426 lines
53 KiB
XML
426 lines
53 KiB
XML
<document id="857908D5F5BB6143C16DCA1F7A83D0E5" ID-CLB-Dataset="5616" ID-DOI="10.11646/zootaxa.3794.1.2" ID-GBIF-Dataset="1451533a-1819-4fed-aa53-c45a8f594844" ID-ISSN="1175-5326" ID-Zenodo-Dep="4914649" ID-ZooBank="AE0A99F2-1D71-46CB-89E2-08ED4C010E17" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623183100878" checkinUser="felipe" docAuthor="Christian, Erhard, Isaia, Marco, Paschetta, Mauro & Bruckner, Alexander" docDate="2014" docId="2D3887EF360DDA2AFF4CFA90FF25FC68" docLanguage="en" docName="zt03794p086.pdf" docOrigin="Zootaxa 3794 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Eukoenenia lanai Christian & Isaia & Paschetta & Bruckner 2014, species nova" docType="treatment" docVersion="8" lastPageNumber="82" masterDocId="D101FF973616DA34FFDBFFE8FFE6FFBB" masterDocTitle="Differentiation among cave populations of the Eukoenenia spelaea species-complex (Arachnida: Palpigradi) in the southwestern Alps" masterLastPageNumber="86" masterPageNumber="52" pageNumber="79" updateTime="1699006105339" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="240DADF564BF509C8893A43A194D3257">Differentiation among cave populations of the Eukoenenia spelaea species-complex (Arachnida: Palpigradi) in the southwestern Alps</mods:title>
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<mods:name id="B6D1FB950BB997DF9E87C0A2ED455FC0" type="personal">
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<mods:namePart id="E6ED5B0A766BBF3154D593B29B2A62FD">Christian, Erhard</mods:namePart>
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<mods:affiliation id="A672E852BB5B284BF1B78D6B1BB0E26A">Institute of Zoology, University of Natural Resources and Life Sciences, G. Mendel Str. 33, 1180 Wien, Austria.</mods:affiliation>
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<mods:namePart id="683BF45013B830A18DE96C75C9442E54">Isaia, Marco</mods:namePart>
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<mods:affiliation id="B02E4376CE42638E868FD2C1A921A654">Department of Life Sciences and Systems Biology, University of Turin, Via Accademia Albertina, 13, 10123 Torino, Italy.</mods:affiliation>
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<mods:namePart id="4AF582F6C992169344CD3A5C7E075BC2">Paschetta, Mauro</mods:namePart>
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<mods:affiliation id="17055719910981ED28474A66B9DF592F">Department of Life Sciences and Systems Biology, University of Turin, Via Accademia Albertina, 13, 10123 Torino, Italy.</mods:affiliation>
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<mods:namePart id="39E938C3F51C7D77C720B901ABC81BAE">Bruckner, Alexander</mods:namePart>
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<mods:affiliation id="ED8B90FAB196488FD8DE7A2F77507042">Institute of Zoology, University of Natural Resources and Life Sciences, G. Mendel Str. 33, 1180 Wien, Austria.</mods:affiliation>
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<mods:title id="97AA3A056D644DADC86FE98A8CE1055F">Zootaxa</mods:title>
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<mods:date id="5DBD721E6243B2E169595964B97D5B61">2014</mods:date>
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<treatment id="2D3887EF360DDA2AFF4CFA90FF25FC68" ID-DOI="http://doi.org/10.5281/zenodo.4914655" ID-GBIF-Taxon="183403614" ID-Zenodo-Dep="4914655" LSID="urn:lsid:plazi:treatment:2D3887EF360DDA2AFF4CFA90FF25FC68" httpUri="http://treatment.plazi.org/id/2D3887EF360DDA2AFF4CFA90FF25FC68" lastPageId="30" lastPageNumber="82" pageId="27" pageNumber="79">
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<subSubSection id="ED8B6572360DDA2FFF4CFA90FD73FA29" box="[151,661,1400,1426]" pageId="27" pageNumber="79" type="nomenclature">
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<paragraph id="A52E36F9360DDA2FFF4CFA90FD73FA29" blockId="27.[151,854,1400,1460]" box="[151,661,1400,1426]" pageId="27" pageNumber="79">
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<heading id="FE668195360DDA2FFF4CFA90FD73FA29" bold="true" box="[151,661,1400,1426]" fontSize="11" level="1" pageId="27" pageNumber="79" reason="1">
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<taxonomicName id="62914D7A360DDA2FFF4CFA90FE08FA29" ID-CoL="3C9B5" authority="Christian" authorityName="Christian & Isaia & Paschetta & Bruckner" authorityYear="2014" box="[151,494,1400,1426]" class="Arachnida" family="Eukoeneniidae" genus="Eukoenenia" kingdom="Animalia" order="Palpigradi" pageId="27" pageNumber="79" phylum="Arthropoda" rank="species" species="lanai" status="sp. nov.">
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<emphasis id="97E5EAEB360DDA2FFF4CFA90FE89FA29" bold="true" box="[151,367,1400,1426]" italics="true" pageId="27" pageNumber="79">Eukoenenia lanai</emphasis>
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Christian
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</taxonomicName>
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,
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<taxonomicNameLabel id="8CD65790360DDA2FFE20FA90FD73FA29" box="[507,661,1400,1426]" pageId="27" pageNumber="79" rank="species">species nova</taxonomicNameLabel>
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<subSubSection id="ED8B6572360DDA2FFF4CFA73FCB0FA0F" box="[151,854,1435,1460]" pageId="27" pageNumber="79" type="description">
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<paragraph id="A52E36F9360DDA2FFF4CFA73FCB0FA0F" blockId="27.[151,854,1400,1460]" box="[151,854,1435,1460]" pageId="27" pageNumber="79">
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(
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<figureCitation id="3DAA2A7C360DDA2FFF45FA73FEEFFA0F" box="[158,265,1435,1460]" captionStart="FIGURE 1" captionStartId="2.[151,250,1884,1906]" captionTargetBox="[188,1398,193,1862]" captionTargetId="figure-60@2.[188,1398,193,1862]" captionTargetPageId="2" captionText="FIGURE 1. Habitus of cave palpigrades of the Eukoenenia spelaea complex. From above: E. spelaea from Ardovská, Slovakia; E. strinatii from Bossea; and E. lanai sp. n. from Monfieis. Photographs courtesy of Ľubomír Kováč (top) and Enrico Lana." figureDoi="http://doi.org/10.5281/zenodo.4914651" httpUri="https://zenodo.org/record/4914651/files/figure.png" pageId="27" pageNumber="79">Figures 1</figureCitation>
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,
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<figureCitation id="3DAA2A7C360DDA2FFECCFA73FEC1FA0F" box="[279,295,1435,1460]" captionStart="FIGURE 2" captionStartId="3.[151,250,1110,1132]" captionTargetBox="[151,1436,193,1089]" captionTargetId="figure-443@3.[151,1436,193,1089]" captionTargetPageId="3" captionText="FIGURE 2. Study area and cave populations of palpigrades. B = Bossea, C = Caudano, F = Fornaci, L = Litrôn, M = Monfieis, P = Partigiano. Not sampled for this study: CL = Grotte de la Clue and SV = Grotte de Saint-Vincent-de-Mélan. Type localities and new species are highlighted. Base map © 2012 Google, Cne/Spot Image, Tele Atlas." figureDoi="http://doi.org/10.5281/zenodo.4914657" httpUri="https://zenodo.org/record/4914657/files/figure.png" pageId="27" pageNumber="79">2</figureCitation>
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,
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<figureCitation id="3DAA2A7C360DDA2FFEE8FA73FE82FA09" box="[307,356,1435,1460]" captionStart="FIGURE 5" captionStartId="11.[151,250,1596,1618]" captionTargetBox="[151,1436,481,1575]" captionTargetId="figure-423@11.[151,1436,481,1575]" captionTargetPageId="11" captionText="FIGURE 5. Lateral organ. B: Bossea ♀ (E. strinatii), L: Litrôn ♂ (E. bonadonai), C: Caudano ♀ (E. bonadonai), M: Monfieis ♀ (E. lanai sp. n.), F: Fornaci ♀ (E. roscia sp. n.). Images enlarged to same size to facilitate comparison. The blades are 40–50 µm long." figureDoi="http://doi.org/10.5281/zenodo.4914663" httpUri="https://zenodo.org/record/4914663/files/figure.png" pageId="27" pageNumber="79">5 M</figureCitation>
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,
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<figureCitation id="3DAA2A7C360DDA2FFEB4FA73FE99FA0F" box="[367,383,1435,1460]" captionStart="FIGURE 6" captionStartId="12.[151,250,1190,1212]" captionTargetBox="[213,1378,437,1159]" captionTargetId="figure-245@12.[200,1386,423,1168]" captionTargetPageId="12" captionText="FIGURE 6. Lateral organ and opisthosomal chaetotaxy of Eukoenenia specimens from the six Italian cave populations. The right column shows the values for published specimens and the two comparative females (Ardovská, Griffen). The top line of each box gives the number of blades in the lateral organs (one digit, if symmetric), the three lines below give the numbers of setae a on sternites IV–VI (left ǀ right)." figureDoi="http://doi.org/10.5281/zenodo.4914665" httpUri="https://zenodo.org/record/4914665/files/figure.png" pageId="27" pageNumber="79">6</figureCitation>
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,
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<figureCitation id="3DAA2A7C360DDA2FFE50FA73FE2FFA09" box="[395,457,1435,1460]" captionStart="FIGURE 10" captionStartId="15.[151,250,1468,1490]" captionTargetBox="[174,1418,203,1436]" captionTargetId="figure-349@15.[174,1418,203,1436]" captionTargetPageId="15" captionText="FIGURE 10. Tip of leg I. Modified setae: rs = rod seta; fs = forked seta; m = macroseta; cs = curved seta; the remaining setae are out of focus. B: Bossea ♀ (E. strinatii), C: Caudano ♀ (E. bonadonai), P: Partigiano ♀ (E. spelaea), M: Monfieis ♀ (E. lanai sp. n.), A: Ardovská ♀ (E. spelaea), G: Griffen ♀ (E. spelaea). Scale bar: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.4914673" httpUri="https://zenodo.org/record/4914673/files/figure.png" pageId="27" pageNumber="79">10 M</figureCitation>
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,
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<figureCitation id="3DAA2A7C360DDA2FFE0EFA73FDF5FA09" box="[469,531,1435,1460]" captionStart="FIGURE 12" captionStartId="17.[151,250,1104,1126]" captionTargetBox="[151,1436,187,1093]" captionTargetId="figure-542@17.[151,1436,187,1093]" captionTargetPageId="17" captionText="FIGURE 12. Third and fourth basitarsus of leg I. For better comparison of the shape, the third basitarsi are all enlarged to the same length. B: Bossea ♂ (E. strinatii), C: Caudano ♂ (E. bonadonai), L: Litrôn ♂ (E. bonadonai), M: Monfieis ♂ (E. lanai sp. n.), P: Partigiano ♀ (E. spelaea), F: Fornaci ♀ (E. roscia sp. n.), A: Ardovská ♀ (E. spelaea), G: Griffen ♀ (E. spelaea)." figureDoi="http://doi.org/10.5281/zenodo.4914677" httpUri="https://zenodo.org/record/4914677/files/figure.png" pageId="27" pageNumber="79">12 M</figureCitation>
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,
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<figureCitation id="3DAA2A7C360DDA2FFDC4FA73FDB8FA09" box="[543,606,1435,1460]" captionStart="FIGURE 13" captionStartId="18.[151,250,1960,1982]" captionTargetBox="[174,1418,904,1940]" captionTargetId="figure-351@18.[174,1418,904,1940]" captionTargetPageId="18" captionText="FIGURE 13. Basitarsus of leg IV. The basitarsi are all enlarged to the same length. Specimens as in Fig. 12." figureDoi="http://doi.org/10.5281/zenodo.4914679" httpUri="https://zenodo.org/record/4914679/files/figure.png" pageId="27" pageNumber="79">13 M</figureCitation>
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,
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<figureCitation id="3DAA2A7C360DDA2FFDB2FA73FD4EFA09" box="[617,680,1435,1460]" captionStart="FIGURE 14" captionStartId="19.[151,250,1283,1305]" captionTargetBox="[151,1436,193,1262]" captionTargetId="figure-367@19.[151,1436,193,1262]" captionTargetPageId="19" captionText="FIGURE 14. Setae a on sternite VI. B: Bossea ♂ (E. strinatii), C: Caudano ♂ (E. bonadonai), L: Litrôn ♂ (E. bonadonai), M: Monfieis ♂ (E. lanai sp. n.), P: Partigiano ♀ (E. spelaea), F: Fornaci ♂ (E. roscia sp. n.), A: Ardovská ♀ (E. spelaea with one seta a absent as an individual aberration), G: Griffen ♀ (E. spelaea). Midline = scale bar: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.4914681" httpUri="https://zenodo.org/record/4914681/files/figure.png" pageId="27" pageNumber="79">14 M</figureCitation>
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,
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<figureCitation id="3DAA2A7C360DDA2FFD68FA73FD37FA0F" box="[691,721,1435,1460]" captionStart="FIGURE 20" captionStartId="24.[151,250,1942,1964]" captionTargetBox="[415,1171,1354,1921]" captionTargetId="figure-114@24.[415,1171,1354,1921]" captionTargetPageId="24" captionText="FIGURE 20. Fusules (f1, f2) on first genital lobe of a male from Monfieis (E. lanai sp. n.). The fusules are inserted on domeshaped bases. Scale bar: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.4914693" httpUri="https://zenodo.org/record/4914693/files/figure.png" pageId="27" pageNumber="79">20</figureCitation>
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,
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<figureCitation id="3DAA2A7C360DDA2FFD06FA73FD1DFA0F" box="[733,763,1435,1460]" captionStart="FIGURE 21" captionStartId="25.[151,250,1172,1194]" captionTargetBox="[151,1436,193,1150]" captionTargetId="figure-446@25.[151,1436,193,1150]" captionTargetPageId="25" captionText="FIGURE 21. Flagellum. Left: articles IV–VI of a Monfieis ♂ (E. lanai sp. n.; scale bar: 100 µm). Right: the same articles of a E. mirabilis ♀ (soil-dwelling) from Bergeggi, Liguria, for comparison (scale bar: 20 µm). Note apical crown of cuticular spines on articles III (at the very top of the images) and V." figureDoi="http://doi.org/10.5281/zenodo.4914695" httpUri="https://zenodo.org/record/4914695/files/figure.png" pageId="27" pageNumber="79">21</figureCitation>
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,
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<figureCitation id="3DAA2A7C360DDA2FFCDCFA73FCA8FA0F" box="[775,846,1435,1460]" captionStart-0="FIGURE 23" captionStart-1="FIGURE 24" captionStart-2="FIGURE 25" captionStartId-0="27.[151,250,817,839]" captionStartId-1="28.[151,250,1953,1975]" captionStartId-2="29.[151,250,687,709]" captionTargetBox-0="[200,1393,201,786]" captionTargetBox-1="[221,1358,208,1896]" captionTargetBox-2="[186,1398,206,657]" captionTargetId-0="figure-546@27.[185,1402,193,796]" captionTargetId-1="figure-58@28.[202,1370,193,1910]" captionTargetId-2="figure-209@29.[177,1410,193,666]" captionTargetPageId-0="27" captionTargetPageId-1="28" captionTargetPageId-2="29" captionText-0="FIGURE 23. Principal component analysis based on measurements of 29 Eukoenenia individuals (females and males) from six Italian and two comparative populations (A, G). G = Griffen, A = Ardovská, P = Partigiano, C = Caudano, L = Litrôn, F = Fornaci, B = Bossea, M = Monfieis." captionText-1="FIGURE 24. Graphic representation of the measurements used for the morphometric analysis (data from Tables 1 & 2, in µm). Only females included. Initials of caves as in Fig. 23 (non-Italian comparative populations in brackets)." captionText-2="FIGURE 25. Graphic representation of the values of commonly used morphological indices (data from Tables 1 & 2). Only females included. Initials of caves as in Fig. 23." figureDoi-0="http://doi.org/10.5281/zenodo.4914699" figureDoi-1="http://doi.org/10.5281/zenodo.4914701" figureDoi-2="http://doi.org/10.5281/zenodo.4914703" httpUri-0="https://zenodo.org/record/4914699/files/figure.png" httpUri-1="https://zenodo.org/record/4914701/files/figure.png" httpUri-2="https://zenodo.org/record/4914703/files/figure.png" pageId="27" pageNumber="79">23–25</figureCitation>
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)
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</paragraph>
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<subSubSection id="ED8B6572360DDA2FFF4CFA0BFC44F937" pageId="27" pageNumber="79" type="materials_examined">
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<emphasis id="97E5EAEB360DDA2FFF4CFA0BFE67FA47" bold="true" box="[151,385,1507,1532]" inLineHeading="true" pageId="27" pageNumber="79" reason="1">Material examined.</emphasis>
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<materialsCitation id="15F93CA4360DDA2FFE53FA0BFE2DF9FF" ID-GBIF-Occurrence="3319538303" collectingDate="2010-09-12" collectorName="di Monfieis & E. Lana. Two" country="Italy" county="Province of Cueno" elevation="1750" latitude="44.36222" location="Miniera" longLatPrecision="19" longitude="7.266667" municipality="Demonte" pageId="27" pageNumber="79" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Piemonte" typeStatus="holotype">
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<typeStatus id="7A2A885B360DDA2FFE53FA0BFE16FA47" box="[392,496,1507,1532]" pageId="27" pageNumber="79" type="holotype">Holotype</typeStatus>
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male:
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<collectingCountry id="DD867669360DDA2FFDE6FA0BFD97FA47" box="[573,625,1507,1532]" name="Italy" pageId="27" pageNumber="79">Italy</collectingCountry>
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,
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<collectingRegion id="6755F81B360DDA2FFDA6FA0BFD0FFA47" box="[637,745,1507,1532]" country="Italy" name="Piemonte" pageId="27" pageNumber="79">Piemonte</collectingRegion>
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,
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<collectingCounty id="4C4F4E75360DDA2FFD2EFA0BFC2CFA47" box="[757,970,1507,1532]" pageId="27" pageNumber="79">Province of Cueno</collectingCounty>
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,
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<collectingMunicipality id="454AAC83360DDA2FFC0DFA0BFBD9FA41" box="[982,1087,1507,1530]" pageId="27" pageNumber="79">Demonte</collectingMunicipality>
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,
|
||
<location id="A04E6022360DDA2FFB91FA0BFB44FA47" LSID="urn:lsid:plazi:treatment:2D3887EF360DDA2AFF4CFA90FF25FC68:A04E6022360DDA2FFB91FA0BFB44FA47" box="[1098,1186,1507,1532]" country="Italy" county="Province of Cueno" latitude="44.36222" longLatPrecision="19" longitude="7.266667" municipality="Demonte" name="Miniera" pageId="27" pageNumber="79" stateProvince="Piemonte">Miniera</location>
|
||
superiore
|
||
<collectorName id="0864532F360DDA2FFAC1FA0BFA7AFA47" box="[1306,1436,1507,1532]" pageId="27" pageNumber="79">di Monfieis</collectorName>
|
||
(
|
||
<geoCoordinate id="C0A5503E360DDA2FFF44F9EFFECEF99B" box="[159,296,1543,1568]" degrees="44" direction="north" minutes="21" orientation="latitude" pageId="27" pageNumber="79" precision="15" seconds="44" value="44.36222">44°21′44″N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="C0A5503E360DDA2FFEE8F9EFFE51F99B" box="[307,439,1543,1568]" degrees="07" direction="east" minutes="16" orientation="longitude" pageId="27" pageNumber="79" precision="15" seconds="00" value="7.266667">07°16′00″E</geoCoordinate>
|
||
,
|
||
<elevation id="2EBCD1CA360DDA2FFE18F9EFFDB2F99B" box="[451,596,1543,1568]" metricMagnitude="3" metricUnit="m" metricValue="1.75" pageId="27" pageNumber="79" unit="m" value="1750.0">
|
||
<quantity id="62699B1C360DDA2FFE18F9EFFDFEF9A4" box="[451,536,1543,1568]" metricMagnitude="3" metricUnit="m" metricValue="1.75" pageId="27" pageNumber="79" unit="m" value="1750.0">1750 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
);
|
||
<date id="D12F1039360DDA2FFDB0F9EFFCAEF99B" box="[619,840,1543,1568]" pageId="27" pageNumber="79" value="2010-09-12">
|
||
<collectingDate id="C16BE9D1360DDA2FFDB0F9EFFCAEF99B" box="[619,840,1543,1568]" pageId="27" pageNumber="79" value="2010-09-12">12 September 2010</collectingDate>
|
||
</date>
|
||
, leg.
|
||
<collectorName id="0864532F360DDA2FFC59F9E0FBFFF9A4" box="[898,1049,1544,1567]" pageId="27" pageNumber="79">E. Lana. Two</collectorName>
|
||
<typeStatus id="7A2A885B360DDA2FFBFBF9E0FB76F99B" box="[1056,1168,1544,1568]" pageId="27" pageNumber="79" type="paratype">paratypes</typeStatus>
|
||
:
|
||
<specimenCount id="B397FD70360DDA2FFB42F9EFFB18F99B" box="[1177,1278,1543,1568]" pageId="27" pageNumber="79" type="female">1 female</specimenCount>
|
||
,
|
||
<specimenCount id="B397FD70360DDA2FFAD2F9EFFAB1F99B" box="[1289,1367,1543,1568]" pageId="27" pageNumber="79" type="male">1 male</specimenCount>
|
||
; same locality, date, and collector.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="A52E36F9360DDA2FFF1CF9A7FC44F937" blockId="27.[151,1437,1507,2036]" pageId="27" pageNumber="79">
|
||
<emphasis id="97E5EAEB360DDA2FFF1CF9A7FEA8F9D3" bold="true" box="[199,334,1615,1640]" inLineHeading="true" pageId="27" pageNumber="79" reason="1">Deposition.</emphasis>
|
||
<materialsCitation id="15F93CA4360DDA2FFE83F9B8FC78F937" ID-GBIF-Occurrence="3319538301" country="Austria" location="Museum of Natural History" pageId="27" pageNumber="79" specimenCount="3" stateProvince="Wien" typeStatus="holotype">
|
||
<location id="A04E6022360DDA2FFE83F9B8FD7DF9D3" LSID="urn:lsid:plazi:treatment:2D3887EF360DDA2AFF4CFA90FF25FC68:A04E6022360DDA2FFE83F9B8FD7DF9D3" box="[344,667,1615,1640]" country="Austria" name="Museum of Natural History" pageId="27" pageNumber="79" stateProvince="Wien">Museum of Natural History</location>
|
||
,
|
||
<collectingRegion id="6755F81B360DDA2FFD71F9A7FD1AF9D3" box="[682,764,1615,1640]" country="Austria" name="Wien" pageId="27" pageNumber="79">Vienna</collectingRegion>
|
||
,
|
||
<collectingCountry id="DD867669360DDA2FFCD7F9A7FC87F9D3" box="[780,865,1615,1640]" name="Austria" pageId="27" pageNumber="79">Austria</collectingCountry>
|
||
, Arachnological Collection. Acquisition numbers 21.878 (
|
||
<typeStatus id="7A2A885B360DDA2FFF28F99BFEBDF937" box="[243,347,1651,1676]" pageId="27" pageNumber="79" type="holotype">holotype</typeStatus>
|
||
), 21.879 (
|
||
<typeStatus id="7A2A885B360DDA2FFE12F99BFDCDF937" box="[457,555,1651,1676]" pageId="27" pageNumber="79" type="paratype">paratype</typeStatus>
|
||
female), 21.880 (
|
||
<typeStatus id="7A2A885B360DDA2FFD2AF99BFCB5F937" box="[753,851,1651,1676]" pageId="27" pageNumber="79" type="paratype">paratype</typeStatus>
|
||
male)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="ED8B6572360DDA2FFF1CF97FFCF7F96F" pageId="27" pageNumber="79" type="etymology">
|
||
<paragraph id="A52E36F9360DDA2FFF1CF97FFCF7F96F" blockId="27.[151,1437,1507,2036]" pageId="27" pageNumber="79">
|
||
<emphasis id="97E5EAEB360DDA2FFF1CF97FFEA8F90B" bold="true" box="[199,334,1687,1712]" inLineHeading="true" pageId="27" pageNumber="79" reason="1">Etymology.</emphasis>
|
||
The specific epithet is a patronym in honour of Enrico Lana, eminent speleobiologist, photographer and collector of rare subterranean animals.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="ED8B6572360DDA2FFF1CF937FE91F833" pageId="27" pageNumber="79" type="diagnosis">
|
||
<paragraph id="A52E36F9360DDA2FFF1CF937FE91F833" blockId="27.[151,1437,1507,2036]" pageId="27" pageNumber="79">
|
||
<emphasis id="97E5EAEB360DDA2FFF1CF937FEA7F943" bold="true" box="[199,321,1759,1784]" inLineHeading="true" pageId="27" pageNumber="79" reason="1">Diagnosis.</emphasis>
|
||
A species with all characters of the
|
||
<taxonomicName id="62914D7A360DDA2FFD07F908FCB4F943" baseAuthorityName="Peyerimhoff" baseAuthorityYear="1902" box="[732,850,1759,1784]" class="Arachnida" family="Eukoeneniidae" genus="Eukoenenia" kingdom="Animalia" order="Palpigradi" pageId="27" pageNumber="79" phylum="Arthropoda" rank="species" species="spelaea">
|
||
<emphasis id="97E5EAEB360DDA2FFD07F908FCB4F943" box="[732,850,1759,1784]" italics="true" pageId="27" pageNumber="79">E. spelaea</emphasis>
|
||
</taxonomicName>
|
||
complex as described above. Body length
|
||
<quantity id="62699B1C360DDA2FFAE3F937FA7AF943" box="[1336,1436,1759,1784]" metricMagnitude="0" metricUnit="m" metricValue="2.1" pageId="27" pageNumber="79" unit="mm" value="2100.0">2100 µm</quantity>
|
||
or more; each finger of chelicera with 8 teeth; lateral organ with 3 blades; 5 deuto-tritosternal setae; forked seta absent on I bta4; proximal forked seta on I ta3 only half length of rod seta; some or all of sternites IV–VI with more than 3+3 setae
|
||
<emphasis id="97E5EAEB360DDA2FFE9EF8A4FEB5F8D8" box="[325,339,1868,1891]" italics="true" pageId="27" pageNumber="79">a</emphasis>
|
||
; segment XI of opisthosoma with (9–)10 setae; fusules on first genital lobe of male inserted on dome-shaped bases.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="ED8B6572360DDA2AFF1CF87BFF25FC68" lastPageId="30" lastPageNumber="82" pageId="27" pageNumber="79" type="description">
|
||
<paragraph id="A52E36F9360DDA2FFF1CF87BFB3FF817" blockId="27.[151,1437,1507,2036]" box="[199,1241,1939,1964]" pageId="27" pageNumber="79">
|
||
<emphasis id="97E5EAEB360DDA2FFF1CF87BFEBFF817" bold="true" box="[199,345,1939,1964]" inLineHeading="true" pageId="27" pageNumber="79" reason="1">Description.</emphasis>
|
||
Morphometric data are given in
|
||
<tableCitation id="E8130342360DDA2FFD13F87BFCFDF817" box="[712,795,1939,1964]" captionStart="TABLE 1" captionStartId="5.[147,233,520,542]" captionTargetBox="[147,1921,571,1685]" captionText="TABLE 1. Measurements and indices of palpigrade individuals from SW Alpine caves and from two external comparative populations (A, G) of the Eukoenenia spelaea" httpUri="http://table.plazi.org/id/F1EE66713613DA31FF48FDE0F8AEFDA5" pageId="27" pageNumber="79" tableUuid="F1EE66713613DA31FF48FDE0F8AEFDA5">Table 1</tableCitation>
|
||
under
|
||
<specimenCount id="B397FD70360DDA2FFCB2F87BFC98F811" box="[873,894,1939,1962]" pageId="27" pageNumber="79" type="female">♀</specimenCount>
|
||
05,
|
||
<specimenCount id="B397FD70360DDA2FFC74F87BFC22F811" box="[943,964,1939,1962]" pageId="27" pageNumber="79" type="male">♂</specimenCount>
|
||
06 and
|
||
<specimenCount id="B397FD70360DDA2FFBC5F87BFBD5F811" box="[1054,1075,1939,1962]" pageId="27" pageNumber="79" type="male">♂</specimenCount>
|
||
07 (
|
||
<typeStatus id="7A2A885B360DDA2FFBBEF87BFB2BF817" box="[1125,1229,1939,1964]" pageId="27" pageNumber="79" type="holotype">holotype</typeStatus>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="A52E36F9360DDA2FFF1CF85FFA77F84F" blockId="27.[151,1437,1507,2036]" pageId="27" pageNumber="79">
|
||
<emphasis id="97E5EAEB360DDA2FFF1CF85FFE4BF874" box="[199,429,1975,2000]" italics="true" pageId="27" pageNumber="79">General appearance</emphasis>
|
||
. Large
|
||
<taxonomicName id="62914D7A360DDA2FFDD8F85FFD61F86B" authorityName="Borner" authorityYear="1901" box="[515,647,1975,2000]" class="Arachnida" family="Eukoeneniidae" genus="Eukoenenia" kingdom="Animalia" order="Palpigradi" pageId="27" pageNumber="79" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="97E5EAEB360DDA2FFDD8F85FFD61F86B" box="[515,647,1975,2000]" italics="true" pageId="27" pageNumber="79">Eukoenenia</emphasis>
|
||
</taxonomicName>
|
||
, with a body length of
|
||
<quantity id="62699B1C360DDA2FFC51F85FFBD0F86B" box="[906,1078,1975,2000]" metricMagnitude="0" metricUnit="m" metricValue="2.135" metricValueMax="2.17" metricValueMin="2.1" pageId="27" pageNumber="79" unit="mm" value="2135.0" valueMax="2170.0" valueMin="2100.0">2100–2170 µm</quantity>
|
||
. Shape and pubescence of trunk as in other members of the
|
||
<taxonomicName id="62914D7A360DDA2FFE13F834FDDAF84F" baseAuthorityName="Peyerimhoff" baseAuthorityYear="1902" box="[456,572,2011,2036]" class="Arachnida" family="Eukoeneniidae" genus="Eukoenenia" kingdom="Animalia" order="Palpigradi" pageId="27" pageNumber="79" phylum="Arthropoda" rank="species" species="spelaea">
|
||
<emphasis id="97E5EAEB360DDA2FFE13F834FDDAF84F" box="[456,572,2011,2036]" italics="true" pageId="27" pageNumber="79">E. spelaea</emphasis>
|
||
</taxonomicName>
|
||
complex. Trunk of live individuals yellowish, particularly the opisthosoma.
|
||
</paragraph>
|
||
<caption id="F1EE6671360ADA28FF4CF849FB48F86D" ID-DOI="http://doi.org/10.5281/zenodo.4914701" ID-Zenodo-Dep="4914701" httpUri="https://zenodo.org/record/4914701/files/figure.png" pageId="28" pageNumber="80" startId="28.[151,250,1953,1975]" targetBox="[221,1358,208,1896]" targetPageId="28">
|
||
<paragraph id="A52E36F9360ADA28FF4CF849FB48F86D" blockId="28.[151,1436,1953,2006]" pageId="28" pageNumber="80">
|
||
<emphasis id="97E5EAEB360ADA28FF4CF849FEF9F80D" bold="true" box="[151,287,1953,1975]" pageId="28" pageNumber="80">FIGURE 24.</emphasis>
|
||
Graphic representation of the measurements used for the morphometric analysis (data from Tables 1 & 2, in µm). Only females included. Initials of caves as in Fig. 23 (non-Italian comparative populations in brackets).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="F1EE6671360BDA29FF4CFD47FD9CFD5F" ID-DOI="http://doi.org/10.5281/zenodo.4914703" ID-Zenodo-Dep="4914703" httpUri="https://zenodo.org/record/4914703/files/figure.png" pageId="29" pageNumber="81" startId="29.[151,250,687,709]" targetBox="[186,1398,206,657]" targetPageId="29">
|
||
<paragraph id="A52E36F9360BDA29FF4CFD47FD9CFD5F" blockId="29.[151,1436,687,740]" pageId="29" pageNumber="81">
|
||
<emphasis id="97E5EAEB360BDA29FF4CFD47FEC7FD7F" bold="true" box="[151,289,687,709]" pageId="29" pageNumber="81">FIGURE 25.</emphasis>
|
||
Graphic representation of the values of commonly used morphological indices (data from Tables 1 & 2). Only females included. Initials of caves as in Fig. 23.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="F1EE6671360BDA29FF4CF965FEB5F979" ID-DOI="http://doi.org/10.5281/zenodo.4914705" ID-Zenodo-Dep="4914705" httpUri="https://zenodo.org/record/4914705/files/figure.png" pageId="29" pageNumber="81" startId="29.[151,250,1677,1699]" targetBox="[188,1399,774,1656]" targetPageId="29">
|
||
<paragraph id="A52E36F9360BDA29FF4CF965FEB5F979" blockId="29.[151,1436,1677,1730]" pageId="29" pageNumber="81">
|
||
<emphasis id="97E5EAEB360BDA29FF4CF965FEC7F919" bold="true" box="[151,289,1677,1699]" pageId="29" pageNumber="81">FIGURE 26.</emphasis>
|
||
General appearance of a larva (left) and an adult female of
|
||
<taxonomicName id="62914D7A360BDA29FC50F966FC1EF918" authorityName="Conde" authorityYear="1977" box="[907,1016,1678,1699]" class="Arachnida" family="Eukoeneniidae" genus="Eukoenenia" kingdom="Animalia" order="Palpigradi" pageId="29" pageNumber="81" phylum="Arthropoda" rank="species" species="strinatii">
|
||
<emphasis id="97E5EAEB360BDA29FC50F966FC1EF918" box="[907,1016,1678,1699]" italics="true" pageId="29" pageNumber="81">E. strinatii</emphasis>
|
||
</taxonomicName>
|
||
from Bossea. Specimens enlarged to the same trunk length.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="A52E36F9360BDA2AFF1CF902FDBEFEFD" blockId="29.[151,1437,1769,2015]" lastBlockId="30.[151,1436,151,979]" lastPageId="30" lastPageNumber="82" pageId="29" pageNumber="81">
|
||
<emphasis id="97E5EAEB360BDA29FF1CF902FECDF8BA" box="[199,299,1770,1793]" italics="true" pageId="29" pageNumber="81">Prosoma</emphasis>
|
||
. Frontal organ
|
||
<quantity id="62699B1C360BDA29FE05F901FDCEF8B9" box="[478,552,1769,1794]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="29" pageNumber="81" unit="mm" value="40.0">40 µm</quantity>
|
||
long (
|
||
<typeStatus id="7A2A885B360BDA29FDAFF901FD3AF8B9" box="[628,732,1769,1794]" pageId="29" pageNumber="81" type="holotype">holotype</typeStatus>
|
||
), with two lanceolate, pointed branches. Lateral organ with 3 reticulated, 45–49 µm-long blades that are parallel-sided with a pointed tip (
|
||
<figureCitation id="3DAA2A7C360BDA29FBDBF8E5FB8CF89D" box="[1024,1130,1805,1830]" captionStart="FIGURE 5" captionStartId="11.[151,250,1596,1618]" captionTargetBox="[151,1436,481,1575]" captionTargetId="figure-423@11.[151,1436,481,1575]" captionTargetPageId="11" captionText="FIGURE 5. Lateral organ. B: Bossea ♀ (E. strinatii), L: Litrôn ♂ (E. bonadonai), C: Caudano ♀ (E. bonadonai), M: Monfieis ♀ (E. lanai sp. n.), F: Fornaci ♀ (E. roscia sp. n.). Images enlarged to same size to facilitate comparison. The blades are 40–50 µm long." figureDoi="http://doi.org/10.5281/zenodo.4914663" httpUri="https://zenodo.org/record/4914663/files/figure.png" pageId="29" pageNumber="81">Fig. 5 M</figureCitation>
|
||
). Propeltidium with 10+10 setae, the shortest (
|
||
<quantity id="62699B1C360BDA29FEB6F8D9FE07F8F1" box="[365,481,1841,1866]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" metricValueMax="3.1" metricValueMin="2.9" pageId="29" pageNumber="81" unit="mm" value="30.0" valueMax="31.0" valueMin="29.0">29–31 µm</quantity>
|
||
) in central position, the longest (
|
||
<quantity id="62699B1C360BDA29FC88F8D9FC2EF8F1" box="[851,968,1841,1866]" metricMagnitude="-2" metricUnit="m" metricValue="5.5" metricValueMax="5.7" metricValueMin="5.3" pageId="29" pageNumber="81" unit="mm" value="55.0" valueMax="57.0" valueMin="53.0">53–57 µm</quantity>
|
||
) laterally in posterior half. Setae
|
||
<emphasis id="97E5EAEB360BDA29FAE4F8DAFAA9F8F5" box="[1343,1359,1842,1870]" italics="true" pageId="29" pageNumber="81">
|
||
t
|
||
<subScript id="391534BC360BDA29FA9CF8A8FAA9F8F5" attach="left" box="[1351,1359,1856,1870]" fontSize="6" pageId="29" pageNumber="81">1</subScript>
|
||
</emphasis>
|
||
,
|
||
<emphasis id="97E5EAEB360BDA29FA86F8DAFA8AF8F5" box="[1373,1388,1842,1870]" italics="true" pageId="29" pageNumber="81">
|
||
t
|
||
<subScript id="391534BC360BDA29FABFF8A8FA8AF8F5" attach="left" box="[1380,1388,1856,1870]" fontSize="6" pageId="29" pageNumber="81">2</subScript>
|
||
</emphasis>
|
||
and
|
||
<emphasis id="97E5EAEB360BDA29FF4CF8B1FF40F8CE" box="[151,166,1881,1909]" italics="true" pageId="29" pageNumber="81">
|
||
t
|
||
<subScript id="391534BC360BDA29FF45F88FFF40F8CE" attach="left" box="[158,166,1895,1909]" fontSize="6" pageId="29" pageNumber="81">3</subScript>
|
||
</emphasis>
|
||
of metapeltidium 74–85, 132–145 and
|
||
<quantity id="62699B1C360BDA29FD84F8B0FD37F8CA" box="[607,721,1880,1905]" metricMagnitude="-2" metricUnit="m" metricValue="9.0" metricValueMax="9.8" metricValueMin="8.2" pageId="29" pageNumber="81" unit="mm" value="90.0" valueMax="98.0" valueMin="82.0">82–98 µm</quantity>
|
||
long, respectively. Labrum with usual cuticular pattern and 5+5 short setae. Deuto-tritosternum with 5 setae of
|
||
<quantity id="62699B1C360BDA29FD18F896FCDDF82C" box="[707,827,1918,1943]" metricMagnitude="-2" metricUnit="m" metricValue="5.05" metricValueMax="5.4" metricValueMin="4.7" pageId="29" pageNumber="81" unit="mm" value="50.5" valueMax="54.0" valueMin="47.0">47–54 µm</quantity>
|
||
, arranged in a wide V. Chaetotaxy of chelicera and dentition of fingers as in
|
||
<taxonomicName id="62914D7A360BDA29FE6BF84CFDF3F800" authorityName="Christian & Isaia & Paschetta & Bruckner" authorityYear="2014" box="[432,533,1956,1979]" class="Arachnida" family="Eukoeneniidae" genus="Eukoenenia" kingdom="Animalia" order="Palpigradi" pageId="29" pageNumber="81" phylum="Arthropoda" rank="species" species="roscia">
|
||
<emphasis id="97E5EAEB360BDA29FE6BF84CFDF3F800" box="[432,533,1956,1979]" italics="true" pageId="29" pageNumber="81">E. roscia</emphasis>
|
||
</taxonomicName>
|
||
. Coxal chaetotaxy as described above for
|
||
<taxonomicName id="62914D7A360BDA29FC2BF84BFBA0F807" baseAuthorityName="Peyerimhoff" baseAuthorityYear="1902" box="[1008,1094,1955,1980]" class="Arachnida" family="Eukoeneniidae" genus="Eukoenenia" kingdom="Animalia" order="Palpigradi" pageId="29" pageNumber="81" phylum="Arthropoda" rank="species" species="spelaea">
|
||
<emphasis id="97E5EAEB360BDA29FC2BF84BFBA0F807" box="[1008,1094,1955,1980]" italics="true" pageId="29" pageNumber="81">spelaea</emphasis>
|
||
</taxonomicName>
|
||
group. Curved seta near tip of pedipalp weakly and asymmetrically forked, the typical
|
||
<emphasis id="97E5EAEB360BDA29FCCBF82EFCC5F864" box="[784,803,1990,2015]" italics="true" pageId="29" pageNumber="81">fs</emphasis>
|
||
is inserted somewhat behind. Leg I with 7 trichobothria in usual arrangement and a total of 7 forked setae. The ta3 of leg I has 4
|
||
<emphasis id="97E5EAEB3608DA2AFC3FFF7FFC11FF0B" box="[996,1015,151,176]" italics="true" pageId="30" pageNumber="82">fs</emphasis>
|
||
in distal half, 1–1–2:
|
||
<emphasis id="97E5EAEB3608DA2AFB2DFF7FFAF7FF0F" box="[1270,1297,151,180]" italics="true" pageId="30" pageNumber="82">
|
||
fs
|
||
<subScript id="391534BC3608DA2AFAD2FF4EFAF7FF0F" attach="left" box="[1289,1297,166,180]" fontSize="6" pageId="30" pageNumber="82">1</subScript>
|
||
</emphasis>
|
||
is only half length of the nearby
|
||
<emphasis id="97E5EAEB3608DA2AFEA6FF28FE75FF6C" box="[381,403,192,215]" italics="true" pageId="30" pageNumber="82">rs</emphasis>
|
||
; in all specimens
|
||
<emphasis id="97E5EAEB3608DA2AFD87FF56FD90FF60" box="[604,630,190,219]" italics="true" pageId="30" pageNumber="82">
|
||
fs
|
||
<subScript id="391534BC3608DA2AFDB5FF25FD90FF60" attach="left" box="[622,630,205,219]" fontSize="6" pageId="30" pageNumber="82">2</subScript>
|
||
</emphasis>
|
||
is inserted much closer to
|
||
<emphasis id="97E5EAEB3608DA2AFC7BFF28FC52FF6C" box="[928,948,192,215]" italics="true" pageId="30" pageNumber="82">m</emphasis>
|
||
than to
|
||
<emphasis id="97E5EAEB3608DA2AFBD4FF28FBC0FF6C" box="[1039,1062,192,215]" italics="true" pageId="30" pageNumber="82">cs</emphasis>
|
||
(
|
||
<figureCitation id="3DAA2A7C3608DA2AFBEEFF56FB4FFF6C" box="[1077,1193,190,215]" captionStart="FIGURE 10" captionStartId="15.[151,250,1468,1490]" captionTargetBox="[174,1418,203,1436]" captionTargetId="figure-349@15.[174,1418,203,1436]" captionTargetPageId="15" captionText="FIGURE 10. Tip of leg I. Modified setae: rs = rod seta; fs = forked seta; m = macroseta; cs = curved seta; the remaining setae are out of focus. B: Bossea ♀ (E. strinatii), C: Caudano ♀ (E. bonadonai), P: Partigiano ♀ (E. spelaea), M: Monfieis ♀ (E. lanai sp. n.), A: Ardovská ♀ (E. spelaea), G: Griffen ♀ (E. spelaea). Scale bar: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.4914673" httpUri="https://zenodo.org/record/4914673/files/figure.png" pageId="30" pageNumber="82">Fig. 10 M</figureCitation>
|
||
). I bta3 not as slender as in
|
||
<taxonomicName id="62914D7A3608DA2AFF0EFF0EFEA9FF46" authorityName="Conde" authorityYear="1977" box="[213,335,230,253]" class="Arachnida" family="Eukoeneniidae" genus="Eukoenenia" kingdom="Animalia" order="Palpigradi" pageId="30" pageNumber="82" phylum="Arthropoda" rank="species" species="strinatii">
|
||
<emphasis id="97E5EAEB3608DA2AFF0EFF0EFEA9FF46" box="[213,335,230,253]" italics="true" pageId="30" pageNumber="82">E. strinatii</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="62914D7A3608DA2AFE53FF0EFE08FF46" authorityName="Christian & Isaia & Paschetta & Bruckner" authorityYear="2014" box="[392,494,230,253]" class="Arachnida" family="Eukoeneniidae" genus="Eukoenenia" kingdom="Animalia" order="Palpigradi" pageId="30" pageNumber="82" phylum="Arthropoda" rank="species" species="roscia">
|
||
<emphasis id="97E5EAEB3608DA2AFE53FF0EFE08FF46" box="[392,494,230,253]" italics="true" pageId="30" pageNumber="82">E. roscia</emphasis>
|
||
</taxonomicName>
|
||
, length/width 3.06–3.19 (
|
||
<figureCitation id="3DAA2A7C3608DA2AFCCEFF0DFC6BFF45" box="[789,909,229,254]" captionStart="FIGURE 12" captionStartId="17.[151,250,1104,1126]" captionTargetBox="[151,1436,187,1093]" captionTargetId="figure-542@17.[151,1436,187,1093]" captionTargetPageId="17" captionText="FIGURE 12. Third and fourth basitarsus of leg I. For better comparison of the shape, the third basitarsi are all enlarged to the same length. B: Bossea ♂ (E. strinatii), C: Caudano ♂ (E. bonadonai), L: Litrôn ♂ (E. bonadonai), M: Monfieis ♂ (E. lanai sp. n.), P: Partigiano ♀ (E. spelaea), F: Fornaci ♀ (E. roscia sp. n.), A: Ardovská ♀ (E. spelaea), G: Griffen ♀ (E. spelaea)." figureDoi="http://doi.org/10.5281/zenodo.4914677" httpUri="https://zenodo.org/record/4914677/files/figure.png" pageId="30" pageNumber="82">Fig. 12 M</figureCitation>
|
||
); seta
|
||
<emphasis id="97E5EAEB3608DA2AFC0DFF0EFC07FF46" box="[982,993,230,253]" italics="true" pageId="30" pageNumber="82">r</emphasis>
|
||
inserted at 48–53% of article length. I bta4 without
|
||
<emphasis id="97E5EAEB3608DA2AFEF6FEE1FEA6FE99" box="[301,320,265,290]" italics="true" pageId="30" pageNumber="82">fs</emphasis>
|
||
. IV bta slender, all ratios very similar to those of
|
||
<taxonomicName id="62914D7A3608DA2AFCA3FEE2FC38FE9A" authorityName="Christian & Isaia & Paschetta & Bruckner" authorityYear="2014" box="[888,990,266,289]" class="Arachnida" family="Eukoeneniidae" genus="Eukoenenia" kingdom="Animalia" order="Palpigradi" pageId="30" pageNumber="82" phylum="Arthropoda" rank="species" species="roscia">
|
||
<emphasis id="97E5EAEB3608DA2AFCA3FEE2FC38FE9A" box="[888,990,266,289]" italics="true" pageId="30" pageNumber="82">E. roscia</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<emphasis id="97E5EAEB3608DA2AFC36FEE2FC1EFE9A" box="[1005,1016,266,289]" italics="true" pageId="30" pageNumber="82">r</emphasis>
|
||
<
|
||
<emphasis id="97E5EAEB3608DA2AFBC2FEE2FBDCFE9A" box="[1049,1082,266,289]" italics="true" pageId="30" pageNumber="82">grt</emphasis>
|
||
<
|
||
<emphasis id="97E5EAEB3608DA2AFB82FEE1FB98FE99" box="[1113,1150,265,290]" italics="true" pageId="30" pageNumber="82">esd</emphasis>
|
||
; tip of
|
||
<emphasis id="97E5EAEB3608DA2AFB0FFEE2FB39FE9A" box="[1236,1247,266,289]" italics="true" pageId="30" pageNumber="82">r</emphasis>
|
||
does not project beyond distal end of article (
|
||
<figureCitation id="3DAA2A7C3608DA2AFE0CFEC5FDADFEFD" box="[471,587,301,326]" captionStart="FIGURE 13" captionStartId="18.[151,250,1960,1982]" captionTargetBox="[174,1418,904,1940]" captionTargetId="figure-351@18.[174,1418,904,1940]" captionTargetPageId="18" captionText="FIGURE 13. Basitarsus of leg IV. The basitarsi are all enlarged to the same length. Specimens as in Fig. 12." figureDoi="http://doi.org/10.5281/zenodo.4914679" httpUri="https://zenodo.org/record/4914679/files/figure.png" pageId="30" pageNumber="82">Fig. 13 M</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="A52E36F93608DA2AFF1CFEB9FD91FD47" blockId="30.[151,1436,151,979]" pageId="30" pageNumber="82">
|
||
<emphasis id="97E5EAEB3608DA2AFF1CFEB9FEBDFED1" box="[199,347,337,362]" italics="true" pageId="30" pageNumber="82">Opisthosoma</emphasis>
|
||
. Anterior tergites with
|
||
<emphasis id="97E5EAEB3608DA2AFDBAFEBAFD97FED5" box="[609,625,338,366]" italics="true" pageId="30" pageNumber="82">
|
||
t
|
||
<subScript id="391534BC3608DA2AFDB2FE88FD97FED5" attach="left" box="[617,625,352,366]" fontSize="6" pageId="30" pageNumber="82">1</subScript>
|
||
</emphasis>
|
||
,
|
||
<emphasis id="97E5EAEB3608DA2AFDA5FEBAFD68FED5" box="[638,654,338,366]" italics="true" pageId="30" pageNumber="82">
|
||
t
|
||
<subScript id="391534BC3608DA2AFD5DFE88FD68FED5" attach="left" box="[646,654,352,366]" fontSize="6" pageId="30" pageNumber="82">3</subScript>
|
||
</emphasis>
|
||
,
|
||
<emphasis id="97E5EAEB3608DA2AFD47FEBAFD41FED2" box="[668,679,338,361]" italics="true" pageId="30" pageNumber="82">s</emphasis>
|
||
on segment II and
|
||
<emphasis id="97E5EAEB3608DA2AFCA4FEBAFC69FED5" box="[895,911,338,366]" italics="true" pageId="30" pageNumber="82">
|
||
t
|
||
<subScript id="391534BC3608DA2AFC5CFE88FC69FED5" attach="left" box="[903,911,352,366]" fontSize="6" pageId="30" pageNumber="82">1</subScript>
|
||
</emphasis>
|
||
,
|
||
<emphasis id="97E5EAEB3608DA2AFC46FEBAFC4AFED5" box="[925,940,338,366]" italics="true" pageId="30" pageNumber="82">
|
||
t
|
||
<subScript id="391534BC3608DA2AFC7FFE88FC4AFED5" attach="left" box="[932,940,352,366]" fontSize="6" pageId="30" pageNumber="82">2</subScript>
|
||
</emphasis>
|
||
,
|
||
<emphasis id="97E5EAEB3608DA2AFC60FEBAFC2CFED5" box="[955,970,338,366]" italics="true" pageId="30" pageNumber="82">
|
||
t
|
||
<subScript id="391534BC3608DA2AFC19FE88FC2CFED5" attach="left" box="[962,970,352,366]" fontSize="6" pageId="30" pageNumber="82">3</subScript>
|
||
</emphasis>
|
||
,
|
||
<emphasis id="97E5EAEB3608DA2AFC03FEBAFC05FED2" box="[984,995,338,361]" italics="true" pageId="30" pageNumber="82">s</emphasis>
|
||
on III–VI. Sternites IV–VI each with a pair of submedian circular structures and lateral setae
|
||
<emphasis id="97E5EAEB3608DA2AFD2FFE91FCE0FE2E" box="[756,774,377,405]" italics="true" pageId="30" pageNumber="82">
|
||
s
|
||
<subScript id="391534BC3608DA2AFD25FE6FFCE0FE2E" attach="left" box="[766,774,391,405]" fontSize="6" pageId="30" pageNumber="82">1</subScript>
|
||
</emphasis>
|
||
and
|
||
<emphasis id="97E5EAEB3608DA2AFCE7FE91FCA9FE2E" box="[828,847,377,405]" italics="true" pageId="30" pageNumber="82">
|
||
s
|
||
<subScript id="391534BC3608DA2AFC9CFE6FFCA9FE2E" attach="left" box="[839,847,391,405]" fontSize="6" pageId="30" pageNumber="82">2</subScript>
|
||
</emphasis>
|
||
; more than 3+3 submedian setae
|
||
<emphasis id="97E5EAEB3608DA2AFB13FE91FB30FE2B" box="[1224,1238,377,400]" italics="true" pageId="30" pageNumber="82">a</emphasis>
|
||
on at least one of the sternites IV–VI, the maximum observed (N = 3) was 4 per side (
|
||
<figureCitation id="3DAA2A7C3608DA2AFC19FE77FBEAFE03" box="[962,1036,415,440]" captionStart="FIGURE 6" captionStartId="12.[151,250,1190,1212]" captionTargetBox="[213,1378,437,1159]" captionTargetId="figure-245@12.[200,1386,423,1168]" captionTargetPageId="12" captionText="FIGURE 6. Lateral organ and opisthosomal chaetotaxy of Eukoenenia specimens from the six Italian cave populations. The right column shows the values for published specimens and the two comparative females (Ardovská, Griffen). The top line of each box gives the number of blades in the lateral organs (one digit, if symmetric), the three lines below give the numbers of setae a on sternites IV–VI (left ǀ right)." figureDoi="http://doi.org/10.5281/zenodo.4914665" httpUri="https://zenodo.org/record/4914665/files/figure.png" pageId="30" pageNumber="82">Figs 6</figureCitation>
|
||
&
|
||
<figureCitation id="3DAA2A7C3608DA2AFBE3FE77FB9DFE03" box="[1080,1147,415,440]" captionStart="FIGURE 14" captionStartId="19.[151,250,1283,1305]" captionTargetBox="[151,1436,193,1262]" captionTargetId="figure-367@19.[151,1436,193,1262]" captionTargetPageId="19" captionText="FIGURE 14. Setae a on sternite VI. B: Bossea ♂ (E. strinatii), C: Caudano ♂ (E. bonadonai), L: Litrôn ♂ (E. bonadonai), M: Monfieis ♂ (E. lanai sp. n.), P: Partigiano ♀ (E. spelaea), F: Fornaci ♂ (E. roscia sp. n.), A: Ardovská ♀ (E. spelaea with one seta a absent as an individual aberration), G: Griffen ♀ (E. spelaea). Midline = scale bar: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.4914681" httpUri="https://zenodo.org/record/4914681/files/figure.png" pageId="30" pageNumber="82">14 M</figureCitation>
|
||
).
|
||
<taxonomicName id="62914D7A3608DA2AFB49FE48FB09FE03" authorityName="Christian & Isaia & Paschetta & Bruckner" authorityYear="2014" box="[1170,1263,415,440]" class="Arachnida" family="Eukoeneniidae" genus="Eukoenenia" kingdom="Animalia" order="Palpigradi" pageId="30" pageNumber="82" phylum="Arthropoda" rank="species" species="lanai">
|
||
<emphasis id="97E5EAEB3608DA2AFB49FE48FB09FE03" box="[1170,1263,415,440]" italics="true" pageId="30" pageNumber="82">E. lanai</emphasis>
|
||
</taxonomicName>
|
||
has by far the relatively longest setae
|
||
<emphasis id="97E5EAEB3608DA2AFE7DFE2CFE52FE60" box="[422,436,452,475]" italics="true" pageId="30" pageNumber="82">a</emphasis>
|
||
among all populations investigated (see above). The setation of the terminal segments (
|
||
<tableCitation id="E81303423608DA2AFF45FE0FFF11FDBB" box="[158,247,487,512]" captionStart="TABLE 3" captionStartId="16.[151,241,436,459]" captionTargetPageId="16" captionText="TABLE 3. Number of setae on opisthosomal segments VII–XI. C = Caudano; other abbreviations of caves as in Table 1." httpUri="http://table.plazi.org/id/F1EE66713606DA24FF4CFE5CFA7AFE70" pageId="30" pageNumber="82" tableUuid="F1EE66713606DA24FF4CFE5CFA7AFE70">Table 3</tableCitation>
|
||
) is partly variable; segment VII carries 13–17 setae, segment XI has 9 (female) or 10 setae (the
|
||
<specimenCount id="B397FD703608DA2AFAA9FE00FF3BFD9F" pageId="30" pageNumber="82" type="male">two males</specimenCount>
|
||
). The 2 pairs of setae on the intermediate ring of the flagellum are of equal length. In the
|
||
<typeStatus id="7A2A885B3608DA2AFADCFDE3FA8AFD9F" box="[1287,1388,523,548]" pageId="30" pageNumber="82" type="holotype">holotype</typeStatus>
|
||
, the flagellum (detached, under separate coverslip) is preserved from the third article to the tip (
|
||
<figureCitation id="3DAA2A7C3608DA2AFB57FDC7FB39FDF3" box="[1164,1247,559,584]" captionStart="FIGURE 21" captionStartId="25.[151,250,1172,1194]" captionTargetBox="[151,1436,193,1150]" captionTargetId="figure-446@25.[151,1436,193,1150]" captionTargetPageId="25" captionText="FIGURE 21. Flagellum. Left: articles IV–VI of a Monfieis ♂ (E. lanai sp. n.; scale bar: 100 µm). Right: the same articles of a E. mirabilis ♀ (soil-dwelling) from Bergeggi, Liguria, for comparison (scale bar: 20 µm). Note apical crown of cuticular spines on articles III (at the very top of the images) and V." figureDoi="http://doi.org/10.5281/zenodo.4914695" httpUri="https://zenodo.org/record/4914695/files/figure.png" pageId="30" pageNumber="82">Fig. 21</figureCitation>
|
||
). Length (in µm) and number of setae (in brackets; “+” indicates the presence of an apical crown) are for article 3: 217 (10+), 4: 214 (9), 5: 176 (8+), 6: 212 (8), 7: 171 (8+), 8: 227 (8), 9: 184 (8+), 10: 249 (7), 11: 207 (7), 12: 239 (7), 13: 179 (7), 14: 123 (6 and 1 terminal seta). On article 3, the longest seta measures
|
||
<quantity id="62699B1C3608DA2AFC5BFD73FC31FD08" box="[896,983,667,692]" metricMagnitude="-1" metricUnit="m" metricValue="2.77" pageId="30" pageNumber="82" unit="mm" value="277.0">277 µm</quantity>
|
||
, the longest spine of the apical crown
|
||
<quantity id="62699B1C3608DA2AFA5BFD73FF59FD63" metricMagnitude="-2" metricUnit="m" metricValue="6.8" pageId="30" pageNumber="82" unit="mm" value="68.0">68 µm</quantity>
|
||
. Photographs of two live individuals show 14 flagellar articles and the relatively longest flagellum observed in this study (1.9–2.0× trunk length,
|
||
<tableCitation id="E81303423608DA2AFDCFFD0BFD8CFD47" box="[532,618,739,764]" captionStart="TABLE 4" captionStartId="18.[151,239,581,604]" captionTargetBox="[159,1362,696,842]" captionText="TABLE 4. Number of flagellar articles and length ratio trunk/flagellum. Data taken from photographs of live animals (courtesy of Ľubomír Kováč and Enrico Lana). Sex could not be determined in two individuals. Abbreviations of caves as in Table 3." httpUri="http://table.plazi.org/id/F1EE66713604DA26FF4CFDADFEC2FD27" pageId="30" pageNumber="82" tableUuid="F1EE66713604DA26FF4CFDADFEC2FD27">Table 4</tableCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="A52E36F93608DA2AFF1CFCEFFAB8FCFF" blockId="30.[151,1436,151,979]" pageId="30" pageNumber="82">
|
||
<emphasis id="97E5EAEB3608DA2AFF1CFCEFFE4EFCA4" box="[199,424,775,800]" italics="true" pageId="30" pageNumber="82">Female genital area</emphasis>
|
||
. First lobe has 7+3 setae on each side; the innermost of the 3 apical setae measures
|
||
<quantity id="62699B1C3608DA2AFA95FCEFFA71FC9B" box="[1358,1431,775,800]" metricMagnitude="-2" metricUnit="m" metricValue="2.4" pageId="30" pageNumber="82" unit="mm" value="24.0">24 µm</quantity>
|
||
, it is somewhat shorter than the others. Each half of second lobe possesses 3 setae. The total formula is 7+3/3.
|
||
</paragraph>
|
||
<paragraph id="A52E36F93608DA2AFF1CFCA7FF25FC68" blockId="30.[151,1436,151,979]" pageId="30" pageNumber="82">
|
||
<emphasis id="97E5EAEB3608DA2AFF1CFCA7FE77FCDC" box="[199,401,847,872]" italics="true" pageId="30" pageNumber="82">Male genital area</emphasis>
|
||
. Chaetotaxy (2+9+2f/3/4) and general shape of genital lobes as in
|
||
<taxonomicName id="62914D7A3608DA2AFB50FCB8FAE2FCDC" authorityName="Conde" authorityYear="1977" box="[1163,1284,848,871]" class="Arachnida" family="Eukoeneniidae" genus="Eukoenenia" kingdom="Animalia" order="Palpigradi" pageId="30" pageNumber="82" phylum="Arthropoda" rank="species" species="strinatii">
|
||
<emphasis id="97E5EAEB3608DA2AFB50FCB8FAE2FCDC" box="[1163,1284,848,871]" italics="true" pageId="30" pageNumber="82">E. strinatii</emphasis>
|
||
</taxonomicName>
|
||
(cf.
|
||
<figureCitation id="3DAA2A7C3608DA2AFAE0FCA7FA69FCD3" box="[1339,1423,847,872]" captionStart="FIGURE 19" captionStartId="24.[151,250,1249,1271]" captionTargetBox="[151,1436,193,1227]" captionTargetId="figure-105@24.[151,1436,193,1227]" captionTargetPageId="24" captionText="FIGURE 19. Male genitalia in ventral view. One side of the first lobe (left), tips of the second lobe (above), and of the third lobe (below) of a E. strinatii ♂ from Bossea. Fusules are marked f1, f2. Scale bar: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.4914691" httpUri="https://zenodo.org/record/4914691/files/figure.png" pageId="30" pageNumber="82">Fig. 19</figureCitation>
|
||
). The fusules, however, are inserted on dome-shaped bases (
|
||
<figureCitation id="3DAA2A7C3608DA2AFCE5FC9BFC75FC37" box="[830,915,883,908]" captionStart="FIGURE 20" captionStartId="24.[151,250,1942,1964]" captionTargetBox="[415,1171,1354,1921]" captionTargetId="figure-114@24.[415,1171,1354,1921]" captionTargetPageId="24" captionText="FIGURE 20. Fusules (f1, f2) on first genital lobe of a male from Monfieis (E. lanai sp. n.). The fusules are inserted on domeshaped bases. Scale bar: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.4914693" httpUri="https://zenodo.org/record/4914693/files/figure.png" pageId="30" pageNumber="82">Fig. 20</figureCitation>
|
||
), a character that separates
|
||
<taxonomicName id="62914D7A3608DA2AFB0FFC9CFAC9FC37" authorityName="Christian & Isaia & Paschetta & Bruckner" authorityYear="2014" box="[1236,1327,883,908]" class="Arachnida" family="Eukoeneniidae" genus="Eukoenenia" kingdom="Animalia" order="Palpigradi" pageId="30" pageNumber="82" phylum="Arthropoda" rank="species" species="lanai">
|
||
<emphasis id="97E5EAEB3608DA2AFB0FFC9CFAC9FC37" box="[1236,1327,883,908]" italics="true" pageId="30" pageNumber="82">E. lanai</emphasis>
|
||
</taxonomicName>
|
||
from the other species of the present study. In the
|
||
<typeStatus id="7A2A885B3608DA2AFD84FC7FFD22FC0B" box="[607,708,919,944]" pageId="30" pageNumber="82" type="holotype">holotype</typeStatus>
|
||
, the longest seta on the first lobe measures
|
||
<quantity id="62699B1C3608DA2AFB71FC7FFB13FC0B" box="[1194,1269,919,944]" metricMagnitude="-2" metricUnit="m" metricValue="9.8" pageId="30" pageNumber="82" unit="mm" value="98.0">98 µm</quantity>
|
||
, the fusules
|
||
<quantity id="62699B1C3608DA2AFA5BFC7FFF59FC68" metricMagnitude="-2" metricUnit="m" metricValue="6.0" pageId="30" pageNumber="82" unit="mm" value="60.0">60 µm</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |