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<document id="2C322DCED1C2CCC477F92DE7140EB334" ID-CLB-Dataset="24105" ID-DOI="10.5852/ejt.2019.585" ID-GBIF-Dataset="dca608e8-fccf-4c1c-b8df-ef0c28e1d518" ID-Zenodo-Dep="3660259" ID-ZooBank="181B6FBA-ED75-4BB4-84C4-FB512B794749" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1581251975682" checkinUser="plazi" docAuthor="Ebersole, Jun A., Cicimurri, David J. &amp; Stringer, Gary L." docDate="2019" docId="18174D41FFB6FF9BFD8A9D7B481C0A3E" docLanguage="en" docName="ejt-585_ebersole_cicimurri_stinger.pdf.imf" docOrigin="European Journal of Taxonomy 585" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Odontaspis winkleri Leriche 1905" docType="treatment" docVersion="18" lastPageNumber="53" masterDocId="E42E3539FF85FFAFFFC69F344D4B0F37" masterDocTitle="Taxonomy and biostratigraphy of the elasmobranchs and bony fishes (Chondrichthyes and Osteichthyes) of the lower-to-middle Eocene (Ypresian to Bartonian) Claiborne Group in Alabama, USA, including an analysis of otoliths" masterLastPageNumber="274" masterPageNumber="1" pageNumber="52" updateTime="1698783570407" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="479DFBBF7FB3D1DE1A9F042BE6885752">Taxonomy and biostratigraphy of the elasmobranchs and bony fishes (Chondrichthyes and Osteichthyes) of the lower-to-middle Eocene (Ypresian to Bartonian) Claiborne Group in Alabama, USA, including an analysis of otoliths</mods:title>
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<mods:namePart id="8069236753A810D55864741C124447BC">Ebersole, Jun A.</mods:namePart>
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<mods:namePart id="56DCA829AEC5BAEB3AD6079F8A0CB598">Cicimurri, David J.</mods:namePart>
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<taxonomicName id="57BE87D4FFB6FF9CFD8A9D7B4EA30D5D" authority="Leriche, 1905" authorityName="Leriche" authorityYear="1905" box="[588,1000,591,618]" class="Elasmobranchii" family="Odontaspididae" genus="Odontaspis" kingdom="Animalia" order="Lamniformes" pageId="51" pageNumber="52" phylum="Chordata" rank="species" species="winkleri">
<emphasis id="A2CA2045FFB6FF9CFD8A9D7B4E760D5D" bold="true" box="[588,829,591,618]" italics="true" pageId="51" pageNumber="52">Odontaspis winkleri</emphasis>
<bibRefCitation id="F42F81A6FFB6FF9CFC839D7B4EA30D5D" author="Leriche M." box="[837,1000,591,618]" pageId="51" pageNumber="52" pagination="49 - 228" refId="ref144901" refString="Leriche M. 1905. Les poissons eocenes de la Belgique. Memoires du Musee royal d'Histoire naturelle de Belgique 3 (11): 49 - 228." type="journal article" year="1905">Leriche, 1905</bibRefCitation>
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</heading>
</paragraph>
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<figureCitation id="0885E0D2FFB6FF9CFD369D474E080DBA" box="[752,835,627,653]" captionStart="Fig" captionStartId="51.[189,232,1861,1887]" captionTargetBox="[201,1389,1224,1820]" captionTargetId="figure@51.[189,1398,1216,1837]" captionTargetPageId="51" captionText="Fig. 18. Odontaspis winkleri Leriche, 1905, teeth. AC. MSC 33380, anterior tooth, lower Tallahatta Formation. A. Labial view. B. Lingual view. C. Mesial view. DF. SC2012.47.162, anterior tooth, basal Lisbon Formation. D. Labial view. E. Lingual view. F. Mesial view. GI. MSC 35764, lateral tooth, lower Tallahatta Formation. G. Labial view. H. Lingual view. I. Mesial view. JL. WSU CC535.1, lateral tooth, basal Lisbon Formation. J. Labial view. K. Lingual view. L. Mesial view. Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3660295" httpUri="https://zenodo.org/record/3660295/files/figure.png" pageId="51" pageNumber="52">Fig. 18</figureCitation>
</paragraph>
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<subSubSection id="D8A4AFDCFFB6FF9CFF7B9D804E6D0C60" pageId="51" pageNumber="52" type="reference_group">
<paragraph id="9001FC57FFB6FF9CFF7B9D804E330DF9" blockId="51.[189,888,692,719]" box="[189,888,692,719]" pageId="51" pageNumber="52">
<treatmentCitationGroup id="B0AEDB79FFB6FF9CFF7B9D804E330DF9" box="[189,888,692,719]" pageId="51" pageNumber="52">
<taxonomicName id="57BE87D4FFB6FF9CFF7B9D804CE10DF9" authority="Ebersole &amp; Cicimurri &amp; Stringer, 2019" authorityName="Ebersole &amp; Cicimurri &amp; Stringer" authorityYear="2019" box="[189,426,692,718]" class="Elasmobranchii" family="Odontaspididae" genus="Odontaspis" kingdom="Animalia" order="Lamniformes" pageId="51" pageNumber="52" phylum="Chordata" rank="species" species="winkleri" status="sp. nov.">
<emphasis id="A2CA2045FFB6FF9CFF7B9D804CE10DF9" box="[189,426,692,718]" italics="true" pageId="51" pageNumber="52">Odontaspis winkleri</emphasis>
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<taxonomicNameLabel id="B9F99D3EFFB6FF9CFE779D814F470DF8" box="[433,524,693,719]" pageId="51" pageNumber="52" rank="species">sp. nov.</taxonomicNameLabel>
<treatmentCitation id="111FDA46FFB6FF9CFDD59D804FAE0DF9" author="Leriche M." box="[531,741,692,719]" page="74" pageId="51" pageNumber="52" year="1905">
<bibRefCitation id="F42F81A6FFB6FF9CFDD59D804FAE0DF9" author="Leriche M." box="[531,741,692,719]" pageId="51" pageNumber="52" pagination="49 - 228" refId="ref144901" refString="Leriche M. 1905. Les poissons eocenes de la Belgique. Memoires du Musee royal d'Histoire naturelle de Belgique 3 (11): 49 - 228." type="journal article" year="1905">Leriche, 1905: 74</bibRefCitation>
</treatmentCitation>
, pl. 6, fig. 8.
</treatmentCitationGroup>
</paragraph>
<paragraph id="9001FC57FFB6FF9CFF7B9DC149370C27" blockId="51.[189,1148,757,855]" box="[189,1148,757,784]" pageId="51" pageNumber="52">
<treatmentCitationGroup id="B0AEDB79FFB6FF9CFF7B9DC149370C27" box="[189,1148,757,784]" pageId="51" pageNumber="52">
<taxonomicName id="57BE87D4FFB6FF9CFF7B9DC14ED70C27" authority="- Arambourg 1935: 425" authorityName="Arambourg" authorityPageNumber="425" authorityYear="1935" box="[189,924,757,784]" class="Elasmobranchii" family="Odontaspididae" genus="Odontaspis" kingdom="Animalia" order="Lamniformes" pageId="51" pageNumber="52" phylum="Chordata" rank="species" species="winkleri" subGenus="Odontaspis">
<emphasis id="A2CA2045FFB6FF9CFF7B9DC14C080C38" box="[189,323,757,783]" italics="true" pageId="51" pageNumber="52">Odontaspis</emphasis>
(
<emphasis id="A2CA2045FFB6FF9CFE929DC14C910C38" box="[340,474,757,783]" italics="true" pageId="51" pageNumber="52">Odontaspis</emphasis>
) aff.
<emphasis id="A2CA2045FFB6FF9CFDDF9DC14F320C38" box="[537,633,757,783]" italics="true" pageId="51" pageNumber="52">winkleri</emphasis>
<treatmentCitation id="111FDA46FFB6FF9CFD539DC14ED70C27" author="Arambourg C." box="[661,924,757,784]" page="425" pageId="51" pageNumber="52" year="1935">
<bibRefCitation id="F42F81A6FFB6FF9CFD539DC14ED70C27" author="Arambourg C." box="[661,924,757,784]" pageId="51" pageNumber="52" pagination="413 - 439" refId="ref136151" refString="Arambourg C. 1935. Note preliminaire sur les vertebres fossiles des phosphates du Maroc. Bulletin de la Societe geologique de France 5 (5): 413 - 439." type="journal article" year="1935">Arambourg 1935: 425</bibRefCitation>
</treatmentCitation>
</taxonomicName>
, pl. 29, figs 2022.
</treatmentCitationGroup>
</paragraph>
<paragraph id="9001FC57FFB6FF9CFF7B9C2D4F960C04" blockId="51.[189,1148,757,855]" box="[189,733,792,819]" pageId="51" pageNumber="52">
<emphasis id="A2CA2045FFB6FF9CFF7B9C2D4C3E0C04" box="[189,373,793,819]" italics="true" pageId="51" pageNumber="52">Synodontaspis?</emphasis>
<taxonomicName id="57BE87D4FFB6FF9CFEBA9C2D4F910C04" authority="- Herman 1977: 245" authorityName="Herman" authorityPageNumber="245" authorityYear="1977" box="[380,730,792,819]" class="Elasmobranchii" family="Odontaspididae" genus="Odontaspis" kingdom="Animalia" order="Lamniformes" pageId="51" pageNumber="52" phylum="Chordata" rank="species" species="winkleri">
<emphasis id="A2CA2045FFB6FF9CFEBA9C2D4C970C04" box="[380,476,793,819]" italics="true" pageId="51" pageNumber="52">winkleri</emphasis>
Herman 1977: 245
</taxonomicName>
.
</paragraph>
<paragraph id="9001FC57FFB6FF9CFF7B9C084E6D0C60" blockId="51.[189,1148,757,855]" box="[189,806,828,855]" pageId="51" pageNumber="52">
<taxonomicName id="57BE87D4FFB6FF9CFF7B9C084E690C60" authority="- Krukow &amp; Thies 1990: 35" authorityName="Krukow &amp; Thies" authorityPageNumber="35" authorityYear="1990" box="[189,802,828,855]" class="Elasmobranchii" family="Odontaspididae" genus="Eugomphodus" higherTaxonomySource="CoL" kingdom="Animalia" order="Lamniformes" pageId="51" pageNumber="52" phylum="Chordata" rank="species" species="winkleri">
<emphasis id="A2CA2045FFB6FF9CFF7B9C084C870C61" box="[189,460,828,854]" italics="true" pageId="51" pageNumber="52">Eugomphodus winkleri</emphasis>
Krukow &amp; Thies 1990: 35
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="D8A4AFDCFFB6FF9CFF7B9C494E5B0CDD" pageId="51" pageNumber="52" type="materials_examined">
<paragraph id="9001FC57FFB6FF9CFF7B9C494CE10CA0" blockId="51.[189,426,893,919]" box="[189,426,893,919]" pageId="51" pageNumber="52">
<heading id="CB494B3BFFB6FF9CFF7B9C494CE10CA0" bold="true" box="[189,426,893,919]" fontSize="11" level="3" pageId="51" pageNumber="52" reason="3">
<emphasis id="A2CA2045FFB6FF9CFF7B9C494CE10CA0" bold="true" box="[189,426,893,919]" pageId="51" pageNumber="52">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="9001FC57FFB6FF9CFF7B9C984E5B0CDD" blockId="51.[189,1398,940,1002]" pageId="51" pageNumber="52">
<collectingCountry id="E8A9BCC7FFB6FF9CFF7B9C984F060CF1" box="[189,589,940,966]" name="United States of America" pageId="51" pageNumber="52">UNITED STATES OF AMERICA</collectingCountry>
<emphasis id="A2CA2045FFB6FF9CFDAE9C984F910CF1" bold="true" box="[616,730,940,966]" pageId="51" pageNumber="52">
<collectingRegion id="527A32B5FFB6FF9CFDAE9C984F910CF1" box="[616,730,940,966]" country="United States of America" name="Alabama" pageId="51" pageNumber="52">Alabama</collectingRegion>
</emphasis>
• 5 isolated teeth; Claiborne Group;
<collectionCode id="F6AF6492FFB6FF9CFB459C9849ED0CF1" box="[1155,1190,940,966]" pageId="51" pageNumber="52">SC</collectionCode>
2012.47.92,
<collectionCode id="F6AF6492FFB6FF9CFAFE9C98483D0CF1" box="[1336,1398,940,966]" pageId="51" pageNumber="52">MSC</collectionCode>
33380,
<collectionCode id="F6AF6492FFB6FF9CFED39CFB4C180CDE" box="[277,339,975,1001]" pageId="51" pageNumber="52">MSC</collectionCode>
35764,
<collectionCode id="F6AF6492FFB6FF9CFE749CFB4CBB0CDE" box="[434,496,975,1001]" pageId="51" pageNumber="52">MSC</collectionCode>
38477,
<collectionCode id="F6AF6492FFB6FF9CFD899CFB4FDB0CDE" box="[591,656,975,1001]" country="USA" httpUri="http://grbio.org/cool/1raa-g3is" name="Weber State University, Bird and Mammal Collection" pageId="51" pageNumber="52">WSU</collectionCode>
CC 535.1.
</paragraph>
</subSubSection>
<subSubSection id="D8A4AFDCFFB6FF9BFF7B9B244EF90EE1" lastPageId="52" lastPageNumber="53" pageId="51" pageNumber="52" type="description">
<paragraph id="9001FC57FFB6FF9CFF7B9B244C050B1D" blockId="51.[189,334,1040,1066]" box="[189,334,1040,1066]" pageId="51" pageNumber="52">
<heading id="CB494B3BFFB6FF9CFF7B9B244C050B1D" bold="true" box="[189,334,1040,1066]" fontSize="11" level="3" pageId="51" pageNumber="52" reason="3">
<emphasis id="A2CA2045FFB6FF9CFF7B9B244C050B1D" bold="true" box="[189,334,1040,1066]" pageId="51" pageNumber="52">Description</emphasis>
</heading>
</paragraph>
<paragraph id="9001FC57FFB6FF9BFF7B9B0B4EF90EE1" blockId="51.[189,1399,1087,1184]" lastBlockId="52.[189,1399,267,470]" lastPageId="52" lastPageNumber="53" pageId="51" pageNumber="52">Teeth small, generally not exceeding 1.0 cm in overall height. Upper teeth slightly sigmoidal; lower teeth with slight lingual bend. Teeth with tall and thin cusp, with characteristically tall, erect, conical and sharply pointed lateral cusplets. Lingual face of main cusp strongly convex; labial face may be nearly flat to convex (particularly at the base). Mesial and distal cutting edges absent or restricted to the upper two-thirds of the main cusp. Lingual and labial cusp faces of anterior teeth smooth, but lateral teeth with distinct folding at labial crown foot. Anterior teeth with single pair of lateral cusplets; lateral teeth with two to three pairs of lateral cusplets. Cusplets divergent and decrease in size laterally. Root bilobate with long, thin, divergent, and rounded lobes; lobes separated by deep U-shaped interlobe area. Deep nutritive groove located on prominent lingual root protuberance.</paragraph>
</subSubSection>
<caption id="C4C1ACDFFFB6FF9CFF7B9871481908DB" ID-DOI="http://doi.org/10.5281/zenodo.3660295" ID-Zenodo-Dep="3660295" httpUri="https://zenodo.org/record/3660295/files/figure.png" pageId="51" pageNumber="52" startId="51.[189,232,1861,1887]" targetBox="[201,1389,1224,1820]" targetPageId="51">
<paragraph id="9001FC57FFB6FF9CFF7B9871481908DB" blockId="51.[189,1398,1861,2029]" pageId="51" pageNumber="52">
<emphasis id="A2CA2045FFB6FF9CFF7B98714C500868" bold="true" box="[189,283,1861,1887]" pageId="51" pageNumber="52">Fig. 18.</emphasis>
<taxonomicName id="57BE87D4FFB6FF9CFEE298714F8F0857" authority="Leriche, 1905" authorityName="Leriche" authorityYear="1905" box="[292,708,1861,1888]" class="Elasmobranchii" family="Odontaspididae" genus="Odontaspis" kingdom="Animalia" order="Lamniformes" pageId="51" pageNumber="52" phylum="Chordata" rank="species" species="winkleri">
<emphasis id="A2CA2045FFB6FF9CFEE298714F580868" box="[292,531,1861,1887]" italics="true" pageId="51" pageNumber="52">Odontaspis winkleri</emphasis>
<bibRefCitation id="F42F81A6FFB6FF9CFDDB98714F8F0857" author="Leriche M." box="[541,708,1861,1888]" pageId="51" pageNumber="52" pagination="49 - 228" refId="ref144901" refString="Leriche M. 1905. Les poissons eocenes de la Belgique. Memoires du Musee royal d'Histoire naturelle de Belgique 3 (11): 49 - 228." type="journal article" year="1905">Leriche, 1905</bibRefCitation>
</taxonomicName>
, teeth.
<emphasis id="A2CA2045FFB6FF9CFCDA98714E1E0868" bold="true" box="[796,853,1861,1887]" pageId="51" pageNumber="52">AC</emphasis>
.
<collectionCode id="F6AF6492FFB6FF9CFCA398714EE80868" box="[869,931,1861,1887]" pageId="51" pageNumber="52">MSC</collectionCode>
33380, anterior tooth, lower Tallahatta Formation.
<emphasis id="A2CA2045FFB6FF9CFE83985C4C1108B5" bold="true" box="[325,346,1896,1922]" pageId="51" pageNumber="52">A</emphasis>
. Labial view.
<emphasis id="A2CA2045FFB6FF9CFE39985C4F5808B5" bold="true" box="[511,531,1896,1922]" pageId="51" pageNumber="52">B</emphasis>
. Lingual view.
<emphasis id="A2CA2045FFB6FF9CFD01985C4F9708B5" bold="true" box="[711,732,1896,1922]" pageId="51" pageNumber="52">C</emphasis>
. Mesial view.
<emphasis id="A2CA2045FFB6FF9CFC40985C4EF708B5" bold="true" box="[902,956,1896,1922]" pageId="51" pageNumber="52">DF</emphasis>
.
<collectionCode id="F6AF6492FFB6FF9CFC0C985C4EA608B5" box="[970,1005,1896,1922]" pageId="51" pageNumber="52">SC</collectionCode>
2012.47.162, anterior tooth, basal Lisbon Formation.
<emphasis id="A2CA2045FFB6FF9CFE6598B84CF30891" bold="true" box="[419,440,1932,1958]" pageId="51" pageNumber="52">D</emphasis>
. Labial view.
<emphasis id="A2CA2045FFB6FF9CFDA098B84F310891" bold="true" box="[614,634,1932,1958]" pageId="51" pageNumber="52">E</emphasis>
. Lingual view.
<emphasis id="A2CA2045FFB6FF9CFCFE98B84E010891" bold="true" box="[824,842,1932,1958]" pageId="51" pageNumber="52">F</emphasis>
. Mesial view.
<emphasis id="A2CA2045FFB6FF9CFC3A98BF49660892" bold="true" box="[1020,1069,1931,1957]" pageId="51" pageNumber="52">GI</emphasis>
.
<collectionCode id="F6AF6492FFB6FF9CFBF898B849370891" box="[1086,1148,1932,1958]" pageId="51" pageNumber="52">MSC</collectionCode>
35764, lateral tooth, lower Tallahatta Formation.
<emphasis id="A2CA2045FFB6FF9CFDC2989A4F5008FF" bold="true" box="[516,539,1966,1992]" pageId="51" pageNumber="52">G</emphasis>
. Labial view.
<emphasis id="A2CA2045FFB6FF9CFD7D989B4F9908FE" bold="true" box="[699,722,1967,1993]" pageId="51" pageNumber="52">H</emphasis>
. Lingual view.
<emphasis id="A2CA2045FFB6FF9CFC45989B4EC508FE" bold="true" box="[899,910,1967,1993]" pageId="51" pageNumber="52">I</emphasis>
. Mesial view.
<emphasis id="A2CA2045FFB6FF9CFBF2989B492E08FE" bold="true" box="[1076,1125,1967,1993]" pageId="51" pageNumber="52">JL</emphasis>
.
<collectionCode id="F6AF6492FFB6FF9CFBB7989B49F908FE" box="[1137,1202,1967,1993]" pageId="51" pageNumber="52">WSU</collectionCode>
CC535.1, lateral tooth, basal Lisbon Formation.
<emphasis id="A2CA2045FFB6FF9CFDEB98E64F7708DB" bold="true" box="[557,572,2002,2028]" pageId="51" pageNumber="52">J</emphasis>
. Labial view.
<emphasis id="A2CA2045FFB6FF9CFD2598E64FB108DB" bold="true" box="[739,762,2002,2028]" pageId="51" pageNumber="52">K</emphasis>
. Lingual view.
<emphasis id="A2CA2045FFB6FF9CFC7698E64E8F08DB" bold="true" box="[944,964,2002,2028]" pageId="51" pageNumber="52">L</emphasis>
. Mesial view. Scale bars = 5 mm.
</paragraph>
</caption>
<subSubSection id="D8A4AFDCFFB1FF9BFF7B9EC9481C0A3E" pageId="52" pageNumber="53" type="discussion">
<paragraph id="9001FC57FFB1FF9BFF7B9EC94C640D20" blockId="52.[189,303,509,535]" box="[189,303,509,535]" pageId="52" pageNumber="53">
<heading id="CB494B3BFFB1FF9BFF7B9EC94C640D20" bold="true" box="[189,303,509,535]" fontSize="11" level="3" pageId="52" pageNumber="53" reason="3">
<emphasis id="A2CA2045FFB1FF9BFF7B9EC94C640D20" bold="true" box="[189,303,509,535]" pageId="52" pageNumber="53">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="9001FC57FFB1FF9BFF7B9D184F890CFD" blockId="52.[189,1399,556,970]" pageId="52" pageNumber="53">
Three species of Paleogene
<taxonomicName id="57BE87D4FFB1FF9BFE3D9D184FCA0D71" authorityName="Agassiz" authorityYear="1843" box="[507,641,556,582]" class="Elasmobranchii" family="Odontaspididae" genus="Odontaspis" kingdom="Animalia" order="Lamniformes" pageId="52" pageNumber="53" phylum="Chordata" rank="genus">
<emphasis id="A2CA2045FFB1FF9BFE3D9D184FCA0D71" box="[507,641,556,582]" italics="true" pageId="52" pageNumber="53">Odontaspis</emphasis>
</taxonomicName>
have been recognized in North America including
<taxonomicName id="57BE87D4FFB1FF9BFB019D184C9D0D5E" authority="Case &amp; Borodin, 2000" authorityName="Case &amp; Borodin" authorityYear="2000" class="Elasmobranchii" family="Odontaspididae" genus="Odontaspis" kingdom="Animalia" order="Lamniformes" pageId="52" pageNumber="53" phylum="Chordata" rank="species" species="carolinensis">
<emphasis id="A2CA2045FFB1FF9BFB019D18483D0D71" box="[1223,1398,556,582]" italics="true" pageId="52" pageNumber="53">O. carolinensis</emphasis>
Case &amp; Borodin, 2000
</taxonomicName>
,
<taxonomicName id="57BE87D4FFB1FF9BFE209D644EF80D5E" authority="Dartevelle &amp; Casier, 1943" authorityName="Dartevelle &amp; Casier" authorityYear="1943" box="[486,947,591,618]" class="Elasmobranchii" family="Odontaspididae" genus="Odontaspis" kingdom="Animalia" order="Lamniformes" pageId="52" pageNumber="53" phylum="Chordata" rank="species" species="speyeri">
<emphasis id="A2CA2045FFB1FF9BFE209D644F2F0D5E" box="[486,612,592,618]" italics="true" pageId="52" pageNumber="53">O. speyeri</emphasis>
<bibRefCitation id="F42F81A6FFB1FF9BFDA99D7B4EF80D5E" author="Dartevelle E. &amp; Casier E." box="[623,947,591,617]" pageId="52" pageNumber="53" pagination="1 - 200" refId="ref140771" refString="Dartevelle E. &amp; Casier E. 1943. Les Poissons fossils du Bas-Congo et des regions voisines. Annals of the Museum of the Congo belge Series A (Mineralogie Geologie, Paleontologie, Serie 3 2 (2): 1 - 200." type="journal article" year="1943">Dartevelle &amp; Casier, 1943</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="57BE87D4FFB1FF9BFC3C9D6448720D5D" authority="Leriche, 1905" authorityName="Leriche" authorityYear="1905" box="[1018,1337,591,618]" class="Elasmobranchii" family="Odontaspididae" genus="Odontaspis" kingdom="Animalia" order="Lamniformes" pageId="52" pageNumber="53" phylum="Chordata" rank="species" species="winkleri">
<emphasis id="A2CA2045FFB1FF9BFC3C9D6449C90D5E" box="[1018,1154,591,618]" italics="true" pageId="52" pageNumber="53">O. winkleri</emphasis>
<bibRefCitation id="F42F81A6FFB1FF9BFB489D7B48720D5D" author="Leriche M." box="[1166,1337,591,618]" pageId="52" pageNumber="53" pagination="49 - 228" refId="ref144901" refString="Leriche M. 1905. Les poissons eocenes de la Belgique. Memoires du Musee royal d'Histoire naturelle de Belgique 3 (11): 49 - 228." type="journal article" year="1905">Leriche, 1905</bibRefCitation>
</taxonomicName>
. The
<taxonomicName id="57BE87D4FFB1FF9BFF7B9D464C080DBB" authorityName="Agassiz" authorityYear="1843" box="[189,323,626,652]" class="Elasmobranchii" family="Odontaspididae" genus="Odontaspis" kingdom="Animalia" order="Lamniformes" pageId="52" pageNumber="53" phylum="Chordata" rank="genus">
<emphasis id="A2CA2045FFB1FF9BFF7B9D464C080DBB" box="[189,323,626,652]" italics="true" pageId="52" pageNumber="53">Odontaspis</emphasis>
</taxonomicName>
teeth in our sample differ from those of
<taxonomicName id="57BE87D4FFB1FF9BFCEE9D474E900DBB" authorityName="Case &amp; Borodin" authorityYear="2000" box="[808,987,626,653]" class="Elasmobranchii" family="Odontaspididae" genus="Odontaspis" kingdom="Animalia" order="Lamniformes" pageId="52" pageNumber="53" phylum="Chordata" rank="species" species="carolinensis">
<emphasis id="A2CA2045FFB1FF9BFCEE9D474E900DBB" box="[808,987,626,653]" italics="true" pageId="52" pageNumber="53">O. carolinensis</emphasis>
</taxonomicName>
by having a less robust main cusp on the anterior teeth and cylindrical, not labiolingually flattened, lateral cusplets on the lateral teeth. The teeth of
<taxonomicName id="57BE87D4FFB1FF9BFE989D8D4C900DE4" authorityName="Dartevelle &amp; Casier" authorityYear="1943" box="[350,475,697,723]" class="Elasmobranchii" family="Odontaspididae" genus="Odontaspis" kingdom="Animalia" order="Lamniformes" pageId="52" pageNumber="53" phylum="Chordata" rank="species" species="speyeri">
<emphasis id="A2CA2045FFB1FF9BFE989D8D4C900DE4" box="[350,475,697,723]" italics="true" pageId="52" pageNumber="53">O. speyeri</emphasis>
</taxonomicName>
are much more robust and have smaller cusplets than those in our sample, and
<bibRefCitation id="F42F81A6FFB1FF9BFF379DE84CFB0DC1" author="Cappetta H." box="[241,432,732,758]" pageId="52" pageNumber="53" refId="ref137948" refString="Cappetta H. 2012. Chondrichthyes (Mesozoic and Cenozoic Elasmobranchii, Teeth). Handbook of Palaeoichthyology. Verlag Friedrich Pfeil, Munchen." type="book" year="2012">Cappetta (2012)</bibRefCitation>
referred this species to
<taxonomicName id="57BE87D4FFB1FF9BFD149DE84E200DC1" authorityName="Menner" authorityYear="1928" box="[722,875,732,758]" class="Chondrichthyes" genus="Jaekelotodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lamniformes" pageId="52" pageNumber="53" phylum="Chordata" rank="genus">
<emphasis id="A2CA2045FFB1FF9BFD149DE84E200DC1" box="[722,875,732,758]" italics="true" pageId="52" pageNumber="53">Jaekelotodus</emphasis>
</taxonomicName>
. Although
<bibRefCitation id="F42F81A6FFB1FF9BFC359DE848470DC1" author="Holman J. A. &amp; Case G. R." box="[1011,1292,732,758]" pageId="52" pageNumber="53" pagination="328 - 333" refId="ref143500" refString="Holman J. A. &amp; Case G. R. 1988. Reptiles from the Eocene Tallahatta Formation of Alabama. Journal of Vertebrate Paleontology 8 (3): 328 - 333. https: // doi. org / 10.1080 / 02724634.1988.10011716" type="journal article" year="1988">Holman &amp; Case (1988)</bibRefCitation>
reported
<taxonomicName id="57BE87D4FFB1FF9BFF7B9C344C720C2E" authorityName="Dartevelle &amp; Casier" authorityYear="1943" box="[189,313,768,794]" class="Elasmobranchii" family="Odontaspididae" genus="Odontaspis" kingdom="Animalia" order="Lamniformes" pageId="52" pageNumber="53" phylum="Chordata" rank="species" species="speyeri">
<emphasis id="A2CA2045FFB1FF9BFF7B9C344C720C2E" box="[189,313,768,794]" italics="true" pageId="52" pageNumber="53">O. speyeri</emphasis>
</taxonomicName>
from the ACov-11 locality, this was likely a misidentification as no such teeth have been identified within our exceptionally large sample of teeth from this locality, nor have they been reported by
<bibRefCitation id="F42F81A6FFB1FF9BFF219C724CA00C57" author="Clayton A. A. &amp; Ciampaglio, C. N. &amp; Cicimurri, D. J." box="[231,491,838,864]" pageId="52" pageNumber="53" pagination="60 - 73" refId="ref139906" refString="Clayton A. A., Ciampaglio, C. N. &amp; Cicimurri, D. J. 2013. An inquiry into the stratigraphic occurrence of a Claibornian (Eocene) vertebrate fauna from Covington County, Alabama. Bulletin Alabama Museum of Natural History 31 (2): 60 - 73." type="journal article" year="2013">
Clayton
<emphasis id="A2CA2045FFB1FF9BFE979C734CDA0C57" box="[337,401,838,864]" italics="true" pageId="52" pageNumber="53">et al.</emphasis>
(2013)
</bibRefCitation>
or
<bibRefCitation id="F42F81A6FFB1FF9BFDDB9C724E050C56" author="Cappetta H. &amp; Case G. R." box="[541,846,838,865]" pageId="52" pageNumber="53" pagination="43 - 103" refId="ref137973" refString="Cappetta H. &amp; Case G. R. 2016. A selachian fauna from the middle Eocene (Lutetian, Lisbon Formation) of Andalusia, Covington County, Alabama, USA. Palaeontographica Abteilung A 307 (1 - 6): 43 - 103." type="journal article" year="2016">Cappetta &amp; Case (2016)</bibRefCitation>
. Furthermore,
<taxonomicName id="57BE87D4FFB1FF9BFBC29C7249C90C57" authorityName="Dartevelle &amp; Casier" authorityYear="1943" box="[1028,1154,838,864]" class="Elasmobranchii" family="Odontaspididae" genus="Odontaspis" kingdom="Animalia" order="Lamniformes" pageId="52" pageNumber="53" phylum="Chordata" rank="species" species="speyeri">
<emphasis id="A2CA2045FFB1FF9BFBC29C7249C90C57" box="[1028,1154,838,864]" italics="true" pageId="52" pageNumber="53">O. speyeri</emphasis>
</taxonomicName>
is a taxon that has generally been reported from Paleocene deposits elsewhere (see
<bibRefCitation id="F42F81A6FFB1FF9BFC779C5D49160CB3" author="Siverson M." box="[945,1117,873,900]" pageId="52" pageNumber="53" pagination="1 - 12" refId="ref150420" refString="Siverson M. 1995. Revision of the Danian cow sharks, sand tiger sharks, and goblin sharks (Hexanchidae, Odontaspididae, and Mitsukurinidae) from southern Sweden. Journal of Vertebrate Paleontology 15 (1): 1 - 12. https: // doi. org / 10.1080 / 02724634.1995.10011203" type="journal article" year="1995">Siverson 1995</bibRefCitation>
;
<bibRefCitation id="F42F81A6FFB1FF9BFBA19C5D48390CB4" author="Yarkov A. A. &amp; Popov E. V." box="[1127,1394,873,900]" pageId="52" pageNumber="53" pagination="59 - 65" refId="ref154505" refString="Yarkov A. A. &amp; Popov E. V. 1998. [A new chondrichthyan fauna from the Beriozovaya beds (lower Paleocene) of the Volgograd Volga River Basin, preliminary data]. Vopr. paleontologii i stratigrafii 1998 (1): 59 - 65. [In Russian.]" type="journal article" year="1998">Yarkov &amp; Popov 1998</bibRefCitation>
;
<bibRefCitation id="F42F81A6FFB1FF9BFF7B9CB84CA90C90" author="Adolfssen J. S. &amp; Ward D. J." box="[189,482,908,935]" pageId="52" pageNumber="53" pagination="313 - 338" refId="ref135956" refString="Adolfssen J. S. &amp; Ward D. J. 2015. Neoselachians from the Danian (early Paleocene) of Denmark. Acta Palaeontologica Polonica 60 (2): 313 - 338. https: // doi. org / 10.4202 / app. 2012.0123" type="journal article" year="2015">Adolfssen &amp; Ward 2015</bibRefCitation>
). Unfortunately,
<bibRefCitation id="F42F81A6FFB1FF9BFD6F9CB84EF40C90" author="Holman J. A. &amp; Case G. R." box="[681,959,908,935]" pageId="52" pageNumber="53" pagination="328 - 333" refId="ref143500" refString="Holman J. A. &amp; Case G. R. 1988. Reptiles from the Eocene Tallahatta Formation of Alabama. Journal of Vertebrate Paleontology 8 (3): 328 - 333. https: // doi. org / 10.1080 / 02724634.1988.10011716" type="journal article" year="1988">Holman &amp; Case (1988)</bibRefCitation>
did not figure their specimens so the identity of these teeth remains unconfirmed.
</paragraph>
<paragraph id="9001FC57FFB1FF9BFF7B9CC54E0A0B42" blockId="52.[189,1399,1009,1142]" pageId="52" pageNumber="53">
The teeth in our sample appear to be conspecific with
<taxonomicName id="57BE87D4FFB1FF9BFCA09CC5491C0B3C" authorityName="Leriche" authorityYear="1905" box="[870,1111,1009,1035]" class="Elasmobranchii" family="Odontaspididae" genus="Odontaspis" kingdom="Animalia" order="Lamniformes" pageId="52" pageNumber="53" phylum="Chordata" rank="species" species="winkleri">
<emphasis id="A2CA2045FFB1FF9BFCA09CC5491C0B3C" box="[870,1111,1009,1035]" italics="true" pageId="52" pageNumber="53">Odontaspis winkleri</emphasis>
</taxonomicName>
as originally described by
<bibRefCitation id="F42F81A6FFB1FF9BFF239B204CD10B18" author="Leriche M." box="[229,410,1044,1071]" pageId="52" pageNumber="53" pagination="49 - 228" refId="ref144901" refString="Leriche M. 1905. Les poissons eocenes de la Belgique. Memoires du Musee royal d'Histoire naturelle de Belgique 3 (11): 49 - 228." type="journal article" year="1905">Leriche (1905)</bibRefCitation>
. These teeth are differentiated from other odontaspids in our sample by their tall, cylindrical lateral cusplets, reduced or absent cutting edges on anterior teeth, and presence of distinctive folds at the base of the labial cusp face on lateral teeth.
</paragraph>
<paragraph id="9001FC57FFB1FF9BFF7B9BA84E5F0B81" blockId="52.[189,788,1180,1207]" box="[189,788,1180,1207]" pageId="52" pageNumber="53">
<heading id="CB494B3BFFB1FF9BFF7B9BA84E5F0B81" bold="true" box="[189,788,1180,1207]" fontSize="11" level="3" pageId="52" pageNumber="53" reason="3">
<emphasis id="A2CA2045FFB1FF9BFF7B9BA84E5F0B81" bold="true" box="[189,788,1180,1207]" pageId="52" pageNumber="53">
Stratigraphic and geographic range in
<collectingRegion id="527A32B5FFB1FF9BFD649BA84E5F0B81" box="[674,788,1180,1206]" country="United States of America" name="Alabama" pageId="52" pageNumber="53">Alabama</collectingRegion>
</emphasis>
</heading>
</paragraph>
<paragraph id="9001FC57FFB1FF9BFF7B9BFF481C0A3E" blockId="52.[189,1399,1227,1289]" pageId="52" pageNumber="53">
The specimens in our sample were collected from the lower Tallahatta Formation at site ADl-1 and the basal
<collectingRegion id="527A32B5FFB1FF9BFEC79BDA4C190A3F" box="[257,338,1262,1288]" country="Portugal" name="Lisboa" pageId="52" pageNumber="53">Lisbon</collectingRegion>
Formation at site ACov-11. Upper Ypresian to middle Lutetian, zones NP14 and NP15.
</paragraph>
</subSubSection>
</treatment>
</document>