426 lines
51 KiB
XML
426 lines
51 KiB
XML
<document ID-CLB-Dataset="20172" ID-DOI="http://dx.doi.org/10.3897/subtbiol.42.79190" ID-GBIF-Dataset="395bc8e5-add1-4fa2-9511-01d66d2f587a" ID-Pensoft-Pub="1314-2615-42-125" ID-Pensoft-UUID="2C89BCC134285666A0E0C110A5F51949" ID-ZooBank="C9CD999BE57147F7830F21081B0E71A6" ModsDocID="1314-2615-42-125" checkinTime="1647696006388" checkinUser="pensoft" docAuthor="Prado 1, Guilherme C., Preez 2, Gerhard C. Du & Ferreira 1, Rodrigo L." docDate="2022" docId="9AE27A8763E8152D3BB79ADAA508B3C6" docLanguage="en" docName="SubterBiol 42: 125-138" docOrigin="Subterranean Biology 42" docPubDate="2022-03-18" docSource="http://dx.doi.org/10.3897/subtbiol.42.79190" docTitle="Selachochthonius naledi Prado 1 & Preez 2 & Ferreira 1 2022, sp. nov." docType="treatment" docUuid="8BD115F5-1B22-4EC7-919F-92007DCD91D1" docUuidSource="ZooBank" docVersion="5" id="2C89BCC134285666A0E0C110A5F51949" lastPageNumber="125" masterDocId="2C89BCC134285666A0E0C110A5F51949" masterDocTitle="Selachochthonius naledi sp. nov. (Pseudoscorpiones, Pseudotyrannochthoniidae), a new troglobitic species from South Africa" masterLastPageNumber="138" masterPageNumber="125" pageNumber="125" updateTime="1732860181774" updateUser="ExternalLinkService">
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<mods:mods id="C8639A4F97DC362891D724A1E33DE1AA" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="F51900080C6E37C0E2786DE3C28362C8">
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<mods:title id="ADA35804E994F6BCE7079F43AC1AF32F">Selachochthonius naledi sp. nov. (Pseudoscorpiones, Pseudotyrannochthoniidae), a new troglobitic species from South Africa</mods:title>
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<mods:name id="81AE858205630565FEB197185C46DF0E" type="personal">
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<mods:role id="C5DAA4E35D1919A544776C2AFD7A7B7C">
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<mods:roleTerm id="D6893A5A5D5C59B40F281A964BAAD6AA">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="49A9388963947664D9DD10C8A701CF1F">Prado 1, Guilherme C.</mods:namePart>
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<mods:affiliation id="FD589A14E4315AE7555BC5CD2568C4F2">Centro de Estudos em Biologia Subterranea, Departamento de Ecologia e Conservacao, Universidade Federal de Lavras, Lavras, MG. CEP 37200 - 900, Brazil</mods:affiliation>
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</mods:name>
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<mods:name id="205CE7578307A2274611C43C67A3C42C" type="personal">
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<mods:role id="28B827DC340D31F0DD94DF7F13FAFBC7">
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<mods:roleTerm id="2BFC6D402C1C549471C89B375DAC9101">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="33FC5B8A028D69C7CF1226F9ED9762F1">Preez 2, Gerhard C. Du</mods:namePart>
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<mods:affiliation id="FA14D144E5C9CFD4E657B2C0D5E91E8B">Centro de Estudos em Biologia Subterranea, Departamento de Ecologia e Conservacao, Universidade Federal de Lavras, Lavras, MG. CEP 37200 - 900, Brazil</mods:affiliation>
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</mods:name>
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<mods:name id="EA2E17C72E2BF67542954AA6F7B9178F" type="personal">
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<mods:roleTerm id="49403A5C45065CEC246B7710E541F47D">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="3AD810109CF16D27C659823A84F53A79">Ferreira 1, Rodrigo L.</mods:namePart>
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<mods:affiliation id="367417A481F3B497E6E6DF3FDC294CC8">Centro de Estudos em Biologia Subterranea, Departamento de Ecologia e Conservacao, Universidade Federal de Lavras, Lavras, MG. CEP 37200 - 900, Brazil</mods:affiliation>
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</mods:name>
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<mods:typeOfResource id="3EDBDC2A563E5246DF5BE36BFD16412D">text</mods:typeOfResource>
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<mods:titleInfo id="1A75EC6E97DE7D437D70B57201E0D65C">
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<mods:title id="0FA4BF150BCA778BBF499B8EBCC6593E">Subterranean Biology</mods:title>
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</mods:titleInfo>
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<mods:part id="92CF576590EDC5E0F22DDFE4B947B2DC">
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<mods:date id="B71FC9A51060191D28A7391C9358CA8C">2022</mods:date>
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<mods:detail id="97F290E06F153B41FB23F40E772D218B" type="pubDate">
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<mods:number id="B4F1049230DA1FFD01B817CBDD7A1475">2022-03-18</mods:number>
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<mods:detail id="027655E20EFF5ED5883B6BF31210F628" type="volume">
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<mods:number id="FD021C7DB77A73A2AE05A779ED6B1C04">42</mods:number>
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</mods:detail>
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<mods:start id="06319B2D16A96C9CF788BE00A1DD2F5F">125</mods:start>
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<mods:end id="54D1C1BF622A3B1CF15E25F0205C9F9E">138</mods:end>
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<mods:location id="6329ACF9CE590683004E8D500CB81A5B">
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<mods:url id="93AB4B26BD46B80B37F9C54378F46D9D">http://dx.doi.org/10.3897/subtbiol.42.79190</mods:url>
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</mods:location>
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<mods:classification id="74665956AFB9B3B22C989AF0896CB506">journal article</mods:classification>
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<mods:identifier id="37391A8D808AD1532D4CFA0B28CF133A" type="DOI">http://dx.doi.org/10.3897/subtbiol.42.79190</mods:identifier>
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<mods:identifier id="E8703F8FBC44CBBD0A66D9E34FD6753E" type="Pensoft-Pub">1314-2615-42-125</mods:identifier>
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<mods:identifier id="1BCC579DF530ECC656614F45685645F8" type="ZooBank">C9CD999BE57147F7830F21081B0E71A6</mods:identifier>
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<mods:identifier id="FF4A28981D1CCDB37B7A47F3D0C1CC47" type="Pensoft-UUID">2C89BCC134285666A0E0C110A5F51949</mods:identifier>
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</mods:mods>
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<treatment id="9AE27A8763E8152D3BB79ADAA508B3C6" ID-GBIF-Taxon="194176178" LSID="urn:lsid:zoobank.org:act:8BD115F5-1B22-4EC7-919F-92007DCD91D1" httpUri="http://treatment.plazi.org/id/9AE27A8763E8152D3BB79ADAA508B3C6" lastPageNumber="125" pageId="0" pageNumber="125" scope_family="Pseudotyrannochthoniidae" scope_order="Pseudoscorpiones">
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<subSubSection id="12A3582D4B640EFE1F132F24AFC3E2CA" pageId="0" pageNumber="125" type="nomenclature">
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<paragraph id="D60A056628C4B68720EF23271C0B5316" pageId="0" pageNumber="125">
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<taxonomicName id="60E70858E56172A4D8261119B7BFE40B" LSID="http://zoobank.org/8BD115F5-1B22-4EC7-919F-92007DCD91D1" authority="Prado 1 & Preez 2 & Ferreira 1, 2022" authorityName="Prado 1 & Preez 2 & Ferreira 1" authorityYear="2022" class="Arachnida" family="Pseudotyrannochthoniidae" genus="Selachochthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Selachochthonius naledi" order="Pseudoscorpiones" pageId="0" pageNumber="125" phylum="Arthropoda" rank="species" species="naledi" status="sp. nov.">Selachochthonius naledi</taxonomicName>
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<taxonomicNameLabel id="799DD5D9C3F8832993B576CC1A0CA190" pageId="0" pageNumber="125">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="9DA5B8A532C788B070ACDB81BA986644" pageId="0" pageNumber="125" type="materials_examined">
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<paragraph id="203CC55532E89E94BA4858D56C07A8F9" pageId="0" pageNumber="125">Material examined.</paragraph>
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<paragraph id="6B1ED2EAAEDC07CB8ACB2DBF1F0348E9" pageId="0" pageNumber="125">
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<materialsCitation id="98584435B1042C7BA9290D989B583A2B" collectingDate="2019-10-09" collectorName="R. L Ferreira." country="South Africa" county="Villa Louisa cave" latitude="-26.02375" location="Cradle of Humankind" longLatPrecision="1" longitude="27.711945" municipality="Maquassi Hills Municipality" specimenCount="1" typeStatus="Holotype">
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<emphasis id="770A8CBE804DCA7B5B1EBF0FBB2A13FB" bold="true" italics="true" pageId="0" pageNumber="125">
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<typeStatus id="97747C5A5F63D5052E42F5DB18D288AB">Holotype</typeStatus>
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</emphasis>
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female (NCA 2021/1351), preserved in ethanol:
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<collectingCountry id="EC5AFEAD9434762FAFCF7B716F41F32E" name="South Africa">South Africa</collectingCountry>
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,
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<location id="2871798B9E31EF6C36E40E9AF5642819" LSID="urn:lsid:plazi:treatment:9AE27A8763E8152D3BB79ADAA508B3C6:2871798B9E31EF6C36E40E9AF5642819" country="South Africa" county="Villa Louisa cave" latitude="-26.02375" longLatPrecision="1" longitude="27.711945" municipality="Maquassi Hills Municipality" name="Cradle of Humankind">Cradle of Humankind</location>
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(UNESCO WHS),
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<collectingMunicipality id="9D6349E9A672D22632D1736A764A5E14">Maquassi Hills Municipality</collectingMunicipality>
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,
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<collectingCounty id="7CBE745B5732B2C0AC5F275992D29A9A">Villa Louisa cave</collectingCounty>
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(
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<geoCoordinate id="2A25E4A886A3042B35539F18E3317582" degrees="26" direction="south" minutes="01" orientation="latitude" precision="1" seconds="25.5" value="-26.02375">26°01'25.5"S</geoCoordinate>
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,
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<geoCoordinate id="50D9D6E39A29FF227604DF2AA53F63F4" degrees="27" direction="east" minutes="42" orientation="longitude" precision="1" seconds="43.0" value="27.711945">27°42'43.0"E</geoCoordinate>
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),
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<collectingDate id="2E34E7AB2F531A275CE1282758810E03" value="2019-10-09">09 October 2019</collectingDate>
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, leg.
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<collectorName id="773B138505AEFD4C3910F854F91D06E7">R.L Ferreira.</collectorName>
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</materialsCitation>
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<materialsCitation id="B83901DD2A646DD5031CDC631C811592" collectingDate="2019-10-17" collectorName="R. L Ferreira." country="South Africa" county="Yom Tov Cave" latitude="-26.020811" location="Cradle of Humankind" longLatPrecision="1" longitude="27.714851" municipality="Maquassi Hills Municipality" specimenCount="1" typeStatus="Paratype">
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<emphasis id="7D8029F56F1B8563D0F3ECD805DCD7EF" bold="true" italics="true" pageId="0" pageNumber="125">
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<typeStatus id="7D825859EDACD03D061A284934179333">Paratype</typeStatus>
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</emphasis>
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male (NCA 2021/1352),
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<location id="0DF6D00013DCA41C5215189ED10F940C" LSID="urn:lsid:plazi:treatment:9AE27A8763E8152D3BB79ADAA508B3C6:0DF6D00013DCA41C5215189ED10F940C" country="South Africa" county="Yom Tov Cave" latitude="-26.020811" longLatPrecision="1" longitude="27.714851" municipality="Maquassi Hills Municipality" name="Cradle of Humankind">Cradle of Humankind</location>
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,
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<collectingMunicipality id="5A4E3DB997674B3A3D52BD7A0B121AF7">Maquassi Hills Municipality</collectingMunicipality>
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,
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<collectingCounty id="0B3F6393E019EE300338770702671D7B">Yom Tov Cave</collectingCounty>
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(
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<geoCoordinate id="5C2BD68CF8B0B499828C18631CCE6784" degrees="26" direction="south" minutes="01" orientation="latitude" precision="1" seconds="14.92" value="-26.020811">26°01'14.92"S</geoCoordinate>
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,
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<geoCoordinate id="80ABEABBA2E335070ACD5359013C7905" degrees="27" direction="east" minutes="42" orientation="longitude" precision="1" seconds="53.46" value="27.714851">27°42'53.46"E</geoCoordinate>
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),
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<collectingDate id="7B1F7E8C72D26DB95972311E79C450E6" value="2019-10-17">17 October 2019</collectingDate>
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, leg.
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<collectorName id="9820F65FA1D39AA7419F475AF3AEF833">R.L Ferreira.</collectorName>
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="8A96F777084AE540234085CEF0B50B77" pageId="0" pageNumber="125" type="etymology">
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<paragraph id="4982EC7B93C74DDD0B2866DF9A4AADA1" pageId="0" pageNumber="125">Etymology.</paragraph>
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<paragraph id="F0430485E923DD0AA63F2DD7B9CD86D6" pageId="0" pageNumber="125">
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The epithet
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<emphasis id="793D11DA211BFC0D0140D7F41F537BD9" italics="true" pageId="0" pageNumber="125">naledi</emphasis>
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refers to
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<taxonomicName id="2647FC117022358967BE30FAA087544F" class="Arachnida" family="Hominidae" genus="Homo" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Homo naledi" order="Primates" pageId="0" pageNumber="125" phylum="Arthropoda" rank="species" species="naledi">
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<emphasis id="36AA942090D874EC96B00644EE07DB22" italics="true" pageId="0" pageNumber="125">Homo naledi</emphasis>
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</taxonomicName>
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, an extinct hominid species discovered within the Dinaledi chamber. This chamber is located in the Rising Star Cave (Fig.
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<figureCitation id="6AEF3F886BF544C30F8665009B80712B" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Distribution map of Selachochthonius species in South Africa A Africa continent, detail on South Africa country B South Africa, exhibiting distribution of Selachochthonius species with detail on S. naledi sp. nov. type localities C Selachochthonius naledi sp. nov. distribution area, exhibiting Yom Tov, Villa Louise and Rising Star Caves (Homo naledi type locality), detail on quarry activity occurring nearby." figureDoi="10.3897/subtbiol.42.79190.figure1" httpUri="https://binary.pensoft.net/fig/660669" pageId="0" pageNumber="125">1C</figureCitation>
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), a World Heritage site located approximately 50 km from Johannesburg. The caves where the pseudoscorpions were found are located on the same hill as Rising Star Cave. The distances between the entrance of Rising Star Cave and the entrances of Vila Louise and Yom Tov caves are approximately 425 and 220 meters, respectively. Considering the fissure system associated with the carbonatic rock in the area, it is likely that the pseudoscorpions are able to disperse through small fissures to other macrocaves in the hill, including Rising Star Cave.
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</paragraph>
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</subSubSection>
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<subSubSection id="3CFCC1CE77D046DCB9340C2F5A15C4D2" pageId="0" pageNumber="125" type="diagnosis">
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<paragraph id="ADBD4F5817505513143B5FC531A8A1EB" pageId="0" pageNumber="125">Diagnosis.</paragraph>
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<paragraph id="13572115306FCCCA29CE2A369BBB2545" pageId="0" pageNumber="125">
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<taxonomicName id="D5BEF6879B8A263C6F36B747BB93FCF6" authorityName="Prado 1 & Preez 2 & Ferreira 1" authorityYear="2022" class="Arachnida" family="Pseudotyrannochthoniidae" genus="Selachochthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Selachochthonius naledi" order="Pseudoscorpionidea" pageId="0" pageNumber="125" phylum="Arthropoda" rank="species" species="naledi">
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<emphasis id="CF615F394A8603CD28BD3DB6642DD423" italics="true" pageId="0" pageNumber="125">Selachochthonius naledi</emphasis>
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</taxonomicName>
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sp. nov. differs from other members of the genus by the following combination of characters: absence of eyes or eyespots (
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<taxonomicName id="BD781007E6CF22C7389EB5A0F39112E3" lsidName="S. heterodentatus" pageId="0" pageNumber="125" rank="species" species="heterodentatus">
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<emphasis id="D5FF37EF33B45460F5238D06B4C06F62" italics="true" pageId="0" pageNumber="125">S. heterodentatus</emphasis>
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</taxonomicName>
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Beier, 1995 with four well-developed eyes and
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<taxonomicName id="44CAD34CDE98741FCE768B0EC25E4589" lsidName="S. serratidentatus" pageId="0" pageNumber="125" rank="species" species="serratidentatus">
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<emphasis id="87679A8EC75882ADB422F313B3206837" italics="true" pageId="0" pageNumber="125">S. serratidentatus</emphasis>
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</taxonomicName>
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Ellingsen, 1912 bearing four small eyes) (
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<bibRefCitation id="487C9621A17548480F7C4B1D26F8D00D" DOI="https://doi.org/10.5962/bhl.part.9312" author="Ellingsen, E" journalOrPublisher="Annals of the South African Museum" pageId="0" pageNumber="125" pagination="75 - 128" refId="B12" refString="Ellingsen, E, 1912. The pseudoscorpions of South Africa, based on the collections of the South African Museum, Cape Town. Annals of the South African Museum 10: 75 - 128, DOI: https://doi.org/10.5962/bhl.part.9312" title="The pseudoscorpions of South Africa, based on the collections of the South African Museum, Cape Town." url="https://doi.org/10.5962/bhl.part.9312" volume="10" year="1912">Ellingsen 1912</bibRefCitation>
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;
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<bibRefCitation id="57E68BBE2108253467FBECAF22DD8A08" author="Beier, M" editor="Hanstrom, B" journalOrPublisher="Results of the Lund Expedition in 1950 - 1951. Almquist and Wiksell, Stockholm" pageId="0" pageNumber="125" pagination="1263 - 1328" refId="B3" refString="Beier, M, 1955. Pseudoscorpionidea. In: Hanstrom, B, Brinck, P, Rudebeck, G, Eds., South African animal life. Results of the Lund Expedition in 1950-1951. Almquist and Wiksell, Stockholm: 1263 - 1328" title="Pseudoscorpionidea." volumeTitle="South African animal life." year="1955">Beier 1955</bibRefCitation>
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); epistome triangular and strongly dentate (
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<taxonomicName id="D44E5B5AFA932335CC209582D74BC374" lsidName="S. cavernicola" pageId="0" pageNumber="125" rank="species" species="cavernicola">
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<emphasis id="9BC7C7F58F5FE056FE96285C1A6DC89E" italics="true" pageId="0" pageNumber="125">S. cavernicola</emphasis>
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</taxonomicName>
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Lawrence, 1935 presents a flattened apex and slightly dentated epistome) (
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<bibRefCitation id="CB247680C9B8CD67AEA067DE7D1BADB7" DOI="https://dx.doi.org/10.1080/00222933508654998" author="Lawrence, RF" journalOrPublisher="Annals and Magazine of Natural History" pageId="0" pageNumber="125" pagination="549 - 555" refId="B19" refString="Lawrence, RF, 1935. A cavernicolous false scorpion from Table Mountain, Cape Town. Annals and Magazine of Natural History 10 (15): 549 - 555, DOI: https://dx.doi.org/10.1080/00222933508654998" title="A cavernicolous false scorpion from Table Mountain, Cape Town." url="https://dx.doi.org/10.1080/00222933508654998" volume="10" year="1935">Lawrence 1935</bibRefCitation>
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); palpal femur with length of 0.65-0.66 mm or 6.2 times longer than wide (4.0 times in
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<taxonomicName id="CB5219E0C18D92DB87944665810F0700" lsidName="S. serratidentatus" pageId="0" pageNumber="125" rank="species" species="serratidentatus">
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<emphasis id="CF1F2871A8E2DF9C6823A63289FA1223" italics="true" pageId="0" pageNumber="125">S. serratidentatus</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="B4DD7EA3794C43C2B56AC224FAA96F06" lsidName="S. heterodentatus" pageId="0" pageNumber="125" rank="species" species="heterodentatus">
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<emphasis id="1AB1381B115214E4FFCB02661E1C0C06" italics="true" pageId="0" pageNumber="125">S. heterodentatus</emphasis>
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</taxonomicName>
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with femur length of 1.0 mm); chela 6.1-6.2 times longer than wide (3.8 times in
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<taxonomicName id="57EABC7C08F2C5E9DC4D631C3897E733" lsidName="S. serratidentatus" pageId="0" pageNumber="125" rank="species" species="serratidentatus">
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<emphasis id="D3C3DC5161256334F884ECAEA17F5139" italics="true" pageId="0" pageNumber="125">S. serratidentatus</emphasis>
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</taxonomicName>
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and 5.0 times in
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<taxonomicName id="862C35D24FEE31E6131FAD297EF24500" lsidName="S. heterodentatus" pageId="0" pageNumber="125" rank="species" species="heterodentatus">
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<emphasis id="DB7D47DC19C943697357549AA68F0748" italics="true" pageId="0" pageNumber="125">S. heterodentatus</emphasis>
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</taxonomicName>
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) (
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<bibRefCitation id="E8D80111060A5968D26A7B181AB3C616" DOI="https://doi.org/10.5962/bhl.part.9312" author="Ellingsen, E" journalOrPublisher="Annals of the South African Museum" pageId="0" pageNumber="125" pagination="75 - 128" refId="B12" refString="Ellingsen, E, 1912. The pseudoscorpions of South Africa, based on the collections of the South African Museum, Cape Town. Annals of the South African Museum 10: 75 - 128, DOI: https://doi.org/10.5962/bhl.part.9312" title="The pseudoscorpions of South Africa, based on the collections of the South African Museum, Cape Town." url="https://doi.org/10.5962/bhl.part.9312" volume="10" year="1912">Ellingsen 1912</bibRefCitation>
|
|
;
|
|
<bibRefCitation id="7C66200F3382B3F18AA8E5E32C089800" author="Beier, M" editor="Hanstrom, B" journalOrPublisher="Results of the Lund Expedition in 1950 - 1951. Almquist and Wiksell, Stockholm" pageId="0" pageNumber="125" pagination="1263 - 1328" refId="B3" refString="Beier, M, 1955. Pseudoscorpionidea. In: Hanstrom, B, Brinck, P, Rudebeck, G, Eds., South African animal life. Results of the Lund Expedition in 1950-1951. Almquist and Wiksell, Stockholm: 1263 - 1328" title="Pseudoscorpionidea." volumeTitle="South African animal life." year="1955">Beier 1955</bibRefCitation>
|
|
); fixed finger with 32-34 simple, sparse, acute and triangular teeth (
|
|
<taxonomicName id="AF4B3607837519FA98B8F988C1B03B4A" lsidName="S. cavernicola" pageId="0" pageNumber="125" rank="species" species="cavernicola">
|
|
<emphasis id="2092C5F14B56DF76B060D5AD6C1FB11F" italics="true" pageId="0" pageNumber="125">S. cavernicola</emphasis>
|
|
</taxonomicName>
|
|
with 23 sparse, simple and triangular teeth, presented in only 2/3 of the segment;
|
|
<taxonomicName id="10047C91F4368226D18B93AC36DF752C" lsidName="S. heterodentatus" pageId="0" pageNumber="125" rank="species" species="heterodentatus">
|
|
<emphasis id="9DBEA0DD743E85FDFE7442081BCCA0FD" italics="true" pageId="0" pageNumber="125">S. heterodentatus</emphasis>
|
|
</taxonomicName>
|
|
with 28 acute teeth intercalated by 24 smaller teeth and
|
|
<taxonomicName id="28AC70241D6B2AA973B39247F46CA226" lsidName="S. serratidentatus" pageId="0" pageNumber="125" rank="species" species="serratidentatus">
|
|
<emphasis id="550D4884085AAB5104228A81FD5D563C" italics="true" pageId="0" pageNumber="125">S. serratidentatus</emphasis>
|
|
</taxonomicName>
|
|
with sparse, acute and complex teeth, each one intercalated by one or two smaller teeth) (
|
|
<bibRefCitation id="7AA25045FF970E9B39C61F3B7CCA1A56" DOI="https://doi.org/10.5962/bhl.part.9312" author="Ellingsen, E" journalOrPublisher="Annals of the South African Museum" pageId="0" pageNumber="125" pagination="75 - 128" refId="B12" refString="Ellingsen, E, 1912. The pseudoscorpions of South Africa, based on the collections of the South African Museum, Cape Town. Annals of the South African Museum 10: 75 - 128, DOI: https://doi.org/10.5962/bhl.part.9312" title="The pseudoscorpions of South Africa, based on the collections of the South African Museum, Cape Town." url="https://doi.org/10.5962/bhl.part.9312" volume="10" year="1912">Ellingsen 1912</bibRefCitation>
|
|
;
|
|
<bibRefCitation id="1C4635AD2E89E1334F25854A9D840020" DOI="https://dx.doi.org/10.1080/00222933508654998" author="Lawrence, RF" journalOrPublisher="Annals and Magazine of Natural History" pageId="0" pageNumber="125" pagination="549 - 555" refId="B19" refString="Lawrence, RF, 1935. A cavernicolous false scorpion from Table Mountain, Cape Town. Annals and Magazine of Natural History 10 (15): 549 - 555, DOI: https://dx.doi.org/10.1080/00222933508654998" title="A cavernicolous false scorpion from Table Mountain, Cape Town." url="https://dx.doi.org/10.1080/00222933508654998" volume="10" year="1935">Lawrence 1935</bibRefCitation>
|
|
;
|
|
<bibRefCitation id="DED415A091EB3EED73991072B3816FA0" author="Beier, M" editor="Hanstrom, B" journalOrPublisher="Results of the Lund Expedition in 1950 - 1951. Almquist and Wiksell, Stockholm" pageId="0" pageNumber="125" pagination="1263 - 1328" refId="B3" refString="Beier, M, 1955. Pseudoscorpionidea. In: Hanstrom, B, Brinck, P, Rudebeck, G, Eds., South African animal life. Results of the Lund Expedition in 1950-1951. Almquist and Wiksell, Stockholm: 1263 - 1328" title="Pseudoscorpionidea." volumeTitle="South African animal life." year="1955">Beier 1955</bibRefCitation>
|
|
); movable finger with 29-31 simple, sparse acute and triangular teeth (
|
|
<taxonomicName id="2C086D309F0499376D022D293FD44E18" lsidName="S. cavernicola" pageId="0" pageNumber="125" rank="species" species="cavernicola">
|
|
<emphasis id="46B75A157985D5D99059A58E1A982E44" italics="true" pageId="0" pageNumber="125">S. cavernicola</emphasis>
|
|
</taxonomicName>
|
|
with 17 simple, sparse, acute and triangular teeth;
|
|
<taxonomicName id="9B11C43FE37EB87C77C255E749D1560E" lsidName="S. heterodentatus" pageId="0" pageNumber="125" rank="species" species="heterodentatus">
|
|
<emphasis id="7397C77C39CD3DD6C699D61CFEB537F4" italics="true" pageId="0" pageNumber="125">S. heterodentatus</emphasis>
|
|
</taxonomicName>
|
|
with 16 acute and triangular, intercalated by 12 smaller teeth) (
|
|
<bibRefCitation id="36B42A03BF659D6342870ADAF37227A0" DOI="https://dx.doi.org/10.1080/00222933508654998" author="Lawrence, RF" journalOrPublisher="Annals and Magazine of Natural History" pageId="0" pageNumber="125" pagination="549 - 555" refId="B19" refString="Lawrence, RF, 1935. A cavernicolous false scorpion from Table Mountain, Cape Town. Annals and Magazine of Natural History 10 (15): 549 - 555, DOI: https://dx.doi.org/10.1080/00222933508654998" title="A cavernicolous false scorpion from Table Mountain, Cape Town." url="https://dx.doi.org/10.1080/00222933508654998" volume="10" year="1935">Lawrence 1935</bibRefCitation>
|
|
;
|
|
<bibRefCitation id="51BFDD8E2D646488A03B5C97708503EC" author="Beier, M" editor="Hanstrom, B" journalOrPublisher="Results of the Lund Expedition in 1950 - 1951. Almquist and Wiksell, Stockholm" pageId="0" pageNumber="125" pagination="1263 - 1328" refId="B3" refString="Beier, M, 1955. Pseudoscorpionidea. In: Hanstrom, B, Brinck, P, Rudebeck, G, Eds., South African animal life. Results of the Lund Expedition in 1950-1951. Almquist and Wiksell, Stockholm: 1263 - 1328" title="Pseudoscorpionidea." volumeTitle="South African animal life." year="1955">Beier 1955</bibRefCitation>
|
|
); presence of a small tubercle in both chelae between the 13th and 14th teeth of the female movable finger and between the 10th and 11th teeth of the male movable finger (
|
|
<taxonomicName id="EC99C296F72FD396EDFA51AD14CF4F27" lsidName="S. cavernicola" pageId="0" pageNumber="125" rank="species" species="cavernicola">
|
|
<emphasis id="A28878BD90FDB0809A0109D9B7401FED" italics="true" pageId="0" pageNumber="125">S. cavernicola</emphasis>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName id="DB7A517293781978B3B284F4E4CF6689" lsidName="S. heterodentatus" pageId="0" pageNumber="125" rank="species" species="heterodentatus">
|
|
<emphasis id="12CA40F81068E2207AA56C89F40C2851" italics="true" pageId="0" pageNumber="125">S. heterodentatus</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="1F53531BE24234A3D6F1753AFB891CDD" lsidName="S. serratidentatus" pageId="0" pageNumber="125" rank="species" species="serratidentatus">
|
|
<emphasis id="DE826FEAE964295D8D3B6A7A5CBAFD49" italics="true" pageId="0" pageNumber="125">S. serratidentatus</emphasis>
|
|
</taxonomicName>
|
|
lacks tubercle); 7-11 coxal spines tripinnate arranged in a single row on the anterior portion of the coxa I (
|
|
<taxonomicName id="8143834195C0510FAFF1EC56FB9C53B9" lsidName="S. cavernicola" pageId="0" pageNumber="125" rank="species" species="cavernicola">
|
|
<emphasis id="5E64EF4743C4E40551B0D8FB6A5B1602" italics="true" pageId="0" pageNumber="125">S. cavernicola</emphasis>
|
|
</taxonomicName>
|
|
with 12 bipinnate coxal spines) (
|
|
<bibRefCitation id="99BDB96730BE149238AA25C71F8C1C58" DOI="https://dx.doi.org/10.1080/00222933508654998" author="Lawrence, RF" journalOrPublisher="Annals and Magazine of Natural History" pageId="0" pageNumber="125" pagination="549 - 555" refId="B19" refString="Lawrence, RF, 1935. A cavernicolous false scorpion from Table Mountain, Cape Town. Annals and Magazine of Natural History 10 (15): 549 - 555, DOI: https://dx.doi.org/10.1080/00222933508654998" title="A cavernicolous false scorpion from Table Mountain, Cape Town." url="https://dx.doi.org/10.1080/00222933508654998" volume="10" year="1935">Lawrence 1935</bibRefCitation>
|
|
).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="516CE91F74AA68E5A69FBF587A25471A" pageId="0" pageNumber="125" type="description">
|
|
<paragraph id="4D2C97D4305EFD800BD56C3314FBABB5" pageId="0" pageNumber="125">Description.</paragraph>
|
|
<paragraph id="FB2485F926D21FBF41146466950512D4" pageId="0" pageNumber="125">
|
|
(Fig.
|
|
<figureCitation id="018E12BFBFD5B92BAECA6E203A8E6A87" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Type locality and habitat of Selachochthonius naledi sp. nov. A Villa Louise cave entrance B Villa Louise cave interior general aspect C Villa Louise cave interior, showing altered floor D Yom Tov cave entrance E live female holotype F live male paratype." figureDoi="10.3897/subtbiol.42.79190.figure5" httpUri="https://binary.pensoft.net/fig/660673" pageId="0" pageNumber="125">5E-F</figureCitation>
|
|
). Body pale yellowish, mostly translucent; chelicerae slightly reddish orange, abdomen beige. Some parts of the body scaly. Vestitural setae sharp and anteriorly projected.
|
|
</paragraph>
|
|
<paragraph id="A45A062F5B6CB2CBA41DEEE4A10C564A" pageId="0" pageNumber="125">
|
|
<emphasis id="4736D82D18C9BBB684F0FBFFCEC0E923" bold="true" pageId="0" pageNumber="125">
|
|
<emphasis id="00992967BB65716C2B8934D633C4A703" italics="true" pageId="0" pageNumber="125">Carapace</emphasis>
|
|
.
|
|
</emphasis>
|
|
(Fig.
|
|
<figureCitation id="64B2085874D36FB7F0F32AE4D854A5D1" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Selachochthonius naledi sp. nov. A left chelicera of female holotype, antiaxial view B female holotype carapace, showing chaetotaxy of carapace C detailed anterior margin of female paratype carapace D detailed anterior margin of male paratype carapace E leg IV of female holotype, showing tubercle location. Scale bars: 0.25 mm (A, C-D); 0.5 mm (B, E)." figureDoi="10.3897/subtbiol.42.79190.figure2" httpUri="https://binary.pensoft.net/fig/660670" pageId="0" pageNumber="125">2B-D</figureCitation>
|
|
). Ratio length/width near 1.0, strongly constricted posteriorly showing a difference between ocular width and posterior width of 0.08-0.10 mm; anterior margin smooth; absence of eyes or eyespots; epistome strongly dentate and saw-like; posterior margin of carapace smooth; chaetotaxy 6: 4: 4: 2: 2 (18), lateral pre ocular setae are shorter in female (Fig.
|
|
<figureCitation id="E960E9E9BE5CFD2B6D46FC8B625B89CD" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Selachochthonius naledi sp. nov. A left chelicera of female holotype, antiaxial view B female holotype carapace, showing chaetotaxy of carapace C detailed anterior margin of female paratype carapace D detailed anterior margin of male paratype carapace E leg IV of female holotype, showing tubercle location. Scale bars: 0.25 mm (A, C-D); 0.5 mm (B, E)." figureDoi="10.3897/subtbiol.42.79190.figure2" httpUri="https://binary.pensoft.net/fig/660670" pageId="0" pageNumber="125">2C</figureCitation>
|
|
).
|
|
</paragraph>
|
|
<caption id="80B048C282BD6A8A19F8952D922CAE45" doi="10.3897/subtbiol.42.79190.figure2" httpUri="https://binary.pensoft.net/fig/660670" pageId="0" pageNumber="125" start="Figure 2" startId="F2">
|
|
<paragraph id="8691F269197AE08A27669109CAA12434" pageId="0" pageNumber="125">
|
|
<emphasis id="9CFFB34A9EB6E6E12987F1AC0898F62A" bold="true" pageId="0" pageNumber="125">Figure 2.</emphasis>
|
|
<taxonomicName id="284E638418F4CC7D3925D53E7F92FB23" authorityName="Prado 1 & Preez 2 & Ferreira 1" authorityYear="2022" class="Arachnida" family="Pseudotyrannochthoniidae" genus="Selachochthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Selachochthonius naledi" order="Pseudoscorpionidea" pageId="0" pageNumber="125" phylum="Arthropoda" rank="species" species="naledi">
|
|
<emphasis id="59FFAA7C2CE97684662CF526BC8D564D" italics="true" pageId="0" pageNumber="125">Selachochthonius naledi</emphasis>
|
|
</taxonomicName>
|
|
sp. nov.
|
|
<emphasis id="C242C28755CD62AB3E9EB0308FE11C80" bold="true" pageId="0" pageNumber="125">A</emphasis>
|
|
left chelicera of female holotype, antiaxial view
|
|
<emphasis id="0B098326693992C69A5B504ED14600BC" bold="true" pageId="0" pageNumber="125">B</emphasis>
|
|
female holotype carapace, showing chaetotaxy of carapace
|
|
<emphasis id="CD8B21EB697B6EBA8765A378734C486E" bold="true" pageId="0" pageNumber="125">C</emphasis>
|
|
detailed anterior margin of female paratype carapace
|
|
<emphasis id="4915C1CBB5D566D6798D9CE42CC39C4A" bold="true" pageId="0" pageNumber="125">D</emphasis>
|
|
detailed anterior margin of male paratype carapace
|
|
<emphasis id="2224D3D10C413D1D07A7C41E84ADE613" bold="true" pageId="0" pageNumber="125">E</emphasis>
|
|
leg IV of female holotype, showing tubercle location. Scale bars: 0.25 mm (
|
|
<emphasis id="7A8F239581415ADB493ECD42AEC36C86" bold="true" pageId="0" pageNumber="125">A, C-D</emphasis>
|
|
); 0.5 mm (
|
|
<emphasis id="56DE6F387EDE8A96461A4EAFB284C32B" bold="true" pageId="0" pageNumber="125">B, E</emphasis>
|
|
).
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph id="DBA1B926959950308E9B2C0D609B5A68" pageId="0" pageNumber="125">
|
|
<emphasis id="A7F24F0ACE2DC8423214AFBC82AEDF96" bold="true" pageId="0" pageNumber="125">
|
|
<emphasis id="547DB6B157644EFBEECB951761F227EC" italics="true" pageId="0" pageNumber="125">Chelicera</emphasis>
|
|
.
|
|
</emphasis>
|
|
(Fig.
|
|
<figureCitation id="E8B64EE7DF652E08A53F77063907147B" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Selachochthonius naledi sp. nov. A left chelicera of female holotype, antiaxial view B female holotype carapace, showing chaetotaxy of carapace C detailed anterior margin of female paratype carapace D detailed anterior margin of male paratype carapace E leg IV of female holotype, showing tubercle location. Scale bars: 0.25 mm (A, C-D); 0.5 mm (B, E)." figureDoi="10.3897/subtbiol.42.79190.figure2" httpUri="https://binary.pensoft.net/fig/660670" pageId="0" pageNumber="125">2A</figureCitation>
|
|
). Hand with 7-8 setae; movable finger with 1 subdistal seta and fixed finger with 1 subdistal setae as well; galea absent; fixed finger with 11-12 acute teeth; movable finger with 8-9 acute teeth including one large distal tooth; rallum with 11 blades, middle blades long; serrula exterior with 22-25 blades, serrula interior with 19-21 blades.
|
|
</paragraph>
|
|
<paragraph id="EA1C7766A4353859CECF2FDE65F4B897" pageId="0" pageNumber="125">
|
|
<emphasis id="83AC807B4B5B89BEF5F1B886395C0F64" bold="true" italics="true" pageId="0" pageNumber="125">Tergites</emphasis>
|
|
. Not divided; surface smooth; chaetotaxy uniseriate, I-XI 2: 2: 4: 4: 4-6: 6-8: 6: 4-6: 4: 4: 4. Anal operculum with two dorsal setae. Pleural membrane smooth.
|
|
</paragraph>
|
|
<paragraph id="EA022668D1CD5FFD0B22F842FD978648" pageId="0" pageNumber="125">
|
|
<emphasis id="F53E351903E0C870B9EFFD14AA702638" bold="true" pageId="0" pageNumber="125">
|
|
<emphasis id="136781C96DA933B4A30EFDFF3939C98F" italics="true" pageId="0" pageNumber="125">Coxae</emphasis>
|
|
.
|
|
</emphasis>
|
|
(Fig.
|
|
<figureCitation id="C1F11B5D1A07E482A34FD1B7EBF586A6" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Selachochthonius naledi sp. nov. A female holotype palp and pedal coxae B detail of female holotype coxa I C detail of female holotype coxal spines D leg I of female holotype E male paratype genitalia, showing arrangement of setae F right pedipalp of female holotype, dorsal view. Scale bars: 0.3 mm (A); 0.1 mm (B-C); 0.5 mm (D); 0.05 mm (E); 1.0 mm (F)." figureDoi="10.3897/subtbiol.42.79190.figure3" httpUri="https://binary.pensoft.net/fig/660671" pageId="0" pageNumber="125">3A-C</figureCitation>
|
|
). Manducatory process with two apical setae (the distal one about half of the length of the proximal one); rest of palp coxae with 3 setae arranged in a triangle; delicate lamellae outlined by 15 small spines. Pedal coxae (Fig.
|
|
<figureCitation id="EF553F4153600965BD742DBE030D1905" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Selachochthonius naledi sp. nov. A female holotype palp and pedal coxae B detail of female holotype coxa I C detail of female holotype coxal spines D leg I of female holotype E male paratype genitalia, showing arrangement of setae F right pedipalp of female holotype, dorsal view. Scale bars: 0.3 mm (A); 0.1 mm (B-C); 0.5 mm (D); 0.05 mm (E); 1.0 mm (F)." figureDoi="10.3897/subtbiol.42.79190.figure3" httpUri="https://binary.pensoft.net/fig/660671" pageId="0" pageNumber="125">3A-C</figureCitation>
|
|
): coxal spines tripinnate, with smooth tips arranged in a single oblique row in coxae I (7-11), chaetotaxy: I 4, II 4, III 4-5, IV 5 intercoxal tubercle present reduced between coxae III and IV, female bearing two setae and the male one.
|
|
</paragraph>
|
|
<caption id="E8AE90AA62397B175A8042111E2860C6" doi="10.3897/subtbiol.42.79190.figure3" httpUri="https://binary.pensoft.net/fig/660671" pageId="0" pageNumber="125" start="Figure 3" startId="F3">
|
|
<paragraph id="C487F4BA20AF319769CD44C0CBE5C456" pageId="0" pageNumber="125">
|
|
<emphasis id="D17D817E6C31012ECAFA03AB973502BE" bold="true" pageId="0" pageNumber="125">Figure 3.</emphasis>
|
|
<taxonomicName id="225E29292E8E38FBAE8A6862C36AE857" authorityName="Prado 1 & Preez 2 & Ferreira 1" authorityYear="2022" class="Arachnida" family="Pseudotyrannochthoniidae" genus="Selachochthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Selachochthonius naledi" order="Pseudoscorpionidea" pageId="0" pageNumber="125" phylum="Arthropoda" rank="species" species="naledi">
|
|
<emphasis id="DCCC4048D1A8F203FF4F8546BB29431A" italics="true" pageId="0" pageNumber="125">Selachochthonius naledi</emphasis>
|
|
</taxonomicName>
|
|
sp. nov.
|
|
<emphasis id="A0EECBDAA51EEA37583586F13A61AD4F" bold="true" pageId="0" pageNumber="125">A</emphasis>
|
|
female holotype palp and pedal coxae
|
|
<emphasis id="51352125882D10DF73D7E304016C3B9F" bold="true" pageId="0" pageNumber="125">B</emphasis>
|
|
detail of female holotype coxa I
|
|
<emphasis id="BAB34DB6053AC4A1BB229EADB4833ABA" bold="true" pageId="0" pageNumber="125">C</emphasis>
|
|
detail of female holotype coxal spines
|
|
<emphasis id="47F751A7C38543C21558DC8CB8B6CDAB" bold="true" pageId="0" pageNumber="125">D</emphasis>
|
|
leg I of female holotype
|
|
<emphasis id="088CC31ACCE0A1C48410D1F8BCBA6C33" bold="true" pageId="0" pageNumber="125">E</emphasis>
|
|
male paratype genitalia, showing arrangement of setae
|
|
<emphasis id="B9ACB7C86EF37EA5EF3A4750FAC0F83E" bold="true" pageId="0" pageNumber="125">F</emphasis>
|
|
right pedipalp of female holotype, dorsal view. Scale bars: 0.3 mm (
|
|
<emphasis id="EA4DBE33BF10BD42006958244169AA30" bold="true" pageId="0" pageNumber="125">A</emphasis>
|
|
); 0.1 mm (
|
|
<emphasis id="D7EBF53F21B98732DEE67399C6257D0B" bold="true" pageId="0" pageNumber="125">B-C</emphasis>
|
|
); 0.5 mm (
|
|
<emphasis id="2BA8AE266B5DAB9E8C3A8E98FB943689" bold="true" pageId="0" pageNumber="125">D</emphasis>
|
|
); 0.05 mm (
|
|
<emphasis id="51AFCA00ABF0CDDB174B52B2A6B5C29B" bold="true" pageId="0" pageNumber="125">E</emphasis>
|
|
); 1.0 mm (
|
|
<emphasis id="BEF3315B7B59E7D9DA3AD5C135FB5CFB" bold="true" pageId="0" pageNumber="125">F</emphasis>
|
|
).
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph id="1BB159A7A8DE5BAA7D920FABED95EEAF" pageId="0" pageNumber="125">Genital operculum of female: setae distributed in three horizontal rows: 2: 2: 3, genital opening not bifurcated (horizontally contiguous).</paragraph>
|
|
<paragraph id="F352C95684102EB28978D4E0D47E18AF" pageId="0" pageNumber="125">
|
|
Genital operculum of male (Fig.
|
|
<figureCitation id="3C9EF10791D01042E3FB95686E23A01A" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Selachochthonius naledi sp. nov. A female holotype palp and pedal coxae B detail of female holotype coxa I C detail of female holotype coxal spines D leg I of female holotype E male paratype genitalia, showing arrangement of setae F right pedipalp of female holotype, dorsal view. Scale bars: 0.3 mm (A); 0.1 mm (B-C); 0.5 mm (D); 0.05 mm (E); 1.0 mm (F)." figureDoi="10.3897/subtbiol.42.79190.figure3" httpUri="https://binary.pensoft.net/fig/660671" pageId="0" pageNumber="125">3D</figureCitation>
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): setae distributed in two horizontal rows: 2: 2; Anterior genital operculum with 10 setae; genital opening with 8-9 valvular genital setae.
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</paragraph>
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|
<paragraph id="C7437D2764827CAAB9A381A334BE53A1" pageId="0" pageNumber="125">
|
|
<emphasis id="5E3B78BE7A78A517F3E0FF0A789C0D0A" bold="true" pageId="0" pageNumber="125">
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<emphasis id="388E37707A27C3B08F028896C108250B" italics="true" pageId="0" pageNumber="125">Sternites</emphasis>
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|
.
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</emphasis>
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chaetotaxy IV-XI: 10-12: 12-14: 6-12: 6-8: 6: 6: 6: 4. Anal operculum with two ventral setae.
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</paragraph>
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|
<paragraph id="246801E2DFCC2AD8D7D42FA50B838ACA" pageId="0" pageNumber="125">
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|
<emphasis id="B29DECFC346DAD85D704F36B4E80B3C1" bold="true" italics="true" pageId="0" pageNumber="125">Palp</emphasis>
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|
(Fig.
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|
<figureCitation id="FC33D8E8D040FFD63E802A0C9F95A175" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Selachochthonius naledi sp. nov., female holotype A right pedipalp chela, showing lyrifissures and trichobothrial pattern, dorsal view B right pedipalp chela, showing lyrifissures arrangement, ventral view C left pedipalp chela, showing teeth morphology, trichobothrial pattern and tubercle location, antiaxial view D right pedipalp chela, antiaxial view. See Material and methods for abbreviations. Scale bar: 1.0 mm." figureDoi="10.3897/subtbiol.42.79190.figure4" httpUri="https://binary.pensoft.net/fig/660672" pageId="0" pageNumber="125">4A-D</figureCitation>
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|
). Trochanter 1.4-1.8 times longer than wide, patella 2.3-2.4 times longer than wide, femur 6.2-6.6 times longer than wide. Femoral chaetotaxy 3: 4: 4-5: 2: 7: 1. Trichobothrial pattern:
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<emphasis id="CBB4D1D5059C2FE4523923A6DBFC220B" italics="true" pageId="0" pageNumber="125">ib</emphasis>
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|
and
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<emphasis id="45885B68449B7709CA90AB91A3AF2403" italics="true" pageId="0" pageNumber="125">isb</emphasis>
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|
located at the distal portion of the hand on a tiny hump (Fig.
|
|
<figureCitation id="8CA2A86A0B8A8046553995326D7B2FB9" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Selachochthonius naledi sp. nov., female holotype A right pedipalp chela, showing lyrifissures and trichobothrial pattern, dorsal view B right pedipalp chela, showing lyrifissures arrangement, ventral view C left pedipalp chela, showing teeth morphology, trichobothrial pattern and tubercle location, antiaxial view D right pedipalp chela, antiaxial view. See Material and methods for abbreviations. Scale bar: 1.0 mm." figureDoi="10.3897/subtbiol.42.79190.figure4" httpUri="https://binary.pensoft.net/fig/660672" pageId="0" pageNumber="125">4C-D</figureCitation>
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), adjacent to each other,
|
|
<emphasis id="37CD4817A9132CBA85111BFA4D2F319D" italics="true" pageId="0" pageNumber="125">eb</emphasis>
|
|
proximad to
|
|
<emphasis id="58FDBFE1486986002AACBCCB2175F83C" italics="true" pageId="0" pageNumber="125">esb</emphasis>
|
|
,
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|
<emphasis id="2710B0DE9F68DEF38DCF83C416A464FD" italics="true" pageId="0" pageNumber="125">ist</emphasis>
|
|
distad to
|
|
<emphasis id="A47E10A024D2123BE658D7C655E8A974" italics="true" pageId="0" pageNumber="125">esb</emphasis>
|
|
(about the same distance between both),
|
|
<emphasis id="C7837A738926AD120149DE5790537E79" italics="true" pageId="0" pageNumber="125">eb</emphasis>
|
|
-
|
|
<emphasis id="DAB9833D48C15C1F150C8BB5B5ABFD62" italics="true" pageId="0" pageNumber="125">esb</emphasis>
|
|
-
|
|
<emphasis id="072C476992351F676622543D8EF7A71E" italics="true" pageId="0" pageNumber="125">ist</emphasis>
|
|
located at the base of fixed finger,
|
|
<emphasis id="3B641FB96811A1A383C23D6D253A300A" italics="true" pageId="0" pageNumber="125">it</emphasis>
|
|
distad to
|
|
<emphasis id="ECF486CFD9828E819234BE37D4CBDAAE" italics="true" pageId="0" pageNumber="125">est</emphasis>
|
|
,
|
|
<emphasis id="8476BECE6984EA9D4EDC775FAE3E86D6" italics="true" pageId="0" pageNumber="125">et</emphasis>
|
|
distad from
|
|
<emphasis id="9E1DB36E409C3F3738424E1F4DD61D9C" italics="true" pageId="0" pageNumber="125">it</emphasis>
|
|
. Trichobothrium
|
|
<emphasis id="AB2FEA519140E2FD8A92DC48A2DDAE69" italics="true" pageId="0" pageNumber="125">st</emphasis>
|
|
located at third teeth level,
|
|
<emphasis id="EB1B108FAB61C85E8F587AA1387A63F9" italics="true" pageId="0" pageNumber="125">sb</emphasis>
|
|
proximad to
|
|
<emphasis id="2501D9A697DD0143157D633571C973E8" italics="true" pageId="0" pageNumber="125">b</emphasis>
|
|
,
|
|
<emphasis id="04D17160B8A59733C4C48ACCD34E4E57" italics="true" pageId="0" pageNumber="125">t</emphasis>
|
|
distad to
|
|
<emphasis id="8164B74CD6F46E216646D9D7E41C9EB1" italics="true" pageId="0" pageNumber="125">b</emphasis>
|
|
,
|
|
<emphasis id="3B32D7E43CE4A78029BA5E6D6931219B" italics="true" pageId="0" pageNumber="125">b</emphasis>
|
|
at the same distance from
|
|
<emphasis id="55859C79F42E6258657AEA31BDEDEF90" italics="true" pageId="0" pageNumber="125">t</emphasis>
|
|
and
|
|
<emphasis id="E647F87D07F4C46D4D9A5046D4A79EC8" italics="true" pageId="0" pageNumber="125">sb</emphasis>
|
|
(Fig.
|
|
<figureCitation id="BADCEFCDB3D9EFC4A4822095FAB48A5C" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Selachochthonius naledi sp. nov., female holotype A right pedipalp chela, showing lyrifissures and trichobothrial pattern, dorsal view B right pedipalp chela, showing lyrifissures arrangement, ventral view C left pedipalp chela, showing teeth morphology, trichobothrial pattern and tubercle location, antiaxial view D right pedipalp chela, antiaxial view. See Material and methods for abbreviations. Scale bar: 1.0 mm." figureDoi="10.3897/subtbiol.42.79190.figure4" httpUri="https://binary.pensoft.net/fig/660672" pageId="0" pageNumber="125">4C-D</figureCitation>
|
|
). Fixed finger almost straight, movable finger slightly bent (Fig.
|
|
<figureCitation id="DA94EF07BEC0E70975DA4F18D37E9FBF" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Selachochthonius naledi sp. nov. A female holotype palp and pedal coxae B detail of female holotype coxa I C detail of female holotype coxal spines D leg I of female holotype E male paratype genitalia, showing arrangement of setae F right pedipalp of female holotype, dorsal view. Scale bars: 0.3 mm (A); 0.1 mm (B-C); 0.5 mm (D); 0.05 mm (E); 1.0 mm (F)." figureDoi="10.3897/subtbiol.42.79190.figure3" httpUri="https://binary.pensoft.net/fig/660671" pageId="0" pageNumber="125">3A-D</figureCitation>
|
|
). Chelal fixed finger with 32-34 acute, triangular, and widely spaced teeth. Movable finger with 29-31 acute teeth, growing from basal to distal, tubercle present between the 13th and 14th teeth of the female movable finger and between the 10th and 11th teeth of the male movable finger (Fig.
|
|
<figureCitation id="E1AC1418DC046AA382C1AEF51E684F14" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Selachochthonius naledi sp. nov., female holotype A right pedipalp chela, showing lyrifissures and trichobothrial pattern, dorsal view B right pedipalp chela, showing lyrifissures arrangement, ventral view C left pedipalp chela, showing teeth morphology, trichobothrial pattern and tubercle location, antiaxial view D right pedipalp chela, antiaxial view. See Material and methods for abbreviations. Scale bar: 1.0 mm." figureDoi="10.3897/subtbiol.42.79190.figure4" httpUri="https://binary.pensoft.net/fig/660672" pageId="0" pageNumber="125">4C-D</figureCitation>
|
|
).
|
|
</paragraph>
|
|
<caption id="785E9433511CE80411D2EA8E5469E767" doi="10.3897/subtbiol.42.79190.figure4" httpUri="https://binary.pensoft.net/fig/660672" pageId="0" pageNumber="125" start="Figure 4" startId="F4">
|
|
<paragraph id="09ECB288D1FE878624958A80C3AD9DEB" pageId="0" pageNumber="125">
|
|
<emphasis id="319D1473DAF4F62B9EBC50A4A28287EA" bold="true" pageId="0" pageNumber="125">Figure 4.</emphasis>
|
|
<taxonomicName id="B7045881E73910F7A2E9F9BE67088B25" authorityName="Prado 1 & Preez 2 & Ferreira 1" authorityYear="2022" class="Arachnida" family="Pseudotyrannochthoniidae" genus="Selachochthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Selachochthonius naledi" order="Pseudoscorpionidea" pageId="0" pageNumber="125" phylum="Arthropoda" rank="species" species="naledi">
|
|
<emphasis id="E8C538EDD50B867AC9EBC0E155278DB5" italics="true" pageId="0" pageNumber="125">Selachochthonius naledi</emphasis>
|
|
</taxonomicName>
|
|
sp. nov., female holotype
|
|
<emphasis id="37E2DB75D8FDF3CD4A7C96343353E554" bold="true" pageId="0" pageNumber="125">A</emphasis>
|
|
right pedipalp chela, showing lyrifissures and trichobothrial pattern, dorsal view
|
|
<emphasis id="710534635D4E3A5D09AD95F0038D9A34" bold="true" pageId="0" pageNumber="125">B</emphasis>
|
|
right pedipalp chela, showing lyrifissures arrangement, ventral view
|
|
<emphasis id="4126772ABFB7B01DD1EE90F068B42628" bold="true" pageId="0" pageNumber="125">C</emphasis>
|
|
left pedipalp chela, showing teeth morphology, trichobothrial pattern and tubercle location, antiaxial view
|
|
<emphasis id="576801B257FAA22CF457790E89F14C8E" bold="true" pageId="0" pageNumber="125">D</emphasis>
|
|
right pedipalp chela, antiaxial view. See Material and methods for abbreviations. Scale bar: 1.0 mm.
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph id="5CFD55B68296512BAA231B1F63690423" pageId="0" pageNumber="125">
|
|
<emphasis id="8CFDC7D8A84D2CC0CB5AA706968C7412" bold="true" italics="true" pageId="0" pageNumber="125">Leg.</emphasis>
|
|
IV (Fig.
|
|
<figureCitation id="89E0D5CAB5778BEEF0A849261DD5407A" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Selachochthonius naledi sp. nov. A left chelicera of female holotype, antiaxial view B female holotype carapace, showing chaetotaxy of carapace C detailed anterior margin of female paratype carapace D detailed anterior margin of male paratype carapace E leg IV of female holotype, showing tubercle location. Scale bars: 0.25 mm (A, C-D); 0.5 mm (B, E)." figureDoi="10.3897/subtbiol.42.79190.figure2" httpUri="https://binary.pensoft.net/fig/660670" pageId="0" pageNumber="125">2E</figureCitation>
|
|
). Arolia same length as claws; a tiny protuberance near end of tarsus.
|
|
</paragraph>
|
|
<paragraph id="F31781BD9B8479D1D0FF55E5A4495F48" pageId="0" pageNumber="125">
|
|
<emphasis id="1FF457F834DFF3DFBE56124FA12774E6" bold="true" pageId="0" pageNumber="125">Measurements.</emphasis>
|
|
(length/width or depth in mm; ratios in parenthesis calculated by using three significant digits): Female holotype and male paratype range. Body length 2.33 [1.86]. Carapace 0.55-0.70/0.52-0.70 (1.0). Palps: trochanter 0.21-0.35/0.15-0.20 (1.4-1.8), femur 0.93-1.22/0.14-0.20 (6.2-6.6), patella 0.38-0.51/0.16-0.21 (2.3-2.4), chela 1.39-1.8/0.30-0.23 (6.1-6.2), movable finger length 0.92-1.14. Leg I: trochanter 0.13-0.23/0.10-0.18 (1.3), femur 0.44-0.67/0.08-0.10 (5.9-6.8), patella 0.29-0.38/0.07-0.09 (4.1-4.4), femur/patella 1.5-1.8, tibia 0.23-0.35/0.05-0.07 (4.3-5.1), tarsus 0.54-0.65/0.05-0.06 (10.7-11.3). Leg IV: Trochanter: 0.24-0.27/0.13-0.17 (1.6-1.8), femur + patella 0.77-1.30/0.19-0.27 (4.1-4.8), tibia 0.50-0.68/0.10-0.12 (5.2-5.4), basitarsus 0.25-0.30/0.07-0.09 (3.4-3.6), telotarsus 0.55-0.73/0.05-0.06 (10.9-12.1).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="47ED4BDA7A79470E70E2D2DD7E163C71" pageId="0" pageNumber="125" type="habitat">
|
|
<paragraph id="382D882AFB08A5A5EDEFB746C404E925" pageId="0" pageNumber="125">Habitat and ecological remarks.</paragraph>
|
|
<paragraph id="A75D75188B5C2EB1DEE21FEFF54A228C" pageId="0" pageNumber="125">
|
|
Despite extensive efforts in search of invertebrates in the caves, only two specimens of
|
|
<taxonomicName id="61B630AA788A793BAA8331366A474FCE" authorityName="Prado 1 & Preez 2 & Ferreira 1" authorityYear="2022" class="Arachnida" family="Pseudotyrannochthoniidae" genus="Selachochthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Selachochthonius naledi" order="Pseudoscorpionidea" pageId="0" pageNumber="125" phylum="Arthropoda" rank="species" species="naledi">
|
|
<emphasis id="C64BDE0103747E79B86C5CD304A226E9" italics="true" pageId="0" pageNumber="125">Selachochthonius naledi</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. were found, thus indicating their low population density. In both Vila Louise and Yom Tov caves, the specimens were found in the deeper, moist and aphotic areas. The two individuals were collected from under block rocks in the cave floor. Neither specimens exhibited any phototactic behaviour. An interesting behaviour was observed for the individual from Vila Louise cave (female holotype): when gently touched by the brush bristles, the individual responded aggressively, repeatedly grasping the bristles. Even though one of the authors (RLF) has been collecting cave pseudoscorpions for the last 30 years (especially
|
|
<taxonomicName id="C424DD11C4788B991355E2AA6FB57A52" authorityName="J. C. Chamberlin" authorityYear="1929" class="Arachnida" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Pseudoscorpionidea" pageId="0" pageNumber="125" phylum="Arthropoda" rank="superFamily" superFamily="Chthonioidea">Chthonioidea</taxonomicName>
|
|
), this
|
|
<normalizedToken id="F1CBDF3B8A1AA4B527C15C7E444595D5" originalValue="“aggressive”">"aggressive"</normalizedToken>
|
|
behaviour has never been recorded by him before. Potential prey in both caves include springtails, juvenile crickets and isopods (
|
|
<taxonomicName id="30242AFBF440366D5F6F918EFACCCF37" authorityName="Vandel" authorityYear="1952" class="Malacostraca" family="Styloniscidae" kingdom="Animalia" lsidName="" order="Isopoda" pageId="0" pageNumber="125" phylum="Arthropoda" rank="family">Styloniscidae</taxonomicName>
|
|
).
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |