treatments-xml/data/C4/31/86/C431860C5837DD32B4DAFE69B5BFFB3E.xml
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<document id="18115461CB2E1738DFE37ED02E03919B" ID-CLB-Dataset="46726" ID-DOI="10.11646/zootaxa.3753.4.4" ID-GBIF-Dataset="59df87e1-3da4-43ef-8c4c-b8a7cdd580ce" ID-ISSN="1175-5326" ID-Zenodo-Dep="249874" ID-ZooBank="394ADA8F-E7A8-4751-941F-FE8E6A3921A0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460365347958" checkinUser="plazi" docAuthor="Ribeiro, Guilherme C. &amp; Lukashevich, Elena D." docDate="2014" docId="C431860C5837DD32B4DAFE69B5BFFB3E" docLanguage="en" docName="zt03753p363.pdf" docOrigin="Zootaxa 3753 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Leptotarsus grimaldii Ribeiro &amp; Lukashevich, 2014, sp. nov." docType="treatment" docVersion="9" lastPageNumber="349" masterDocId="3808FE745835DD30B44DFFE1B570FFD9" masterDocTitle="New Leptotarsus from the Early Cretaceous of Brazil and Spain: the oldest members of the family Tipulidae (Diptera)" masterLastPageNumber="363" masterPageNumber="347" pageNumber="349" updateTime="1698295164094" updateUser="ExternalLinkService">
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<mods:title id="C223A0300567A66E3D399A8D0D258991">New Leptotarsus from the Early Cretaceous of Brazil and Spain: the oldest members of the family Tipulidae (Diptera)</mods:title>
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<mods:namePart id="5CD2A0A6D2F54A486B277C8A70D5D71B">Ribeiro, Guilherme C.</mods:namePart>
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<mods:namePart id="FF3F47D794654E407E53A17B700296B8">Lukashevich, Elena D.</mods:namePart>
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<mods:date id="FAFA6EF2502E4A008D20F980BC63A9D9">2014</mods:date>
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<treatment id="C431860C5837DD32B4DAFE69B5BFFB3E" ID-DOI="http://doi.org/10.5281/zenodo.6143093" ID-GBIF-Taxon="119410704" ID-Zenodo-Dep="6143093" LSID="urn:lsid:plazi:treatment:C431860C5837DD32B4DAFE69B5BFFB3E" httpUri="http://treatment.plazi.org/id/C431860C5837DD32B4DAFE69B5BFFB3E" lastPageNumber="349" pageId="2" pageNumber="349">
<subSubSection id="048264915837DD32B4DAFE69B42EFE1D" pageId="2" pageNumber="349" type="nomenclature">
<paragraph id="4C27371A5837DD32B4DAFE69B49FFE79" blockId="2.[151,495,391,452]" box="[151,495,391,416]" pageId="2" pageNumber="349">
<heading id="176F80765837DD32B4DAFE69B49FFE79" bold="true" box="[151,495,391,416]" fontSize="10" level="2" pageId="2" pageNumber="349" reason="2">
<emphasis id="7EECEB085837DD32B4DAFE69B49FFE79" bold="true" box="[151,495,391,416]" pageId="2" pageNumber="349">
<taxonomicName id="8B984C995837DD32B4DAFE69B4FFFE79" box="[151,399,391,416]" class="Insecta" family="Tipulidae" genus="Leptotarsus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="349" phylum="Arthropoda" rank="species" species="grimaldii" status="sp. nov.">
<emphasis id="7EECEB085837DD32B4DAFE69B4FFFE79" bold="true" box="[151,399,391,416]" italics="true" pageId="2" pageNumber="349">Leptotarsus grimaldii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="65DF56735837DD32B5D8FE66B49FFE79" box="[405,495,391,416]" pageId="2" pageNumber="349" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
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<paragraph id="4C27371A5837DD32B4DAFE4AB42EFE1D" blockId="2.[151,495,391,452]" box="[151,350,427,452]" pageId="2" pageNumber="349">
(
<figureCitation id="D4A32B9F5837DD32B4D2FE4AB473FE1D" box="[159,259,427,452]" captionStart="FIGURES 1 4" captionStartId="3.[151,264,1955,1977]" captionTargetBox="[156,1422,198,1926]" captionTargetId="figure@3.[153,1433,195,1933]" captionTargetPageId="3" captionText="FIGURES 1 4. Wing venation of Leptotarsus spp. from Santana Formation, Early Cretaceous, Brazil. 1 — L. grimaldii sp. nov., holotype AMNH SA 46358, right wing. 2 — L. grimaldii sp. nov., paratype AMNH SA 46366, left wing. 3 — L. cretaceus sp. nov., holotype AMNH SA 46354, right wing. 4 — L. martinsnetoi sp. nov., holotype MZUSP 1580, left wing." httpUri="https://zenodo.org/record/249875/files/figure.png" pageId="2" pageNumber="349">Figs 1, 2</figureCitation>
,
<figureCitation id="D4A32B9F5837DD32B542FE4DB427FE1D" box="[271,343,428,452]" captionStart-0="FIGURES 10 11" captionStart-1="FIGURES 12 13" captionStartId-0="5.[151,264,1899,1921]" captionStartId-1="6.[151,264,1903,1925]" captionTargetBox-0="[153,1433,193,1877]" captionTargetBox-1="[153,1433,193,1880]" captionTargetId-0="figure@5.[153,1433,193,1877]" captionTargetId-1="figure@6.[153,1433,193,1881]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="FIGURES 10 11. Leptotarsus grimaldii, sp. nov., from Santana Formation, Early Cretaceous, Brazil, holotype AMNH SA 46358. 10 — Total view. 11 — Detail of abdomen and genitalia." captionText-1="FIGURES 12 13. Leptotarsus grimaldii, sp. nov., from Santana Formation, Early Cretaceous, Brazil, paratype AMNH SA 46366. 12 — Total view. 13 — Detail of head. Abbreviations: comp eye: compound eye; lbl: labella; na: nasus; ped: pedicel; plp: palpus; scp: scape." httpUri-0="https://zenodo.org/record/249877/files/figure.png" httpUri-1="https://zenodo.org/record/249878/files/figure.png" pageId="2" pageNumber="349">1013</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="048264915837DD32B4DAFE12B67FFD45" pageId="2" pageNumber="349" type="materials_examined">
<paragraph id="4C27371A5837DD32B4DAFE12B67FFD45" blockId="2.[151,1436,499,1255]" pageId="2" pageNumber="349">
<emphasis id="7EECEB085837DD32B4DAFE12B435FDD5" bold="true" box="[151,325,499,524]" pageId="2" pageNumber="349">
<typeStatus id="932389B85837DD32B4DAFE12B5A2FDD5" box="[151,210,499,524]" pageId="2" pageNumber="349">Type</typeStatus>
material.
</emphasis>
<typeStatus id="932389B85837DD32B506FE12B4C1FDD5" box="[331,433,499,524]" pageId="2" pageNumber="349" type="holotype">
<emphasis id="7EECEB085837DD32B506FE12B4C1FDD5" box="[331,433,499,524]" italics="true" pageId="2" pageNumber="349">Holotype</emphasis>
</typeStatus>
:
<collectionCode id="2A89AFDF5837DD32B58CFE15B767FDD5" LSID="urn:lsid:biocol.org:col:34925" box="[449,535,500,524]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="2" pageNumber="349">AMNH</collectionCode>
SA 46358 (entire male specimen, preserved in dorsal view; preserved structures include the long antennae, left front leg and terminalia).
<emphasis id="7EECEB085837DD32B77AFDF9B6D7FDF6" box="[823,935,536,559]" italics="true" pageId="2" pageNumber="349">
<typeStatus id="932389B85837DD32B77AFDF9B6D1FDF6" box="[823,929,536,559]" pageId="2" pageNumber="349" type="paratype">Paratype</typeStatus>
:
</emphasis>
<collectionCode id="2A89AFDF5837DD32B7FEFDF9B178FDE9" LSID="urn:lsid:biocol.org:col:34925" box="[947,1032,536,560]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="2" pageNumber="349">AMNH</collectionCode>
SA 46366 (entire male specimen, preserved in lateral view; preserved structures include head with mouthparts, the long antennae, and middle legs). NE
<collectingCountry id="348F778A5837DD32B488FDBEB47CFDA1" box="[197,268,607,632]" name="Brazil" pageId="2" pageNumber="349">Brazil</collectingCountry>
, Santana Fm., Aptian/Albian, Early Cretaceous. Both housed in the Division of Invertebrate Zoology, American Museum of Natural History, New York,
<collectingCountry id="348F778A5837DD32B682FD65B67BFD45" box="[719,779,644,668]" name="United States of America" pageId="2" pageNumber="349">USA</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="048264915837DD32B48AFD46B68FFD3D" pageId="2" pageNumber="349" type="etymology">
<paragraph id="4C27371A5837DD32B48AFD46B68FFD3D" blockId="2.[151,1436,499,1255]" pageId="2" pageNumber="349">
<emphasis id="7EECEB085837DD32B48AFD46B43EFD19" bold="true" box="[199,334,679,704]" pageId="2" pageNumber="349">Etymology.</emphasis>
The species is named after Dr. David Grimaldi, curator at the American Museum of Natural History in New York, for his great contribution to the paleontology of insects.
</paragraph>
</subSubSection>
<subSubSection id="048264915837DD32B48AFD0EB6AAFCF5" pageId="2" pageNumber="349" type="diagnosis">
<paragraph id="4C27371A5837DD32B48AFD0EB6AAFCF5" blockId="2.[151,1436,499,1255]" pageId="2" pageNumber="349">
<emphasis id="7EECEB085837DD32B48AFD0EB431FCD1" bold="true" box="[199,321,751,776]" pageId="2" pageNumber="349">Diagnosis.</emphasis>
This species differs from the other Cretaceous species mostly by having the distal sections of veins M3 and M4 very similar in length and shape, running parallel to each other.
</paragraph>
</subSubSection>
<subSubSection id="048264915837DD32B48AFCDBB5BFFB3E" pageId="2" pageNumber="349" type="description">
<paragraph id="4C27371A5837DD32B48AFCDBB788FB46" blockId="2.[151,1436,499,1255]" pageId="2" pageNumber="349">
<emphasis id="7EECEB085837DD32B48AFCDBB429FC8A" bold="true" box="[199,345,826,851]" pageId="2" pageNumber="349">Description.</emphasis>
Male.
<emphasis id="7EECEB085837DD32B5E5FCDBB49CFC8A" box="[424,492,826,851]" italics="true" pageId="2" pageNumber="349">Head.</emphasis>
Head capsule with an anterodorsal protuberance; antenna long and slender, bearing long and slender flagellomeres (total number of flagellomeres unknown); scape stout and long, ca. 3.5x longer than pedicel; nasus distinct; rostrum ca. 0.5x the total length of head.
<emphasis id="7EECEB085837DD32B72BFC63B6C7FC42" box="[870,951,898,923]" italics="true" pageId="2" pageNumber="349">Thorax</emphasis>
stout, ca. 1.8x longer than wide and almost as long as high.
<emphasis id="7EECEB085837DD32B506FC46B4F3FC67" box="[331,387,935,958]" italics="true" pageId="2" pageNumber="349">Wing</emphasis>
transparent except for dark pterostigma and costal field. Sc reaching wing margin at the level of first bifurcation of Rs and beyond the level of the tip of Cu; sc-r present, positioned near the tip of Sc; Rs curved, subequal in length with R2+3; R1 reaching wing margin at the level of bifurcation of R2+3; r-r oblique, linking R1 to R2; R2 oblique; medial vein four-branched; discal cell small, ca. 1.23 to 1.5x longer than wide; veins M3 and M4 running parallel to each other, very similar in length and shape; m-cu linking Cu to the base of M4.
<emphasis id="7EECEB085837DD32B0A0FBDAB027FB8D" box="[1261,1367,1083,1108]" italics="true" pageId="2" pageNumber="349">Abdomen</emphasis>
stout, ca. 1.7x longer than thorax; gonocoxite conical, ca. 2.4x longer than wide; gonostylus divided with clasper of gonostylus slightly longer than the lobe of gonostylus.
</paragraph>
<paragraph id="4C27371A5837DD32B48AFB4BB5BFFB3E" blockId="2.[151,1436,499,1255]" pageId="2" pageNumber="349">
<emphasis id="7EECEB085837DD32B48AFB4BB406FB1A" bold="true" box="[199,374,1194,1219]" pageId="2" pageNumber="349">Measurements</emphasis>
. Male. Antenna length, at least 16.49. Body length, ca. 7.2. Wing length/width ca. 6.9/ 1.78 1.89.
</paragraph>
</subSubSection>
</treatment>
</document>