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<document id="E4F9727927484C02347DC3FE176A9DC8" ID-DOI="10.11646/zootaxa.3990.2.1" ID-GBIF-Dataset="2dd793f4-d2f7-46c2-abbf-9952384cb55b" ID-ISSN="1175-5326" ID-Zenodo-Dep="245442" ID-ZooBank="C3F0F9F4-2879-402C-8504-D12810018E81" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461267390392" checkinUser="plazi" docAuthor="Cai, Yan-Peng &amp; Zhou, Hong-Zhang" docDate="2015" docId="03CC2578FF90146FFF04F280FE27E181" docLanguage="en" docName="zt03990p196.pdf" docOrigin="Zootaxa 3990 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Quedius (Raphirus) torrentum Smetana 2002" docType="treatment" docVersion="8" lastPageNumber="188" masterDocId="FFF55D00FFB5144AFF93F70FFFA8E62F" masterDocTitle="Taxonomy of the subgenus Quedius (Raphirus) Stephens (Coleoptera: Staphylinidae: Staphylinini: Quediina) with descriptions of four new species from China" masterLastPageNumber="196" masterPageNumber="151" pageNumber="188" updateTime="1698610850963" updateUser="plazi">
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<mods:title id="66C3E9A4A269331B8711ED7269676133">Taxonomy of the subgenus Quedius (Raphirus) Stephens (Coleoptera: Staphylinidae: Staphylinini: Quediina) with descriptions of four new species from China</mods:title>
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<treatment id="03CC2578FF90146FFF04F280FE27E181" ID-DOI="http://doi.org/10.5281/zenodo.6115668" ID-GBIF-Taxon="119638124" ID-Zenodo-Dep="6115668" LSID="urn:lsid:plazi:treatment:03CC2578FF90146FFF04F280FE27E181" httpUri="http://treatment.plazi.org/id/03CC2578FF90146FFF04F280FE27E181" lastPageNumber="188" pageId="37" pageNumber="188">
<subSubSection id="C37FC7E5FF90146FFF04F280FB8FE043" pageId="37" pageNumber="188" type="nomenclature">
<paragraph id="8BDA946EFF90146FFF04F280FD6FE385" blockId="37.[151,711,1423,1483]" box="[151,711,1423,1450]" pageId="37" pageNumber="188">
<heading id="D0922302FF90146FFF04F280FD6FE385" bold="true" box="[151,711,1423,1450]" fontSize="11" level="1" pageId="37" pageNumber="188" reason="1">
<taxonomicName id="4C65EFEDFF90146FFF04F280FD6FE385" ID-CoL="4R3BT" authority="Smetana, 2002" authorityName="Smetana" authorityYear="2002" box="[151,711,1423,1450]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="188" phylum="Arthropoda" rank="species" species="torrentum" subGenus="Raphirus">
<emphasis id="B911487CFF90146FFF04F280FD6FE385" bold="true" box="[151,711,1423,1450]" pageId="37" pageNumber="188">
<emphasis id="B911487CFF90146FFF04F280FF54E386" bold="true" box="[151,252,1423,1449]" italics="true" pageId="37" pageNumber="188">Quedius</emphasis>
(
<emphasis id="B911487CFF90146FFE9EF280FED5E386" bold="true" box="[269,381,1423,1449]" italics="true" pageId="37" pageNumber="188">Raphirus</emphasis>
)
<emphasis id="B911487CFF90146FFE1DF280FDA0E386" bold="true" box="[398,520,1423,1449]" italics="true" pageId="37" pageNumber="188">torrentum</emphasis>
<bibRefCitation id="EFF4E99FFF90146FFD83F29FFD6FE385" author="Smetana" box="[528,711,1424,1450]" pageId="37" pageNumber="188" refString="Smetana, A. (2002) Contributions to the knowledge of the Quediina (Coleoptera, Staphylinidae, Staphylinini) of China. Part 21. Genus Quedius Stephens, 1829. Subgenus Raphirus Stephens, 1829. Section 4. Elytra, 30 (1), 119 - 135." type="journal article" year="2002">Smetana, 2002</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BDA946EFF90146FFF04F2BDFEECE3E4" blockId="37.[151,711,1423,1483]" box="[151,324,1458,1483]" pageId="37" pageNumber="188">
(
<figureCitation id="135E88EBFF90146FFF0CF2BDFE93E3E4" box="[159,315,1458,1483]" captionStart="FIGURE 16" captionStartId="40.[151,250,1877,1899]" captionTargetBox="[180,1418,224,1855]" captionTargetId="figure@40.[164,1422,193,1856]" captionTargetPageId="40" captionText="FIGURE 16. Quedius (Raphirus) torrentum Smetana, 2002: A. head (1.0 mm); B. pronotum (1.0 mm); C. elytra (1.0 mm); D. underside of paramere (0.2 mm); E. apical portion of median lobe, parameral view (0.2 mm). (Scale in parentheses)." httpUri="https://zenodo.org/record/245470/files/figure.png" pageId="37" pageNumber="188">Figs. 16, 16-1</figureCitation>
)
</paragraph>
<paragraph id="8BDA946EFF90146FFF04F2F5FB8FE043" blockId="37.[151,1436,1530,1645]" pageId="37" pageNumber="188">
<bibRefCitation id="EFF4E99FFF90146FFF04F2F5FE9AE03F" author="Smetana" box="[151,306,1530,1552]" pageId="37" pageNumber="188" refString="Smetana, A. (2002) Contributions to the knowledge of the Quediina (Coleoptera, Staphylinidae, Staphylinini) of China. Part 21. Genus Quedius Stephens, 1829. Subgenus Raphirus Stephens, 1829. Section 4. Elytra, 30 (1), 119 - 135." type="journal article" year="2002">Smetana, 2002</bibRefCitation>
: 124 (
<taxonomicName id="4C65EFEDFF90146FFEE3F2F5FE6CE03F" box="[368,452,1530,1552]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="188" phylum="Arthropoda" rank="genus">
<emphasis id="B911487CFF90146FFEE3F2F5FE6CE03F" box="[368,452,1530,1552]" italics="true" pageId="37" pageNumber="188">Quedius</emphasis>
</taxonomicName>
; subgenus
<taxonomicName id="4C65EFEDFF90146FFDA4F2F5FD3CE03F" box="[567,660,1530,1552]" class="Insecta" family="Staphylinidae" genus="Raphirus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="188" phylum="Arthropoda" rank="genus">
<emphasis id="B911487CFF90146FFDA4F2F5FD3CE03F" box="[567,660,1530,1552]" italics="true" pageId="37" pageNumber="188">Raphirus</emphasis>
</taxonomicName>
;
<typeStatus id="54DE2ACCFF90146FFD32F2F4FD7BE03F" box="[673,723,1531,1552]" pageId="37" pageNumber="188">Type</typeStatus>
locality:
<collectingCountry id="F372D4FEFF90146FFCA7F2F5FCDBE03F" box="[820,883,1530,1552]" name="China" pageId="37" pageNumber="188">China</collectingCountry>
, W-Hubei Daba Shan crk. valley
<quantity id="4C9D398BFF90146FFB54F2F5FB53E03F" box="[1223,1275,1530,1552]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="37" pageNumber="188" unit="km" value="8.0">8 km</quantity>
NW Muyuping
<geoCoordinate id="EE51F2A9FF90146FFF54F116FEB0E000" box="[199,280,1561,1583]" direction="north" orientation="latitude" pageId="37" pageNumber="188" precision="925" value="31.483334">31°29'N</geoCoordinate>
/
<geoCoordinate id="EE51F2A9FF90146FFEB3F116FED6E000" box="[288,382,1561,1583]" direction="east" orientation="longitude" pageId="37" pageNumber="188" precision="925" value="110.36667">110°22'E</geoCoordinate>
<quantity id="4C9D398BFF90146FFE17F116FDBCE000" box="[388,532,1561,1583]" metricMagnitude="3" metricUnit="m" metricValue="1.6" metricValueMax="1.65" metricValueMin="1.55" pageId="37" pageNumber="188" unit="m" value="1600.0" valueMax="1650.0" valueMin="1550.0">15501650 m</quantity>
);
<bibRefCitation id="EFF4E99FFF90146FFDB5F116FD69E000" author="Smetana" box="[550,705,1561,1583]" pageId="37" pageNumber="188" refString="Smetana, A. (2009) Contributions to the knowledge of the Quediina (Coleoptera: Staphylinidae: Staphylinini) of China. Part 3. Genus Quedius Stephens, 1829. Subgenus Raphirus Stephens, 1829. Section 7. Studies and Reports of District Museum Prague-East Taxonomical Series, 5 (1 - 2), 279 - 294." type="journal article" year="2009">Smetana, 2009</bibRefCitation>
: 280 (
<collectingCountry id="F372D4FEFF90146FFD6EF116FC93E000" box="[765,827,1561,1583]" name="China" pageId="37" pageNumber="188">China</collectingCountry>
(Yunnan));
<bibRefCitation id="EFF4E99FFF90146FFC26F116FBF4E000" author="Smetana" box="[949,1116,1561,1583]" pageId="37" pageNumber="188" refString="Smetana, A. (2014 b) Contributions to the knowledge of the Quediini (Coleoptera: Staphylinidae: Staphylinini) of China. Part 49. Genus Quedius Stephens, 1829. Subgenus Raphirus Stephens, 1829. Section 13. Studies and Reports Taxonomical Series, 10 (2), 595 - 609." type="journal article" year="2014" yearSuffix="b">Smetana, 2014b</bibRefCitation>
: 596 (
<collectingCountry id="F372D4FEFF90146FFB0AF116FB7FE000" box="[1177,1239,1561,1583]" name="China" pageId="37" pageNumber="188">China</collectingCountry>
(Yunnan)).
<taxonomicName id="4C65EFEDFF90146FFF04F137FDF1E061" authority="Zheng, 2002b: 106" authorityName="Zheng" authorityPageNumber="106" authorityYear="2002" box="[151,601,1592,1614]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="188" phylum="Arthropoda" rank="species" species="luae" subGenus="Raphirus">
<emphasis id="B911487CFF90146FFF04F137FF43E061" box="[151,235,1592,1614]" italics="true" pageId="37" pageNumber="188">Quedius</emphasis>
(
<emphasis id="B911487CFF90146FFF69F137FEFFE061" box="[250,343,1592,1614]" italics="true" pageId="37" pageNumber="188">Raphirus</emphasis>
)
<emphasis id="B911487CFF90146FFEF5F137FE3AE061" box="[358,402,1592,1614]" italics="true" pageId="37" pageNumber="188">luae</emphasis>
<bibRefCitation id="EFF4E99FFF90146FFE0BF137FD81E061" author="Zheng" box="[408,553,1592,1614]" pageId="37" pageNumber="188" refString="Zheng, F. K. (2002 b) A New Species of the Group of Quedius (Raphirus) multipunctatus (Coleoptera, Staphylinidae) from Sichuan Province, China. Journal of Sichuan Teachers College (Natural Science), 23 (2), 106 - 109." type="journal article" year="2002" yearSuffix="b">Zheng, 2002b</bibRefCitation>
: 106
</taxonomicName>
(
<taxonomicName id="4C65EFEDFF90146FFDFBF137FD14E061" box="[616,700,1592,1614]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="188" phylum="Arthropoda" rank="genus">
<emphasis id="B911487CFF90146FFDFBF137FD14E061" box="[616,700,1592,1614]" italics="true" pageId="37" pageNumber="188">Quedius</emphasis>
</taxonomicName>
; subgenus
<taxonomicName id="4C65EFEDFF90146FFCBDF137FC23E061" box="[814,907,1592,1614]" class="Insecta" family="Staphylinidae" genus="Raphirus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="188" phylum="Arthropoda" rank="genus">
<emphasis id="B911487CFF90146FFCBDF137FC23E061" box="[814,907,1592,1614]" italics="true" pageId="37" pageNumber="188">Raphirus</emphasis>
</taxonomicName>
;
<typeStatus id="54DE2ACCFF90146FFC0AF136FC63E061" box="[921,971,1593,1614]" pageId="37" pageNumber="188">Type</typeStatus>
locality:
<collectingCountry id="F372D4FEFF90146FFBB8F137FBC2E061" box="[1067,1130,1592,1614]" name="China" pageId="37" pageNumber="188">China</collectingCountry>
, Sichuan, Yujialiu forest land, Daba Shan, Nanjiang County,
<quantity id="4C9D398BFF90146FFE69F159FDE2E043" box="[506,586,1622,1644]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="37" pageNumber="188" unit="m" value="1600.0">1600 m</quantity>
);
<bibRefCitation id="EFF4E99FFF90146FFDCEF159FD5EE043" author="Smetana" box="[605,758,1622,1644]" pageId="37" pageNumber="188" refString="Smetana, A. (2009) Contributions to the knowledge of the Quediina (Coleoptera: Staphylinidae: Staphylinini) of China. Part 3. Genus Quedius Stephens, 1829. Subgenus Raphirus Stephens, 1829. Section 7. Studies and Reports of District Museum Prague-East Taxonomical Series, 5 (1 - 2), 279 - 294." type="journal article" year="2009">Smetana, 2009</bibRefCitation>
: 279 (synonym of
<taxonomicName id="4C65EFEDFF90146FFC27F157FBB0E042" box="[948,1048,1624,1645]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="188" phylum="Arthropoda" rank="species" species="torrentum">
<emphasis id="B911487CFF90146FFC27F157FBB0E042" box="[948,1048,1624,1645]" italics="true" pageId="37" pageNumber="188">torrentum</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C37FC7E5FF90146FFF04F196FF72E0D5" pageId="37" pageNumber="188" type="materials_examined">
<paragraph id="8BDA946EFF90146FFF04F196FF72E0D5" blockId="37.[151,1436,1689,1966]" pageId="37" pageNumber="188">
<emphasis id="B911487CFF90146FFF04F196FDC9E09D" bold="true" box="[151,609,1689,1714]" pageId="37" pageNumber="188">
Material examined.
<collectingCountry id="F372D4FEFF90146FFE1CF196FE59E09D" box="[399,497,1689,1714]" name="China" pageId="37" pageNumber="188">CHINA</collectingCountry>
: Sichuan
</emphasis>
Province:
<specimenCount id="9D635FE7FF90146FFD70F195FCBBE09E" box="[739,787,1690,1714]" pageId="37" pageNumber="188" type="generic">1 ♂</specimenCount>
, Emeishan City, Emei Mountain,
<quantity id="4C9D398BFF90146FFB31F195FA8FE09E" box="[1186,1319,1690,1714]" metricMagnitude="2" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" pageId="37" pageNumber="188" unit="m" value="850.0" valueMax="900.0" valueMin="800.0">800900 m</quantity>
,
<date id="FFDBB2AEFF90146FFAA4F195FF7AE0F9" pageId="37" pageNumber="188" value="2005-08-28">28. VIII. 2005</date>
, coll. Hongzhang Zhou;
<specimenCount id="9D635FE7FF90146FFE74F1B1FDBBE0FA" box="[487,531,1726,1750]" pageId="37" pageNumber="188" type="generic">1 ♂</specimenCount>
, Luding County, Hailuogou, Qingshibangou,
<quantity id="4C9D398BFF90146FFB80F1B1FBC3E0FA" box="[1043,1131,1726,1750]" metricMagnitude="3" metricUnit="m" metricValue="2.471" pageId="37" pageNumber="188" unit="m" value="2471.0">2471 m</quantity>
,
<date id="FFDBB2AEFF90146FFBE5F1B1FB52E0F9" box="[1142,1274,1726,1750]" pageId="37" pageNumber="188" value="2009-05-17">17. V. 2009</date>
, coll. Yulingzi Zhou.
</paragraph>
</subSubSection>
<subSubSection id="C37FC7E5FF90146FFF54F00AFB1BE16D" pageId="37" pageNumber="188" type="description">
<paragraph id="8BDA946EFF90146FFF54F00AFB1BE16D" blockId="37.[151,1436,1689,1966]" pageId="37" pageNumber="188">
<emphasis id="B911487CFF90146FFF54F00AFED5E131" bold="true" box="[199,381,1797,1822]" pageId="37" pageNumber="188">Measurements.</emphasis>
BL =
<quantity id="4C9D398BFF90146FFE5BF00AFD8AE132" box="[456,546,1797,1822]" metricMagnitude="-3" metricUnit="m" metricValue="6.9" pageId="37" pageNumber="188" unit="mm" value="6.9">6.9 mm</quantity>
, BW =
<quantity id="4C9D398BFF90146FFDE8F009FD7CE132" box="[635,724,1798,1822]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="37" pageNumber="188" unit="mm" value="1.4">1.4 mm</quantity>
, HL/PL/EL = 1.00: 1.19: 1.48, HW/PW/EW/AW = 1.00: 1.14: 1.36: 1.19, HW/HL = 1.06, HEL/HTL = 6.49, PW/PL = 1.02, EW/EL = 0.98, ESL/EL = 0.59.
</paragraph>
</subSubSection>
<subSubSection id="C37FC7E5FF90146FFF54F042FE27E181" pageId="37" pageNumber="188" type="distribution">
<paragraph id="8BDA946EFF90146FFF54F042FE27E181" blockId="37.[151,1436,1689,1966]" pageId="37" pageNumber="188">
<emphasis id="B911487CFF90146FFF54F042FEC8E149" bold="true" box="[199,352,1869,1894]" pageId="37" pageNumber="188">Distribution.</emphasis>
<taxonomicName id="4C65EFEDFF90146FFEFFF042FD1AE149" authority="Smetana" authorityName="Smetana" box="[364,690,1869,1894]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="188" phylum="Arthropoda" rank="species" species="torrentum">
<emphasis id="B911487CFF90146FFEFFF042FDEDE14A" box="[364,581,1869,1893]" italics="true" pageId="37" pageNumber="188">Quedius torrentum</emphasis>
Smetana
</taxonomicName>
is distributed in the mountains of southwestern
<collectingCountry id="F372D4FEFF90146FFB6BF042FA93E149" box="[1272,1339,1869,1894]" name="China" pageId="37" pageNumber="188">China</collectingCountry>
(Hubei, Sichuan and Yunnan Province), at altitudes from
<quantity id="4C9D398BFF90146FFD28F07EFCB8E1A6" box="[699,784,1905,1930]" metricMagnitude="3" metricUnit="m" metricValue="1.55" pageId="37" pageNumber="188" unit="m" value="1550.0">1550 m</quantity>
to
<quantity id="4C9D398BFF90146FFCA0F07DFC22E1A6" box="[819,906,1906,1930]" metricMagnitude="3" metricUnit="m" metricValue="2.8899999999999997" pageId="37" pageNumber="188" unit="m" value="2890.0">2890 m</quantity>
. The examined specimens were collected at the edges of small creeks.
</paragraph>
</subSubSection>
</treatment>
</document>