336 lines
36 KiB
XML
336 lines
36 KiB
XML
<document id="3E7D54D0D4ACE43CBA6CF6041171AC08" ID-DOI="10.1649/0010-065X-72.2.331" ID-ISSN="1938-4394" ID-Zenodo-Dep="5382545" ID-ZooBank="urn:lsid:zoobank.org:pub:11BE44A2-4C73-4D36-A761-C9059C9C4222" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630295096312" checkinUser="diego" docAuthor="López-García, Margarita M. & Deloya, Cuauhtémoc" docDate="2018" docId="03F187ED6F1FFF8ADBDBFED1FE0AFD45" docLanguage="en" docName="0010-065X-72.2.331.pdf" docOrigin="The Coleopterists Bulletin 72 (2)" docSource="http://dx.doi.org/10.1649/0010-065x-72.2.331" docStyle="DocumentStyle:F824235CA89CB83023C7D5789122F870.3:TheColeopBull.2017-.journal_article.1cover" docStyleId="F824235CA89CB83023C7D5789122F870" docStyleName="TheColeopBull.2017-.journal_article.1cover" docStyleVersion="3" docTitle="Tomarus pumilus" docType="treatment" docVersion="4" lastPageNumber="334" masterDocId="FFC8FF956F1BFF8FDB2BFF98FFFBFFF0" masterDocTitle="Description of the Female of Tomarus rostratus Dupuis and New Synonyms and Records for the Genus Tomarus Erichson (Coleoptera: Scarabaeidae: Dynastinae)" masterLastPageNumber="338" masterPageNumber="331" pageNumber="334" updateTime="1699218372114" updateUser="plazi">
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<mods:title id="07236FD9987C9016F437AA8E90BB52BF">Description of the Female of Tomarus rostratus Dupuis and New Synonyms and Records for the Genus Tomarus Erichson (Coleoptera: Scarabaeidae: Dynastinae)</mods:title>
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<mods:namePart id="EC535AA431BD5E508D614CE64790F297">López-García, Margarita M.</mods:namePart>
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<mods:namePart id="C12D3B4B0838B289D45F9017103C5515">Deloya, Cuauhtémoc</mods:namePart>
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<mods:title id="40E283591D1172A3E2A75F00DA9CF655">The Coleopterists Bulletin</mods:title>
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<mods:part id="F756BAD8B7527296B96A86127AA0BF7C">
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<mods:date id="F6F668397BF83EB212DEF58349FFDC15">2018</mods:date>
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<mods:number id="C5309CEAAD848FE9E70B003C67B0EF26">2018-06-20</mods:number>
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<mods:number id="8E4C1C059AD6B5A13E7AE099EBE927BF">72</mods:number>
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<mods:url id="03A7A18839E8A0BDD0B1B2A93197B825">http://dx.doi.org/10.1649/0010-065x-72.2.331</mods:url>
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<mods:identifier id="0328385B8158E2E268C23BB6219D58A4" type="DOI">10.1649/0010-065X-72.2.331</mods:identifier>
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<mods:identifier id="6BCE9EFFC4FD126FDD915EAA5C227DC9" type="ISSN">1938-4394</mods:identifier>
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<subSubSection id="C34265706F1FFF8BDBDBFED1FDDEFEAE" box="[240,549,329,350]" pageId="4" pageNumber="334" type="nomenclature">
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<paragraph id="8BE736FB6F1FFF8BDBDBFED1FDDEFEAE" blockId="4.[240,549,329,377]" box="[240,549,329,350]" pageId="4" pageNumber="334">
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<heading id="D0AF81976F1FFF8BDBDBFED1FDDEFEAE" box="[240,549,329,350]" centered="true" fontSize="9" level="2" pageId="4" pageNumber="334" reason="2">
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<taxonomicName id="4C584D786F1FFF8BDBDBFED1FDDEFEAE" authority="(Prell, 1937)" baseAuthorityName="Prell" baseAuthorityYear="1937" box="[240,549,329,350]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="334" phylum="Arthropoda" rank="species" species="pumilus">
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<emphasis id="B92CEAE96F1FFF8BDBDBFED1FDDEFEAE" bold="true" box="[240,549,329,350]" pageId="4" pageNumber="334">
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<emphasis id="B92CEAE96F1FFF8BDBDBFED1FE5AFEAE" bold="true" box="[240,417,329,350]" italics="true" pageId="4" pageNumber="334">Tomarus pumilus</emphasis>
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(
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<bibRefCitation id="EFC94B0A6F1FFF8BDA84FED1FDE5FEAE" author="Prell, H." box="[431,542,329,350]" pageId="4" pageNumber="334" pagination="89 - 90" refId="ref5303" refString="Prell, H. 1937. Beitrage zur Kenntnis der Dynastiden. XV. Neue Arten und Rassen. Entomologische Zeitschrift 51: 89 - 90." type="journal article" year="1937">Prell, 1937</bibRefCitation>
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)
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C34265706F1FFF8BDA61FEFCFE30FE89" box="[330,459,355,377]" pageId="4" pageNumber="334" type="description">
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||
<paragraph id="8BE736FB6F1FFF8BDA61FEFCFE30FE89" blockId="4.[240,549,329,377]" box="[330,459,355,377]" pageId="4" pageNumber="334">
|
||
(
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<figureCitation id="13632A7E6F1FFF8BDA7AFEFCFE3FFE89" box="[337,452,355,377]" captionStart="Figs" captionStartId="4.[175,214,1663,1682]" captionTargetBox="[157,1166,816,1644]" captionTargetId="figure-356@4.[154,1169,814,1646]" captionTargetPageId="4" captionText="Figs. 14–19. Tomarus pumilus, lectotype: 14) Head and pronotum; 15) Parameres, frontal view; 16) Parameres, lateral view, and labels. Tomarus rosettae, holotype: 17) Head and pronotum; 18) Parameres, frontal view; 19) Parameres, lateral view, and labels." figureDoi="http://doi.org/10.5281/zenodo.5382551" httpUri="https://zenodo.org/record/5382551/files/figure.png" pageId="4" pageNumber="334">Figs. 14–19</figureCitation>
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)
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C34265706F1FFF8BDBBCFE16FD85FE4E" pageId="4" pageNumber="334" type="reference_group">
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||
<paragraph id="8BE736FB6F1FFF8BDBBCFE16FE3FFE53" blockId="4.[151,638,398,446]" box="[151,452,398,419]" pageId="4" pageNumber="334">
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<treatmentCitationGroup id="AB4811D56F1FFF8BDBBCFE16FE3FFE53" box="[151,452,398,419]" pageId="4" pageNumber="334">
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<treatmentCitation id="0AF910EA6F1FFF8BDBBCFE16FE3FFE53" author="Prell, H." box="[151,452,398,419]" page="90" pageId="4" pageNumber="334" year="1937">
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<taxonomicName id="4C584D786F1FFF8BDBBCFE16FE3FFE53" authority="Prell 1937: 90" authorityName="Prell, Endrody" authorityPageNumber="90" authorityYear="1937" box="[151,452,398,419]" class="Insecta" family="Dynastidae" genus="Ligyrus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="334" phylum="Arthropoda" rank="species" species="pumilus">
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<emphasis id="B92CEAE96F1FFF8BDBBCFE16FEC8FE53" box="[151,307,398,419]" italics="true" pageId="4" pageNumber="334">Ligyrus pumilus</emphasis>
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||
<bibRefCitation id="EFC94B0A6F1FFF8BDA11FE16FE3FFE53" author="Prell, H." box="[314,452,398,419]" pageId="4" pageNumber="334" pagination="89 - 90" refId="ref5303" refString="Prell, H. 1937. Beitrage zur Kenntnis der Dynastiden. XV. Neue Arten und Rassen. Entomologische Zeitschrift 51: 89 - 90." type="journal article" year="1937">Prell 1937: 90</bibRefCitation>
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</taxonomicName>
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||
</treatmentCitation>
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||
</treatmentCitationGroup>
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||
</paragraph>
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||
<paragraph id="8BE736FB6F1FFF8BDBBCFE31FD85FE4E" blockId="4.[151,638,398,446]" box="[151,638,425,446]" pageId="4" pageNumber="334">
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||
<treatmentCitationGroup id="AB4811D56F1FFF8BDBBCFE31FD85FE4E" box="[151,638,425,446]" pageId="4" pageNumber="334">
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||
<treatmentCitation id="0AF910EA6F1FFF8BDBBCFE31FE43FE4E" author="Endrodi, S." box="[151,440,425,446]" page="68" pageId="4" pageNumber="334" year="1968">
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||
<taxonomicName id="4C584D786F1FFF8BDBBCFE31FE43FE4E" authority="Endrodi 1968: 68" authorityName="Endrodi" authorityPageNumber="68" authorityYear="1968" box="[151,440,425,446]" class="Insecta" family="Dynastidae" genus="Ligyrus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="334" phylum="Arthropoda" rank="species" species="rosettae">
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<emphasis id="B92CEAE96F1FFF8BDBBCFE31FEE4FE4E" box="[151,287,425,446]" italics="true" pageId="4" pageNumber="334">Ligyrus rosettae</emphasis>
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<bibRefCitation id="EFC94B0A6F1FFF8BDA09FE31FE43FE4E" author="Endrodi, S." box="[290,440,425,446]" pageId="4" pageNumber="334" pagination="161 - 177" refId="ref4904" refString="Endrodi, S. 1968. Neue Arten der Pentodontini. (Col. Dynastinae). Folia Entomologica Hungarica 21: 161 - 177." type="journal article" year="1968">Endrödi 1968:68</bibRefCitation>
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</taxonomicName>
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||
</treatmentCitation>
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||
.
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<emphasis id="B92CEAE96F1FFF8BDA95FE31FD85FE4E" bold="true" box="[446,638,425,446]" pageId="4" pageNumber="334">NEW SYNONYMY.</emphasis>
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||
</treatmentCitationGroup>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C34265706F1FFF8BDB84FE4BFBA8FDEA" pageId="4" pageNumber="334" type="materials_examined">
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||
<paragraph id="8BE736FB6F1FFF8BDB84FE4BFBA8FDEA" blockId="4.[151,638,467,754]" lastBlockId="4.[685,1173,193,754]" pageId="4" pageNumber="334">
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||
<emphasis id="B92CEAE96F1FFF8BDB84FE4BFDC3FE18" bold="true" box="[175,568,467,488]" pageId="4" pageNumber="334">
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||
Examined Material.
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||
<taxonomicName id="4C584D786F1FFF8BDAA9FE4BFDCBFE18" baseAuthorityName="Prell" baseAuthorityYear="1937" box="[386,560,467,488]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="334" phylum="Arthropoda" rank="species" species="pumilus">
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||
<emphasis id="B92CEAE96F1FFF8BDAA9FE4BFDCBFE18" bold="true" box="[386,560,467,488]" italics="true" pageId="4" pageNumber="334">Tomarus pumilus</emphasis>
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||
</taxonomicName>
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||
:
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||
</emphasis>
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||
<materialsCitation id="3B303CA66F1FFF8BD969FE4BFE6AFDA2" determinerName="B. C. Ratcliffe" pageId="4" pageNumber="334" specimenCount="6" typeStatus="lectotype">
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||
<typeStatus id="54E388596F1FFF8BD969FE4BFF44FDF3" pageId="4" pageNumber="334" type="lectotype">Lectotype</typeStatus>
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||
deposited in ZMHU (16): “Corralitos //
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||
<taxonomicName id="4C584D786F1FFF8BD91DFE76FF1FFDED" authorityName="Prell, Endrody" authorityYear="1937" class="Insecta" family="Dynastidae" genus="Ligyrus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="334" phylum="Arthropoda" rank="species" species="pumilus">Ligyrus pumilus</taxonomicName>
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||
,
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||
<specimenCount id="9D5EFD726F1FFF8BDBC7FD90FED1FDED" box="[236,298,520,542]" pageId="4" pageNumber="334" type="generic">6Type</specimenCount>
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||
//
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||
<typeStatus id="54E388596F1FFF8BDA6BFD90FE52FDED" box="[320,425,520,541]" pageId="4" pageNumber="334" type="lectotype">Lectotypus</typeStatus>
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||
,
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||
<taxonomicName id="4C584D786F1FFF8BDA9AFD90FF1EFDC8" authority="Prell, Endrody" authorityName="Prell, Endrody" authorityYear="1937" class="Insecta" family="Dynastidae" genus="Ligyrus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="334" phylum="Arthropoda" rank="species" species="pumilus">Ligyrus pumilus Prell, Endrody</taxonomicName>
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||
//
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||
<taxonomicName id="4C584D786F1FFF8BDBD0FDBBFE2AFDC8" authority="(Prell)" baseAuthorityName="Prell" baseAuthorityYear="1937" box="[251,465,547,568]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="334" phylum="Arthropoda" rank="species" species="pumilus">
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||
<emphasis id="B92CEAE96F1FFF8BDBD0FDBBFE6DFDC8" box="[251,406,547,568]" italics="true" pageId="4" pageNumber="334">Tomarus pumilus</emphasis>
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(Prell)
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</taxonomicName>
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/ det.
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<determinerName id="7E3BDA3F6F1FFF8BD929FDBBFD85FDC8" box="[514,638,547,568]" pageId="4" pageNumber="334">B.C. Ratcliffe</determinerName>
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||
2011”.
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||
<taxonomicName id="4C584D786F1FFF8BDBCAFDA5FE6AFDA2" baseAuthorityName="Endrodi" baseAuthorityYear="1968" box="[225,401,573,594]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="334" phylum="Arthropoda" rank="species" species="rosettae">Tomarus rosettae</taxonomicName>
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||
</materialsCitation>
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||
<emphasis id="B92CEAE96F1FFF8BDAB9FDA5FE61FDA2" bold="true" box="[402,410,573,594]" pageId="4" pageNumber="334">:</emphasis>
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||
<materialsCitation id="3B303CA66F1FFF8BDA88FDA5FE98FD02" collectionCode="HNHM" collectorName="Prov. Bs. Aires" pageId="4" pageNumber="334" specimenCount="3" typeStatus="holotype">
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||
<typeStatus id="54E388596F1FFF8BDA88FDA5FE00FDA2" box="[419,507,573,594]" pageId="4" pageNumber="334" type="holotype">Holotype</typeStatus>
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||
deposited in NHMB (16,
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<figureCitation id="13632A7E6F1FFF8BDA32FDC0FE75FD9D" box="[281,398,600,622]" captionStart="Figs" captionStartId="4.[175,214,1663,1682]" captionTargetBox="[157,1166,816,1644]" captionTargetId="figure-356@4.[154,1169,814,1646]" captionTargetPageId="4" captionText="Figs. 14–19. Tomarus pumilus, lectotype: 14) Head and pronotum; 15) Parameres, frontal view; 16) Parameres, lateral view, and labels. Tomarus rosettae, holotype: 17) Head and pronotum; 18) Parameres, frontal view; 19) Parameres, lateral view, and labels." figureDoi="http://doi.org/10.5281/zenodo.5382551" httpUri="https://zenodo.org/record/5382551/files/figure.png" pageId="4" pageNumber="334">Figs. 16–17</figureCitation>
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||
): “Proc. San Clemente / del Tuyu, Arg. /
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||
<collectorName id="26AD532D6F1FFF8BDA05FDEBFE40FD78" box="[302,443,627,648]" pageId="4" pageNumber="334">Prov. Bs. Aires</collectorName>
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||
, xii.49 //
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||
<typeStatus id="54E388596F1FFF8BD93BFDEBFD88FD78" box="[528,627,627,648]" pageId="4" pageNumber="334" type="holotype">Holotypus</typeStatus>
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||
/
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||
<taxonomicName id="4C584D786F1FFF8BDBBCFD15FE8AFD52" authority="Endr." authorityName="Endrodi" authorityYear="1968" box="[151,369,653,674]" class="Insecta" family="Dynastidae" genus="Ligyrus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="334" phylum="Arthropoda" rank="species" species="rosettae">Ligyrus rosettae Endr.</taxonomicName>
|
||
//
|
||
<taxonomicName id="4C584D786F1FFF8BDABBFD15FD86FD52" authority="(Prell)" baseAuthorityName="Prell" baseAuthorityYear="1937" box="[400,637,653,674]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="334" phylum="Arthropoda" rank="species" species="pumilus">Tomarus pumilus (Prell)</taxonomicName>
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||
det. M.M.
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<bibRefCitation id="EFC94B0A6F1FFF8BDBD7FD3FFE4FFD4D" author="Lopez-Garc'ia, M. M. & Gasca-Alvarez, R. D." box="[252,436,679,701]" pageId="4" pageNumber="334" pagination="491 - 509" refId="ref5136" refString="Lopez-Garc'ia, M. M., H. J. Gasca-Alvarez, R. D. Cave, and G. Amat-Garc'ia. 2016. An annotated checklist of the New World pentodontine scarab beetles (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini). Zootaxa 4170 (3): 491 - 509." type="journal article" year="2016">López-Garc´ıa 2016</bibRefCitation>
|
||
”.
|
||
<typeStatus id="54E388596F1FFF8BDAE1FD30FDE6FD4D" box="[458,541,680,701]" pageId="4" pageNumber="334" type="paratype">Paratype</typeStatus>
|
||
deposited in
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||
<collectionCode id="ED49AE3E6F1FFF8BDB85FD5AFF0DFD27" box="[174,246,706,727]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="4" pageNumber="334">HNHM</collectionCode>
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||
(16): Proc. San Clemente /del Tuyu, Arg. /
|
||
<collectorName id="26AD532D6F1FFF8BDBBCFD45FEE4FD02" box="[151,287,733,754]" pageId="4" pageNumber="334">Prov.Bs. Aires</collectorName>
|
||
, xii.49”
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B303CA66F1FFF8BDA47FD45FBE6FF27" box="[364,1053,193,754]" collectionCode="HNHM" pageId="4" pageNumber="334" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
|
||
<typeStatus id="54E388596F1FFF8BDA47FD45FE38FD02" box="[364,451,733,754]" pageId="4" pageNumber="334" type="paratype">Paratypes</typeStatus>
|
||
deposited in
|
||
<collectionCode id="ED49AE3E6F1FFF8BD91CFD45FD85FD02" box="[567,638,733,754]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="4" pageNumber="334">HNHM</collectionCode>
|
||
(16,
|
||
<specimenCount id="9D5EFD726F1FFF8BD9FDFF5AFD14FF27" box="[726,751,193,215]" pageId="4" pageNumber="334" type="female">1♀</specimenCount>
|
||
): Mendoza, Rep. Arg., v.26
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||
</materialsCitation>
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||
.
|
||
<materialsCitation id="3B303CA66F1FFF8BDF05FF5AFBB5FDEA" collectingDate="1906-10-20" collectingDateMax="1973-12-22" collectingDateMin="1906-10-20" collectionCode="HNHM, USNM" collectorName="Coll. Jensen-Haarup & de Cordoba & C. S. Reed Coll. & R. Sanzin Coll. & C. M. & O. S. Flint Coll. & A. Wetmore" country="Argentina" county="Arandel" determinerName="Endrodi & Dr. Endrodi" location="Pedregal" municipality="Leg Pena" pageId="4" pageNumber="334" specimenCount="8" specimenCount-female="8" stateProvince="Mendoza">
|
||
Additional material:
|
||
<emphasis id="B92CEAE96F1FFF8BD9D0FF44FC7CFF01" bold="true" box="[763,903,220,241]" pageId="4" pageNumber="334">
|
||
<collectingCountry id="F34F766B6F1FFF8BD9D0FF44FC78FF01" box="[763,899,220,241]" name="Argentina" pageId="4" pageNumber="334">ARGENTINA</collectingCountry>
|
||
:
|
||
</emphasis>
|
||
<collectingRegion id="499CF8196F1FFF8BD8A7FF44FC26FF01" box="[908,989,220,241]" country="Argentina" name="Mendoza" pageId="4" pageNumber="334">Mendoza</collectingRegion>
|
||
,
|
||
<collectorName id="26AD532D6F1FFF8BD8CEFF44FB6AFF01" box="[997,1169,220,241]" pageId="4" pageNumber="334">Coll. Jensen-Haarup</collectorName>
|
||
,
|
||
<taxonomicName id="4C584D786F1FFF8BD986FF6FFCF5FEFC" authorityName="Endrodi" authorityYear="1968" box="[685,782,247,268]" class="Insecta" family="Dynastidae" genus="Ligyrus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="334" phylum="Arthropoda" rank="species" species="rosettae">L. rosettae</taxonomicName>
|
||
, det.
|
||
<determinerName id="7E3BDA3F6F1FFF8BD810FF6FFC73FEFC" box="[827,904,247,268]" pageId="4" pageNumber="334">Endrödi</determinerName>
|
||
, 1967 [
|
||
<specimenCount id="9D5EFD726F1FFF8BD8E2FF6FFC1AFEFD" box="[969,993,247,269]" pageId="4" pageNumber="334" type="female">1♀</specimenCount>
|
||
,
|
||
<collectionCode id="ED49AE3E6F1FFF8BD8C7FF6FFBC3FEFC" box="[1004,1080,247,268]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="4" pageNumber="334">HNHM</collectionCode>
|
||
].
|
||
<collectingCounty id="62864E776F1FFF8BDF6FFF6FFB6BFEFC" box="[1092,1168,247,268]" pageId="4" pageNumber="334">Arandel</collectingCounty>
|
||
,
|
||
<collectingRegion id="499CF8196F1FFF8BD986FE8AFD26FED7" box="[685,733,274,295]" country="Argentina" name="Salta" pageId="4" pageNumber="334">Salta</collectingRegion>
|
||
, 10/11.x.68,
|
||
<collectingMunicipality id="6B83AC816F1FFF8BD879FE8AFC51FED7" box="[850,938,274,295]" pageId="4" pageNumber="334">Leg Pena</collectingMunicipality>
|
||
,
|
||
<taxonomicName id="4C584D786F1FFF8BD898FE8AFBEFFED7" authorityName="Endrodi" authorityYear="1968" box="[947,1044,274,295]" class="Insecta" family="Dynastidae" genus="Ligyrus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="334" phylum="Arthropoda" rank="species" species="rosettae">L. rosettae</taxonomicName>
|
||
, det.
|
||
<determinerName id="7E3BDA3F6F1FFF8BDF68FE8AFB6BFED7" box="[1091,1168,274,295]" pageId="4" pageNumber="334">Endrödi</determinerName>
|
||
, 1970 [16,
|
||
<collectionCode id="ED49AE3E6F1FFF8BD824FEB5FCA0FEB2" box="[783,859,301,322]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="4" pageNumber="334">HNHM</collectionCode>
|
||
].
|
||
<location id="8E8760206F1FFF8BD842FEB5FC40FEB2" LSID="urn:lsid:plazi:treatment:03F187ED6F1FFF8ADBDBFED1FE0AFD45:8E8760206F1FFF8BD842FEB5FC40FEB2" box="[873,955,301,322]" country="Argentina" county="Arandel" municipality="Leg Pena" name="Pedregal" pageId="4" pageNumber="334" stateProvince="Mendoza">Pedregal</location>
|
||
,
|
||
<collectingRegion id="499CF8196F1FFF8BD8EFFEB5FBE7FEB2" box="[964,1052,301,322]" country="Argentina" name="Mendoza" pageId="4" pageNumber="334">Mendoza</collectingRegion>
|
||
,
|
||
<date id="FFE6103B6F1FFF8BDF0DFEB5FB6BFEB2" box="[1062,1168,301,322]" pageId="4" pageNumber="334" value="1906-10-20">
|
||
<collectingDate id="EFA2E9D36F1FFF8BDF0DFEB5FB6BFEB2" box="[1062,1168,301,322]" pageId="4" pageNumber="334" value="1906-10-20">20.10.1906</collectingDate>
|
||
</date>
|
||
,
|
||
<taxonomicName id="4C584D786F1FFF8BD986FED0FCEEFEAD" authorityName="Endrodi" authorityYear="1968" box="[685,789,328,349]" class="Insecta" family="Dynastidae" genus="Ligyrus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="334" phylum="Arthropoda" rank="species" species="rosettae">L. rosettae</taxonomicName>
|
||
, det.
|
||
<determinerName id="7E3BDA3F6F1FFF8BD87BFED0FC3EFEAD" box="[848,965,328,349]" pageId="4" pageNumber="334">Dr. Endrödi</determinerName>
|
||
, 1967 [16,
|
||
<collectionCode id="ED49AE3E6F1FFF8BDF14FED0FB70FEAD" box="[1087,1163,328,349]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="4" pageNumber="334">HNHM</collectionCode>
|
||
]. Prov. Córdoba, Fanti,
|
||
<location id="8E8760206F1FFF8BD855FEFBFBD0FE88" LSID="urn:lsid:plazi:treatment:03F187ED6F1FFF8ADBDBFED1FE0AFD45:8E8760206F1FFF8BD855FEFBFBD0FE88" box="[894,1067,355,376]" country="Argentina" county="Arandel" municipality="Leg Pena" name="Sierra de Cordoba" pageId="4" pageNumber="334" stateProvince="Mendoza">
|
||
Sierra
|
||
<collectorName id="26AD532D6F1FFF8BD897FEFBFBD0FE88" box="[956,1067,355,376]" pageId="4" pageNumber="334">de Córdoba</collectorName>
|
||
</location>
|
||
,
|
||
<date id="FFE6103B6F1FFF8BDF1DFEFBFB6BFE88" box="[1078,1168,355,376]" pageId="4" pageNumber="334" value="1966-01-11">
|
||
<collectingDate id="EFA2E9D36F1FFF8BDF1DFEFBFB6BFE88" box="[1078,1168,355,376]" pageId="4" pageNumber="334" value="1966-01-11">11.I.1966</collectingDate>
|
||
</date>
|
||
, Leg. Mahunka, det: Dr.
|
||
<bibRefCitation id="EFC94B0A6F1FFF8BD882FEE6FBC0FE63" author="Endrodi, S." box="[937,1083,382,403]" pageId="4" pageNumber="334" pagination="161 - 177" refId="ref4904" refString="Endrodi, S. 1968. Neue Arten der Pentodontini. (Col. Dynastinae). Folia Entomologica Hungarica 21: 161 - 177." type="journal article" year="1968">Endrödi, 1968</bibRefCitation>
|
||
[26,
|
||
<specimenCount id="9D5EFD726F1FFF8BDF5EFEE6FB76FE63" box="[1141,1165,381,403]" pageId="4" pageNumber="334" type="female">1♀</specimenCount>
|
||
,
|
||
<collectionCode id="ED49AE3E6F1FFF8BD986FE01FD06FE5E" box="[685,765,409,430]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="4" pageNumber="334">HNHM</collectionCode>
|
||
].
|
||
<collectingRegion id="499CF8196F1FFF8BD83FFE01FC8AFE5E" box="[788,881,409,430]" country="Argentina" name="Mendoza" pageId="4" pageNumber="334">Mendoza</collectingRegion>
|
||
,
|
||
<collectorName id="26AD532D6F1FFF8BD8AFFE01FBD5FE5E" box="[900,1070,409,430]" pageId="4" pageNumber="334">C.S. Reed Coll.</collectorName>
|
||
[26,
|
||
<specimenCount id="9D5EFD726F1FFF8BDF5FFE01FB76FE5E" box="[1140,1165,408,430]" pageId="4" pageNumber="334" type="female">2♀</specimenCount>
|
||
,
|
||
<collectionCode id="ED49AE3E6F1FFF8BD986FE2CFD02FE39" box="[685,761,436,457]" country="USA" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="4" pageNumber="334" type="Museum">USNM</collectionCode>
|
||
].
|
||
<collectingRegion id="499CF8196F1FFF8BD822FE2CFC9DFE39" box="[777,870,436,457]" country="Argentina" name="Mendoza" pageId="4" pageNumber="334">Mendoza</collectingRegion>
|
||
,
|
||
<collectorName id="26AD532D6F1FFF8BD85AFE2CFBF1FE39" box="[881,1034,436,457]" pageId="4" pageNumber="334">R. Sanzin Coll.</collectorName>
|
||
[
|
||
<specimenCount id="9D5EFD726F1FFF8BDF32FE2CFBC9FE39" box="[1049,1074,435,457]" pageId="4" pageNumber="334" type="female">1♀</specimenCount>
|
||
,
|
||
<collectionCode id="ED49AE3E6F1FFF8BDF14FE2CFB70FE39" box="[1087,1163,436,457]" country="USA" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="4" pageNumber="334" type="Museum">USNM</collectionCode>
|
||
].
|
||
<collectingRegion id="499CF8196F1FFF8BD986FE57FCFEFE14" box="[685,773,463,484]" country="Argentina" name="Mendoza" pageId="4" pageNumber="334">Mendoza</collectingRegion>
|
||
,
|
||
<locationDeviation id="9A552FD96F1FFF8BD825FE57FC4FFE14" box="[782,948,463,484]" location="Potrerillos" pageId="4" pageNumber="334">
|
||
<quantity id="4CA09B1E6F1FFF8BD825FE57FCC7FE14" box="[782,828,463,484]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="4" pageNumber="334" unit="km" value="4.0">4 km</quantity>
|
||
E Potrerillos
|
||
</locationDeviation>
|
||
,
|
||
<date id="FFE6103B6F1FFF8BD890FE57FBBDFE14" box="[955,1094,463,484]" pageId="4" pageNumber="334" value="1973-12-20" valueMax="1973-12-22" valueMin="1973-12-20">
|
||
<collectingDate id="EFA2E9D36F1FFF8BD890FE57FBBDFE14" box="[955,1094,463,484]" pageId="4" pageNumber="334" value="1973-12-20" valueMax="1973-12-22" valueMin="1973-12-20">20-22.xii.1973</collectingDate>
|
||
</date>
|
||
.
|
||
<collectorName id="26AD532D6F1FFF8BDF66FE57FCBFFE0F" pageId="4" pageNumber="334">C.M. & O.S. Flint Coll.</collectorName>
|
||
[
|
||
<specimenCount id="9D5EFD726F1FFF8BD87FFE72FC97FE0F" box="[852,876,489,511]" pageId="4" pageNumber="334" type="female">1♀</specimenCount>
|
||
,
|
||
<collectionCode id="ED49AE3E6F1FFF8BD857FE72FC3FFE0F" box="[892,964,490,511]" country="USA" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="4" pageNumber="334" type="Museum">USNM</collectionCode>
|
||
]. R´ıo Negro, General Roca,
|
||
<date id="FFE6103B6F1FFF8BD9C1FD9DFCB4FDEA" box="[746,847,517,538]" pageId="4" pageNumber="334" value="1920-11-24">
|
||
<collectingDate id="EFA2E9D36F1FFF8BD9C1FD9DFCB4FDEA" box="[746,847,517,538]" pageId="4" pageNumber="334" value="1920-11-24">24.xi.1920</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26AD532D6F1FFF8BD871FD9DFC37FDEA" box="[858,972,517,538]" pageId="4" pageNumber="334">A. Wetmore</collectorName>
|
||
[
|
||
<specimenCount id="9D5EFD726F1FFF8BD8F2FD9DFC08FDEA" box="[985,1011,516,538]" pageId="4" pageNumber="334" type="female">2♀</specimenCount>
|
||
,
|
||
<collectionCode id="ED49AE3E6F1FFF8BDF2AFD9DFBB2FDEA" box="[1025,1097,517,538]" country="USA" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="4" pageNumber="334" type="Museum">USNM</collectionCode>
|
||
]
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34265706F1FFF8AD9EEFDB9FE0AFD45" lastPageId="5" lastPageNumber="335" pageId="4" pageNumber="334" type="discussion">
|
||
<paragraph id="8BE736FB6F1FFF8AD9EEFDB9FE0AFD45" blockId="4.[685,1173,193,754]" lastBlockId="5.[173,660,193,693]" lastPageId="5" lastPageNumber="335" pageId="4" pageNumber="334">
|
||
<emphasis id="B92CEAE96F1FFF8BD9EEFDB9FCC3FDC6" bold="true" box="[709,824,545,566]" pageId="4" pageNumber="334">Comments.</emphasis>
|
||
<bibRefCitation id="EFC94B0A6F1FFF8BD86DFDB9FC24FDC6" author="Endrodi, S." box="[838,991,545,566]" pageId="4" pageNumber="334" pagination="1 - 145" refId="ref4930" refString="Endrodi, S. 1969. Monographie der Dynastinae. 4. Tribus: Pentodontini (Coleoptera, Lamellicornia). Entomologische Abhandlungen 37: 1 - 145." type="journal article" year="1969">Endrödi (1969)</bibRefCitation>
|
||
indicated that
|
||
<taxonomicName id="4C584D786F1FFF8BDFA9FDB9FD08FDA0" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="334" phylum="Arthropoda" rank="species" species="pumilis">
|
||
<emphasis id="B92CEAE96F1FFF8BDFA9FDB9FD08FDA0" italics="true" pageId="4" pageNumber="334">T. pumilis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C584D786F1FFF8BD80CFDA3FC70FDA0" authorityName="Endrodi" authorityYear="1968" box="[807,907,571,592]" class="Insecta" family="Dynastidae" genus="Ligyrus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="334" phylum="Arthropoda" rank="species" species="rosettae">
|
||
<emphasis id="B92CEAE96F1FFF8BD80CFDA3FC70FDA0" box="[807,907,571,592]" italics="true" pageId="4" pageNumber="334">T. rosettae</emphasis>
|
||
</taxonomicName>
|
||
were very similar and differentiated only by the size of the elytral punctures, which should not be considered a diagnostic character to separate species in the genus. He kept them as different species because of their allopatric distribution,
|
||
<taxonomicName id="4C584D786F1FFF8BD80DFD5AFC7CFD27" baseAuthorityName="Prell" baseAuthorityYear="1937" box="[806,903,706,727]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="334" phylum="Arthropoda" rank="species" species="pumilus">
|
||
<emphasis id="B92CEAE96F1FFF8BD80DFD5AFC7CFD27" box="[806,903,706,727]" italics="true" pageId="4" pageNumber="334">T. pumilus</emphasis>
|
||
</taxonomicName>
|
||
in
|
||
<collectingCountry id="F34F766B6F1FFF8BD88FFD5AFC17FD27" box="[932,1004,706,727]" name="Mexico" pageId="4" pageNumber="334">Mexico</collectingCountry>
|
||
and
|
||
<taxonomicName id="4C584D786F1FFF8BDF31FD5AFB80FD27" authorityName="Endrodi" authorityYear="1968" box="[1050,1147,706,727]" class="Insecta" family="Dynastidae" genus="Ligyrus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="334" phylum="Arthropoda" rank="species" species="rosettae">
|
||
<emphasis id="B92CEAE96F1FFF8BDF31FD5AFB80FD27" box="[1050,1147,706,727]" italics="true" pageId="4" pageNumber="334">T. rosettae</emphasis>
|
||
</taxonomicName>
|
||
in
|
||
<collectingCountry id="F34F766B6F1FFF8BD986FD45FCF7FD02" box="[685,780,733,754]" name="Argentina" pageId="4" pageNumber="334">Argentina</collectingCountry>
|
||
. However,
|
||
<bibRefCitation id="EFC94B0A6F1FFF8BD85EFD45FBC7FD02" author="Ratcliffe, B. C." box="[885,1084,733,754]" pageId="4" pageNumber="334" pagination="1 - 666" refId="ref5521" refString="Ratcliffe, B. C., R. D. Cave, and E. B. Cano. 2013. The dynastine scarab beetles of Mexico, Guatemala, and Belize (Coleoptera: Scarabaeidae: Dynastinae). Bulletin of the University of Nebraska State Museum 27: 1 - 666." type="journal article" year="2013">
|
||
Ratcliffe
|
||
<emphasis id="B92CEAE96F1FFF8BD8E1FD45FC02FD02" box="[970,1017,733,754]" italics="true" pageId="4" pageNumber="334">et al.</emphasis>
|
||
(2013)
|
||
</bibRefCitation>
|
||
reasoned that the
|
||
<typeStatus id="54E388596F1EFF8ADA2AFF5AFED2FF27" box="[257,297,194,215]" pageId="5" pageNumber="335">type</typeStatus>
|
||
locality of
|
||
<taxonomicName id="4C584D786F1EFF8ADA8AFF5AFDF3FF27" baseAuthorityName="Prell" baseAuthorityYear="1937" box="[417,520,194,215]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="335" phylum="Arthropoda" rank="species" species="pumilus">
|
||
<emphasis id="B92CEAE96F1EFF8ADA8AFF5AFDF3FF27" box="[417,520,194,215]" italics="true" pageId="5" pageNumber="335">T. pumilus</emphasis>
|
||
</taxonomicName>
|
||
(“Corralitos”) must be in
|
||
<collectingCountry id="F34F766B6F1EFF8ADA3EFF44FE8EFF01" box="[277,373,220,241]" name="Argentina" pageId="5" pageNumber="335">Argentina</collectingCountry>
|
||
and not in
|
||
<collectingCountry id="F34F766B6F1EFF8ADACBFF44FDD1FF01" box="[480,554,220,241]" name="Mexico" pageId="5" pageNumber="335">Mexico</collectingCountry>
|
||
as thought by
|
||
<bibRefCitation id="EFC94B0A6F1EFF8ADBE0FF6FFE99FEFC" author="Endrodi, S." box="[203,354,247,268]" pageId="5" pageNumber="335" pagination="1 - 145" refId="ref4930" refString="Endrodi, S. 1969. Monographie der Dynastinae. 4. Tribus: Pentodontini (Coleoptera, Lamellicornia). Entomologische Abhandlungen 37: 1 - 145." type="journal article" year="1969">Endrödi (1969)</bibRefCitation>
|
||
. Supporting evidence is that no specimens of
|
||
<taxonomicName id="4C584D786F1EFF8ADA17FE89FE5DFED6" baseAuthorityName="Prell" baseAuthorityYear="1937" box="[316,422,273,294]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="335" phylum="Arthropoda" rank="species" species="pumilus">
|
||
<emphasis id="B92CEAE96F1EFF8ADA17FE89FE5DFED6" box="[316,422,273,294]" italics="true" pageId="5" pageNumber="335">T. pumilus</emphasis>
|
||
</taxonomicName>
|
||
have been found after intensive sampling and revision of collections in
|
||
<collectingCountry id="F34F766B6F1EFF8ADBE1FEDEFEEDFEAB" box="[202,278,326,347]" name="Mexico" pageId="5" pageNumber="335">Mexico</collectingCountry>
|
||
, and the most closely related species (
|
||
<taxonomicName id="4C584D786F1EFF8ADB9EFEF9FDFFFE86" authority="Neita and Ratcliffe" authorityName="Neita and Ratcliffe" box="[181,516,353,374]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="335" phylum="Arthropoda" rank="species" species="roigjunenti">
|
||
<emphasis id="B92CEAE96F1EFF8ADB9EFEF9FEC0FE86" box="[181,315,353,374]" italics="true" pageId="5" pageNumber="335">T. roigjunenti</emphasis>
|
||
Neita and Ratcliffe
|
||
</taxonomicName>
|
||
and an undescribed species) are only found in southern South America. It is very likely that the
|
||
<typeStatus id="54E388596F1EFF8ADAD6FE0EFDDCFE5B" box="[509,551,406,427]" pageId="5" pageNumber="335">type</typeStatus>
|
||
locality of
|
||
<taxonomicName id="4C584D786F1EFF8ADB86FE29FEEBFE36" baseAuthorityName="Prell" baseAuthorityYear="1937" box="[173,272,433,454]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="335" phylum="Arthropoda" rank="species" species="pumilus">
|
||
<emphasis id="B92CEAE96F1EFF8ADB86FE29FEEBFE36" box="[173,272,433,454]" italics="true" pageId="5" pageNumber="335">T. pumilus</emphasis>
|
||
</taxonomicName>
|
||
is Corralito, a locality in Córdoba founded in 1913, more than a decade before the description of
|
||
<taxonomicName id="4C584D786F1EFF8ADBE3FE7EFED0FE0B" baseAuthorityName="Prell" baseAuthorityYear="1937" box="[200,299,486,507]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="335" phylum="Arthropoda" rank="species" species="pumilus">
|
||
<emphasis id="B92CEAE96F1EFF8ADBE3FE7EFED0FE0B" box="[200,299,486,507]" italics="true" pageId="5" pageNumber="335">T. pumilus</emphasis>
|
||
</taxonomicName>
|
||
. After reviewing the
|
||
<typeStatus id="54E388596F1EFF8ADAD7FE7EFDDFFE0B" box="[508,548,486,507]" pageId="5" pageNumber="335">type</typeStatus>
|
||
material of both species, we conclude that these are the same taxon for the following reasons: 1) There are no differences in body length nor in any other characters; 2) The parameres of the male have the same general shape (
|
||
<figureCitation id="13632A7E6F1EFF8ADBD9FDF3FE43FD70" box="[242,440,619,640]" captionStart="Figs" captionStartId="4.[175,214,1663,1682]" captionTargetBox="[157,1166,816,1644]" captionTargetId="figure-356@4.[154,1169,814,1646]" captionTargetPageId="4" captionText="Figs. 14–19. Tomarus pumilus, lectotype: 14) Head and pronotum; 15) Parameres, frontal view; 16) Parameres, lateral view, and labels. Tomarus rosettae, holotype: 17) Head and pronotum; 18) Parameres, frontal view; 19) Parameres, lateral view, and labels." figureDoi="http://doi.org/10.5281/zenodo.5382551" httpUri="https://zenodo.org/record/5382551/files/figure.png" pageId="5" pageNumber="335">Figs. 15, 16, 18, 19</figureCitation>
|
||
), although the apex is slightly wider in some specimens; and 3) Both species occur in Córdoba,
|
||
<collectingCountry id="F34F766B6F1EFF8ADAA6FD38FE15FD45" box="[397,494,672,693]" name="Argentina" pageId="5" pageNumber="335">Argentina</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |