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<document id="65343DEAE3E1E750E494F7A399868908" ID-DOI="10.11646/zootaxa.3990.2.1" ID-GBIF-Dataset="2dd793f4-d2f7-46c2-abbf-9952384cb55b" ID-ISSN="1175-5326" ID-Zenodo-Dep="245442" ID-ZooBank="C3F0F9F4-2879-402C-8504-D12810018E81" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461267390392" checkinUser="plazi" docAuthor="Cai, Yan-Peng &amp; Zhou, Hong-Zhang" docDate="2015" docId="03CC2578FFB91445FF04F313FE51E31F" docLanguage="en" docName="zt03990p196.pdf" docOrigin="Zootaxa 3990 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Quedius (Raphirus) tenuiculus Cai &amp; Zhou, 2015, sp. nov." docType="treatment" docVersion="8" lastPageNumber="166" masterDocId="FFF55D00FFB5144AFF93F70FFFA8E62F" masterDocTitle="Taxonomy of the subgenus Quedius (Raphirus) Stephens (Coleoptera: Staphylinidae: Staphylinini: Quediina) with descriptions of four new species from China" masterLastPageNumber="196" masterPageNumber="151" pageNumber="163" updateTime="1698610850963" updateUser="plazi">
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<mods:title id="FCDC1B111D81EA1EF5FBF2BCC0493BD8">Taxonomy of the subgenus Quedius (Raphirus) Stephens (Coleoptera: Staphylinidae: Staphylinini: Quediina) with descriptions of four new species from China</mods:title>
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<mods:namePart id="A528A7535E8456EE8F176DB2B008D804">Cai, Yan-Peng</mods:namePart>
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<treatment id="03CC2578FFB91445FF04F313FE51E31F" ID-DOI="http://doi.org/10.5281/zenodo.6115643" ID-GBIF-Taxon="119638130" ID-Zenodo-Dep="6115643" LSID="urn:lsid:plazi:treatment:03CC2578FFB91445FF04F313FE51E31F" httpUri="http://treatment.plazi.org/id/03CC2578FFB91445FF04F313FE51E31F" lastPageId="15" lastPageNumber="166" pageId="12" pageNumber="163">
<subSubSection id="C37FC7E5FFB91446FF04F313FE81E278" pageId="12" pageNumber="163" type="nomenclature">
<paragraph id="8BDA946EFFB91446FF04F313FDDAE219" blockId="12.[151,626,1052,1111]" box="[151,626,1052,1078]" pageId="12" pageNumber="163">
<heading id="D0922302FFB91446FF04F313FDDAE219" bold="true" box="[151,626,1052,1078]" fontSize="11" level="1" pageId="12" pageNumber="163" reason="1">
<emphasis id="B911487CFFB91446FF04F313FDDAE219" bold="true" box="[151,626,1052,1078]" pageId="12" pageNumber="163">
<taxonomicName id="4C65EFEDFFB91446FF04F313FDA5E219" ID-CoL="4R3BM" box="[151,525,1052,1078]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="tenuiculus" status="sp. nov." subGenus="Raphirus">
<emphasis id="B911487CFFB91446FF04F313FF54E219" bold="true" box="[151,252,1052,1078]" italics="true" pageId="12" pageNumber="163">Quedius</emphasis>
(
<emphasis id="B911487CFFB91446FE9EF313FED5E219" bold="true" box="[269,381,1052,1078]" italics="true" pageId="12" pageNumber="163">Raphirus</emphasis>
)
<emphasis id="B911487CFFB91446FE1DF313FDA5E219" bold="true" box="[398,525,1052,1078]" italics="true" pageId="12" pageNumber="163">tenuiculus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A222F507FFB91446FD87F313FDDAE219" box="[532,626,1052,1078]" pageId="12" pageNumber="163" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BDA946EFFB91446FF04F331FE81E278" blockId="12.[151,626,1052,1111]" box="[151,297,1086,1111]" pageId="12" pageNumber="163">
(
<figureCitation id="135E88EBFFB91446FF0CF331FEB7E278" box="[159,287,1086,1111]" captionStart="FIGURE 5" captionStartId="13.[151,250,1877,1899]" captionTargetBox="[165,1433,202,1855]" captionTargetId="figure@13.[151,1436,193,1856]" captionTargetPageId="13" captionText="FIGURE 5. Quedius (Raphirus) tenuiculus sp. nov.: A. head (1.0 mm); B. pronotum (1.0 mm); C. elytra (1.0 mm); D. underside of paramere (0.2 mm); E. apical portion of median lobe, parameral view (0.2 mm). (Scale in parentheses)." httpUri="https://zenodo.org/record/245449/files/figure.png" pageId="12" pageNumber="163">Figs. 5, 5-1</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C37FC7E5FFB91446FF04F389FBC9E2C9" pageId="12" pageNumber="163" type="materials_examined">
<paragraph id="8BDA946EFFB91446FF04F389FBC9E2C9" blockId="12.[151,1436,1158,2011]" pageId="12" pageNumber="163">
<emphasis id="B911487CFFB91446FF04F389FEEFE2B0" bold="true" box="[151,327,1158,1183]" pageId="12" pageNumber="163">
<typeStatus id="54DE2ACCFFB91446FF04F389FF7AE2B0" box="[151,210,1158,1183]" pageId="12" pageNumber="163">Type</typeStatus>
material.
</emphasis>
<typeStatus id="54DE2ACCFFB91446FEC3F389FE10E2B0" box="[336,440,1158,1183]" pageId="12" pageNumber="163" type="holotype">Holotype</typeStatus>
♂,
<emphasis id="B911487CFFB91446FE75F389FD0DE2B0" bold="true" box="[486,677,1158,1183]" pageId="12" pageNumber="163">
<collectingCountry id="F372D4FEFFB91446FE75F389FDE0E2B0" box="[486,584,1158,1183]" name="China" pageId="12" pageNumber="163">CHINA</collectingCountry>
: Fujian
</emphasis>
Province: Nanping City, Jianyang District, Huangkeng Township, Guilin Village,
<quantity id="4C9D398BFFB91446FED6F3A4FE6EE2ED" box="[325,454,1195,1219]" metricMagnitude="2" metricUnit="m" metricValue="3.4" metricValueMax="4.1" metricValueMin="2.7" pageId="12" pageNumber="163" unit="m" value="340.0" valueMax="410.0" valueMin="270.0">270410 m</quantity>
,
<date id="FFDBB2AEFFB91446FE41F3A5FDFAE2EC" box="[466,594,1194,1219]" pageId="12" pageNumber="163" value="1960-04-06">6. IV. 1960</date>
, coll. Yong Zuo.
<typeStatus id="54DE2ACCFFB91446FC80F3A4FC2CE2EC" box="[787,900,1195,1219]" pageId="12" pageNumber="163" type="paratype">Paratypes</typeStatus>
:
<emphasis id="B911487CFFB91446FC1CF3A5FBE4E2EC" bold="true" box="[911,1100,1194,1219]" pageId="12" pageNumber="163">
<collectingCountry id="F372D4FEFFB91446FC1CF3A5FC59E2EC" box="[911,1009,1194,1219]" name="China" pageId="12" pageNumber="163">CHINA</collectingCountry>
: Fujian
</emphasis>
Province:
<specimenCount id="9D635FE7FFB91446FB55F3A4FB5BE2ED" box="[1222,1267,1195,1219]" pageId="12" pageNumber="163" type="generic">1 ♂</specimenCount>
, Nanping City, Jianyang District, Huangkeng Township,
<quantity id="4C9D398BFFB91446FDFBF3C1FD40E2C9" box="[616,744,1230,1255]" metricMagnitude="2" metricUnit="m" metricValue="3.1" metricValueMax="3.5" metricValueMin="2.7" pageId="12" pageNumber="163" unit="m" value="310.0" valueMax="350.0" valueMin="270.0">270350 m</quantity>
,
<date id="FFDBB2AEFFB91446FD67F3C0FC2BE2C8" box="[756,899,1230,1255]" pageId="12" pageNumber="163" value="1960-03-27">27. III. 1960</date>
, coll. Chenglin Ma.
</paragraph>
</subSubSection>
<subSubSection id="C37FC7E5FFB91445FF54F3FDFE38E517" lastPageId="15" lastPageNumber="166" pageId="12" pageNumber="163" type="description">
<paragraph id="8BDA946EFFB91446FF54F3FDFAE2E300" blockId="12.[151,1436,1158,2011]" pageId="12" pageNumber="163">
<emphasis id="B911487CFFB91446FF54F3FDFEF1E324" bold="true" box="[199,345,1266,1291]" pageId="12" pageNumber="163">Description.</emphasis>
Entirely dark brown to black; abdomen moderately iridescent; antennae dark brown, labrum and mandibles dark reddish brown, maxillary and labial palpi dark brown; legs blackish brown, with tarsi paler.
</paragraph>
<paragraph id="8BDA946EFFB91446FF54F234FC56E358" blockId="12.[151,1436,1158,2011]" pageId="12" pageNumber="163">
BL =
<quantity id="4C9D398BFFB91446FE9AF235FEC6E37D" box="[265,366,1338,1363]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" pageId="12" pageNumber="163" unit="mm" value="11.5">11.5 mm</quantity>
, BW =
<quantity id="4C9D398BFFB91446FE55F234FDB6E37D" box="[454,542,1339,1363]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="12" pageNumber="163" unit="mm" value="2.7">2.7 mm</quantity>
, HL/PL/EL = 1.00: 1.47: 1.82, HW/PW/EW/AW = 1.00: 1.44: 1.67: 1.44, HW/ HL = 1.16, HEL/HTL = 3.84, PW/PL = 1.14, EW/EL = 1.07, ESL/EL = 0.49.
</paragraph>
<paragraph id="8BDA946EFFB91446FF54F28DFC28E0B8" blockId="12.[151,1436,1158,2011]" pageId="12" pageNumber="163">
Head (
<figureCitation id="135E88EBFFB91446FE80F28DFEFEE3B4" box="[275,342,1410,1435]" captionStart="FIGURE 5" captionStartId="13.[151,250,1877,1899]" captionTargetBox="[165,1433,202,1855]" captionTargetId="figure@13.[151,1436,193,1856]" captionTargetPageId="13" captionText="FIGURE 5. Quedius (Raphirus) tenuiculus sp. nov.: A. head (1.0 mm); B. pronotum (1.0 mm); C. elytra (1.0 mm); D. underside of paramere (0.2 mm); E. apical portion of median lobe, parameral view (0.2 mm). (Scale in parentheses)." httpUri="https://zenodo.org/record/245449/files/figure.png" pageId="12" pageNumber="163">Fig. 5</figureCitation>
A) round, wider than long; eye large and convex, in dorsal view tempora shorter than length of eye, markedly narrowed posteriad; no additional setiferous punctures between anterior frontal setiferous punctures; posterior frontal setiferous puncture situated behind level of posteriomedial margin of eye, separated by distance about as long as diameter of puncture; temporal setiferous puncture touching posterior margin of eye, with several small setiferous punctures arranged in oblique curve below it; one basal setiferous puncture situated closer to nuchal constriction than to posterior frontal setiferous puncture; head with very fine and dense microsculpture of transverse waves and meshes. Antenna slender, with segment I longer than segment II or III, segment III slightly longer than segment II, segments IVXI slightly longer than wide.
</paragraph>
<paragraph id="8BDA946EFFB91446FF54F1ACFAFCE108" blockId="12.[151,1436,1158,2011]" pageId="12" pageNumber="163">
Pronotum (
<figureCitation id="135E88EBFFB91446FED7F1ADFE2EE094" box="[324,390,1698,1723]" captionStart="FIGURE 5" captionStartId="13.[151,250,1877,1899]" captionTargetBox="[165,1433,202,1855]" captionTargetId="figure@13.[151,1436,193,1856]" captionTargetPageId="13" captionText="FIGURE 5. Quedius (Raphirus) tenuiculus sp. nov.: A. head (1.0 mm); B. pronotum (1.0 mm); C. elytra (1.0 mm); D. underside of paramere (0.2 mm); E. apical portion of median lobe, parameral view (0.2 mm). (Scale in parentheses)." httpUri="https://zenodo.org/record/245449/files/figure.png" pageId="12" pageNumber="163">Fig. 5</figureCitation>
B) wider than long, narrowed anteriad, posteriolateral and posterior margins continuously and broadly rounded, lateral margins not explanate; three setiferous punctures in each dorsal row and one or two setiferous punctures in each sublateral row, last sublateral row puncture situated distinctly before level of large lateral setiferous puncture; surface of pronotum with microsculpture similar to that of head, but even denser.
</paragraph>
<paragraph id="8BDA946EFFB91446FF54F03DFEF5E141" blockId="12.[151,1436,1158,2011]" pageId="12" pageNumber="163">
Scutellum (
<figureCitation id="135E88EBFFB91446FEDEF03DFE3BE164" box="[333,403,1842,1867]" captionStart="FIGURE 5" captionStartId="13.[151,250,1877,1899]" captionTargetBox="[165,1433,202,1855]" captionTargetId="figure@13.[151,1436,193,1856]" captionTargetPageId="13" captionText="FIGURE 5. Quedius (Raphirus) tenuiculus sp. nov.: A. head (1.0 mm); B. pronotum (1.0 mm); C. elytra (1.0 mm); D. underside of paramere (0.2 mm); E. apical portion of median lobe, parameral view (0.2 mm). (Scale in parentheses)." httpUri="https://zenodo.org/record/245449/files/figure.png" pageId="12" pageNumber="163">Fig. 5</figureCitation>
C) with dense setiferous punctures, surface with very fine and dense microsculpture of transverse waves.
</paragraph>
<paragraph id="8BDA946EFFB91446FF54F075FAD5E1F4" blockId="12.[151,1436,1158,2011]" pageId="12" pageNumber="163">
Elytra (
<figureCitation id="135E88EBFFB91446FE8CF075FECFE1BC" box="[287,359,1914,1939]" captionStart="FIGURE 5" captionStartId="13.[151,250,1877,1899]" captionTargetBox="[165,1433,202,1855]" captionTargetId="figure@13.[151,1436,193,1856]" captionTargetPageId="13" captionText="FIGURE 5. Quedius (Raphirus) tenuiculus sp. nov.: A. head (1.0 mm); B. pronotum (1.0 mm); C. elytra (1.0 mm); D. underside of paramere (0.2 mm); E. apical portion of median lobe, parameral view (0.2 mm). (Scale in parentheses)." httpUri="https://zenodo.org/record/245449/files/figure.png" pageId="12" pageNumber="163">Fig. 5</figureCitation>
C) moderately long and broad, slightly wider than long, vaguely broadened posteriad, each elytron with surface covered with very dense setiferous punctures, transverse interspaces between punctures smaller than diameters of punctures; surface between punctures without microsculpture. Wings fully developed.
</paragraph>
<caption id="DF1AC4E6FFB81447FF04F05AFA99E1A5" httpUri="https://zenodo.org/record/245449/files/figure.png" pageId="13" pageNumber="164" targetBox="[165,1433,202,1855]" targetPageId="13">
<paragraph id="8BDA946EFFB81447FF04F05AFA99E1A5" blockId="13.[151,1436,1877,1930]" pageId="13" pageNumber="164">
<emphasis id="B911487CFFB81447FF04F05AFEBFE144" bold="true" box="[151,279,1877,1899]" pageId="13" pageNumber="164">FIGURE 5.</emphasis>
<taxonomicName id="4C65EFEDFFB81447FEB1F05AFDC9E144" box="[290,609,1877,1899]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="164" phylum="Arthropoda" rank="species" species="tenuiculus" status="sp. nov." subGenus="Raphirus">
<emphasis id="B911487CFFB81447FEB1F05AFEDEE144" box="[290,374,1877,1899]" italics="true" pageId="13" pageNumber="164">Quedius</emphasis>
(
<emphasis id="B911487CFFB81447FE1AF05AFE4EE144" box="[393,486,1877,1899]" italics="true" pageId="13" pageNumber="164">Raphirus</emphasis>
)
<emphasis id="B911487CFFB81447FE6AF05AFDC9E144" box="[505,609,1877,1899]" italics="true" pageId="13" pageNumber="164">tenuiculus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A222F507FFB81447FDFFF059FD69E144" box="[620,705,1878,1899]" pageId="13" pageNumber="164" rank="species">
<emphasis id="B911487CFFB81447FDFFF059FD69E144" bold="true" box="[620,705,1878,1899]" pageId="13" pageNumber="164">sp. nov.</emphasis>
</taxonomicNameLabel>
: A. head (1.0 mm); B. pronotum (1.0 mm); C. elytra (1.0 mm); D. underside of paramere (0.2 mm); E. apical portion of median lobe, parameral view (0.2 mm). (Scale in parentheses).
</paragraph>
</caption>
<caption id="DF1AC4E6FFBB1444FF04F0AEFE6FE1DB" httpUri="https://zenodo.org/record/245450/files/figure.png" pageId="14" pageNumber="165" targetBox="[163,1424,193,1931]" targetPageId="14">
<paragraph id="8BDA946EFFBB1444FF04F0AEFE6FE1DB" blockId="14.[151,1436,1953,2036]" pageId="14" pageNumber="165">
<emphasis id="B911487CFFBB1444FF04F0AEFE83E199" bold="true" box="[151,299,1953,1975]" pageId="14" pageNumber="165">FIGURE 5-1.</emphasis>
<taxonomicName id="4C65EFEDFFBB1444FEA5F0AEFDDDE198" box="[310,629,1953,1975]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="165" phylum="Arthropoda" rank="species" species="tenuiculus" status="sp. nov." subGenus="Raphirus">
<emphasis id="B911487CFFBB1444FEA5F0AEFE22E198" box="[310,394,1953,1975]" italics="true" pageId="14" pageNumber="165">Quedius</emphasis>
(
<emphasis id="B911487CFFBB1444FE0DF0AEFE53E198" box="[414,507,1953,1975]" italics="true" pageId="14" pageNumber="165">Raphirus</emphasis>
)
<emphasis id="B911487CFFBB1444FD9EF0AEFDDDE198" box="[525,629,1953,1975]" italics="true" pageId="14" pageNumber="165">tenuiculus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A222F507FFBB1444FD13F0AEFD7CE199" box="[640,724,1953,1974]" pageId="14" pageNumber="165" rank="species">
<emphasis id="B911487CFFBB1444FD13F0AEFD7CE199" bold="true" box="[640,724,1953,1974]" pageId="14" pageNumber="165">sp. nov.</emphasis>
</taxonomicNameLabel>
: A. male sternite VIII; B. male sternite IX; C. male tergite X; D. aedeagus, lateral view; E. aedeagus, parameral view; F. underside of paramere; G. apical portion of median lobe, parameral view. (Scale bars = 0.20 mm).
</paragraph>
</caption>
<paragraph id="8BDA946EFFBA1445FF54F798FB23E6D8" blockId="15.[151,1437,151,1328]" pageId="15" pageNumber="166">Abdominal tergite II impunctate, with some fine setae along posterior margin; setiferous punctures of other tergites finer and denser than those of elytra, becoming slightly sparser toward posterior margin of each tergite, and in general toward apex of abdomen; tergite VII with whitish apical seam of palisade setae.</paragraph>
<paragraph id="8BDA946EFFBA1445FF54F60BFC83E53A" blockId="15.[151,1437,151,1328]" pageId="15" pageNumber="166">
Male with first four segments of foretarsus strongly dilated, sub-bilobed, each heavily covered with tenent setae ventrally, segment II slightly wider than apex of tibia; sternite VII with very inconspicuous medioapical emargination at apical margin; tergite VIII with basal ridge complete, nearly straight, with one long seta on each side; sternite VIII (
<figureCitation id="135E88EBFFBA1445FEE3F660FE6EE7A7" box="[368,454,367,392]" captionStart="FIGURE 5" captionStartId="13.[151,250,1877,1899]" captionTargetBox="[165,1433,202,1855]" captionTargetId="figure@13.[151,1436,193,1856]" captionTargetPageId="13" captionText="FIGURE 5. Quedius (Raphirus) tenuiculus sp. nov.: A. head (1.0 mm); B. pronotum (1.0 mm); C. elytra (1.0 mm); D. underside of paramere (0.2 mm); E. apical portion of median lobe, parameral view (0.2 mm). (Scale in parentheses)." httpUri="https://zenodo.org/record/245449/files/figure.png" pageId="15" pageNumber="166">Fig. 5-1</figureCitation>
A) with basal ridge complete, nearly straight, with two long setae on each side, apical margin with a wide and deep sub-semicircular medioapical emargination, a large triangular area in front of the emargination impunctate; sternite IX (
<figureCitation id="135E88EBFFBA1445FDC0F6B8FD06E7FF" box="[595,686,439,464]" captionStart="FIGURE 5" captionStartId="13.[151,250,1877,1899]" captionTargetBox="[165,1433,202,1855]" captionTargetId="figure@13.[151,1436,193,1856]" captionTargetPageId="13" captionText="FIGURE 5. Quedius (Raphirus) tenuiculus sp. nov.: A. head (1.0 mm); B. pronotum (1.0 mm); C. elytra (1.0 mm); D. underside of paramere (0.2 mm); E. apical portion of median lobe, parameral view (0.2 mm). (Scale in parentheses)." httpUri="https://zenodo.org/record/245449/files/figure.png" pageId="15" pageNumber="166">Fig. 5-1</figureCitation>
B) with basal portion wide and short, with a small and shallow medioapical emargination apically; tergite X (
<figureCitation id="135E88EBFFBA1445FD04F6D3FD45E7DA" box="[663,749,476,501]" captionStart="FIGURE 5" captionStartId="13.[151,250,1877,1899]" captionTargetBox="[165,1433,202,1855]" captionTargetId="figure@13.[151,1436,193,1856]" captionTargetPageId="13" captionText="FIGURE 5. Quedius (Raphirus) tenuiculus sp. nov.: A. head (1.0 mm); B. pronotum (1.0 mm); C. elytra (1.0 mm); D. underside of paramere (0.2 mm); E. apical portion of median lobe, parameral view (0.2 mm). (Scale in parentheses)." httpUri="https://zenodo.org/record/245449/files/figure.png" pageId="15" pageNumber="166">Fig. 5-1</figureCitation>
C) with basal side broadly and deeply concave, apical margin widely rounded; aedeagus in lateral view (
<figureCitation id="135E88EBFFBA1445FDEFF6F0FD7CE437" box="[636,724,511,536]" captionStart="FIGURE 5" captionStartId="13.[151,250,1877,1899]" captionTargetBox="[165,1433,202,1855]" captionTargetId="figure@13.[151,1436,193,1856]" captionTargetPageId="13" captionText="FIGURE 5. Quedius (Raphirus) tenuiculus sp. nov.: A. head (1.0 mm); B. pronotum (1.0 mm); C. elytra (1.0 mm); D. underside of paramere (0.2 mm); E. apical portion of median lobe, parameral view (0.2 mm). (Scale in parentheses)." httpUri="https://zenodo.org/record/245449/files/figure.png" pageId="15" pageNumber="166">Fig. 5-1</figureCitation>
D) with apex of paramere protruding distinctly beyond that of median lobe, median lobe with tiny hook-shaped process at apex facing parameral side; aedeagus in parameral view (
<figureCitation id="135E88EBFFBA1445FF4DF548FE9CE44F" box="[222,308,583,608]" captionStart="FIGURE 5" captionStartId="13.[151,250,1877,1899]" captionTargetBox="[165,1433,202,1855]" captionTargetId="figure@13.[151,1436,193,1856]" captionTargetPageId="13" captionText="FIGURE 5. Quedius (Raphirus) tenuiculus sp. nov.: A. head (1.0 mm); B. pronotum (1.0 mm); C. elytra (1.0 mm); D. underside of paramere (0.2 mm); E. apical portion of median lobe, parameral view (0.2 mm). (Scale in parentheses)." httpUri="https://zenodo.org/record/245449/files/figure.png" pageId="15" pageNumber="166">Fig. 5-1</figureCitation>
E) with paramere quite narrow and elongate, nearly parallel-sided in about basal 3/4, apical portion gradually narrowed into narrowly arcuate apex, median lobe voluminous, almost evenly narrowed from base to about basal 4/5, then suddenly constricted into narrow apical portion with subacute apex (
<figureCitation id="135E88EBFFBA1445FB04F580FB4EE487" box="[1175,1254,655,680]" captionStart="FIGURE 5" captionStartId="13.[151,250,1877,1899]" captionTargetBox="[165,1433,202,1855]" captionTargetId="figure@13.[151,1436,193,1856]" captionTargetPageId="13" captionText="FIGURE 5. Quedius (Raphirus) tenuiculus sp. nov.: A. head (1.0 mm); B. pronotum (1.0 mm); C. elytra (1.0 mm); D. underside of paramere (0.2 mm); E. apical portion of median lobe, parameral view (0.2 mm). (Scale in parentheses)." httpUri="https://zenodo.org/record/245449/files/figure.png" pageId="15" pageNumber="166">Figs. 5</figureCitation>
E, 5-1G); apical portion of paramere with four apical setae at apical margin, and two similar setae on each lateral margin far below apex, underside with numerous sensory peg setae, forming a small solid apical field extending posteriad as a long, longitudinal group along each lateral side (
<figureCitation id="135E88EBFFBA1445FDE4F5F3FD6CE53A" box="[631,708,764,789]" captionStart="FIGURE 5" captionStartId="13.[151,250,1877,1899]" captionTargetBox="[165,1433,202,1855]" captionTargetId="figure@13.[151,1436,193,1856]" captionTargetPageId="13" captionText="FIGURE 5. Quedius (Raphirus) tenuiculus sp. nov.: A. head (1.0 mm); B. pronotum (1.0 mm); C. elytra (1.0 mm); D. underside of paramere (0.2 mm); E. apical portion of median lobe, parameral view (0.2 mm). (Scale in parentheses)." httpUri="https://zenodo.org/record/245449/files/figure.png" pageId="15" pageNumber="166">Figs. 5</figureCitation>
D, 5-1F).
</paragraph>
<paragraph id="8BDA946EFFBA1445FF54F410FE38E517" blockId="15.[151,1437,151,1328]" box="[199,400,799,824]" pageId="15" pageNumber="166">Female unknown.</paragraph>
</subSubSection>
<subSubSection id="C37FC7E5FFBA1445FF54F44BFA9EE5AF" pageId="15" pageNumber="166" type="distribution">
<paragraph id="8BDA946EFFBA1445FF54F44BFA9EE5AF" blockId="15.[151,1437,151,1328]" pageId="15" pageNumber="166">
<emphasis id="B911487CFFBA1445FF54F44BFEC8E572" bold="true" box="[199,352,836,861]" pageId="15" pageNumber="166">Distribution.</emphasis>
<taxonomicName id="4C65EFEDFFBA1445FEFAF44BFDEAE573" box="[361,578,836,860]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="166" phylum="Arthropoda" rank="species" species="tenuiculus" status="sp. nov.">
<emphasis id="B911487CFFBA1445FEFAF44BFDEAE573" box="[361,578,836,860]" italics="true" pageId="15" pageNumber="166">Quedius tenuiculus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A222F507FFBA1445FDD9F44BFD0EE572" box="[586,678,836,861]" pageId="15" pageNumber="166" rank="species">
<emphasis id="B911487CFFBA1445FDD9F44BFD0EE572" bold="true" box="[586,678,836,861]" pageId="15" pageNumber="166">sp. nov.</emphasis>
</taxonomicNameLabel>
is at present known only from the
<typeStatus id="54DE2ACCFFBA1445FBAAF44AFBC1E572" box="[1081,1129,837,861]" pageId="15" pageNumber="166">type</typeStatus>
locality in northern Fujian Province, at low altitudes between
<quantity id="4C9D398BFFBA1445FD8CF467FDCEE550" box="[543,614,872,896]" metricMagnitude="2" metricUnit="m" metricValue="2.7" pageId="15" pageNumber="166" unit="m" value="270.0">270 m</quantity>
and
<specimenCount id="9D635FE7FFBA1445FD0FF467FD42E550" box="[668,746,872,896]" pageId="15" pageNumber="166" type="generic">
<quantity id="4C9D398BFFBA1445FD0FF467FD4EE550" box="[668,742,872,896]" metricMagnitude="2" metricUnit="m" metricValue="4.1" pageId="15" pageNumber="166" unit="m" value="410.0">410 m</quantity>
.
</specimenCount>
The specimens were collected from March to April.
</paragraph>
</subSubSection>
<subSubSection id="C37FC7E5FFBA1445FF54F483FD65E2C7" pageId="15" pageNumber="166" type="diagnosis">
<paragraph id="8BDA946EFFBA1445FF54F483FD65E2C7" blockId="15.[151,1437,151,1328]" pageId="15" pageNumber="166">
<emphasis id="B911487CFFBA1445FF54F483FEE9E58A" bold="true" box="[199,321,908,933]" pageId="15" pageNumber="166">Diagnosis.</emphasis>
This new species belongs to the
<emphasis id="B911487CFFBA1445FD3DF483FC9DE58B" box="[686,821,908,932]" italics="true" pageId="15" pageNumber="166">himalayicus</emphasis>
group. Generally, it is most similar to
<taxonomicName id="4C65EFEDFFBA1445FB72F483FF7AE5E7" authority="Smetana, 1996" authorityName="Smetana" authorityYear="1996" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="166" phylum="Arthropoda" rank="species" species="tian">
<emphasis id="B911487CFFBA1445FB72F483FA87E58B" box="[1249,1327,908,932]" italics="true" pageId="15" pageNumber="166">Q. tian</emphasis>
Smetana, 1996
</taxonomicName>
, known from Sichuan, in both appearance and male genitalia. It can be distinguished from the latter by having male sternite VII with a very inconspicuous medioapical emargination, sternite IX with a small and shallow medioapical emargination, aedeagus with paramere somewhat more elongate, nearly parallel-sided in about basal 3/4, apical portion gradually narrowed into narrowly arcuate apex, with more sensory peg setae on its underside, median lobe suddenly constricted at about basal 4/5, with tiny hook-shaped process at apex; whereas
<taxonomicName id="4C65EFEDFFBA1445FAD9F34FFF50E253" authority="Smetana" authorityName="Smetana" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="166" phylum="Arthropoda" rank="species" species="tian">
<emphasis id="B911487CFFBA1445FAD9F34FFA34E278" box="[1354,1436,1088,1111]" italics="true" pageId="15" pageNumber="166">Q. tian</emphasis>
Smetana
</taxonomicName>
has male sternite VII with a more distinct medioapical emargination, sternite IX broadly arcuate apically, without medioapical emargination, aedeagus with paramere somewhat less elongate, lancet-shaped, with fewer sensory peg setae on its underside, median lobe almost conically narrowed to apex, without any constriction throughout its length, without any process at apex.
</paragraph>
</subSubSection>
<subSubSection id="C37FC7E5FFBA1445FF54F3FBFE51E31F" pageId="15" pageNumber="166" type="etymology">
<paragraph id="8BDA946EFFBA1445FF54F3FBFE51E31F" blockId="15.[151,1437,151,1328]" pageId="15" pageNumber="166">
<emphasis id="B911487CFFBA1445FF54F3FBFEE6E322" bold="true" box="[199,334,1268,1293]" pageId="15" pageNumber="166">Etymology.</emphasis>
The specific name is from the Latin adjective
<taxonomicName id="4C65EFEDFFBA1445FCEAF3FBFC44E323" box="[889,1004,1268,1292]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="166" phylum="Arthropoda" rank="species" species="tenuiculus">
<emphasis id="B911487CFFBA1445FCEAF3FBFC44E323" box="[889,1004,1268,1292]" italics="true" pageId="15" pageNumber="166">tenuiculus</emphasis>
</taxonomicName>
(rather thin), referring to the narrow paramere of the male aedeagus.
</paragraph>
</subSubSection>
</treatment>
</document>