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<document id="D6929A1285799DC33D4DBC225F33D7E4" ID-CLB-Dataset="295840" ID-DOI="10.5852/ejt.2024.932.2523" ID-GBIF-Dataset="cf4f559a-e7c3-4c31-96a7-b02df9e72f2a" ID-ISSN="2118-9773" ID-Zenodo-Dep="11145103" ID-ZooBank="8049A322-2716-4E9D-A8B7-BF3B5DE82ADC" IM.bibliography_approvedBy="jonas" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="jonas" IM.metadata_approvedBy="jonas" IM.taxonomicNames_approvedBy="jonas" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="jonas" checkinTime="1715161415022" checkinUser="plazi" docAuthor="Villarreal-Blanco, Eduardo, Martínez, Leonel &amp; Eyes-Escalante, Melisa" docDate="2024" docId="C071D04C765DFFF6FDD8FD333A06FE4F" docLanguage="en" docName="EJT.2024.932.204-224.pdf" docOrigin="European Journal of Taxonomy 932" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2523/11319" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Nyetnops madre Villarreal &amp; Martinez 2024, sp. nov." docType="treatment" docVersion="4" lastPageNumber="218" masterDocId="3C48A8347657FFF8FFDBFF983B5CFFAB" masterDocTitle="Update in the Peruvian Caponiidae: New records and new species of Nyetnops Platnick &amp; Lise, 2007 (Araneae: Caponiidae)" masterLastPageNumber="224" masterPageNumber="204" pageNumber="214" updateTime="1715195292513" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="560AB34D2D735283792F2CA43C1CB4D3">Update in the Peruvian Caponiidae: New records and new species of Nyetnops Platnick &amp; Lise, 2007 (Araneae: Caponiidae)</mods:title>
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<mods:namePart id="914CA5CEEFB887130DE5E6F194431A62">Villarreal-Blanco, Eduardo</mods:namePart>
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<mods:affiliation id="1209DA41F1AA0A7E0E035AAA775D0348">Grupo de Investigación Biodiversidad del Caribe Colombiano, Semillero de Investigación Sistemática de Artrópodos Neotropicales, Departamento de Biología, Universidad del Atlántico, Barranquilla, Colombia.</mods:affiliation>
<mods:nameIdentifier id="5FAB040D07E0D47672610EDD1F015940" type="email">ervillarreal95@gmail.com</mods:nameIdentifier>
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<mods:name id="B219F05477E7311C276257D7B06DC6E4" type="personal">
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<mods:namePart id="B79DE067D4BFBA33D56868222611302B">Martínez, Leonel</mods:namePart>
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<mods:affiliation id="A3BE2D2EFEC38215DD83BD077C5702E8">División Aracnología, Museo Argentino de Ciencias Naturales “ Bernardino Rivadavia ” CONICET, Avenida Ángel Gallardo 470, CP: 1405 DJR, C. A. B. A., Buenos Aires, Argentina.</mods:affiliation>
<mods:nameIdentifier id="1B9E15BC72A34BE7D3EDCCCAABE9DE67" type="email">leonelmarbio@gmail.com</mods:nameIdentifier>
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<mods:namePart id="D38A4317C8D0D4D9AF977E47A8B02012">Eyes-Escalante, Melisa</mods:namePart>
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<mods:affiliation id="378E21EABA2DCD76670736A0EBF51E47">Grupo de Investigación Biodiversidad del Caribe Colombiano, Semillero de Investigación Sistemática de Artrópodos Neotropicales, Departamento de Biología, Universidad del Atlántico, Barranquilla, Colombia.</mods:affiliation>
<mods:nameIdentifier id="92629DFA7BD433AAC15B23E53E8DDFC7" type="email">meyese@mail.uniatlantico.edu.co</mods:nameIdentifier>
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<mods:title id="8C6F646F7F736017182EAA8BAB57B821">European Journal of Taxonomy</mods:title>
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<mods:date id="8B1437BF2D0B98D5D5561796F0AC910A">2024</mods:date>
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<mods:number id="1F0D9643A40D8CCB505C00314E40FE76">2024-05-06</mods:number>
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<treatment id="C071D04C765DFFF6FDD8FD333A06FE4F" ID-DOI="http://doi.org/10.5281/zenodo.11145153" ID-Zenodo-Dep="11145153" LSID="urn:lsid:plazi:treatment:C071D04C765DFFF6FDD8FD333A06FE4F" httpUri="http://treatment.plazi.org/id/C071D04C765DFFF6FDD8FD333A06FE4F" lastPageId="14" lastPageNumber="218" pageId="10" pageNumber="214">
<subSubSection id="00C232D1765DFFF2FDD8FD333F6DFD6E" box="[515,1073,682,709]" pageId="10" pageNumber="214" type="nomenclature">
<paragraph id="4867615A765DFFF2FDD8FD333F6DFD6E" blockId="10.[357,1231,682,779]" box="[515,1073,682,709]" pageId="10" pageNumber="214">
<heading id="132FD636765DFFF2FDD8FD333F6DFD6E" box="[515,1073,682,709]" centered="true" fontSize="11" level="2" pageId="10" pageNumber="214" reason="2">
<taxonomicName id="8FD81AD9765DFFF2FDD8FD333893FD6E" authority="Villarreal &amp; Martinez" authorityName="Villarreal &amp; Martinez" authorityYear="2024" box="[515,975,682,709]" class="Arachnida" family="Caponiidae" genus="Nyetnops" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="214" phylum="Arthropoda" rank="species" species="madre" status="sp. nov.">
<emphasis id="7AACBD48765DFFF2FDD8FD333995FD6E" bold="true" box="[515,713,682,709]" italics="true" pageId="10" pageNumber="214">Nyetnops madre</emphasis>
Villarreal &amp; Martínez
</taxonomicName>
<taxonomicNameLabel id="619F0033765DFFF2FC0DFD333F6DFD6E" box="[982,1073,683,709]" pageId="10" pageNumber="214" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
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<subSubSection id="00C232D1765DFFF2FEBEFD563824FCA0" pageId="10" pageNumber="214" type="description">
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urn:lsid:zoobank.org:act:
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</uri>
</paragraph>
<paragraph id="4867615A765DFFF2FD60FD693824FCA0" blockId="10.[357,1231,682,779]" box="[699,888,753,779]" pageId="10" pageNumber="214">
<figureCitation id="D0E37DDF765DFFF2FD60FD693878FCA0" box="[699,804,753,779]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="11.[190,233,1928,1954]" captionStartId-1="12.[189,232,1928,1954]" captionStartId-2="13.[189,232,1892,1918]" captionTargetBox-0="[268,1321,704,1889]" captionTargetBox-1="[271,1315,819,1889]" captionTargetBox-2="[249,1338,265,1855]" captionTargetId-0="figure-142@11.[267,1321,704,1889]" captionTargetId-1="figure-136@12.[271,1317,816,1889]" captionTargetId-2="figure-18@13.[249,1338,265,1855]" captionTargetPageId-0="11" captionTargetPageId-1="12" captionTargetPageId-2="13" captionText-0="Fig. 6. Nyetnops madre Villarreal &amp; Martínez sp. nov. A–B, E, G. ♂, holotype (MUSM–ENT 0503213). C–D, F, H. ♀, paratype (MUSM–ENT 0503206). A–B. Habitus, dorsal view. C–D. Habitus, ventral view. E–F. Prosoma, dorsal view. G–H. Prosoma, ventral view. Scale bars =1 mm." captionText-1="Fig. 7. Nyetnops madre Villarreal &amp; Martínez sp. nov. A–B. ♂, holotype (MUSM–ENT 0503213). C–D. ♀, paratype (MUSM–ENT 0503206). A. Left palp, prolateral view. B. Left palp, retrolateral view. C. External genitalia, ventral view. D. Internal genitalia, dorsal view. Scale bars=0.5 mm." captionText-2="Fig. 8. Nyetnops madre Villarreal &amp; Martínez sp. nov., ♂, paratype (MUSM–ENT 0515830). A, C. Left palp, dorsal view. B, F. Tip of the embolus, prolateral view. D. Tip of the embolus, proventral view E. Copulatory bulb, prolateral view. G. Copulatory bulb, retrolateral view. H. Tip of the embolus, dorsal view. Scale bars: A, C, E, G =0.1, mm; B, D, F, H =0.01 mm." figureDoi-0="http://doi.org/10.5281/zenodo.11145121" figureDoi-1="http://doi.org/10.5281/zenodo.11145123" figureDoi-2="http://doi.org/10.5281/zenodo.11145125" httpUri-0="https://zenodo.org/record/11145121/files/figure.png" httpUri-1="https://zenodo.org/record/11145123/files/figure.png" httpUri-2="https://zenodo.org/record/11145125/files/figure.png" pageId="10" pageNumber="214">Figs 6–8</figureCitation>
,
<figureCitation id="D0E37DDF765DFFF2FCEBFD693824FCA0" box="[816,888,753,779]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="16.[189,232,1998,2024]" captionStartId-1="17.[189,232,1998,2024]" captionTargetBox-0="[189,1392,909,1953]" captionTargetBox-1="[189,1392,1113,1959]" captionTargetId-0="figure-119@16.[189,1398,888,1959]" captionTargetId-1="figure-249@17.[189,1398,1092,1959]" captionTargetPageId-0="16" captionTargetPageId-1="17" captionText-0="Fig. 11. Distribution map of species of Caponiidae Simon, 1890 recorded from Peru." captionText-1="Fig. 12. Distribution map of the genus Nyetnops Platnick &amp; Lise, 2007." figureDoi-0="http://doi.org/10.5281/zenodo.11145133" figureDoi-1="http://doi.org/10.5281/zenodo.11145135" httpUri-0="https://zenodo.org/record/11145133/files/figure.png" httpUri-1="https://zenodo.org/record/11145135/files/figure.png" pageId="10" pageNumber="214">11–12</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="00C232D1765DFFF2FF66FCAA39C2FB3E" pageId="10" pageNumber="214" type="diagnosis">
<paragraph id="4867615A765DFFF2FF66FCAA3A6BFCE7" blockId="10.[189,311,818,844]" box="[189,311,818,844]" pageId="10" pageNumber="214">
<heading id="132FD636765DFFF2FF66FCAA3A6BFCE7" bold="true" box="[189,311,818,844]" fontSize="11" level="3" pageId="10" pageNumber="214" reason="3">
<emphasis id="7AACBD48765DFFF2FF66FCAA3A6BFCE7" bold="true" box="[189,311,818,844]" pageId="10" pageNumber="214">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="4867615A765DFFF2FF66FCF939C2FB3E" blockId="10.[188,1399,865,1173]" pageId="10" pageNumber="214">
Males of
<taxonomicName id="8FD81AD9765DFFF2FEEBFCFA39A6FCD0" authority="Villarreal &amp; Martinez" authorityName="Villarreal &amp; Martinez" authorityYear="2024" box="[304,762,865,891]" class="Arachnida" family="Caponiidae" genus="Nyetnops" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="214" phylum="Arthropoda" rank="species" species="madre" status="sp. nov.">
<emphasis id="7AACBD48765DFFF2FEEBFCFA3AADFCD0" box="[304,497,865,891]" italics="true" pageId="10" pageNumber="214">Nyetnops madre</emphasis>
Villarreal &amp; Martínez
</taxonomicName>
<taxonomicNameLabel id="619F0033765DFFF2FCD8FCFA3803FCD7" box="[771,863,866,892]" pageId="10" pageNumber="214" rank="species">sp. nov.</taxonomicNameLabel>
resemble those of
<taxonomicName id="8FD81AD9765DFFF2FB98FCFA3F93FCD0" authorityName="Sanchez-Ruiz, Brescovit &amp; Bonaldo" authorityYear="2020" box="[1091,1231,865,891]" class="Arachnida" family="Caponiidae" genus="Nyetnops" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="214" phylum="Arthropoda" rank="species" species="lachonta">
<emphasis id="7AACBD48765DFFF2FB98FCFA3F93FCD0" box="[1091,1231,865,891]" italics="true" pageId="10" pageNumber="214">N. lachonta</emphasis>
</taxonomicName>
and
<taxonomicName id="8FD81AD9765DFFF2FAD1FCFA3E2AFCD0" authorityName="Sanchez-Ruiz, Brescovit &amp; Bonaldo" authorityYear="2020" box="[1290,1398,865,891]" class="Arachnida" family="Caponiidae" genus="Nyetnops" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="214" phylum="Arthropoda" rank="species" species="buruti">
<emphasis id="7AACBD48765DFFF2FAD1FCFA3E2AFCD0" box="[1290,1398,865,891]" italics="true" pageId="10" pageNumber="214">N. buruti</emphasis>
</taxonomicName>
by having a copulatory bulb (tegulum+embolus), more than 2.2 × as long as palpal tibia but differ from those of
<taxonomicName id="8FD81AD9765DFFF2FEBEFC303A8CFC6A" authorityName="Sanchez-Ruiz, Brescovit &amp; Bonaldo" authorityYear="2020" box="[357,464,935,961]" class="Arachnida" family="Caponiidae" genus="Nyetnops" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="214" phylum="Arthropoda" rank="species" species="buruti">
<emphasis id="7AACBD48765DFFF2FEBEFC303A8CFC6A" box="[357,464,935,961]" italics="true" pageId="10" pageNumber="214">N. buruti</emphasis>
</taxonomicName>
and
<taxonomicName id="8FD81AD9765DFFF2FDD1FC3039CAFC6A" authorityName="Sanchez-Ruiz, Brescovit &amp; Bonaldo" authorityYear="2020" box="[522,662,935,961]" class="Arachnida" family="Caponiidae" genus="Nyetnops" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="214" phylum="Arthropoda" rank="species" species="lachonta">
<emphasis id="7AACBD48765DFFF2FDD1FC3039CAFC6A" box="[522,662,935,961]" italics="true" pageId="10" pageNumber="214">N. lachonta</emphasis>
</taxonomicName>
by having the tegulum basal section of almost the same width as the distal section across the entire length of the bulb (
<figureCitation id="D0E37DDF765DFFF2FCB3FC5338B6FC4E" box="[872,1002,971,997]" captionStart="Fig" captionStartId="12.[189,232,1928,1954]" captionTargetBox="[271,1315,819,1889]" captionTargetId="figure-136@12.[271,1317,816,1889]" captionTargetPageId="12" captionText="Fig. 7. Nyetnops madre Villarreal &amp; Martínez sp. nov. A–B. ♂, holotype (MUSM–ENT 0503213). C–D. ♀, paratype (MUSM–ENT 0503206). A. Left palp, prolateral view. B. Left palp, retrolateral view. C. External genitalia, ventral view. D. Internal genitalia, dorsal view. Scale bars=0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11145123" httpUri="https://zenodo.org/record/11145123/files/figure.png" pageId="10" pageNumber="214">Fig. 7A–B</figureCitation>
), instead of a distinctly narrower basal section as in
<taxonomicName id="8FD81AD9765DFFF2FE44FC773957FBA3" authorityName="Sanchez-Ruiz, Brescovit &amp; Bonaldo" authorityYear="2020" box="[415,523,1006,1032]" class="Arachnida" family="Caponiidae" genus="Nyetnops" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="214" phylum="Arthropoda" rank="species" species="buruti">
<emphasis id="7AACBD48765DFFF2FE44FC773957FBA3" box="[415,523,1006,1032]" italics="true" pageId="10" pageNumber="214">N. buruti</emphasis>
</taxonomicName>
and the narrowing in the anterior section of the tegulum apical section in
<taxonomicName id="8FD81AD9765DFFF2FF65FB8A3A1BFB80" authorityName="Sanchez-Ruiz, Brescovit &amp; Bonaldo" authorityYear="2020" box="[190,327,1041,1067]" class="Arachnida" family="Caponiidae" genus="Nyetnops" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="214" phylum="Arthropoda" rank="species" species="lachonta">
<emphasis id="7AACBD48765DFFF2FF65FB8A3A1BFB80" box="[190,327,1041,1067]" italics="true" pageId="10" pageNumber="214">N. lachonta</emphasis>
</taxonomicName>
; additionally, the males of this species have an embolar subapical dorsal and ventral serrated keel unique in the genus. Females are similar to those of
<taxonomicName id="8FD81AD9765DFFF2FC86FBAD3881FBE5" authorityName="Platnick &amp; Lise" authorityYear="2007" box="[861,989,1077,1102]" class="Arachnida" family="Caponiidae" genus="Nyetnops" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="214" phylum="Arthropoda" rank="species" species="guarani">
<emphasis id="7AACBD48765DFFF2FC86FBAD3881FBE5" box="[861,989,1077,1102]" italics="true" pageId="10" pageNumber="214">N. guarani</emphasis>
</taxonomicName>
by having the posterior plate wide and a triangular distal margin of the receptaculum but can be distinguished by the less sharp dorsal margin of the receptaculum (
<figureCitation id="D0E37DDF765DFFF2FDCBFBE339CEFB3E" box="[528,658,1147,1173]" captionStart="Fig" captionStartId="12.[189,232,1928,1954]" captionTargetBox="[271,1315,819,1889]" captionTargetId="figure-136@12.[271,1317,816,1889]" captionTargetPageId="12" captionText="Fig. 7. Nyetnops madre Villarreal &amp; Martínez sp. nov. A–B. ♂, holotype (MUSM–ENT 0503213). C–D. ♀, paratype (MUSM–ENT 0503206). A. Left palp, prolateral view. B. Left palp, retrolateral view. C. External genitalia, ventral view. D. Internal genitalia, dorsal view. Scale bars=0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11145123" httpUri="https://zenodo.org/record/11145123/files/figure.png" pageId="10" pageNumber="214">Fig. 7C–D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="00C232D1765DFFF2FF66FB243FE1FAAF" box="[189,1213,1212,1285]" pageId="10" pageNumber="214" type="etymology">
<paragraph id="4867615A765DFFF2FF66FB243A19FB7D" blockId="10.[189,325,1212,1238]" box="[189,325,1212,1238]" pageId="10" pageNumber="214">
<heading id="132FD636765DFFF2FF66FB243A19FB7D" bold="true" box="[189,325,1212,1238]" fontSize="11" level="3" pageId="10" pageNumber="214" reason="3">
<emphasis id="7AACBD48765DFFF2FF66FB243A19FB7D" bold="true" box="[189,325,1212,1238]" pageId="10" pageNumber="214">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="4867615A765DFFF2FF66FB723FE1FAAF" blockId="10.[189,1213,1258,1285]" box="[189,1213,1258,1285]" pageId="10" pageNumber="214">
The specific epithet is a noun in apposition taken from the
<typeStatus id="9763DFF8765DFFF2FCB6FB7338FCFAAE" box="[877,928,1259,1285]" pageId="10" pageNumber="214">type</typeStatus>
locality
<collectingRegion id="8A1CAFB8765DFFF2FBDCFB723FE5FAAF" box="[1031,1209,1258,1284]" country="Peru" name="Madre de Dios" pageId="10" pageNumber="214">Madre de Dios</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="00C232D1765DFFF2FF66FAB43838F89E" pageId="10" pageNumber="214" type="materials_examined">
<paragraph id="4867615A765DFFF2FF66FAB43A31FAED" blockId="10.[189,365,1324,1350]" box="[189,365,1324,1350]" pageId="10" pageNumber="214">
<heading id="132FD636765DFFF2FF66FAB43A31FAED" bold="true" box="[189,365,1324,1350]" fontSize="11" level="3" pageId="10" pageNumber="214" reason="3">
<emphasis id="7AACBD48765DFFF2FF66FAB43A31FAED" bold="true" box="[189,365,1324,1350]" pageId="10" pageNumber="214">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="4867615A765DFFF2FF31FAC338C8FA17" blockId="10.[189,1399,1371,1468]" pageId="10" pageNumber="214">
<materialsCitation id="F8B06B07765DFFF2FF31FAC338C8FA17" collectingDate="2010-06-09" collectingDateMax="2010-06-13" collectingDateMin="2010-06-09" collectionCode="MUSM" collectorName="H. Silva" country="Peru" elevation="270" latitude="-12.569166" location="CICRA" longLatPrecision="15" longitude="-70.10011" municipality="Boca Rio Los Amigos" pageId="10" pageNumber="214" specimenCode="ENT 0503213" specimenCount="1" specimenCount-male="1" stateProvince="Madre de Dios" typeStatus="holotype">
<emphasis id="7AACBD48765DFFF2FF31FAC33A01FADE" bold="true" box="[234,349,1371,1397]" pageId="10" pageNumber="214">
<typeStatus id="9763DFF8765DFFF2FF31FAC33A01FADE" box="[234,349,1371,1397]" pageId="10" pageNumber="214" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="30CF21CA765DFFF2FF66FAE63A54FA33" box="[189,264,1406,1432]" name="Peru" pageId="10" pageNumber="214">PERU</collectingCountry>
•
<specimenCount id="5EDEAAD3765DFFF2FEF9FAE63A67FA33" box="[290,315,1406,1432]" count="1" pageId="10" pageNumber="214" type="male">♂</specimenCount>
;
<collectingRegion id="8A1CAFB8765DFFF2FE93FAE63AA4FA33" box="[328,504,1406,1432]" country="Peru" name="Madre de Dios" pageId="10" pageNumber="214">Madre de Dios</collectingRegion>
department,
<collectingMunicipality id="A803FB20765DFFF2FD48FAE638C0FA33" box="[659,924,1406,1432]" pageId="10" pageNumber="214">Boca Río Los Amigos</collectingMunicipality>
,
<location id="4D073781765DFFF2FC72FAE63F5AFA33" LSID="urn:lsid:plazi:treatment:C071D04C765DFFF6FDD8FD333A06FE4F:4D073781765DFFF2FC72FAE63F5AFA33" box="[937,1030,1406,1432]" country="Peru" latitude="-12.569166" longLatPrecision="15" longitude="-70.10011" municipality="Boca Rio Los Amigos" name="CICRA" pageId="10" pageNumber="214" stateProvince="Madre de Dios">CICRA</location>
;
<geoCoordinate id="2DEC079D765DFFF2FBC9FAE63FC8FA33" box="[1042,1172,1404,1432]" degrees="12" direction="south" minutes="34" orientation="latitude" pageId="10" pageNumber="214" precision="15" seconds="9" value="-12.569166">12°34′9″ S</geoCoordinate>
,
<geoCoordinate id="2DEC079D765DFFF2FB79FAE63E19FA33" box="[1186,1349,1404,1432]" degrees="70" direction="west" minutes="6" orientation="longitude" pageId="10" pageNumber="214" precision="1" seconds="0.40" value="-70.10011">70°6′0.40″ W</geoCoordinate>
; alt.
<quantity id="8F20CCBF765DFFF2FF65FA393A57FA10" box="[190,267,1441,1467]" metricMagnitude="2" metricUnit="m" metricValue="2.7" pageId="10" pageNumber="214" unit="m" value="270.0">
<elevation id="C3F58669765DFFF2FF65FA393A57FA10" box="[190,267,1441,1467]" metricMagnitude="2" metricUnit="m" metricValue="2.7" pageId="10" pageNumber="214" unit="m" value="270.0">270 m</elevation>
</quantity>
;
<date id="3C66479A765DFFF2FECCFA393A93FA10" box="[279,463,1441,1467]" pageId="10" pageNumber="214" value="2010-06-09" valueMax="2010-06-13" valueMin="2010-06-09">
<collectingDate id="2C22BE72765DFFF2FECCFA393A93FA10" box="[279,463,1441,1467]" pageId="10" pageNumber="214" value="2010-06-09" valueMax="2010-06-13" valueMin="2010-06-09">9–13 Jun. 2010</collectingDate>
</date>
;
<collectorName id="E52D048C765DFFF2FE00FA393967FA10" box="[475,571,1441,1467]" pageId="10" pageNumber="214">H. Silva</collectorName>
leg.;
<collectionCode id="2EC9F99F765DFFF2FDA6FA39398AFA10" box="[637,726,1441,1467]" pageId="10" pageNumber="214">MUSM</collectionCode>
–
<specimenCode id="187EC921765DFFF2FD3EFA3938CCFA17" box="[741,912,1441,1468]" collectionCode="MUSM" pageId="10" pageNumber="214">ENT 0503213</specimenCode>
.
</materialsCitation>
</paragraph>
<paragraph id="4867615A765DFFF2FF31FA7A3A70F921" blockId="10.[189,1399,1506,1674]" pageId="10" pageNumber="214">
<materialsCitation id="F8B06B07765DFFF2FF31FA7A3F71F98B" collectingDate="2010-06-09" collectingDateMax="2010-06-13" collectingDateMin="2010-06-09" collectionCode="MUSM" collectorName="H. Silva" country="Peru" elevation="270" latitude="-12.569166" location="Rio Los Amigos" longLatPrecision="15" longitude="-70.10011" municipality="Boca Rio Los Amigos" pageId="10" pageNumber="214" specimenCode="ENT 0503206" specimenCount="1" specimenCount-female="1" stateProvince="Madre de Dios" typeStatus="paratype">
<emphasis id="7AACBD48765DFFF2FF31FA7A3A35FA57" bold="true" box="[234,361,1506,1532]" pageId="10" pageNumber="214">
<typeStatus id="9763DFF8765DFFF2FF31FA7A3A35FA57" box="[234,361,1506,1532]" pageId="10" pageNumber="214" type="paratype">Paratypes</typeStatus>
</emphasis>
<collectingCountry id="30CF21CA765DFFF2FF66F99E3A54F98B" box="[189,264,1542,1568]" name="Peru" pageId="10" pageNumber="214">PERU</collectingCountry>
•
<specimenCount id="5EDEAAD3765DFFF2FEF8F99E3A0DF98B" box="[291,337,1542,1568]" count="1" pageId="10" pageNumber="214" type="female">1 ♀</specimenCount>
; same collection data as for holotype;
<collectionCode id="2EC9F99F765DFFF2FCC6F99E382AF98B" box="[797,886,1542,1568]" pageId="10" pageNumber="214">MUSM</collectionCode>
–
<specimenCode id="187EC921765DFFF2FC5EF99E3F71F98B" box="[901,1069,1541,1568]" collectionCode="MUSM" pageId="10" pageNumber="214">ENT 0503206</specimenCode>
</materialsCitation>
•
<materialsCitation id="F8B06B07765DFFF2FB9CF99E39C4F9EF" collectingDate="2010-06-09" collectingDateMax="2010-06-13" collectingDateMin="2010-06-09" collectionCode="MUSM" collectorName="H. Silva" country="Peru" elevation="270" latitude="-12.569166" location="Rio Los Amigos" longLatPrecision="15" longitude="-70.10011" municipality="Boca Rio Los Amigos" pageId="10" pageNumber="214" specimenCode="ENT 0515831" specimenCount="1" specimenCount-male="1" stateProvince="Madre de Dios" typeStatus="paratype">
<specimenCount id="5EDEAAD3765DFFF2FB9CF99E3F2BF98B" box="[1095,1143,1542,1568]" count="1" pageId="10" pageNumber="214" type="male">1 ♂</specimenCount>
; same collection data as for holotype;
<collectionCode id="2EC9F99F765DFFF2FE5EF9B13A82F9E8" box="[389,478,1577,1603]" pageId="10" pageNumber="214">MUSM</collectionCode>
–
<specimenCode id="187EC921765DFFF2FE36F9B139C4F9EF" box="[493,664,1577,1604]" collectionCode="MUSM" pageId="10" pageNumber="214">ENT 0515831</specimenCode>
</materialsCitation>
•
<materialsCitation id="F8B06B07765DFFF2FD6DF9B13A78F9CC" collectingDate="2010-06-09" collectingDateMax="2010-06-13" collectingDateMin="2010-06-09" collectionCode="MUSM" collectorName="H. Silva" country="Peru" elevation="270" latitude="-12.569166" location="Rio Los Amigos" longLatPrecision="15" longitude="-70.10011" municipality="Boca Rio Los Amigos" pageId="10" pageNumber="214" specimenCode="ENT 0515830" specimenCount="2" specimenCount-male="2" stateProvince="Madre de Dios" typeStatus="paratype">
<specimenCount id="5EDEAAD3765DFFF2FD6DF9B139A2F9E8" box="[694,766,1577,1603]" count="2" pageId="10" pageNumber="214" type="male">2 ♂♂</specimenCount>
; same collection data as for holotype;
<collectionCode id="2EC9F99F765DFFF2FB0DF9B13E73F9E8" box="[1238,1327,1577,1603]" pageId="10" pageNumber="214">MUSM</collectionCode>
–
<specimenCode id="187EC921765DFFF2FAE6F9B13A78F9CC" collectionCode="MUSM" pageId="10" pageNumber="214">ENT 0515830</specimenCode>
</materialsCitation>
•
<materialsCitation id="F8B06B07765DFFF2FE9FF9D43A74F921" collectingDate="2010-06-14" collectingDateMax="2010-06-13" collectingDateMin="2010-06-09" collectionCode="MUSM" collectorName="M. Vicchez" country="Peru" elevation="270" latitude="-12.569166" location="Rio Los Amigos" longLatPrecision="15" longitude="-70.10011" municipality="Boca Rio Los Amigos" pageId="10" pageNumber="214" specimenCode="ENT 0503223" specimenCount="1" specimenCount-male="1" stateProvince="Madre de Dios" typeStatus="paratype">
<specimenCount id="5EDEAAD3765DFFF2FE9FF9D43A2AF9CC" box="[324,374,1612,1639]" count="1" pageId="10" pageNumber="214" type="male">1 ♂</specimenCount>
; same collection data as for holotype;
<date id="3C66479A765DFFF2FC89F9D438AFF9CD" box="[850,1011,1612,1639]" pageId="10" pageNumber="214" value="2010-06-14">
<collectingDate id="2C22BE72765DFFF2FC89F9D438AFF9CD" box="[850,1011,1612,1639]" pageId="10" pageNumber="214" value="2010-06-14">14 Jun. 2010</collectingDate>
</date>
;
<collectorName id="E52D048C765DFFF2FBD9F9D53FD0F9CD" box="[1026,1164,1612,1639]" pageId="10" pageNumber="214">M. Vicchez</collectorName>
leg.;
<collectionCode id="2EC9F99F765DFFF2FB0DF9D43E73F9CD" box="[1238,1327,1612,1638]" pageId="10" pageNumber="214">MUSM</collectionCode>
–
<specimenCode id="187EC921765DFFF2FAE6F9D43A74F921" collectionCode="MUSM" pageId="10" pageNumber="214">ENT 0503223</specimenCode>
</materialsCitation>
.
</paragraph>
<paragraph id="4867615A765DFFF2FF31F9283AF4F961" blockId="10.[189,1398,1712,1845]" box="[234,424,1712,1738]" pageId="10" pageNumber="214">
<emphasis id="7AACBD48765DFFF2FF31F9283AF4F961" bold="true" box="[234,424,1712,1738]" pageId="10" pageNumber="214">Other material</emphasis>
</paragraph>
<paragraph id="4867615A765DFFF2FF66F94C3838F89E" blockId="10.[189,1398,1712,1845]" pageId="10" pageNumber="214">
<materialsCitation id="F8B06B07765DFFF2FF66F94C3842F8B9" collectingDate="1998-05-07" collectingDateMax="1998-05-28" collectingDateMin="1998-05-07" collectionCode="MUSM" collectorName="S. Cordova" country="Peru" elevation="465" latitude="-11.70625" location="Pagoreni" longLatPrecision="1" longitude="-72.90298" municipality="Rio Camisea" pageId="10" pageNumber="214" specimenCode="ENT 0506898" specimenCount="1" specimenCount-male="1" stateProvince="Cusco">
<collectingCountry id="30CF21CA765DFFF2FF66F94C3A54F945" box="[189,264,1748,1774]" name="Peru" pageId="10" pageNumber="214">PERU</collectingCountry>
•
<specimenCount id="5EDEAAD3765DFFF2FEC5F94C3A10F945" box="[286,332,1748,1774]" count="1" pageId="10" pageNumber="214" type="male">1 ♂</specimenCount>
;
<collectingRegion id="8A1CAFB8765DFFF2FE8CF94C3AFCF945" box="[343,416,1748,1774]" country="Peru" name="Cusco" pageId="10" pageNumber="214">Cusco</collectingRegion>
department,
<collectingMunicipality id="A803FB20765DFFF2FDE3F94C398CF945" box="[568,720,1748,1774]" pageId="10" pageNumber="214">Río Camisea</collectingMunicipality>
,
<location id="4D073781765DFFF2FD01F94C381AF945" LSID="urn:lsid:plazi:treatment:C071D04C765DFFF6FDD8FD333A06FE4F:4D073781765DFFF2FD01F94C381AF945" box="[730,838,1748,1774]" country="Peru" latitude="-11.70625" longLatPrecision="1" longitude="-72.90298" municipality="Rio Camisea" name="Pagoreni" pageId="10" pageNumber="214" stateProvince="Cusco">Pagoreni</location>
;
<geoCoordinate id="2DEC079D765DFFF2FC8AF94A38A8F945" box="[849,1012,1746,1774]" degrees="11" direction="south" minutes="42" orientation="latitude" pageId="10" pageNumber="214" precision="1" seconds="22.5" value="-11.70625">11°42′22.5″ S</geoCoordinate>
,
<geoCoordinate id="2DEC079D765DFFF2FBDBF94A3FF3F945" box="[1024,1199,1746,1774]" degrees="72" direction="west" minutes="54" orientation="longitude" pageId="10" pageNumber="214" precision="1" seconds="10.7" value="-72.90298">72°54′10.7″ W</geoCoordinate>
; alt.
<quantity id="8F20CCBF765DFFF2FB3EF94C3E6DF945" box="[1253,1329,1748,1775]" metricMagnitude="2" metricUnit="m" metricValue="4.65" pageId="10" pageNumber="214" unit="m" value="465.0">
<elevation id="C3F58669765DFFF2FB3EF94C3E6DF945" box="[1253,1329,1748,1775]" metricMagnitude="2" metricUnit="m" metricValue="4.65" pageId="10" pageNumber="214" unit="m" value="465.0">465 m</elevation>
</quantity>
;
<date id="3C66479A765DFFF2FAE7F94C3A65F8BA" pageId="10" pageNumber="214" value="1998-05-07" valueMax="1998-05-28" valueMin="1998-05-07">
<collectingDate id="2C22BE72765DFFF2FAE7F94C3A65F8BA" pageId="10" pageNumber="214" value="1998-05-07" valueMax="1998-05-28" valueMin="1998-05-07">7–28 May 1998</collectingDate>
</date>
;
<collectorName id="E52D048C765DFFF2FE9DF96F3A96F8BA" box="[326,458,1783,1809]" pageId="10" pageNumber="214">S. Cordova</collectorName>
leg.;
<collectionCode id="2EC9F99F765DFFF2FDD5F96F393BF8BA" box="[526,615,1783,1809]" pageId="10" pageNumber="214">MUSM</collectionCode>
–
<specimenCode id="187EC921765DFFF2FDADF96F3842F8B9" box="[630,798,1783,1810]" collectionCode="MUSM" pageId="10" pageNumber="214">ENT 0506898</specimenCode>
</materialsCitation>
•
<materialsCitation id="F8B06B07765DFFF2FCE3F96F383CF89E" collectingDate="1998-05-07" collectingDateMax="1998-05-29" collectingDateMin="1998-05-07" collectionCode="MUSM" collectorName="S. Cordova" country="Peru" elevation="465" latitude="-11.70625" location="Pagoreni" longLatPrecision="1" longitude="-72.90298" municipality="Rio Camisea" pageId="10" pageNumber="214" specimenCode="ENT 0515832" specimenCount="1" specimenCount-male="1" stateProvince="Cusco">
<specimenCount id="5EDEAAD3765DFFF2FCE3F96F383BF8B9" box="[824,871,1783,1810]" count="1" pageId="10" pageNumber="214" type="male">1 ♂</specimenCount>
; same collection data as for preceding;
<date id="3C66479A765DFFF2FAE7F96F3A64F89F" pageId="10" pageNumber="214" value="1998-05-07" valueMax="1998-05-29" valueMin="1998-05-07">
<collectingDate id="2C22BE72765DFFF2FAE7F96F3A64F89F" pageId="10" pageNumber="214" value="1998-05-07" valueMax="1998-05-29" valueMin="1998-05-07">7–29 May 1998</collectingDate>
</date>
;
<collectorName id="E52D048C765DFFF2FE9FF8823A94F89F" box="[324,456,1818,1844]" pageId="10" pageNumber="214">S. Cordova</collectorName>
<emphasis id="7AACBD48765DFFF2FE0BF8833957F89F" box="[464,523,1818,1844]" italics="true" pageId="10" pageNumber="214">et al.</emphasis>
leg.;
<collectionCode id="2EC9F99F765DFFF2FD96F88239FAF89F" box="[589,678,1818,1844]" pageId="10" pageNumber="214">MUSM</collectionCode>
–
<specimenCode id="187EC921765DFFF2FD6EF882383CF89E" box="[693,864,1818,1845]" collectionCode="MUSM" pageId="10" pageNumber="214">ENT 0515832</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="00C232D1765DFFF6FF66F8C3389DFEFA" lastPageId="14" lastPageNumber="218" pageId="10" pageNumber="214" type="description">
<paragraph id="4867615A765DFFF2FF66F8C33A13F8DE" blockId="10.[189,335,1883,1909]" box="[189,335,1883,1909]" pageId="10" pageNumber="214">
<heading id="132FD636765DFFF2FF66F8C33A13F8DE" bold="true" box="[189,335,1883,1909]" fontSize="11" level="3" pageId="10" pageNumber="214" reason="3">
<emphasis id="7AACBD48765DFFF2FF66F8C33A13F8DE" bold="true" box="[189,335,1883,1909]" pageId="10" pageNumber="214">Description</emphasis>
</heading>
</paragraph>
<paragraph id="4867615A765DFFF2FF31F8123996F80E" blockId="10.[234,714,1930,1957]" box="[234,714,1930,1957]" pageId="10" pageNumber="214">
<emphasis id="7AACBD48765DFFF2FF31F8123A76F80F" bold="true" box="[234,298,1930,1956]" pageId="10" pageNumber="214">Male</emphasis>
(
<typeStatus id="9763DFF8765DFFF2FEE1F8123AF9F80F" box="[314,421,1930,1956]" pageId="10" pageNumber="214" type="holotype">holotype</typeStatus>
, MUSM–ENT 0503213)
</paragraph>
<paragraph id="4867615A765CFFF3FF66FE913E36FEC1" blockId="11.[188,1399,265,362]" pageId="11" pageNumber="215">
<smallCapsWord id="4E81F786765CFFF3FF66FE913A14FE8A" baselines="284,284" box="[189,328,265,291]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Coloration" pageId="11" pageNumber="215">COLORATION</smallCapsWord>
(
<figureCitation id="D0E37DDF765CFFF3FE83FE913947FE88" box="[344,539,265,291]" captionStart="Fig" captionStartId="11.[190,233,1928,1954]" captionTargetBox="[268,1321,704,1889]" captionTargetId="figure-142@11.[267,1321,704,1889]" captionTargetPageId="11" captionText="Fig. 6. Nyetnops madre Villarreal &amp; Martínez sp. nov. A–B, E, G. ♂, holotype (MUSM–ENT 0503213). C–D, F, H. ♀, paratype (MUSM–ENT 0503206). A–B. Habitus, dorsal view. C–D. Habitus, ventral view. E–F. Prosoma, dorsal view. G–H. Prosoma, ventral view. Scale bars =1 mm." figureDoi="http://doi.org/10.5281/zenodo.11145121" httpUri="https://zenodo.org/record/11145121/files/figure.png" pageId="11" pageNumber="215">Fig. 6A–B, E, G</figureCitation>
). Carapace orange. Chelicerae, endites, labium, and sternum light orange. Legs light orange. Abdomen: dorsally dark gray, without noticeable pattern (
<figureCitation id="D0E37DDF765CFFF3FB8EFEB43FEBFEED" box="[1109,1207,300,326]" captionStart="Fig" captionStartId="11.[190,233,1928,1954]" captionTargetBox="[268,1321,704,1889]" captionTargetId="figure-142@11.[267,1321,704,1889]" captionTargetPageId="11" captionText="Fig. 6. Nyetnops madre Villarreal &amp; Martínez sp. nov. A–B, E, G. ♂, holotype (MUSM–ENT 0503213). C–D, F, H. ♀, paratype (MUSM–ENT 0503206). A–B. Habitus, dorsal view. C–D. Habitus, ventral view. E–F. Prosoma, dorsal view. G–H. Prosoma, ventral view. Scale bars =1 mm." figureDoi="http://doi.org/10.5281/zenodo.11145121" httpUri="https://zenodo.org/record/11145121/files/figure.png" pageId="11" pageNumber="215">Fig. 6A</figureCitation>
). Ventrally gray yellowish, darker on epiandric area. Spinnerets beige. Anterior legs without arolium, crista or gladius.
</paragraph>
<paragraph id="4867615A765CFFF3FF66FE093F26FE59" blockId="11.[189,1399,401,498]" pageId="11" pageNumber="215">
<smallCapsWord id="4E81F786765CFFF3FF66FE093A32FE02" baselines="420,420" box="[189,366,401,427]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Measurements" pageId="11" pageNumber="215">MEASUREMENTS</smallCapsWord>
. Total length 3.51; carapace length 1.54; width 1.30; height 0.68. Clypeus height 0.31. Eye diameters and interdistances: PME 0.17; PME–PME 0.39. Chelicerae length 0.47. Sternum length 0.98; width 0.85. Legs: I: 5.21; II: 4.70; III: 4.29; IV: 6.37. Abdomen length 1.95.
</paragraph>
<paragraph id="4867615A765CFFF4FF66FD81396AFEED" blockId="11.[189,1399,537,598]" lastBlockId="12.[188,1398,265,326]" lastPageId="12" lastPageNumber="216" pageId="11" pageNumber="215">
<smallCapsWord id="4E81F786765CFFF3FF66FD813A6DFD9A" baselines="556,556" box="[189,305,537,563]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Genitalia" pageId="11" pageNumber="215">GENITALIA</smallCapsWord>
. Very elongated copulatory bulb, tegulum basal section slightly swollen and almost as width as apical section, tegulum surface smooth, embolus surface with sparse longitudinal grooves, embolus with dorsal and ventral dentated keel and tip with very long projections, dorsally petal-like and ventrally rooster-like (
<figureCitation id="D0E37DDF765BFFF4FE89FEB43A8AFEED" box="[338,470,300,326]" captionStart="Fig" captionStartId="12.[189,232,1928,1954]" captionTargetBox="[271,1315,819,1889]" captionTargetId="figure-136@12.[271,1317,816,1889]" captionTargetPageId="12" captionText="Fig. 7. Nyetnops madre Villarreal &amp; Martínez sp. nov. A–B. ♂, holotype (MUSM–ENT 0503213). C–D. ♀, paratype (MUSM–ENT 0503206). A. Left palp, prolateral view. B. Left palp, retrolateral view. C. External genitalia, ventral view. D. Internal genitalia, dorsal view. Scale bars=0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11145123" httpUri="https://zenodo.org/record/11145123/files/figure.png" pageId="12" pageNumber="216">Figs 7A–B</figureCitation>
,
<figureCitation id="D0E37DDF765BFFF4FE39FEB43976FEED" box="[482,554,300,326]" captionStart="Fig" captionStartId="13.[189,232,1892,1918]" captionTargetBox="[249,1338,265,1855]" captionTargetId="figure-18@13.[249,1338,265,1855]" captionTargetPageId="13" captionText="Fig. 8. Nyetnops madre Villarreal &amp; Martínez sp. nov., ♂, paratype (MUSM–ENT 0515830). A, C. Left palp, dorsal view. B, F. Tip of the embolus, prolateral view. D. Tip of the embolus, proventral view E. Copulatory bulb, prolateral view. G. Copulatory bulb, retrolateral view. H. Tip of the embolus, dorsal view. Scale bars: A, C, E, G =0.1, mm; B, D, F, H =0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.11145125" httpUri="https://zenodo.org/record/11145125/files/figure.png" pageId="12" pageNumber="216">8A–F</figureCitation>
).
</paragraph>
<caption id="1CA731D2765CFFF3FF65F8103FD6F842" ID-DOI="http://doi.org/10.5281/zenodo.11145121" ID-Zenodo-Dep="11145121" httpUri="https://zenodo.org/record/11145121/files/figure.png" pageId="11" pageNumber="215" startId="11.[190,233,1928,1954]" targetBox="[268,1321,704,1889]" targetPageId="11">
<paragraph id="4867615A765CFFF3FF65F8103FD6F842" blockId="11.[189,1399,1928,2025]" pageId="11" pageNumber="215">
<emphasis id="7AACBD48765CFFF3FF65F8103A57F809" bold="true" box="[190,267,1928,1954]" pageId="11" pageNumber="215">Fig. 6.</emphasis>
<taxonomicName id="8FD81AD9765CFFF3FED4F8113993F809" authority="Villarreal &amp; Martinez" authorityName="Villarreal &amp; Martinez" authorityYear="2024" box="[271,719,1928,1954]" class="Arachnida" family="Caponiidae" genus="Nyetnops" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="215" phylum="Arthropoda" rank="species" species="madre" status="sp. nov.">
<emphasis id="7AACBD48765CFFF3FED4F8113A91F809" box="[271,461,1928,1954]" italics="true" pageId="11" pageNumber="215">Nyetnops madre</emphasis>
Villarreal &amp; Martínez
</taxonomicName>
<taxonomicNameLabel id="619F0033765CFFF3FD0EF8103872F809" box="[725,814,1928,1954]" pageId="11" pageNumber="215" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="7AACBD48765CFFF3FCE9F81038ECF809" bold="true" box="[818,944,1928,1954]" pageId="11" pageNumber="215">A–B, E, G</emphasis>
. ♂, holotype (MUSM–ENT 0503213).
<emphasis id="7AACBD48765CFFF3FF66F8333A1CF86E" bold="true" box="[189,320,1963,1989]" pageId="11" pageNumber="215">C–D, F, H</emphasis>
. ♀, paratype (MUSM–ENT 0503206).
<emphasis id="7AACBD48765CFFF3FCC5F833380AF86E" bold="true" box="[798,854,1963,1989]" pageId="11" pageNumber="215">A–B</emphasis>
. Habitus, dorsal view.
<emphasis id="7AACBD48765CFFF3FBB5F8333FFBF86E" bold="true" box="[1134,1191,1963,1989]" pageId="11" pageNumber="215">C–D</emphasis>
. Habitus, ventral view.
<emphasis id="7AACBD48765CFFF3FED8F8563A64F843" bold="true" box="[259,312,1998,2024]" pageId="11" pageNumber="215">E–F</emphasis>
. Prosoma, dorsal view.
<emphasis id="7AACBD48765CFFF3FD8BF85639D1F843" bold="true" box="[592,653,1998,2024]" pageId="11" pageNumber="215">G–H</emphasis>
. Prosoma, ventral view. Scale bars =1 mm.
</paragraph>
</caption>
<paragraph id="4867615A765BFFF4FF31FEF639B8FE23" blockId="12.[189,1276,365,427]" box="[234,740,365,392]" pageId="12" pageNumber="216">
<emphasis id="7AACBD48765BFFF4FF31FEF63A1AFE23" bold="true" box="[234,326,366,392]" pageId="12" pageNumber="216">Female</emphasis>
(
<typeStatus id="9763DFF8765BFFF4FE8DFEF63A9CFE23" box="[342,448,366,392]" pageId="12" pageNumber="216" type="paratype">paratype</typeStatus>
, MUSM–ENT 0503206)
</paragraph>
<paragraph id="4867615A765BFFF4FF66FE093FA0FE00" blockId="12.[189,1276,365,427]" box="[189,1276,401,427]" pageId="12" pageNumber="216">
<smallCapsWord id="4E81F786765BFFF4FF66FE093A14FE02" baselines="420,420" box="[189,328,401,427]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Coloration" pageId="12" pageNumber="216">COLORATION</smallCapsWord>
<smallCapsWord id="4E81F786765BFFF4FE94FE0E3A20FE02" baselines="420" box="[335,380,406,425]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="and" pageId="12" pageNumber="216">AND</smallCapsWord>
<smallCapsWord id="4E81F786765BFFF4FE58FE0E395EFE02" baselines="420" box="[387,514,406,425]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="abdominal" pageId="12" pageNumber="216">ABDOMINAL</smallCapsWord>
<smallCapsWord id="4E81F786765BFFF4FDD1FE0E393FFE02" baselines="420" box="[522,611,406,425]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="pattern" pageId="12" pageNumber="216">PATTERN</smallCapsWord>
. As male, but with lighter color tones (
<figureCitation id="D0E37DDF765BFFF4FBF5FE093FB3FE00" box="[1070,1263,401,427]" captionStart="Fig" captionStartId="11.[190,233,1928,1954]" captionTargetBox="[268,1321,704,1889]" captionTargetId="figure-142@11.[267,1321,704,1889]" captionTargetPageId="11" captionText="Fig. 6. Nyetnops madre Villarreal &amp; Martínez sp. nov. A–B, E, G. ♂, holotype (MUSM–ENT 0503213). C–D, F, H. ♀, paratype (MUSM–ENT 0503206). A–B. Habitus, dorsal view. C–D. Habitus, ventral view. E–F. Prosoma, dorsal view. G–H. Prosoma, ventral view. Scale bars =1 mm." figureDoi="http://doi.org/10.5281/zenodo.11145121" httpUri="https://zenodo.org/record/11145121/files/figure.png" pageId="12" pageNumber="216">Fig. 6C–D, F, H</figureCitation>
).
</paragraph>
<paragraph id="4867615A765BFFF4FF66FE4B3F25FD98" blockId="12.[189,1400,466,563]" pageId="12" pageNumber="216">
<smallCapsWord id="4E81F786765BFFF4FF66FE4B3A32FE40" baselines="486,486" box="[189,366,467,493]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Measurements" pageId="12" pageNumber="216">MEASUREMENTS</smallCapsWord>
. Total length 5.08, carapace length 1.65, width 1.50, height 0.44. Clypeus height 0.33. Eye diameters and interdistances: PME 0.16, PME–PME 0.38. Chelicerae length 0.51. Sternum length 1.16, width 1.05. Legs: I: 6.20; II: 5.76; III: 5.37; IV: 7.55. Abdomen length 3.24.
</paragraph>
<paragraph id="4867615A765BFFF4FF66FDC23A6DFD10" blockId="12.[187,1399,602,699]" pageId="12" pageNumber="216">
<smallCapsWord id="4E81F786765BFFF4FF66FDC23A6DFDD8" baselines="621,622" box="[189,305,602,628]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Genitalia" pageId="12" pageNumber="216">GENITALIA</smallCapsWord>
. External genital area with weakly sclerotized anterior plate, lep and ess, posterior plate narrow (
<figureCitation id="D0E37DDF765BFFF4FF1FFDE53A7BFD3C" box="[196,295,637,663]" captionStart="Fig" captionStartId="12.[189,232,1928,1954]" captionTargetBox="[271,1315,819,1889]" captionTargetId="figure-136@12.[271,1317,816,1889]" captionTargetPageId="12" captionText="Fig. 7. Nyetnops madre Villarreal &amp; Martínez sp. nov. A–B. ♂, holotype (MUSM–ENT 0503213). C–D. ♀, paratype (MUSM–ENT 0503206). A. Left palp, prolateral view. B. Left palp, retrolateral view. C. External genitalia, ventral view. D. Internal genitalia, dorsal view. Scale bars=0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11145123" httpUri="https://zenodo.org/record/11145123/files/figure.png" pageId="12" pageNumber="216">Fig. 7C</figureCitation>
). Internal genitalia with triangular margin of receptaculum, posterior plate well developed (
<figureCitation id="D0E37DDF765BFFF4FF1DFD383A78FD10" box="[198,292,672,699]" captionStart="Fig" captionStartId="12.[189,232,1928,1954]" captionTargetBox="[271,1315,819,1889]" captionTargetId="figure-136@12.[271,1317,816,1889]" captionTargetPageId="12" captionText="Fig. 7. Nyetnops madre Villarreal &amp; Martínez sp. nov. A–B. ♂, holotype (MUSM–ENT 0503213). C–D. ♀, paratype (MUSM–ENT 0503206). A. Left palp, prolateral view. B. Left palp, retrolateral view. C. External genitalia, ventral view. D. Internal genitalia, dorsal view. Scale bars=0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11145123" httpUri="https://zenodo.org/record/11145123/files/figure.png" pageId="12" pageNumber="216">Fig. 7D</figureCitation>
).
</paragraph>
<caption id="1CA731D2765BFFF4FF66F8103F82F842" ID-DOI="http://doi.org/10.5281/zenodo.11145123" ID-Zenodo-Dep="11145123" httpUri="https://zenodo.org/record/11145123/files/figure.png" pageId="12" pageNumber="216" startId="12.[189,232,1928,1954]" targetBox="[271,1315,819,1889]" targetPageId="12">
<paragraph id="4867615A765BFFF4FF66F8103F82F842" blockId="12.[188,1399,1928,2025]" pageId="12" pageNumber="216">
<emphasis id="7AACBD48765BFFF4FF66F8103A53F809" bold="true" box="[189,271,1928,1954]" pageId="12" pageNumber="216">Fig. 7.</emphasis>
<taxonomicName id="8FD81AD9765BFFF4FEC0F81139ACF809" authority="Villarreal &amp; Martinez" authorityName="Villarreal &amp; Martinez" authorityYear="2024" box="[283,752,1928,1954]" class="Arachnida" family="Caponiidae" genus="Nyetnops" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="216" phylum="Arthropoda" rank="species" species="madre" status="sp. nov.">
<emphasis id="7AACBD48765BFFF4FEC0F8113A82F809" box="[283,478,1928,1954]" italics="true" pageId="12" pageNumber="216">Nyetnops madre</emphasis>
Villarreal &amp; Martínez
</taxonomicName>
<taxonomicNameLabel id="619F0033765BFFF4FD20F8103805F809" box="[763,857,1928,1954]" pageId="12" pageNumber="216" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="7AACBD48765BFFF4FCBFF81038C0F809" bold="true" box="[868,924,1928,1954]" pageId="12" pageNumber="216">A–B</emphasis>
. ♂, holotype (MUSM–ENT 0503213).
<emphasis id="7AACBD48765BFFF4FF66F8333BAAF86E" bold="true" box="[189,246,1963,1989]" pageId="12" pageNumber="216">C–D</emphasis>
. ♀, paratype (MUSM–ENT 0503206).
<emphasis id="7AACBD48765BFFF4FD19F833398BF86E" bold="true" box="[706,727,1963,1989]" pageId="12" pageNumber="216">A</emphasis>
. Left palp, prolateral view.
<emphasis id="7AACBD48765BFFF4FBCCF8333F77F86E" bold="true" box="[1047,1067,1963,1989]" pageId="12" pageNumber="216">B</emphasis>
. Left palp, retrolateral view.
<emphasis id="7AACBD48765BFFF4FF67F8563B8DF843" bold="true" box="[188,209,1998,2024]" pageId="12" pageNumber="216">C</emphasis>
. External genitalia, ventral view.
<emphasis id="7AACBD48765BFFF4FD85F856392FF843" bold="true" box="[606,627,1998,2024]" pageId="12" pageNumber="216">D</emphasis>
. Internal genitalia, dorsal view. Scale bars=0.5 mm.
</paragraph>
</caption>
<caption id="1CA731D2765AFFF5FF66F8FC38D0F842" ID-DOI="http://doi.org/10.5281/zenodo.11145125" ID-Zenodo-Dep="11145125" httpUri="https://zenodo.org/record/11145125/files/figure.png" pageId="13" pageNumber="217" startId="13.[189,232,1892,1918]" targetBox="[249,1338,265,1855]" targetPageId="13">
<paragraph id="4867615A765AFFF5FF66F8FC38D0F842" blockId="13.[188,1399,1892,2025]" pageId="13" pageNumber="217">
<emphasis id="7AACBD48765AFFF5FF66F8FC3A56F8D4" bold="true" box="[189,266,1892,1919]" pageId="13" pageNumber="217">Fig. 8.</emphasis>
<taxonomicName id="8FD81AD9765AFFF5FECAF8FE3988F8D4" authority="Villarreal &amp; Martinez" authorityName="Villarreal &amp; Martinez" authorityYear="2024" box="[273,724,1893,1919]" class="Arachnida" family="Caponiidae" genus="Nyetnops" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="217" phylum="Arthropoda" rank="species" species="madre" status="sp. nov.">
<emphasis id="7AACBD48765AFFF5FECAF8FE3A8CF8D4" box="[273,464,1893,1919]" italics="true" pageId="13" pageNumber="217">Nyetnops madre</emphasis>
Villarreal &amp; Martínez
</taxonomicName>
<taxonomicNameLabel id="619F0033765AFFF5FD01F8FD386AF8D4" box="[730,822,1893,1919]" pageId="13" pageNumber="217" rank="species">sp. nov.</taxonomicNameLabel>
, ♂, paratype (MUSM–ENT 0515830).
<emphasis id="7AACBD48765AFFF5FB24F8FC3E6BF8D5" bold="true" box="[1279,1335,1892,1918]" pageId="13" pageNumber="217">A, C</emphasis>
. Left palp, dorsal view.
<emphasis id="7AACBD48765AFFF5FE40F8103A8EF809" bold="true" box="[411,466,1928,1954]" pageId="13" pageNumber="217">B, F</emphasis>
. Tip of the embolus, prolateral view.
<emphasis id="7AACBD48765AFFF5FC46F81038EEF809" bold="true" box="[925,946,1928,1954]" pageId="13" pageNumber="217">D</emphasis>
. Tip of the embolus, proventral view
<emphasis id="7AACBD48765AFFF5FF66F8333B8DF86E" bold="true" box="[189,209,1963,1989]" pageId="13" pageNumber="217">E</emphasis>
. Copulatory bulb, prolateral view.
<emphasis id="7AACBD48765AFFF5FDBEF8333920F86E" bold="true" box="[613,636,1963,1989]" pageId="13" pageNumber="217">G</emphasis>
. Copulatory bulb, retrolateral view.
<emphasis id="7AACBD48765AFFF5FBFAF8333F64F86E" bold="true" box="[1057,1080,1963,1989]" pageId="13" pageNumber="217">H</emphasis>
. Tip of the embolus, dorsal view. Scale bars: A, C, E, G =0.1, mm; B, D, F, H =0.01 mm.
</paragraph>
</caption>
<paragraph id="4867615A7659FFF6FF66FE903A68FE89" blockId="14.[189,308,264,290]" box="[189,308,264,290]" pageId="14" pageNumber="218">
<heading id="132FD6367659FFF6FF66FE903A68FE89" bold="true" box="[189,308,264,290]" fontSize="11" level="3" pageId="14" pageNumber="218" reason="3">
<emphasis id="7AACBD487659FFF6FF66FE903A68FE89" bold="true" box="[189,308,264,290]" pageId="14" pageNumber="218">Variation</emphasis>
</heading>
</paragraph>
<paragraph id="4867615A7659FFF6FF66FEAF389DFEFA" blockId="14.[189,961,311,338]" box="[189,961,311,338]" pageId="14" pageNumber="218">Males (n=3): total length: 3.51–3.96; carapace length: 1.49–1.83.</paragraph>
</subSubSection>
<subSubSection id="00C232D17659FFF6FF66FEE03A06FE4F" pageId="14" pageNumber="218" type="distribution">
<paragraph id="4867615A7659FFF6FF66FEE03A04FE39" blockId="14.[189,344,376,402]" box="[189,344,376,402]" pageId="14" pageNumber="218">
<heading id="132FD6367659FFF6FF66FEE03A04FE39" bold="true" box="[189,344,376,402]" fontSize="11" level="3" pageId="14" pageNumber="218" reason="3">
<emphasis id="7AACBD487659FFF6FF66FEE03A04FE39" bold="true" box="[189,344,376,402]" pageId="14" pageNumber="218">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="4867615A7659FFF6FF66FE3F3A06FE4F" blockId="14.[189,1399,423,484]" pageId="14" pageNumber="218">
Widespread distribution is known from one locality in
<collectingRegion id="8A1CAFB87659FFF6FC87FE3F3F4DFE6A" box="[860,1041,423,449]" country="Peru" name="Madre de Dios" pageId="14" pageNumber="218">Madre de Dios</collectingRegion>
and
<collectingRegion id="8A1CAFB87659FFF6FB89FE3F3FC7FE6A" box="[1106,1179,423,449]" country="Peru" name="Cusco" pageId="14" pageNumber="218">Cusco</collectingRegion>
departments each (
<figureCitation id="D0E37DDF7659FFF6FF1EFE523A11FE4F" box="[197,333,458,484]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="16.[189,232,1998,2024]" captionStartId-1="17.[189,232,1998,2024]" captionTargetBox-0="[189,1392,909,1953]" captionTargetBox-1="[189,1392,1113,1959]" captionTargetId-0="figure-119@16.[189,1398,888,1959]" captionTargetId-1="figure-249@17.[189,1398,1092,1959]" captionTargetPageId-0="16" captionTargetPageId-1="17" captionText-0="Fig. 11. Distribution map of species of Caponiidae Simon, 1890 recorded from Peru." captionText-1="Fig. 12. Distribution map of the genus Nyetnops Platnick &amp; Lise, 2007." figureDoi-0="http://doi.org/10.5281/zenodo.11145133" figureDoi-1="http://doi.org/10.5281/zenodo.11145135" httpUri-0="https://zenodo.org/record/11145133/files/figure.png" httpUri-1="https://zenodo.org/record/11145135/files/figure.png" pageId="14" pageNumber="218">Figs 11–12</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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