121 lines
12 KiB
XML
121 lines
12 KiB
XML
<document id="C07D8DC5D2E6D1FDE95A1F7F02F70DCE" ID-CLB-Dataset="46726" ID-DOI="10.11646/zootaxa.3753.4.4" ID-GBIF-Dataset="59df87e1-3da4-43ef-8c4c-b8a7cdd580ce" ID-ISSN="1175-5326" ID-Zenodo-Dep="249874" ID-ZooBank="394ADA8F-E7A8-4751-941F-FE8E6A3921A0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460365347958" checkinUser="plazi" docAuthor="Ribeiro, Guilherme C. & Lukashevich, Elena D." docDate="2014" docId="C431860C583FDD3AB4DAFEAAB1ABFBB1" docLanguage="en" docName="zt03753p363.pdf" docOrigin="Zootaxa 3753 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Leptotarsus buscalioniae Ribeiro & Lukashevich, 2014, sp. nov." docType="treatment" docVersion="9" lastPageNumber="357" masterDocId="3808FE745835DD30B44DFFE1B570FFD9" masterDocTitle="New Leptotarsus from the Early Cretaceous of Brazil and Spain: the oldest members of the family Tipulidae (Diptera)" masterLastPageNumber="363" masterPageNumber="347" pageNumber="357" updateTime="1698295164094" updateUser="ExternalLinkService">
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<mods:mods id="1E2F757B77130F736826234FD13E2383" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="CDC7D668A7D4979E00E965A12A6A7F83">
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<mods:title id="5BA533127F0597291234D62F29C16B1B">New Leptotarsus from the Early Cretaceous of Brazil and Spain: the oldest members of the family Tipulidae (Diptera)</mods:title>
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<mods:name id="9B150C2CD85741493701A2D63D5CAF00" type="personal">
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<mods:roleTerm id="9BB3E1DE5D06EEC7BC6747237B1A437E">Author</mods:roleTerm>
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<mods:namePart id="EF13ABC0D56C72ED04908CA123E03D81">Ribeiro, Guilherme C.</mods:namePart>
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</mods:name>
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<mods:name id="D099DA4584CB02CCB7CE4AF93486A039" type="personal">
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<mods:role id="BC1F1043985CD0AEA3C3E4FDD1772766">
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<mods:roleTerm id="BEC11D244DCE5C2DDAA4CD670849E004">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="0CB816AF89D8812F5E30B62CE22D9309">Lukashevich, Elena D.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="E451C4760B568B020DFEEAD9C731270F">text</mods:typeOfResource>
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<mods:titleInfo id="09953AF37D95C29F648EF55175CC1BBC">
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<mods:title id="B62187F6F0D7A8CAC211E28FA1D65AEA">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="0AD284DCC0708976BEB1772C202D97AB">
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<mods:date id="72F038F0053D2A7AFD53A86876DE6EAE">2014</mods:date>
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<mods:detail id="F7EA8C04B63767E5CD91A96474F1E1DE" type="volume">
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<mods:number id="C1D4466AD4AB91E6B8F3632CFE6CCBC8">3753</mods:number>
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</mods:detail>
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<mods:detail id="A6D5C822D35416DB625D465F5C6C0307" type="issue">
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<mods:number id="1F4FE114F2A114C551276B5DC87018EE">4</mods:number>
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<mods:start id="13E8412FB07992DB999147BC75B1FCAF">347</mods:start>
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<mods:classification id="589CDEDE39503CC1AB85C6BC2160116F">journal article</mods:classification>
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<mods:identifier id="1C9861C7F6384DD2B5DBE7D8A9098AAA" type="CLB-Dataset">46726</mods:identifier>
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<mods:identifier id="2B9BB64EC86FB60F15E29BFB9861271C" type="DOI">10.11646/zootaxa.3753.4.4</mods:identifier>
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<mods:identifier id="51189A91D8DCF2F94B1D1659AC505295" type="GBIF-Dataset">59df87e1-3da4-43ef-8c4c-b8a7cdd580ce</mods:identifier>
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<mods:identifier id="9257108F7F1D53E3FB848ABB33438374" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="43083AD849CD35AA2434345765DB0072" type="Zenodo-Dep">249874</mods:identifier>
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</mods:mods>
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<treatment id="C431860C583FDD3AB4DAFEAAB1ABFBB1" ID-DOI="http://doi.org/10.5281/zenodo.6143101" ID-GBIF-Taxon="119410700" ID-Zenodo-Dep="6143101" LSID="urn:lsid:plazi:treatment:C431860C583FDD3AB4DAFEAAB1ABFBB1" httpUri="http://treatment.plazi.org/id/C431860C583FDD3AB4DAFEAAB1ABFBB1" lastPageNumber="357" pageId="10" pageNumber="357">
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<subSubSection id="04826491583FDD3AB4DAFEAAB42EFE5E" pageId="10" pageNumber="357" type="nomenclature">
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<paragraph id="4C27371A583FDD3AB4DAFEAAB759FEBF" blockId="10.[151,553,331,391]" box="[151,553,331,358]" pageId="10" pageNumber="357">
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<heading id="176F8076583FDD3AB4DAFEAAB759FEBF" bold="true" box="[151,553,331,358]" fontSize="11" level="1" pageId="10" pageNumber="357" reason="1">
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<emphasis id="7EECEB08583FDD3AB4DAFEAAB759FEBF" bold="true" box="[151,553,331,358]" pageId="10" pageNumber="357">
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<taxonomicName id="8B984C99583FDD3AB4DAFEAAB4B4FEBC" box="[151,452,331,357]" class="Insecta" family="Tipulidae" genus="Leptotarsus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="357" phylum="Arthropoda" rank="species" species="buscalioniae" status="sp. nov.">
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<emphasis id="7EECEB08583FDD3AB4DAFEAAB4B4FEBC" bold="true" box="[151,452,331,357]" italics="true" pageId="10" pageNumber="357">Leptotarsus buscalioniae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="65DF5673583FDD3AB581FEADB759FEBF" box="[460,553,332,358]" pageId="10" pageNumber="357" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="4C27371A583FDD3AB4DAFE8FB42EFE5E" blockId="10.[151,553,331,391]" box="[151,350,366,391]" pageId="10" pageNumber="357">
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(
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<figureCitation id="D4A32B9F583FDD3AB4D2FE8FB473FE5E" box="[159,259,366,391]" captionStart="FIGURES 5 – 9" captionStartId="4.[151,264,1935,1957]" captionTargetBox="[290,1291,196,1902]" captionTargetId="figure@4.[286,1297,193,1905]" captionTargetPageId="4" captionText="FIGURES 5 – 9. Wing venation of Leptotarsus spp. from Las Hoyas, Early Cretaceous, Spain. 5 — L. buscalioniae sp. nov., holotype LH 35088, right wing. 6 — L. buscalioniae sp. nov., holotype LH 35088, left wing. 7 — L. ibericus sp. nov., holotype LH 28206. 8 — L. ibericus sp. nov., paratype LH 29995. 9 — L. contractus sp. nov., holotype LH 15419. Scale bar 1 mm." httpUri="https://zenodo.org/record/249876/files/figure.png" pageId="10" pageNumber="357">Figs 5, 6</figureCitation>
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,
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<figureCitation id="D4A32B9F583FDD3AB542FE8EB427FE5E" box="[271,343,367,391]" captionStart="FIGURES 19 – 21" captionStartId="11.[151,264,1962,1984]" captionTargetBox="[196,1391,195,1940]" captionTargetId="figure@11.[196,1392,195,1940]" captionTargetPageId="11" captionText="FIGURES 19 – 21. Leptotarsus buscalioniae sp. nov., from Las Hoyas, Early Cretaceous, Spain, holotype LH 35088. 19 — Total view (part). 20 — Forebody (counterpart). 21 — Distal portion of wing (part)." httpUri="https://zenodo.org/record/249882/files/figure.png" pageId="10" pageNumber="357">19–21</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="04826491583FDD3AB4DAFE57B122FDCE" pageId="10" pageNumber="357" type="materials_examined">
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<paragraph id="4C27371A583FDD3AB4DAFE57B122FDCE" blockId="10.[151,1437,438,1128]" pageId="10" pageNumber="357">
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<emphasis id="7EECEB08583FDD3AB4DAFE57B436FE16" bold="true" box="[151,326,438,463]" pageId="10" pageNumber="357">
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<typeStatus id="932389B8583FDD3AB4DAFE57B5A2FE16" box="[151,210,438,463]" pageId="10" pageNumber="357">Type</typeStatus>
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material.
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</emphasis>
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<typeStatus id="932389B8583FDD3AB500FE57B4C3FE16" box="[333,435,438,463]" pageId="10" pageNumber="357" type="holotype">
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<emphasis id="7EECEB08583FDD3AB500FE57B4C3FE16" box="[333,435,438,463]" italics="true" pageId="10" pageNumber="357">Holotype</emphasis>
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</typeStatus>
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: LH 35088± (part and counterpart of entire male specimen; preserved structures include the long antenna and terminalia). East-central
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<collectingCountry id="348F778A583FDD3AB688FE3BB678FE2A" box="[709,776,474,499]" name="Spain" pageId="10" pageNumber="357">Spain</collectingCountry>
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, Las Hoyas; La Huérguina Limestone Formation, Late Barremian, Early Cretaceous. Housed in Museo de Ciencias de Castillia–La Mancha.
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</paragraph>
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</subSubSection>
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<subSubSection id="04826491583FDD3AB48AFDC3B491FD86" pageId="10" pageNumber="357" type="etymology">
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<paragraph id="4C27371A583FDD3AB48AFDC3B491FD86" blockId="10.[151,1437,438,1128]" pageId="10" pageNumber="357">
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<emphasis id="7EECEB08583FDD3AB48AFDC3B43EFDE2" bold="true" box="[199,334,546,571]" pageId="10" pageNumber="357">Etymology.</emphasis>
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The species is named after Dr. Angela Buscalioni, the main organizer and motor of the excavation in Las Hoyas for many years.
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</paragraph>
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</subSubSection>
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<subSubSection id="04826491583FDD3AB48AFD8BB04DFD17" pageId="10" pageNumber="357" type="diagnosis">
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<paragraph id="4C27371A583FDD3AB48AFD8BB04DFD17" blockId="10.[151,1437,438,1128]" pageId="10" pageNumber="357">
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<emphasis id="7EECEB08583FDD3AB48AFD8BB431FD5A" bold="true" box="[199,321,618,643]" pageId="10" pageNumber="357">Diagnosis.</emphasis>
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This species differs from all known Cretaceous species by relatively long abdomen. It differs from other species of
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<taxonomicName id="8B984C99583FDD3AB514FD6EB4ADFD7F" box="[345,477,655,678]" class="Insecta" family="Tipulidae" genus="Leptotarsus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="357" phylum="Arthropoda" rank="genus">
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<emphasis id="7EECEB08583FDD3AB514FD6EB4ADFD7F" box="[345,477,655,678]" italics="true" pageId="10" pageNumber="357">Leptotarsus</emphasis>
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</taxonomicName>
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from the Early Cretaceous of
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<collectingCountry id="348F778A583FDD3AB705FD6FB6F8FD7E" box="[840,904,654,679]" name="Spain" pageId="10" pageNumber="357">Spain</collectingCountry>
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by long distal section of vein M1+2 and distal sections of veins M3 and M4 strongly divergent, and from Brazilian species by absence of dark costal field.
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</paragraph>
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</subSubSection>
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<subSubSection id="04826491583FDD3AB48AFD3DB1ABFBB1" pageId="10" pageNumber="357" type="description">
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<paragraph id="4C27371A583FDD3AB48AFD3DB060FC0C" blockId="10.[151,1437,438,1128]" pageId="10" pageNumber="357">
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<emphasis id="7EECEB08583FDD3AB48AFD3DB429FD2C" bold="true" box="[199,345,732,757]" pageId="10" pageNumber="357">Description.</emphasis>
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<emphasis id="7EECEB08583FDD3AB528FD3DB4D9FD2C" box="[357,425,732,757]" italics="true" pageId="10" pageNumber="357">Head.</emphasis>
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Antennae long and slender, bearing long and slender flagellomeres (total number of flagellomeres unknown); rostrum ca. 0.5x the total length of head.
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<emphasis id="7EECEB08583FDD3AB7CAFCE1B6A8FCC0" box="[903,984,768,793]" italics="true" pageId="10" pageNumber="357">Thorax</emphasis>
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stout, as long as wide.
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<emphasis id="7EECEB08583FDD3AB092FCE0B067FCC1" box="[1247,1303,769,792]" italics="true" pageId="10" pageNumber="357">Wing</emphasis>
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transparent except for dark pterostigma; Sc reaching wing margin at the level of first bifurcation of Rs and beyond level of the tip of Cu; sc-r present, positioned near the tip of Sc; Rs 1.2–1.4 times longer than R2+3 stem; R1 reaching wing margin distal to the level of R2+3 bifurcation; r-r oblique, linking R1 to R2; R2 oblique; medial vein four-branched; discal cell as long as wide; M3 and M4 strongly divergent; M3 sinuous; M4 arcuated and shorter; long m-cu linking Cu to the base of M4.
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<emphasis id="7EECEB08583FDD3AB5C7FC5DB484FC0C" box="[394,500,956,981]" italics="true" pageId="10" pageNumber="357">Abdomen</emphasis>
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stout, ca. 3.0x longer than thorax; gonocoxite conical, as long as wide.
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</paragraph>
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<paragraph id="4C27371A583FDD3AB48AFC02B017FC25" blockId="10.[151,1437,438,1128]" box="[199,1383,995,1020]" pageId="10" pageNumber="357">
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<emphasis id="7EECEB08583FDD3AB48AFC02B406FC25" bold="true" box="[199,374,995,1020]" pageId="10" pageNumber="357">Measurements</emphasis>
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. Antenna length, at least 10.2. Body length, ca. 9.5. Wing length/width, 9.4–10.0/2.2–2.7.
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</paragraph>
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<paragraph id="4C27371A583FDD3AB48AFBE6B1ABFBB1" blockId="10.[151,1437,438,1128]" pageId="10" pageNumber="357">
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<emphasis id="7EECEB08583FDD3AB48AFBE6B476FBF9" bold="true" box="[199,262,1031,1056]" pageId="10" pageNumber="357">Note.</emphasis>
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In the
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<typeStatus id="932389B8583FDD3AB516FBE6B4B0FBF9" box="[347,448,1031,1056]" pageId="10" pageNumber="357" type="holotype">holotype</typeStatus>
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, one wing is longer and narrower than the other (proportions of length to width about 1:4.5 and 1:3.5 respectively). This shows clearly that the original proportions of the wings have been distorted. Therefore, one must be careful when comparing proportions of wings in Las Hoyas fossil insects.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |