDAFFBBB6918" ID-GBIF-Occurrence="1703062667" collectingDate="1905-02-18" collectionCode="UMMZ" collectorName="Livingston &amp; E. B. Williamson" country="Guatemala" latitude="15.833333" location="Izabal" longLatPrecision="1284" longitude="-88.75" pageId="22" pageNumber="23" specimenCount="1" specimenCount-male="1" stateProvince="Izabal" typeStatus="paratype">
<collectingRegion id="985E55ADFFB84E41FB8CFDAFFB32697A" box="[1094,1162,548,573]" country="Guatemala" name="Izabal" pageId="22" pageNumber="23">Izabal</collectingRegion>
Dept.:
<specimenCount id="4C9C50C6FFB84E41FB28FDAFFAB6697B" box="[1250,1294,548,572]" pageId="22" pageNumber="23" type="male">1 ♂</specimenCount>
(
<emphasis id="68EE475DFFB84E41FAEAFDAFFA33697A" bold="true" box="[1312,1419,548,573]" pageId="22" pageNumber="23">
<typeStatus id="852125EDFFB84E41FAEAFDAFFA33697A" box="[1312,1419,548,573]" pageId="22" pageNumber="23" type="paratype">paratype</typeStatus>
</emphasis>
),
<collectorName id="F76FFE99FFB84E41FF5DFDCCFEA86927" box="[151,272,583,608]" pageId="22" pageNumber="23">Livingston</collectorName>
{
<geoCoordinate id="3FAEFD88FFB84E41FEEBFDCCFE346918" box="[289,396,583,608]" direction="north" orientation="latitude" pageId="22" pageNumber="23" precision="925" value="15.833333">15°50' N</geoCoordinate>
,
<geoCoordinate id="3FAEFD88FFB84E41FE53FDCCFDB26927" box="[409,522,583,608]" direction="west" orientation="longitude" pageId="22" pageNumber="23" precision="925" value="-88.75">88°45' W</geoCoordinate>
},
<date id="2E24BD8FFFB84E41FDD5FDC3FD2E6927" box="[543,662,583,608]" pageId="22" pageNumber="23" value="1905-02-18">
<collectingDate id="3E604467FFB84E41FDD5FDC3FD2E6927" box="[543,662,583,608]" pageId="22" pageNumber="23" value="1905-02-18">18 ii 1905</collectingDate>
</date>
,
<collectorName id="F76FFE99FFB84E41FD68FDC3FCE56927" box="[674,861,583,608]" pageId="22" pageNumber="23">E.B. Williamson</collectorName>
leg. [
<collectionCode id="3C8B038AFFB84E41FC57FDC3FC456918" box="[925,1021,584,607]" country="USA" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="22" pageNumber="23">UMMZ</collectionCode>
]
</materialsCitation>
; 2 ♂, Morales {15°29' N, 88°49' W,
<quantity id="9D6236AAFFB84E41FF5DFDE7FF6E69C2" box="[151,214,620,645]" metricMagnitude="1" metricUnit="m" metricValue="4.0" pageId="22" pageNumber="23" unit="m" value="40.0">40 m</quantity>
},
<date id="2E24BD8FFFB84E41FF21FDE7FEE669C3" box="[235,350,620,644]" pageId="22" pageNumber="23" value="1909-05-27">27 v 1909</date>
, E.B. Williamson leg. [UMMZ].
</paragraph>
<paragraph id="5A259B4FFFB84E41FF0DFD04FD386F1F" blockId="22.[151,1437,151,1941]" pageId="22" pageNumber="23">
<emphasis id="68EE475DFFB84E41FF0DFD04FE2269EF" bold="true" box="[199,410,655,680]" pageId="22" pageNumber="23">Characterization.</emphasis>
Male: Epicranium black with metallic copper and green reflections, thorax dorsum black and orange, dorsum of S12, distal portion of S35, and S610 black; pale colors orange and yellow (
<figureCitation id="C2A187CAFFB84E41FA8CFD3FFA36698B" box="[1350,1422,692,717]" captionStart="FIGURES 58" captionStartId="17.[151,264,1532,1555]" captionTargetBox="[162,1429,524,1500]" captionTargetId="figure@17.[151,1436,519,1510]" captionTargetPageId="17" captionText="FIGURES 58. Male body scan; head in dorsal view, thorax and abdomen in lateral view (not to scale). (5) P. calverti; (6) P. capillaris; (7) P.cara; (8) P.corculum." httpUri="https://zenodo.org/record/1069341/files/figure.png" pageId="22" pageNumber="23">Fig. 8</figureCitation>
). Pronotum black with anterior and lateral margins orange. Mesepisternum orange with middorsal carina and dorsal margin black; mesepimeron black with small orange spot on anterodorsal corner; metepisternum yellow with distal 1/4 black, with an orange spot on anterodorsal corner; metepimeron entirely yellow. Coxa, trochanter, and basal half of femur yellow; remainder of femur brown; tibia and tarsi yellow with brown margins; tibial spurs shorter than twice intervening spaces. Genital ligula lacking lateral lobes and with a straight distal margin, with margins of distal segment spiculate, and laterodistal corners projected anteroventrally (
<figureCitation id="C2A187CAFFB84E41FBC4FC07FBD068E3" box="[1038,1128,908,933]" captionStart="FIGURES 7178" captionStartId="55.[151,264,1923,1946]" captionTargetBox="[158,1426,206,1865]" captionTargetId="figure@55.[155,1440,193,1880]" captionTargetPageId="55" captionText="FIGURES 7178. Genital ligula inectal (7178a) and lateral (7178b) views (to scale). (71) P.ailsa; (72) P.amatoria; (73) P. aurantiaca; (74) P.caligata; (75) P.calverti; (76) P.capillaris; (77) P.cara; (78) P.corculum. All by RWG." httpUri="https://zenodo.org/record/1069365/files/figure.png" pageId="22" pageNumber="23">Fig. 78</figureCitation>
). Cercus shorter than S10 length, shorter than half of paraprocts length, about as long as wide, with a depression on external lateral surface (
<figureCitation id="C2A187CAFFB84E41FF55FC5FFEB568AB" box="[159,269,980,1005]" captionStart="FIGURES 103105" captionStartId="61.[151,264,1910,1933]" captionTargetBox="[180,1444,206,1860]" captionTargetId="figure@61.[164,1449,193,1867]" captionTargetPageId="61" captionText="FIGURES 103105. Male caudal appendages in lateral (103105a), posterodorsal (103105b), and dorsal (103105b) views (not to scale). (103) P.cara; (104)P.corculum; (105)P.cupida." httpUri="https://zenodo.org/record/1069378/files/figure.png" pageId="22" pageNumber="23">Fig. 104a</figureCitation>
); medial surface with a small blunt ventrobasal and a rounded mediobasal tooth; tip slightly bifid, with a stronger outer tooth and a lower blunt medial tooth, visible in dorsal and mediodorsal views (
<figureCitation id="C2A187CAFFB84E41FB3BFC7CFA366F57" box="[1265,1422,1015,1040]" captionStart="FIGURES 103105" captionStartId="61.[151,264,1910,1933]" captionTargetBox="[180,1444,206,1860]" captionTargetId="figure@61.[164,1449,193,1867]" captionTargetPageId="61" captionText="FIGURES 103105. Male caudal appendages in lateral (103105a), posterodorsal (103105b), and dorsal (103105b) views (not to scale). (103) P.cara; (104)P.corculum; (105)P.cupida." httpUri="https://zenodo.org/record/1069378/files/figure.png" pageId="22" pageNumber="23">Figs. 104b, c</figureCitation>
). Paraproct about twice as long as S10 length, at midlength narrowing to half its basal width and with medial surface concave (
<figureCitation id="C2A187CAFFB84E41FEC8FBB4FEC86F1F" box="[258,368,1087,1112]" captionStart="FIGURES 103105" captionStartId="61.[151,264,1910,1933]" captionTargetBox="[180,1444,206,1860]" captionTargetId="figure@61.[164,1449,193,1867]" captionTargetPageId="61" captionText="FIGURES 103105. Male caudal appendages in lateral (103105a), posterodorsal (103105b), and dorsal (103105b) views (not to scale). (103) P.cara; (104)P.corculum; (105)P.cupida." httpUri="https://zenodo.org/record/1069378/files/figure.png" pageId="22" pageNumber="23">Fig. 104a</figureCitation>
). TL 3538; Hw 1618.
</paragraph>
<paragraph id="5A259B4FFFB84E41FF0DFBEFFE056E12" blockId="22.[151,1437,151,1941]" pageId="22" pageNumber="23">
Female: As male but pale colors yellow; mesepisternum black with narrow yellow antehumeral stripe extending along basal 1/2; mesepimeron with narrow yellow humeral stripe along distal 1/2; S110 black dorsally, S37 with narrow yellow basal ring; pale lateral area of S9 extended medioposteriorly to about 1/2 of segment height (
<figureCitation id="C2A187CAFFB84E41FF27FB44FEFA6FAF" box="[237,322,1231,1256]" captionStart="FIGURES 2932" captionStartId="45.[151,264,1190,1213]" captionTargetBox="[163,1426,193,1166]" captionTargetId="figure@45.[151,1436,193,1169]" captionTargetPageId="45" captionText="FIGURES 2932. Female body scan; head in dorsal view, thorax and abdomen in lateral view (not to scale). (29) P. calverti; (30) P.capillaris; (31) P.cara; (32) P.corculum. Insert in Fig. 29: posterior lobe of pronotum in dorsal view." httpUri="https://zenodo.org/record/1069353/files/figure.png" pageId="22" pageNumber="23">Fig. 32</figureCitation>
). Middle lobe of pronotum lacking pronounced lateral depressions; posterior lobe entire, smoothly convex, directed posterodorsally (
<figureCitation id="C2A187CAFFB84E41FDEEFB7FFD336E4A" box="[548,651,1268,1293]" captionStart="FIGURES 5154" captionStartId="51.[151,264,1900,1923]" captionTargetBox="[204,1399,241,1827]" captionTargetId="figure@51.[158,1443,227,1834]" captionText="FIGURES 5154. Female pronotum (54c) and mesostigmal plates (5153; 54a, b), in lateral (51a; 52a; 53a; 54a), dorsal (51b; 52b; 53b; 54b, c), and dorsolateral (51c) views (to scale). (51) P. calverti; (52) P. capillaris; (53) P. cara; (54) P. corculum." figureDoi="http://doi.org/10.5281/zenodo.7319752" httpUri="https://zenodo.org/record/7319752/files/figure.png" pageId="22" pageNumber="23">Fig. 54c</figureCitation>
). Mesostigmal plate upright and foliate, adjacent at posteromedial corner with a transverse tubercle on mesepisternum (
<figureCitation id="C2A187CAFFB84E41FD3AFA9CFCC16E77" box="[752,889,1303,1328]" captionStart="FIGURES 5154" captionStartId="51.[151,264,1900,1923]" captionTargetBox="[204,1399,241,1827]" captionTargetId="figure@51.[158,1443,227,1834]" captionText="FIGURES 5154. Female pronotum (54c) and mesostigmal plates (5153; 54a, b), in lateral (51a; 52a; 53a; 54a), dorsal (51b; 52b; 53b; 54b, c), and dorsolateral (51c) views (to scale). (51) P. calverti; (52) P. capillaris; (53) P. cara; (54) P. corculum." figureDoi="http://doi.org/10.5281/zenodo.7319752" httpUri="https://zenodo.org/record/7319752/files/figure.png" pageId="22" pageNumber="23">Figs. 54a, b</figureCitation>
); no depressions behind mesostigmal plates. TL 31.534.5; Hw 17.519.5.
</paragraph>
</subSubSection>
<subSubSection id="1280C8C4FFB84E41FF0DFAD4FCB46D32" pageId="22" pageNumber="23" type="diagnosis">
<paragraph id="5A259B4FFFB84E41FF0DFAD4FCB46D32" blockId="22.[151,1437,151,1941]" pageId="22" pageNumber="23">
<emphasis id="68EE475DFFB84E41FF0DFAD4FEF96E3F" bold="true" box="[199,321,1375,1400]" pageId="22" pageNumber="23">Diagnosis.</emphasis>
<taxonomicName id="9D9AE0CCFFB84E41FE8DFAEAFD8E6E3F" authorityName="Calvert" authorityYear="1907" box="[327,566,1376,1400]" class="Insecta" family="Protoneuridae" genus="Protoneura" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="corculum">
<emphasis id="68EE475DFFB84E41FE8DFAEAFD8E6E3F" box="[327,566,1376,1400]" italics="true" pageId="22" pageNumber="23">Protoneura corculum</emphasis>
</taxonomicName>
and
<taxonomicName id="9D9AE0CCFFB84E41FDA6FAEAFCB26E30" authorityName="Selys" authorityYear="1886" box="[620,778,1376,1400]" class="Insecta" family="Protoneuridae" genus="Protoneura" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="paucinervis">
<emphasis id="68EE475DFFB84E41FDA6FAEAFCB26E30" box="[620,778,1376,1400]" italics="true" pageId="22" pageNumber="23">P. paucinervis</emphasis>
</taxonomicName>
can be distinguished from all congeners by male paraproct longer than twice cercus length (
<figureCitation id="C2A187CAFFB84E41FDC5FA0FFD326EDB" box="[527,650,1412,1437]" captionStart="FIGURES 103105" captionStartId="61.[151,264,1910,1933]" captionTargetBox="[180,1444,206,1860]" captionTargetId="figure@61.[164,1449,193,1867]" captionTargetPageId="61" captionText="FIGURES 103105. Male caudal appendages in lateral (103105a), posterodorsal (103105b), and dorsal (103105b) views (not to scale). (103) P.cara; (104)P.corculum; (105)P.cupida." httpUri="https://zenodo.org/record/1069378/files/figure.png" pageId="22" pageNumber="23">Figs. 104a</figureCitation>
; 109a) and female upright foliaceous mesostigmal plate with medial corner adjacent to a transverse tubercle on mesepisternum (
<figureCitation id="C2A187CAFFB84E41FCE1FA2CFC086E87" box="[811,944,1447,1472]" captionStart="FIGURES 5154" captionStartId="51.[151,264,1900,1923]" captionTargetBox="[204,1399,241,1827]" captionTargetId="figure@51.[158,1443,227,1834]" captionText="FIGURES 5154. Female pronotum (54c) and mesostigmal plates (5153; 54a, b), in lateral (51a; 52a; 53a; 54a), dorsal (51b; 52b; 53b; 54b, c), and dorsolateral (51c) views (to scale). (51) P. calverti; (52) P. capillaris; (53) P. cara; (54) P. corculum." figureDoi="http://doi.org/10.5281/zenodo.7319752" httpUri="https://zenodo.org/record/7319752/files/figure.png" pageId="22" pageNumber="23">Figs. 54a, b</figureCitation>
;
<figureCitation id="C2A187CAFFB84E41FC71FA2CFBB86E87" box="[955,1024,1447,1472]" captionStart="FIGURES 5559" captionStartId="52.[151,264,1891,1914]" captionTargetBox="[208,1395,236,1822]" captionTargetId="figure@52.[156,1441,223,1829]" captionText="FIGURES 5559. Female pronotum (58b) and mesostigmal plates (5558; 59a, c), in lateral (55a; 56a; 59a) and dorsal (55b; 56b; 57; 58; 59b, c) views (to scale). (55) P. cupida; (56) P. dunklei; (57) P. klugi; (58) P. macintyrei; (59) P. paucinervis." figureDoi="http://doi.org/10.5281/zenodo.7319754" httpUri="https://zenodo.org/record/7319754/files/figure.png" pageId="22" pageNumber="23">59a, c</figureCitation>
). Male of
<taxonomicName id="9D9AE0CCFFB84E41FBB8FA22FB4D6E87" authorityName="Calvert" authorityYear="1907" box="[1138,1269,1448,1472]" class="Insecta" family="Protoneuridae" genus="Protoneura" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="corculum">
<emphasis id="68EE475DFFB84E41FBB8FA22FB4D6E87" box="[1138,1269,1448,1472]" italics="true" pageId="22" pageNumber="23">P. corculum</emphasis>
</taxonomicName>
differs from
<taxonomicName id="9D9AE0CCFFB84E41FA4DFA22FEA26EA3" authorityName="Selys" authorityYear="1886" class="Insecta" family="Protoneuridae" genus="Protoneura" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="paucinervis">
<emphasis id="68EE475DFFB84E41FA4DFA22FEA26EA3" italics="true" pageId="22" pageNumber="23">P. paucinervis</emphasis>
</taxonomicName>
by orange color of mesepisternum (
<figureCitation id="C2A187CAFFB84E41FD7AFA47FD4D6EA3" box="[688,757,1484,1509]" captionStart="FIGURES 58" captionStartId="17.[151,264,1532,1555]" captionTargetBox="[162,1429,524,1500]" captionTargetId="figure@17.[151,1436,519,1510]" captionTargetPageId="17" captionText="FIGURES 58. Male body scan; head in dorsal view, thorax and abdomen in lateral view (not to scale). (5) P. calverti; (6) P. capillaris; (7) P.cara; (8) P.corculum." httpUri="https://zenodo.org/record/1069341/files/figure.png" pageId="22" pageNumber="23">Fig. 8</figureCitation>
; yellow in
<emphasis id="68EE475DFFB84E41FCB9FA46FBA06EA3" box="[883,1048,1485,1508]" italics="true" pageId="22" pageNumber="23">
<taxonomicName id="9D9AE0CCFFB84E41FCB9FA46FBAC6EA3" authorityName="Selys" authorityYear="1886" box="[883,1044,1485,1508]" class="Insecta" family="Protoneuridae" genus="Protoneura" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="paucinervis">P. paucinervis</taxonomicName>
,
</emphasis>
<figureCitation id="C2A187CAFFB84E41FBD5FA47FBC96EA3" box="[1055,1137,1484,1509]" captionStart="FIGURES 1316" captionStartId="30.[151,264,1592,1615]" captionTargetBox="[161,1426,597,1565]" captionTargetId="figure@30.[151,1436,591,1570]" captionTargetPageId="30" captionText="FIGURES 1316. Male body scan; head in dorsal view, thorax and abdomen in lateral view (not to scale).(13) P. paucinervis; (14) P.peramans; (15) P.rojiza; (16) P. romanae." httpUri="https://zenodo.org/record/1069345/files/figure.png" pageId="22" pageNumber="23">Fig. 13</figureCitation>
), and by blunt ventrobasal and mediobasal cercus teeth (104b, c; pointed in
<taxonomicName id="9D9AE0CCFFB84E41FD34FA7AFC1D6D40" authorityName="Selys" authorityYear="1886" box="[766,933,1520,1544]" class="Insecta" family="Protoneuridae" genus="Protoneura" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="paucinervis">
<emphasis id="68EE475DFFB84E41FD34FA7AFC1D6D40" box="[766,933,1520,1544]" italics="true" pageId="22" pageNumber="23">P. paucinervis</emphasis>
</taxonomicName>
,
<figureCitation id="C2A187CAFFB84E41FC71FA64FB8B6D4F" box="[955,1075,1519,1544]" captionStart="FIGURES 109111" captionStartId="63.[151,264,1911,1934]" captionTargetBox="[168,1432,208,1849]" captionTargetId="figure@63.[153,1438,193,1861]" captionTargetPageId="63" captionText="FIGURES 109111. Male caudal appendages in lateral (109111a), posterodorsal (109111b), and dorsal (109111b) views (not to scale). (109) P.paucinervis; (110) P.peramans; (111) P.rojiza." httpUri="https://zenodo.org/record/1069382/files/figure.png" pageId="22" pageNumber="23">Fig. 109b</figureCitation>
). Female differs by black on metepisternum against metepisternal-metepimeral suture extending along distal 1/4 of suture in
<taxonomicName id="9D9AE0CCFFB84E41FB12F99EFAE66D6B" authorityName="Calvert" authorityYear="1907" box="[1240,1374,1556,1580]" class="Insecta" family="Protoneuridae" genus="Protoneura" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="corculum">
<emphasis id="68EE475DFFB84E41FB12F99EFAE66D6B" box="[1240,1374,1556,1580]" italics="true" pageId="22" pageNumber="23">P. corculum</emphasis>
</taxonomicName>
(
<figureCitation id="C2A187CAFFB84E41FAA4F99FFF0E6D17" captionStart="FIGURES 2932" captionStartId="45.[151,264,1190,1213]" captionTargetBox="[163,1426,193,1166]" captionTargetId="figure@45.[151,1436,193,1169]" captionTargetPageId="45" captionText="FIGURES 2932. Female body scan; head in dorsal view, thorax and abdomen in lateral view (not to scale). (29) P. calverti; (30) P.capillaris; (31) P.cara; (32) P.corculum. Insert in Fig. 29: posterior lobe of pronotum in dorsal view." httpUri="https://zenodo.org/record/1069353/files/figure.png" pageId="22" pageNumber="23">Fig. 32</figureCitation>
; along distal 3/
<quantity id="9D6236AAFFB84E41FEAEF9B3FE296D17" box="[356,401,1591,1616]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="22" pageNumber="23" unit="in" value="4.0">4 in</quantity>
<taxonomicName id="9D9AE0CCFFB84E41FE53F9B2FD816D08" authorityName="Selys" authorityYear="1886" box="[409,569,1592,1616]" class="Insecta" family="Protoneuridae" genus="Protoneura" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="paucinervis">
<emphasis id="68EE475DFFB84E41FE53F9B2FD816D08" box="[409,569,1592,1616]" italics="true" pageId="22" pageNumber="23">P. paucinervis</emphasis>
</taxonomicName>
,
<figureCitation id="C2A187CAFFB84E41FD82F9BCFD236D17" box="[584,667,1591,1616]" captionStart="FIGURES 3740" captionStartId="47.[151,264,1190,1213]" captionTargetBox="[166,1416,198,1156]" captionTargetId="figure@47.[151,1436,193,1169]" captionTargetPageId="47" captionText="FIGURES 3740. Female body scan; head in dorsal view, thorax and abdomen in lateral view (not to scale). (37) P. paucinervis; (38) P.peramans; (39) P.rojiza; (40) P.romanae. Insert in Fig. 40: pronotum in dorsal view." httpUri="https://zenodo.org/record/1069357/files/figure.png" pageId="22" pageNumber="23">Fig. 37</figureCitation>
), and by sides of posterior lobe of pronotum smoothly convex in
<taxonomicName id="9D9AE0CCFFB84E41FA4DF9B2FEB86D33" authorityName="Calvert" authorityYear="1907" class="Insecta" family="Protoneuridae" genus="Protoneura" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="corculum">
<emphasis id="68EE475DFFB84E41FA4DF9B2FEB86D33" italics="true" pageId="22" pageNumber="23">P. corculum</emphasis>
</taxonomicName>
(
<figureCitation id="C2A187CAFFB84E41FEC5F9D7FED76D32" box="[271,367,1628,1653]" captionStart="FIGURES 5154" captionStartId="51.[151,264,1900,1923]" captionTargetBox="[204,1399,241,1827]" captionTargetId="figure@51.[158,1443,227,1834]" captionText="FIGURES 5154. Female pronotum (54c) and mesostigmal plates (5153; 54a, b), in lateral (51a; 52a; 53a; 54a), dorsal (51b; 52b; 53b; 54b, c), and dorsolateral (51c) views (to scale). (51) P. calverti; (52) P. capillaris; (53) P. cara; (54) P. corculum." figureDoi="http://doi.org/10.5281/zenodo.7319752" httpUri="https://zenodo.org/record/7319752/files/figure.png" pageId="22" pageNumber="23">Fig. 54c</figureCitation>
; straight in
<taxonomicName id="9D9AE0CCFFB84E41FE38F9D6FD286D33" authorityName="Selys" authorityYear="1886" box="[498,656,1629,1652]" class="Insecta" family="Protoneuridae" genus="Protoneura" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="paucinervis">
<emphasis id="68EE475DFFB84E41FE38F9D6FD286D33" box="[498,656,1629,1652]" italics="true" pageId="22" pageNumber="23">P. paucinervis</emphasis>
</taxonomicName>
,
<figureCitation id="C2A187CAFFB84E41FD54F9D7FD476D32" box="[670,767,1628,1653]" captionStart="FIGURES 5559" captionStartId="52.[151,264,1891,1914]" captionTargetBox="[208,1395,236,1822]" captionTargetId="figure@52.[156,1441,223,1829]" captionText="FIGURES 5559. Female pronotum (58b) and mesostigmal plates (5558; 59a, c), in lateral (55a; 56a; 59a) and dorsal (55b; 56b; 57; 58; 59b, c) views (to scale). (55) P. cupida; (56) P. dunklei; (57) P. klugi; (58) P. macintyrei; (59) P. paucinervis." figureDoi="http://doi.org/10.5281/zenodo.7319754" httpUri="https://zenodo.org/record/7319754/files/figure.png" pageId="22" pageNumber="23">Fig. 59b</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="1280C8C4FFB84E41FF0DF9F4FB5A6C37" pageId="22" pageNumber="23" type="discussion">
<paragraph id="5A259B4FFFB84E41FF0DF9F4FB206DFA" blockId="22.[151,1437,151,1941]" pageId="22" pageNumber="23">
<emphasis id="68EE475DFFB84E41FF0DF9F4FE836DDF" bold="true" box="[199,315,1663,1688]" pageId="22" pageNumber="23">Remarks.</emphasis>
<bibRefCitation id="3E0BE6BEFFB84E41FE8BF9F4FE5C6DDF" author="Calvert" box="[321,484,1663,1688]" pageId="22" pageNumber="23" refString="Calvert, P. P. (1907) Odonata. In: Godman, F. d. C. &amp; Salvin, O. (Eds.), Biologia Centrali Americana: Insecta Neuroptera. R. H. Porter &amp; Dulau Co., London, pp. 309 - 404." type="book chapter" year="1907">Calvert (1907)</bibRefCitation>
was unsure of the pale colors of the male when describing this species. He stated that the thorax was mostly pale blue but this was in error most likely due to postmortem effects.
</paragraph>
<paragraph id="5A259B4FFFB84E41FF0DF94CFB5A6C37" blockId="22.[151,1437,151,1941]" pageId="22" pageNumber="23">
<emphasis id="68EE475DFFB84E41FF0DF94CFE026DA7" bold="true" box="[199,442,1735,1760]" pageId="22" pageNumber="23">Habitat and biology.</emphasis>
Adults found at lagoons, cenotes (DRP, RWG), and open slow moving rivers surrounded by forest (
<bibRefCitation id="3E0BE6BEFFB84E41FEC3F967FE1E6C43" author="Measey" box="[265,422,1772,1796]" pageId="22" pageNumber="23" refString="Measey, G. J. (1994) Some Odonata from Belize, Central America. Notulae odonatologicae, 4 (3), 40 - 46." type="journal article" year="1994">Measey 1994</bibRefCitation>
), where tandem pairs hover over waters surface near edge of shore (R. W. Garrison
<emphasis id="68EE475DFFB84E41FAAEF966FF066C6F" italics="true" pageId="22" pageNumber="23">pers. obs</emphasis>
.). Pairs oviposit on floating stems in usual
<taxonomicName id="9D9AE0CCFFB84E41FD61F89AFC936C6F" authorityName="Selys in Sagra" authorityYear="1857" box="[683,811,1809,1832]" class="Insecta" family="Protoneuridae" genus="Protoneura" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="68EE475DFFB84E41FD61F89AFC936C6F" box="[683,811,1809,1832]" italics="true" pageId="22" pageNumber="23">Protoneura</emphasis>
</taxonomicName>
fashion, with female abdomen bent at approximately a right angle in two places so the posterior part of S6 can be held against S2 between the wings, stabilizing it, presumably an adaptation to the long, slender abdomen in this genus (D. R. Paulson
<emphasis id="68EE475DFFB84E41FB83F8D2FB6A6C37" box="[1097,1234,1881,1904]" italics="true" pageId="22" pageNumber="23">pers. comm.</emphasis>
).
</paragraph>
</subSubSection>
<subSubSection id="1280C8C4FFB84E41FF0DF8F7FC286CD3" box="[199,912,1916,1941]" pageId="22" pageNumber="23" type="distribution">
<paragraph id="5A259B4FFFB84E41FF0DF8F7FC286CD3" blockId="22.[151,1437,151,1941]" box="[199,912,1916,1941]" pageId="22" pageNumber="23">
<emphasis id="68EE475DFFB84E41FF0DF8F7FED86CD2" bold="true" box="[199,352,1916,1941]" pageId="22" pageNumber="23">Distribution.</emphasis>
<collectingCountry id="228DDBDFFFB84E41FEADF8F7FE076CD2" box="[359,447,1916,1941]" name="Mexico" pageId="22" pageNumber="23">
<collectingRegion id="985E55ADFFB84E41FEADF8F7FE076CD2" box="[359,447,1916,1941]" country="Mexico" name="Mexico" pageId="22" pageNumber="23">Mexico</collectingRegion>
</collectingCountry>
,
<collectingCountry id="228DDBDFFFB84E41FE01F8F7FDAD6CD2" box="[459,533,1916,1941]" name="Belize" pageId="22" pageNumber="23">
<collectingRegion id="985E55ADFFB84E41FE01F8F7FDAD6CD2" box="[459,533,1916,1941]" country="Belize" name="Belize" pageId="22" pageNumber="23">Belize</collectingRegion>
</collectingCountry>
,
<collectingCountry id="228DDBDFFFB84E41FDEAF8F7FD246CD2" box="[544,668,1916,1941]" name="Guatemala" pageId="22" pageNumber="23">
<collectingRegion id="985E55ADFFB84E41FDEAF8F7FD246CD2" box="[544,668,1916,1941]" country="Guatemala" name="Guatemala" pageId="22" pageNumber="23">Guatemala</collectingRegion>
</collectingCountry>
, and
<collectingCountry id="228DDBDFFFB84E41FD1DF8F7FCAA6CD2" box="[727,786,1916,1941]" name="Cuba" pageId="22" pageNumber="23">Cuba</collectingCountry>
(
<figureCitation id="C2A187CAFFB84E41FCEBF8F7FC3C6CD3" box="[801,900,1916,1941]" captionStart="FIGURES 121122" captionStartId="67.[151,264,1900,1923]" captionTargetBox="[169,1406,215,1857]" captionTargetId="figure@67.[151,1436,193,1880]" captionTargetPageId="67" captionText="FIGURES 121122. Distribution maps." httpUri="https://zenodo.org/record/1069394/files/figure.png" pageId="22" pageNumber="23">Fig. 121</figureCitation>
).
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