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<document id="7249A3A7525036EA2E59FB14D7DCD4D1" ID-DOI="10.15553/c2012v672a12" ID-ISSN="2235-3658" ID-Zenodo-Dep="5790002" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1637611591098" checkinUser="carolina" docAuthor="Callmander, Martin W., Lowry Ii, Porter P., Forest, Félix, Devey, Dion S., Beentje, Henk &amp; Buerki, Sven" docDate="2012" docId="03F20C491D142B4D5D5DFEEA762452FB" docLanguage="en" docName="Candollea.67.2.323-345.pdf.imf" docOrigin="Candollea 67 (2)" docStyle="DocumentStyle:FB35ED2BCB231A5F446DBDA23BDBADFC.2:Candollea.2012-2014.journal_article" docStyleId="FB35ED2BCB231A5F446DBDA23BDBADFC" docStyleName="Candollea.2012-2014.journal_article" docStyleVersion="2" docTitle="Benstonea biplicata Callm. &amp; Buerki, comb. nova." docType="treatment" docVersion="7" lastPageNumber="332" masterDocId="FFCB74311D1D2B445D3AFFF3771C5134" masterDocTitle="Benstonea Callm. &amp; Buerki (Pandanaceae): characterization, circumscription, and distribution of a new genus of screw-pines, with a synopsis of accepted species" masterLastPageNumber="345" masterPageNumber="323" pageNumber="332" updateTime="1699425018902" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="B96041865EE28C58EE06E34B8F065BC3">Benstonea Callm. &amp; Buerki (Pandanaceae): characterization, circumscription, and distribution of a new genus of screw-pines, with a synopsis of accepted species</mods:title>
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<mods:namePart id="8C457CBC2DA3B262007E4FEE9A816ADF">Callmander, Martin W.</mods:namePart>
<mods:affiliation id="4219601B53E955B46F63098A15E9F77E">Missouri Botanical Garden, P. O. Box 299, St. Louis, MO, 63166 - 0299, U. S. A. and Conservatoire et Jardin botaniques de la Ville de Genève, CP 60, CH- 1292 Chambésy, Switzerland.</mods:affiliation>
<mods:nameIdentifier id="E34C16962660D1D45ED85364C7F20FE4" type="email">martin.callmander@mobot.org</mods:nameIdentifier>
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<mods:roleTerm id="2662AA9900FC03F42BE1AFCCC338A857">Author</mods:roleTerm>
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<mods:namePart id="B20BECB2B01B7B6ED8D4DBB73678D254">Lowry Ii, Porter P.</mods:namePart>
<mods:affiliation id="A8A7DF2D8D704A34478CCDC9116F82DA">Missouri Botanical Garden, P. O. Box 299, St. Louis, MO, 63166 - 0299, U. S. A. and Département Systématique et Evolution (UMR 7205), Muséum National d’Histoire Naturelle, case postale 39, rue Cuvier 57, 75231 Paris (Cedex) 05, France.</mods:affiliation>
</mods:name>
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</mods:role>
<mods:namePart id="580D22BE941D074A3DBBE3EFD255DDF4">Forest, Félix</mods:namePart>
<mods:affiliation id="C90C8DE845E950614640C47964C5887F">JodrellLaboratory, Royal Botanic Gardens, Kew, Richmond, Surrey, TW 9 3 DS, United Kingdom.</mods:affiliation>
</mods:name>
<mods:name id="424552406AE1DF35D04D4033C5CCF4EF" type="personal">
<mods:role id="6CEE9B7D502C7F72B9287AD7E43D68D7">
<mods:roleTerm id="AA7B676AB4CB45FBDF21CA350236F9BF">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D4F837AABD48E0F8C0960C9657FC6896">Devey, Dion S.</mods:namePart>
<mods:affiliation id="86E382FFC563EF52E0C54080A15C3D79">JodrellLaboratory, Royal Botanic Gardens, Kew, Richmond, Surrey, TW 9 3 DS, United Kingdom.</mods:affiliation>
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<mods:roleTerm id="7A30954AA43BECD607C44BA8FCDF2AB3">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7F9CF4F5A07152C6E57157BCAC7B8E84">Beentje, Henk</mods:namePart>
<mods:affiliation id="AC984EFF7DA89DB7EEE53B041FDC0F83">Herbarium, Library, Art &amp; Archives, Royal Botanic Gardens, Kew, Richmond, Surrey, TW 9 3 DS, United Kingdom.</mods:affiliation>
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<mods:namePart id="2EDC76AED66737AC174B4F12EBFADD8B">Buerki, Sven</mods:namePart>
<mods:affiliation id="5FAFA721D4BD405B0116578104FFAE3E">JodrellLaboratory, Royal Botanic Gardens, Kew, Richmond, Surrey, TW 9 3 DS, United Kingdom.</mods:affiliation>
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<mods:typeOfResource id="44FD6C59EF16166918E34F034D45A09F">text</mods:typeOfResource>
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<mods:title id="D377C5A393F33778C577C6DE2FA4F707">Candollea</mods:title>
</mods:titleInfo>
<mods:part id="D9332BCB4FA1701BC028A413D101CDBD">
<mods:date id="3052CE805A43DC315C27E7EC7174ED50">2012</mods:date>
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<mods:number id="5F8EC1E79C0C331A3859FFCB5E3665A3">2012-12-01</mods:number>
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<mods:number id="6C7281ED75260D7C2242DBB25DBB0512">67</mods:number>
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<mods:number id="CF59CED5F0C6B128A520A1F421552579">2</mods:number>
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<mods:classification id="10BB4CEA119953B3CB07338769E0A3E8">journal article</mods:classification>
<mods:identifier id="069F419D7947A266876F2CFFC0E406D2" type="DOI">10.15553/c2012v672a12</mods:identifier>
<mods:identifier id="C75AE52AB5CF31856F3FB33BC57A5CB2" type="ISSN">2235-3658</mods:identifier>
<mods:identifier id="17AB1E283B8A102B6CA7586C7FCBD3ED" type="Zenodo-Dep">5790002</mods:identifier>
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<treatment id="03F20C491D142B4D5D5DFEEA762452FB" ID-DOI="http://doi.org/10.5281/zenodo.6349694" ID-Zenodo-Dep="6349694" LSID="urn:lsid:plazi:treatment:03F20C491D142B4D5D5DFEEA762452FB" httpUri="http://treatment.plazi.org/id/03F20C491D142B4D5D5DFEEA762452FB" lastPageNumber="332" pageId="9" pageNumber="332">
<subSubSection id="C341EED41D142B4D5D5DFEEA77D65065" pageId="9" pageNumber="332" type="nomenclature">
<paragraph id="8BE4BD5F1D142B4D5D5DFEEA77D65065" blockId="9.[103,756,281,408]" pageId="9" pageNumber="332">
8.
<taxonomicName id="4C5BC6DC1D142B4D5DB6FEEA75BF5005" ID-CoL="LHHK" authority="(H. St. John) Callm. &amp; Buerki" authorityName="Callm. &amp; Buerki" baseAuthorityName="H. St. John" box="[140,675,281,305]" class="Liliopsida" family="Pandanaceae" genus="Benstonea" kingdom="Plantae" order="Pandanales" pageId="9" pageNumber="332" phylum="Tracheophyta" rank="species" species="biplicata" status="comb. nova.">
<emphasis id="B92F614D1D142B4D5DB6FEEA767D5005" bold="true" box="[140,353,281,305]" italics="true" pageId="9" pageNumber="332">Benstonea biplicata</emphasis>
(H. St. John) Callm. &amp; Buerki
</taxonomicName>
,
<emphasis id="B92F614D1D142B4D5F97FEEA77D65065" bold="true" pageId="9" pageNumber="332">
<taxonomicNameLabel id="A21CDC361D142B4D5F97FEEA77D65065" pageId="9" pageNumber="332">comb. nova.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C341EED41D142B4D5DB6FEAC77F350AC" pageId="9" pageNumber="332" type="reference_group">
<paragraph id="8BE4BD5F1D142B4D5DB6FEAC77F350AC" blockId="9.[103,756,281,408]" pageId="9" pageNumber="332">
<treatmentCitationGroup id="AB4B9A711D142B4D5DB6FEAC77F350AC" pageId="9" pageNumber="332">
≡
<treatmentCitation id="0AFA9B4E1D142B4D5D88FE9377F350AC" author="H. St. John" journal="Pacific Sci." page="466" pageId="9" pageNumber="332" score="7.816395839474338" volume="17" year="1963">
<taxonomicName id="4C5BC6DC1D142B4D5D88FE9376E3504C" ID-CoL="6TLR4" authority="H. St. John" authorityName="H. St. John" box="[178,511,352,376]" class="Liliopsida" family="Pandanaceae" genus="Pandanus" kingdom="Plantae" order="Pandanales" pageId="9" pageNumber="332" phylum="Tracheophyta" rank="species" species="biplicatus">
<emphasis id="B92F614D1D142B4D5D88FE93769B504C" box="[178,391,352,376]" italics="true" pageId="9" pageNumber="332">Pandanus biplicatus</emphasis>
H. St. John
</taxonomicName>
in Pacific Sci. 17: 466. 1963.
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C341EED41D142B4D5DB6FE5B754A5334" pageId="9" pageNumber="332" type="materials_examined">
<paragraph id="8BE4BD5F1D142B4D5DB6FE5B754A5334" blockId="9.[103,753,424,615]" pageId="9" pageNumber="332">
<materialsCitation id="3B33B7021D142B4D5DB6FE5B765B5334" collectingDate="1917-12-09" collectionCode="SING" collectorName="Md. Haniff &amp; Md. Nur" country="Thailand" location="Kopah" municipality="Kopah" pageId="9" pageNumber="332" specimenCode="SING0032172" specimenCount="1" stateProvince="Jaujau" typeStatus="holotype">
<emphasis id="B92F614D1D142B4D5DB6FE5B764550F4" bold="true" box="[140,345,424,448]" pageId="9" pageNumber="332">
Typus:
<emphasis id="B92F614D1D142B4D5DE5FE5B764550F4" bold="true" box="[223,345,424,448]" pageId="9" pageNumber="332">
<collectingCountry id="F34CFDCF1D142B4D5DE5FE5B764550F4" box="[223,345,424,448]" pageId="9" pageNumber="332">THAILAND</collectingCountry>
</emphasis>
</emphasis>
.
<collectingRegion id="499F73BD1D142B4D5C65FE5B76A450F4" box="[351,440,424,448]" pageId="9" pageNumber="332">Jaujau</collectingRegion>
: lower
<collectingRegion id="499F73BD1D142B4D5F2BFE5B755050F4" box="[529,588,424,448]" pageId="9" pageNumber="332">Siam</collectingRegion>
,
<collectingMunicipality id="6B8027251D142B4D5F6DFE5B75BF50F4" box="[599,675,424,448]" pageId="9" pageNumber="332">Kopah</collectingMunicipality>
,
<date id="FFE59B9F1D142B4D5F97FE5B77DA50D4" pageId="9" pageNumber="332" value="1917-12-09">
<collectingDate id="EFA162771D142B4D5F97FE5B77DA50D4" pageId="9" pageNumber="332" value="1917-12-09">9.XII. 1917</collectingDate>
</date>
,
<emphasis id="B92F614D1D142B4D5DEEFE3B755450D4" box="[212,584,456,480]" italics="true" pageId="9" pageNumber="332">
<collectorName id="26AED8891D142B4D5DEEFE3B764C50D4" box="[212,336,456,480]" pageId="9" pageNumber="332">Md. Haniff</collectorName>
&amp;
<collectorName id="26AED8891D142B4D5C4CFE3B76CA50D4" box="[374,470,456,480]" pageId="9" pageNumber="332">Md. Nur</collectorName>
<specimenCode id="DBFD15241D142B4D5CE4FE3B755450D4" box="[478,584,456,480]" pageId="9" pageNumber="332">SFN 2703</specimenCode>
</emphasis>
(
<typeStatus id="54E003FD1D142B4D5F5EFE3B75B850D4" box="[612,676,456,480]" pageId="9" pageNumber="332" type="holotype">holo-</typeStatus>
:
<collectionCode id="ED4A259A1D142B4D5F8AFE3B75EC50D4" box="[688,752,456,480]" country="Singapore" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15640" lsid="urn:lsid:biocol.org:col:15640" name="Singapore Botanic Gardens" pageId="9" pageNumber="332" type="Herbarium">SING</collectionCode>
[
<specimenCode id="DBFD15241D142B4D5DA9FE1B76205334" box="[147,316,488,512]" collectionCode="SING" country="Singapore" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15640" lsid="urn:lsid:biocol.org:col:15640" name="Singapore Botanic Gardens" pageId="9" pageNumber="332" type="Herbarium">SING0032172</specimenCode>
]!
</materialsCitation>
;
<materialsCitation id="3B33B7021D142B4D5C6BFE1B754A5334" box="[337,598,488,512]" collectingDate="1917-12-09" collectionCode="K" collectorName="Md. Haniff &amp; Md. Nur" country="Thailand" location="Kopah" municipality="Kopah" pageId="9" pageNumber="332" specimenCode="K000781297" specimenCount="1" stateProvince="Jaujau" typeStatus="isotype">
<typeStatus id="54E003FD1D142B4D5C6BFE1B76615334" box="[337,381,488,512]" pageId="9" pageNumber="332" type="isotype">iso-</typeStatus>
:
<collectionCode id="ED4A259A1D142B4D5CBDFE1B76865334" box="[391,410,488,512]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="9" pageNumber="332" type="Herbarium">K</collectionCode>
[
<specimenCode id="DBFD15241D142B4D5C9DFE1B755D5334" box="[423,577,488,512]" collectionCode="K" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="9" pageNumber="332" type="Herbarium">K000781297</specimenCode>
]!).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C341EED41D142B4D5DB6FDFC77A55353" pageId="9" pageNumber="332" type="distribution">
<paragraph id="8BE4BD5F1D142B4D5DB6FDFC77A55353" blockId="9.[103,753,424,615]" pageId="9" pageNumber="332">
<emphasis id="B92F614D1D142B4D5DB6FDFC76975313" box="[140,395,527,551]" italics="true" pageId="9" pageNumber="332">Distribution and ecology</emphasis>
. –
<taxonomicName id="4C5BC6DC1D142B4D5C91FDFC756B5313" box="[427,631,527,551]" class="Liliopsida" family="Pandanaceae" genus="Benstonea" kingdom="Plantae" order="Pandanales" pageId="9" pageNumber="332" phylum="Tracheophyta" rank="species" species="biplicata">
<emphasis id="B92F614D1D142B4D5C91FDFC756B5313" box="[427,631,527,551]" italics="true" pageId="9" pageNumber="332">Benstonea biplicata</emphasis>
</taxonomicName>
is restricted to southern
<collectingCountry id="F34CFDCF1D142B4D5DDCFDDC76555373" box="[230,329,559,583]" name="Thailand" pageId="9" pageNumber="332">Thailand</collectingCountry>
, close to the Malaysian border (
<bibRefCitation id="EFCAC0AE1D142B4D5F99FDDC77B15353" author="STONE" firstAuthor="STONE" pageId="9" pageNumber="332" pagination="411 - 420" refId="ref16102" refString="STONE, B. C. (1968 b). Studies in the Malesian Pandanaceae, II. Two new species of Pandanus Stickm. Sect. Fusiforma St. John. Reinwardtia 7: 411 - 420." type="journal article" year="1968" yearSuffix="b">STONE, 1968b</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C341EED41D142B4D5DB6FD84762452FB" pageId="9" pageNumber="332" type="discussion">
<paragraph id="8BE4BD5F1D142B4D5DB6FD84762452FB" blockId="9.[103,754,631,975]" pageId="9" pageNumber="332">
<emphasis id="B92F614D1D142B4D5DB6FD84760B53BB" box="[140,279,631,655]" italics="true" pageId="9" pageNumber="332">Observations</emphasis>
. – This species was placed in
<taxonomicName id="4C5BC6DC1D142B4D5F69FD8477C6539B" class="Liliopsida" family="Pandanaceae" genus="Pandanus" kingdom="Plantae" order="Pandanales" pageId="9" pageNumber="332" phylum="Tracheophyta" rank="section" section="Fusiforma">
<emphasis id="B92F614D1D142B4D5F69FD8475A053BB" box="[595,700,631,655]" italics="true" pageId="9" pageNumber="332">Pandanus</emphasis>
sect.
<emphasis id="B92F614D1D142B4D5D5DFD6477C6539B" box="[103,218,663,687]" italics="true" pageId="9" pageNumber="332">Fusiforma</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFCAC0AE1D142B4D5C33FD6476A8539B" author="STONE" box="[265,436,663,687]" firstAuthor="STONE" pageId="9" pageNumber="332" pagination="411 - 420" refId="ref16102" refString="STONE, B. C. (1968 b). Studies in the Malesian Pandanaceae, II. Two new species of Pandanus Stickm. Sect. Fusiforma St. John. Reinwardtia 7: 411 - 420." type="journal article" year="1968" yearSuffix="b">STONE (1968b)</bibRefCitation>
, a section that we have not included in
<taxonomicName id="4C5BC6DC1D142B4D5DD8FD44765353FB" box="[226,335,695,719]" class="Liliopsida" family="Pandanaceae" genus="Benstonea" kingdom="Plantae" order="Pandanales" pageId="9" pageNumber="332" phylum="Tracheophyta" rank="genus">
<emphasis id="B92F614D1D142B4D5DD8FD44765353FB" box="[226,335,695,719]" italics="true" pageId="9" pageNumber="332">Benstonea</emphasis>
</taxonomicName>
(see above). Careful examination of the
<typeStatus id="54E003FD1D142B4D5D5DFD24778853DB" box="[103,148,727,751]" pageId="9" pageNumber="332">type</typeStatus>
in SING showed that its leaves (c. 4
<quantity id="4CA310BA1D142B4D5F17FD24756053DB" box="[557,636,727,751]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="9" pageNumber="332" unit="cm" value="100.0">100 cm</quantity>
) and styles (sharp and narrow) are similar to those in all the other species of
<taxonomicName id="4C5BC6DC1D142B4D5DB9FCE477EE521B" box="[131,242,791,815]" class="Liliopsida" family="Pandanaceae" genus="Benstonea" kingdom="Plantae" order="Pandanales" pageId="9" pageNumber="332" phylum="Tracheophyta" rank="genus">
<emphasis id="B92F614D1D142B4D5DB9FCE477EE521B" box="[131,242,791,815]" italics="true" pageId="9" pageNumber="332">Benstonea</emphasis>
</taxonomicName>
.
<taxonomicName id="4C5BC6DC1D142B4D5DC5FCE476C8521B" box="[255,468,791,815]" class="Liliopsida" family="Pandanaceae" genus="Benstonea" kingdom="Plantae" order="Pandanales" pageId="9" pageNumber="332" phylum="Tracheophyta" rank="species" species="biplicata">
<emphasis id="B92F614D1D142B4D5DC5FCE476C8521B" box="[255,468,791,815]" italics="true" pageId="9" pageNumber="332">Benstonea biplicata</emphasis>
</taxonomicName>
is therefore excluded from
<taxonomicName id="4C5BC6DC1D142B4D5D5DFCC47662527B" box="[103,382,823,847]" class="Liliopsida" family="Pandanaceae" genus="Pandanus" kingdom="Plantae" order="Pandanales" pageId="9" pageNumber="332" phylum="Tracheophyta" rank="section" section="Fusiforma">
<emphasis id="B92F614D1D142B4D5D5DFCC477CE527B" box="[103,210,823,847]" italics="true" pageId="9" pageNumber="332">Pandanus</emphasis>
sect.
<emphasis id="B92F614D1D142B4D5C35FCC47662527B" box="[271,382,823,847]" italics="true" pageId="9" pageNumber="332">Fusiforma</emphasis>
</taxonomicName>
and transferred here to
<emphasis id="B92F614D1D142B4D5F41FCC475ED527B" box="[635,753,823,847]" italics="true" pageId="9" pageNumber="332">
<taxonomicName id="4C5BC6DC1D142B4D5F41FCC475F1527B" box="[635,749,823,847]" class="Liliopsida" family="Pandanaceae" genus="Benstonea" kingdom="Plantae" order="Pandanales" pageId="9" pageNumber="332" phylum="Tracheophyta" rank="genus">Benstonea</taxonomicName>
.
</emphasis>
It is morphologically similar to
<taxonomicName id="4C5BC6DC1D142B4D5CFAFCA477A952BB" authority="(Martelli) Callm. &amp; Buerki" authorityName="Callm. &amp; Buerki" baseAuthorityName="Martelli" class="Liliopsida" family="Pandanaceae" genus="Benstonea" kingdom="Plantae" order="Pandanales" pageId="9" pageNumber="332" phylum="Tracheophyta" rank="species" species="nana">
<emphasis id="B92F614D1D142B4D5CFAFCA476CC525B" box="[448,464,855,879]" italics="true" pageId="9" pageNumber="332">B</emphasis>
.
<emphasis id="B92F614D1D142B4D5CE4FCA47508525B" box="[478,532,855,879]" italics="true" pageId="9" pageNumber="332">nana</emphasis>
(Martelli) Callm. &amp; Buerki
</taxonomicName>
, but differs in possessing a larger syncarp. Pending taxonomic revisions in the region should help clarify the status of this taxon.
</paragraph>
</subSubSection>
</treatment>
</document>
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