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<document id="CB9E948FD4D3AD23105045A4061A80A2" ID-DOI="10.11646/zootaxa.3990.2.1" ID-GBIF-Dataset="2dd793f4-d2f7-46c2-abbf-9952384cb55b" ID-ISSN="1175-5326" ID-Zenodo-Dep="245442" ID-ZooBank="C3F0F9F4-2879-402C-8504-D12810018E81" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461267390392" checkinUser="plazi" docAuthor="Cai, Yan-Peng &amp; Zhou, Hong-Zhang" docDate="2015" docId="03CC2578FFA71458FF04F6F0FD1AE29C" docLanguage="en" docName="zt03990p196.pdf" docOrigin="Zootaxa 3990 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Quedius (Raphirus) shunichii Smetana 2010" docType="treatment" docVersion="8" lastPageNumber="169" masterDocId="FFF55D00FFB5144AFF93F70FFFA8E62F" masterDocTitle="Taxonomy of the subgenus Quedius (Raphirus) Stephens (Coleoptera: Staphylinidae: Staphylinini: Quediina) with descriptions of four new species from China" masterLastPageNumber="196" masterPageNumber="151" pageNumber="169" updateTime="1698610850963" updateUser="plazi">
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<mods:title id="7FF159518D6B856DBEAA90CC57E91EF1">Taxonomy of the subgenus Quedius (Raphirus) Stephens (Coleoptera: Staphylinidae: Staphylinini: Quediina) with descriptions of four new species from China</mods:title>
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<mods:namePart id="1832F2AC0F90132ED6B2971659813485">Cai, Yan-Peng</mods:namePart>
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<subSubSection id="C37FC7E5FFA71458FF04F6F0FE80E4B0" pageId="18" pageNumber="169" type="nomenclature">
<paragraph id="8BDA946EFFA71458FF04F6F0FD16E435" blockId="18.[151,702,511,571]" box="[151,702,511,538]" pageId="18" pageNumber="169">
<heading id="D0922302FFA71458FF04F6F0FD16E435" bold="true" box="[151,702,511,538]" fontSize="11" level="1" pageId="18" pageNumber="169" reason="1">
<taxonomicName id="4C65EFEDFFA71458FF04F6F0FD16E435" ID-CoL="78RP9" authority="Smetana, 2010" authorityName="Smetana" authorityYear="2010" box="[151,702,511,538]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="169" phylum="Arthropoda" rank="species" species="shunichii" subGenus="Raphirus">
<emphasis id="B911487CFFA71458FF04F6F0FD16E435" bold="true" box="[151,702,511,538]" pageId="18" pageNumber="169">
<emphasis id="B911487CFFA71458FF04F6F0FF54E436" bold="true" box="[151,252,511,537]" italics="true" pageId="18" pageNumber="169">Quedius</emphasis>
(
<emphasis id="B911487CFFA71458FE9EF6F0FED5E436" bold="true" box="[269,381,511,537]" italics="true" pageId="18" pageNumber="169">Raphirus</emphasis>
)
<emphasis id="B911487CFFA71458FE1DF6F0FDA8E436" bold="true" box="[398,512,511,537]" italics="true" pageId="18" pageNumber="169">shunichii</emphasis>
<bibRefCitation id="EFF4E99FFFA71458FD94F50FFD16E435" author="Smetana" box="[519,702,512,538]" pageId="18" pageNumber="169" refString="Smetana, A. (2010) Contributions to the knowledge of the Quediina (Coleoptera, Staphylinidae, Staphylinini) of China. Part 38. Genus Quedius Stephens, 1829. Subgenus Raphirus Stephens, 1829. Section 8. Elytra, 38 (2), 249 - 256." type="journal article" year="2010">Smetana, 2010</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BDA946EFFA71458FF04F52DFE81E414" blockId="18.[151,702,511,571]" box="[151,297,546,571]" pageId="18" pageNumber="169">
(
<figureCitation id="135E88EBFFA71458FF0CF52DFEB7E414" box="[159,287,546,571]" captionStart="FIGURE 7" captionStartId="20.[151,250,1877,1899]" captionTargetBox="[182,1420,196,1855]" captionTargetId="figure@20.[162,1424,193,1856]" captionTargetPageId="20" captionText="FIGURE 7. Quedius (Raphirus) shunichii Smetana, 2010: A. head (1.0 mm); B. pronotum (1.0 mm); C. elytra (1.0 mm); D. underside of paramere (0.2 mm); E. apical portion of median lobe, parameral view (0.2 mm). (Scale in parentheses)." httpUri="https://zenodo.org/record/245453/files/figure.png" pageId="18" pageNumber="169">Figs. 7, 7-1</figureCitation>
)
</paragraph>
<paragraph id="8BDA946EFFA71458FF04F565FE80E4B0" blockId="18.[151,1436,618,702]" pageId="18" pageNumber="169">
<bibRefCitation id="EFF4E99FFFA71458FF04F565FE9DE4AF" author="Smetana" box="[151,309,618,640]" pageId="18" pageNumber="169" refString="Smetana, A. (2010) Contributions to the knowledge of the Quediina (Coleoptera, Staphylinidae, Staphylinini) of China. Part 38. Genus Quedius Stephens, 1829. Subgenus Raphirus Stephens, 1829. Section 8. Elytra, 38 (2), 249 - 256." type="journal article" year="2010">Smetana, 2010</bibRefCitation>
: 250 (
<taxonomicName id="4C65EFEDFFA71458FEEAF565FE65E4AF" box="[377,461,618,640]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="169" phylum="Arthropoda" rank="genus">
<emphasis id="B911487CFFA71458FEEAF565FE65E4AF" box="[377,461,618,640]" italics="true" pageId="18" pageNumber="169">Quedius</emphasis>
</taxonomicName>
; subgenus
<taxonomicName id="4C65EFEDFFA71458FDD5F565FD0BE4AF" box="[582,675,618,640]" class="Insecta" family="Staphylinidae" genus="Raphirus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="169" phylum="Arthropoda" rank="genus">
<emphasis id="B911487CFFA71458FDD5F565FD0BE4AF" box="[582,675,618,640]" italics="true" pageId="18" pageNumber="169">Raphirus</emphasis>
</taxonomicName>
;
<typeStatus id="54DE2ACCFFA71458FD27F564FD4EE4AF" box="[692,742,619,640]" pageId="18" pageNumber="169">Type</typeStatus>
locality:
<collectingCountry id="F372D4FEFFA71458FCDFF565FC23E4AF" box="[844,907,618,640]" name="China" pageId="18" pageNumber="169">China</collectingCountry>
, Xilingxue Shan Dujuanlin,
<quantity id="4C9D398BFFA71458FB2AF565FAA3E4AF" box="[1209,1291,618,640]" metricMagnitude="3" metricUnit="m" metricValue="2.18" pageId="18" pageNumber="169" unit="m" value="2180.0">2180 m</quantity>
, DayiXian C. Sichuan).
</paragraph>
</subSubSection>
<subSubSection id="C37FC7E5FFA71458FF04F5A7FAAFE491" box="[151,1287,680,702]" pageId="18" pageNumber="169" type="reference_group">
<paragraph id="8BDA946EFFA71458FF04F5A7FAAFE491" blockId="18.[151,1436,618,702]" box="[151,1287,680,702]" pageId="18" pageNumber="169">
<taxonomicName id="4C65EFEDFFA71458FF04F5A7FDF3E491" ID-CoL="4R39K" authority="Smetana, 2007" authorityName="Smetana" authorityYear="2007" box="[151,603,680,702]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="169" phylum="Arthropoda" rank="species" species="pluvialis" subGenus="Raphirus">
<emphasis id="B911487CFFA71458FF04F5A7FF43E491" box="[151,235,680,702]" italics="true" pageId="18" pageNumber="169">Quedius</emphasis>
(
<emphasis id="B911487CFFA71458FF69F5A7FEFFE491" box="[250,343,680,702]" italics="true" pageId="18" pageNumber="169">Raphirus</emphasis>
)
<emphasis id="B911487CFFA71458FEF6F5A7FE14E491" box="[357,444,680,702]" italics="true" pageId="18" pageNumber="169">pluvialis</emphasis>
<bibRefCitation id="EFF4E99FFFA71458FE50F5A7FDF3E491" author="Smetana" box="[451,603,680,702]" pageId="18" pageNumber="169" refString="Smetana, A. (2007) Contributions to the knowledge of the Quediina (Coleoptera, Staphylinidae, Staphylinini) of China. Part 28. Genera Strouhalium Scheerpeltz, 1962 (section 5) and Quedius Stephens, 1829 (subgenus Microsaurus Dejean, 1833, section 16 and Raphirus Stephens, 1829, section 5). Elytra, 35 (2), 461 - 469." type="journal article" year="2007">Smetana, 2007</bibRefCitation>
</taxonomicName>
, 466 (description; misidentification; non.
<taxonomicName id="4C65EFEDFFA71458FB97F5A7FB54E491" ID-CoL="4R39K" authority="Smetana, 1998" authorityName="Smetana" authorityYear="1998" box="[1028,1276,680,702]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="169" phylum="Arthropoda" rank="species" species="pluvialis">
<emphasis id="B911487CFFA71458FB97F5A7FBF3E491" box="[1028,1115,680,702]" italics="true" pageId="18" pageNumber="169">pluvialis</emphasis>
<bibRefCitation id="EFF4E99FFFA71458FBF2F5A7FB54E491" author="Smetana" box="[1121,1276,680,702]" pageId="18" pageNumber="169" refString="Smetana, A. (1998) Contributions to the knowledge of the Quediina (Coleoptera, Staphylinidae, Staphylinini) of China. Part 10. Genus Quedius Stephens, 1829. Subgenus Raphirus Stephens, 1829. Section 3. Elytra, 26 (1), 99 - 113." type="journal article" year="1998">Smetana, 1998</bibRefCitation>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C37FC7E5FFA71458FF04F5E5FA2DE509" pageId="18" pageNumber="169" type="materials_examined">
<paragraph id="8BDA946EFFA71458FF04F5E5FA2DE509" blockId="18.[151,1436,746,1203]" pageId="18" pageNumber="169">
<emphasis id="B911487CFFA71458FF04F5E5FDF2E52C" bold="true" box="[151,602,746,771]" pageId="18" pageNumber="169">
Material examined.
<collectingCountry id="F372D4FEFFA71458FE19F5E5FE44E52C" box="[394,492,746,771]" name="China" pageId="18" pageNumber="169">CHINA</collectingCountry>
: Sichuan
</emphasis>
Province:
<specimenCount id="9D635FE7FFA71458FD45F5E4FCACE52D" box="[726,772,747,771]" pageId="18" pageNumber="169" type="generic">1 ♂</specimenCount>
, 1 ♀, Baoxing County, Muping,
<quantity id="4C9D398BFFA71458FBEDF5E5FB7EE52D" box="[1150,1238,746,771]" metricMagnitude="3" metricUnit="m" metricValue="1.75" pageId="18" pageNumber="169" unit="m" value="1750.0">1750 m</quantity>
,
<date id="FFDBB2AEFFA71458FB77F5E5FA30E52C" box="[1252,1432,746,771]" pageId="18" pageNumber="169" value="2003-08-02" valueMax="2003-08-05" valueMin="2003-08-02">25. VIII. 2003</date>
, coll. Xiaodong Yu;
<specimenCount id="9D635FE7FFA71458FEE1F400FE36E509" box="[370,414,783,807]" pageId="18" pageNumber="169" type="generic">1 ♂</specimenCount>
, 1 ♀, Baoxing County, Muping,
<quantity id="4C9D398BFFA71458FC9EF401FCCCE509" box="[781,868,782,807]" metricMagnitude="3" metricUnit="m" metricValue="1.8649999999999998" pageId="18" pageNumber="169" unit="m" value="1865.0">1865 m</quantity>
,
<date id="FFDBB2AEFFA71458FCE3F401FB89E508" box="[880,1057,782,807]" pageId="18" pageNumber="169" value="2003-08-02" valueMax="2003-08-05" valueMin="2003-08-02">25. VIII. 2003</date>
, coll. Xiaodong Yu and Jie Wu.
</paragraph>
</subSubSection>
<subSubSection id="C37FC7E5FFA71458FF54F43DFE4FE268" pageId="18" pageNumber="169" type="description">
<paragraph id="8BDA946EFFA71458FF54F43DFB1AE540" blockId="18.[151,1436,746,1203]" pageId="18" pageNumber="169">
<emphasis id="B911487CFFA71458FF54F43DFED5E564" bold="true" box="[199,381,818,843]" pageId="18" pageNumber="169">Measurements.</emphasis>
BL =
<quantity id="4C9D398BFFA71458FE55F43CFD84E565" box="[454,556,819,843]" metricMagnitude="-2" metricUnit="m" metricValue="1.33" pageId="18" pageNumber="169" unit="mm" value="13.3">13.3 mm</quantity>
, BW =
<quantity id="4C9D398BFFA71458FD10F43DFD73E565" box="[643,731,818,843]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="18" pageNumber="169" unit="mm" value="2.6">2.6 mm</quantity>
, HL/PL/EL = 1.00: 1.42: 1.31, HW/PW/EW/AW = 1.00: 1.45: 1.47: 1.50, HW/HL = 1.11, HEL/HTL = 3.17, PW/PL = 1.13, EW/EL = 1.25, ESL/EL = 0.49.
</paragraph>
<paragraph id="8BDA946EFFA71458FF54F475FE4FE268" blockId="18.[151,1436,746,1203]" pageId="18" pageNumber="169">
<emphasis id="B911487CFFA71458FF54F475FE78E5BC" bold="true" box="[199,464,890,915]" pageId="18" pageNumber="169">Description of female.</emphasis>
Female with first four segments of foretarsus similar to those of male, but much less dilated, segment II narrower than apex of tibia; tergite VIII with basal ridge complete and straight, surface without long seta; sternite VIII with basal ridge complete and straight, with two long setae on each side; tergite X (
<figureCitation id="135E88EBFFA71458FAC3F4CDFF09E5D0" captionStart="FIGURE 7" captionStartId="20.[151,250,1877,1899]" captionTargetBox="[182,1420,196,1855]" captionTargetId="figure@20.[162,1424,193,1856]" captionTargetPageId="20" captionText="FIGURE 7. Quedius (Raphirus) shunichii Smetana, 2010: A. head (1.0 mm); B. pronotum (1.0 mm); C. elytra (1.0 mm); D. underside of paramere (0.2 mm); E. apical portion of median lobe, parameral view (0.2 mm). (Scale in parentheses)." httpUri="https://zenodo.org/record/245453/files/figure.png" pageId="18" pageNumber="169">Fig. 7- 1</figureCitation>
H) with basal side broadly and shallowly concave, middle and apical portion more strongly sclerotized and pigmented, apex rod-like, distinctly protruded and pointed, with long setae along protruded portion, and some shorter ones a little anteriorly.
</paragraph>
</subSubSection>
<subSubSection id="C37FC7E5FFA71458FF54F35DFD1AE29C" pageId="18" pageNumber="169" type="distribution">
<paragraph id="8BDA946EFFA71458FF54F35DFD1AE29C" blockId="18.[151,1436,746,1203]" pageId="18" pageNumber="169">
<emphasis id="B911487CFFA71458FF54F35DFEC8E244" bold="true" box="[199,352,1106,1131]" pageId="18" pageNumber="169">Distribution.</emphasis>
<taxonomicName id="4C65EFEDFFA71458FEF8F35CFD0DE244" authority="Smetana" authorityName="Smetana" box="[363,677,1107,1131]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="169" phylum="Arthropoda" rank="species" species="shunichii">
<emphasis id="B911487CFFA71458FEF8F35CFD91E244" box="[363,569,1107,1131]" italics="true" pageId="18" pageNumber="169">Quedius shunichii</emphasis>
Smetana
</taxonomicName>
is known only from the mountains in Dayi County and Baoxing County in central Sichuan Province, at altitudes between
<quantity id="4C9D398BFFA71458FCA4F379FC38E2A1" box="[823,912,1142,1167]" metricMagnitude="3" metricUnit="m" metricValue="1.75" pageId="18" pageNumber="169" unit="m" value="1750.0">1750 m</quantity>
and
<quantity id="4C9D398BFFA71458FC5DF378FB81E2A1" box="[974,1065,1143,1167]" metricMagnitude="3" metricUnit="m" metricValue="2.18" pageId="18" pageNumber="169" unit="m" value="2180.0">2180 m</quantity>
. The examined specimens were collected in forests using pitfall traps in August.
</paragraph>
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</treatment>
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