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<document id="5ADCF07C0426CC6755DD528F3103B9EB" ID-CLB-Dataset="59554" ID-DOI="10.5852/ejt.2023.885.2189" ID-GBIF-Dataset="f8509ab7-b884-41aa-a5ee-f8c425a46031" ID-ISSN="2118-9773" ID-Zenodo-Dep="8205500" ID-ZooBank="444CE1D2-8098-4CDE-9B97-927CA13FC39F" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1690898759111" checkinUser="felipe" docAuthor="Means, Jackson C., Bouzan, Rodrigo S., Iniesta, Luiz Felipe Moretti, Martínez-Torres, Daniela, Vasquez-Valverde, Luisa F., Brescovit, Antonio D. &amp; Ivanov, Kaloyan" docDate="2023" docId="03CC1013FFC43C7E9458F9233699F9A3" docLanguage="en" docName="ejt-885_65-85.pdf" docOrigin="European Journal of Taxonomy 885 (1)" docSource="http://dx.doi.org/10.5852/ejt.2023.885.2189" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Dibolostethus sicarius Hoffman 2009" docType="treatment" docVersion="6" lastPageNumber="74" masterDocId="FFF5686BFFC23C77966FFF973518FFDD" masterDocTitle="A review of the previously monotypic tribe Dibolostethini (Chelodesmidae: Chelodesminae) with description of two new species and a summary of the Chelodesmidae of the Tropical Andes Biodiversity Hotspot" masterLastPageNumber="85" masterPageNumber="65" pageNumber="71" updateTime="1699550199751" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="5D8F003ABCB03E2281DAB18E1CD04BE7">A review of the previously monotypic tribe Dibolostethini (Chelodesmidae: Chelodesminae) with description of two new species and a summary of the Chelodesmidae of the Tropical Andes Biodiversity Hotspot</mods:title>
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<mods:namePart id="2B23F808BB18A3D033DBE10B760F46FD">Means, Jackson C.</mods:namePart>
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<mods:affiliation id="57C0826090173784C0F56FF24114D4A4">Pós-graduação em Zoologia, Instituto de Biociências, Universidade de São Paulo, São Paulo, Brazil. &amp; Laboratório Especial de Coleções Zoológicas, Instituto Butantan, Avenida Vital Brasil, 1500, 05503 - 090, São Paulo, Brazil.</mods:affiliation>
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<emphasis id="B9117D17FFC43C719458F923365BF912" bold="true" box="[567,835,1716,1743]" italics="true" pageId="6" pageNumber="71">Dibolostethus sicarius</emphasis>
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<figureCitation id="135EBD80FFC43C7194EBF94F361DF92E" box="[644,773,1752,1779]" captionStart="Fig" captionStartId="7.[189,232,1782,1808]" captionTargetBox="[189,1398,812,1742]" captionTargetId="figure-103@7.[189,1398,806,1742]" captionTargetPageId="7" captionText="Fig. 3. Dibolostethus Hoffman, 2009 somatic characters continued. A, E. Dibolostethus sicarius Hoffman, 2009, holotype, ♂ (VMNH110810). B–D. D. kattani Means, Bouzan, Martínez-Torres &amp; Ivanov sp. nov., holotype, ♂ (ICN-MD-1317-1). A. Coxa 2, mesal view, red arrow = gonopore. B. Left leg 4, posterior view, red arrow = tibial ventro-apical projection. C. Left leg of 5th leg pair showing incrassate femur, red arrow = tibial ventro-apical projection. D. Body ring 5, lateral view, red circle = anterior pair of sternal projections. E. Prefemur 5, red arrow = basal pore. Abbreviations: Cx = coxa; Pf = prefemur." figureDoi="http://doi.org/10.5281/zenodo.8205506" httpUri="https://zenodo.org/record/8205506/files/figure.png" pageId="6" pageNumber="71">Figs 3A, E</figureCitation>
, 
<figureCitation id="135EBD80FFC43C71957EF94F3642F92F" box="[785,858,1752,1778]" captionStart="Fig" captionStartId="8.[189,232,1928,1954]" captionTargetBox="[190,1352,1052,1886]" captionTargetId="figure-271@8.[189,1398,1044,1886]" captionTargetPageId="8" captionText="Fig. 4. Body ring 4, sternal projections. A–B. Dibolostethus sicarius Hoffman, 2009, holotype, ♂ (VMNH110810). C–D. D. inopinatus Means, Bouzan &amp; Ivanov sp. nov., holotype, ♂ (VMNH110813). A, C. Lateral views. B, D. Posterolateral views. Abbreviation: Cx = coxae." figureDoi="http://doi.org/10.5281/zenodo.8205508" httpUri="https://zenodo.org/record/8205508/files/figure.png" pageId="6" pageNumber="71">4A–B</figureCitation>
, 
<figureCitation id="135EBD80FFC43C719509F94F366EF92E" box="[870,886,1752,1779]" captionStart="Fig" captionStartId="9.[189,232,970,996]" captionTargetBox="[189,1397,293,928]" captionTargetId="figure-255@9.[189,1398,287,928]" captionTargetPageId="9" captionText="Fig. 5. Dibolostethus sicarius Hoffman, 2009, holotype, ♂ (VMNH110810), left gonopod. A. Lateral view. B. Anterior view. C. Mesal view. Red circles = cingulum. Dashed lines = prostatic groove. Abbreviations: PfP = prefemoral process; S = solenomere." figureDoi="http://doi.org/10.5281/zenodo.8205510" httpUri="https://zenodo.org/record/8205510/files/figure.png" pageId="6" pageNumber="71">5</figureCitation>
, 
<figureCitation id="135EBD80FFC43C7195ECF94F36B7F92F" box="[899,943,1752,1778]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="13.[189,232,1928,1954]" captionStartId-1="14.[188,231,1892,1918]" captionTargetBox-0="[190,1397,1511,1885]" captionTargetBox-1="[243,1357,497,1838]" captionTargetId-0="figure-343@13.[189,1398,1505,1885]" captionTargetId-1="figure-72@14.[212,1380,484,1853]" captionTargetPageId-0="13" captionTargetPageId-1="14" captionText-0="Fig. 8. Dibolostethus sicarius Hoffman, 2009, paratype, ♀ (VMNH110811), vulvae. A. Left vulva, lateroventral view, with operculum at base. B. Vulvae in situ, held appressed to 2nd leg pair coxae with valve openings facing each other. C. Left vulva, lateral view." captionText-1="Fig. 9. Distribution of Chelodesmidae spp. in the Tropical Andes Biodiversity Hotspot, by country. Note: the type locality of Dibolostethus sicarius Hoffman, 2009 is included for completeness although it falls just outside of the boundaries of the Tropical Andes Biodiversity Hotspot. For information on these taxa see Supp. file 1." figureDoi-0="http://doi.org/10.5281/zenodo.8205516" figureDoi-1="http://doi.org/10.5281/zenodo.8205518" httpUri-0="https://zenodo.org/record/8205516/files/figure.png" httpUri-1="https://zenodo.org/record/8205518/files/figure.png" pageId="6" pageNumber="71">8–9</figureCitation>
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<emphasis id="B9117D17FFC43C7196D2F88E34DCF8EE" box="[189,452,1817,1843]" italics="true" pageId="6" pageNumber="71">Dibolostethus sicarius</emphasis>
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, figs 1–11.
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<emphasis id="B9117D17FFC43C7196D2F8CC342EF8A8" bold="true" box="[189,310,1883,1909]" pageId="6" pageNumber="71">Diagnosis</emphasis>
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Adult males of 
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<emphasis id="B9117D17FFC43C719713F81D3489F87E" box="[380,401,1930,1955]" italics="true" pageId="6" pageNumber="71">D</emphasis>
. 
<emphasis id="B9117D17FFC43C7197CDF81D34E7F87E" box="[418,511,1930,1955]" italics="true" pageId="6" pageNumber="71">sicarius</emphasis>
</taxonomicName>
differ from those of 
<taxonomicName id="4C65DA86FFC43C71956CF81D3663F87E" authority="Means &amp; Bouzan &amp; Iniesta &amp; Martínez-Torres &amp; Vasquez-Valverde &amp; Brescovit &amp; Ivanov, 2023" authorityName="Means &amp; Bouzan &amp; Iniesta &amp; Martínez-Torres &amp; Vasquez-Valverde &amp; Brescovit &amp; Ivanov" authorityYear="2023" box="[771,891,1929,1955]" class="Diplopoda" family="Chelodesmidae" genus="Dibolostethus" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="71" phylum="Arthropoda" rank="species" species="kattani" status="sp. nov.">
<emphasis id="B9117D17FFC43C71956CF81D3663F87E" box="[771,891,1929,1955]" italics="true" pageId="6" pageNumber="71">D. kattani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A222C06CFFC43C7195E9F81D36FBF879" box="[902,995,1930,1956]" pageId="6" pageNumber="71" rank="species">sp. nov.</taxonomicNameLabel>
based on the lack of short, blunt projections on the sternite of the 5 
<superScript id="7C100C4DFFC43C71943CF83B3778F867" attach="left" box="[595,608,1964,1978]" fontSize="6" pageId="6" pageNumber="71">th</superScript>
body ring, and from 
<taxonomicName id="4C65DA86FFC43C719530F83A3118F81B" authority="Means &amp; Bouzan &amp; Iniesta &amp; Martínez-Torres &amp; Vasquez-Valverde &amp; Brescovit &amp; Ivanov, 2023" authorityName="Means &amp; Bouzan &amp; Iniesta &amp; Martínez-Torres &amp; Vasquez-Valverde &amp; Brescovit &amp; Ivanov" authorityYear="2023" box="[863,1024,1965,1990]" class="Diplopoda" family="Chelodesmidae" genus="Dibolostethus" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="71" phylum="Arthropoda" rank="species" species="inopinatus" status="sp. nov.">
<emphasis id="B9117D17FFC43C719530F83A3118F81B" box="[863,1024,1965,1990]" italics="true" pageId="6" pageNumber="71">D. inopinatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A222C06CFFC43C719268F83A317BF81A" box="[1031,1123,1965,1991]" pageId="6" pageNumber="71" rank="species">sp. nov.</taxonomicNameLabel>
and 
<taxonomicName id="4C65DA86FFC43C7192F1F83A300CF81A" authority="Means &amp; Bouzan &amp; Iniesta &amp; Martínez-Torres &amp; Vasquez-Valverde &amp; Brescovit &amp; Ivanov, 2023" authorityName="Means &amp; Bouzan &amp; Iniesta &amp; Martínez-Torres &amp; Vasquez-Valverde &amp; Brescovit &amp; Ivanov" authorityYear="2023" box="[1182,1300,1965,1991]" class="Diplopoda" family="Chelodesmidae" genus="Dibolostethus" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="71" phylum="Arthropoda" rank="species" species="kattani" status="sp. nov.">
<emphasis id="B9117D17FFC43C7192F1F83A300CF81A" box="[1182,1300,1965,1991]" italics="true" pageId="6" pageNumber="71">D. kattani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A222C06CFFC43C719374F83A306FF81A" box="[1307,1399,1965,1991]" pageId="6" pageNumber="71" rank="species">sp. nov.</taxonomicNameLabel>
based on the following combination of gonopodal characteristics: solenomere relatively straight until gently curving dorsally at apex, tip sinuous (
<figureCitation id="135EBD80FFC53C70949AFEBB3642FE9A" box="[757,858,300,327]" captionStart="Fig" captionStartId="9.[189,232,970,996]" captionTargetBox="[189,1397,293,928]" captionTargetId="figure-255@9.[189,1398,287,928]" captionTargetPageId="9" captionText="Fig. 5. Dibolostethus sicarius Hoffman, 2009, holotype, ♂ (VMNH110810), left gonopod. A. Lateral view. B. Anterior view. C. Mesal view. Red circles = cingulum. Dashed lines = prostatic groove. Abbreviations: PfP = prefemoral process; S = solenomere." figureDoi="http://doi.org/10.5281/zenodo.8205510" httpUri="https://zenodo.org/record/8205510/files/figure.png" pageId="7" pageNumber="72">Fig. 5A</figureCitation>
); prefemoral process slightly wider basally, expanded at midpoint and recurved, forming a cup-shaped depression before apex; apex with three lamellate flanges, best seen in anterior view (
<figureCitation id="135EBD80FFC53C7094A2FEE53632FE50" box="[717,810,370,397]" captionStart="Fig" captionStartId="9.[189,232,970,996]" captionTargetBox="[189,1397,293,928]" captionTargetId="figure-255@9.[189,1398,287,928]" captionTargetPageId="9" captionText="Fig. 5. Dibolostethus sicarius Hoffman, 2009, holotype, ♂ (VMNH110810), left gonopod. A. Lateral view. B. Anterior view. C. Mesal view. Red circles = cingulum. Dashed lines = prostatic groove. Abbreviations: PfP = prefemoral process; S = solenomere." figureDoi="http://doi.org/10.5281/zenodo.8205510" httpUri="https://zenodo.org/record/8205510/files/figure.png" pageId="7" pageNumber="72">Fig. 5B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C37FF28EFFC53C7096D2FE2331F9FD75" pageId="7" pageNumber="72" type="materials_examined">
<paragraph id="8BDAA105FFC53C7096D2FE233474FE13" blockId="7.[189,364,436,462]" box="[189,364,436,462]" pageId="7" pageNumber="72">
<heading id="D0921669FFC53C7096D2FE233474FE13" bold="true" box="[189,364,436,462]" fontSize="11" level="3" pageId="7" pageNumber="72" reason="3">
<emphasis id="B9117D17FFC53C7096D2FE233474FE13" bold="true" box="[189,364,436,462]" pageId="7" pageNumber="72">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="8BDAA105FFC53C709685FE7431D9FD9E" blockId="7.[189,1399,483,580]" pageId="7" pageNumber="72">
<materialsCitation id="3B0DAB58FFC53C709685FE7431D9FD9E" ID-GBIF-Occurrence="4167732304" collectingDate="1977-03-21" collectionCode="VMNH" collectorName="J. Reiskind" country="Ecuador" latitude="0.588" location="Rio Palenque Science Center" longLatPrecision="78" longitude="-79.362" pageId="7" pageNumber="72" specimenCode="VMNH110810" specimenCount="1" specimenCount-male="1" stateProvince="Los Rios Province" typeStatus="holotype">
<emphasis id="B9117D17FFC53C709685FE743444FE20" bold="true" box="[234,348,483,509]" pageId="7" pageNumber="72">
<typeStatus id="54DE1FA7FFC53C709685FE743444FE20" box="[234,348,483,509]" pageId="7" pageNumber="72" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="F372E195FFC53C7096D2FD913453FDFD" box="[189,331,518,544]" name="Ecuador" pageId="7" pageNumber="72">ECUADOR</collectingCountry>
• 
<specimenCount id="9D636A8CFFC53C70970FFD913461FDFD" box="[352,377,518,544]" pageId="7" pageNumber="72" type="male">♂</specimenCount>
; 
<collectingRegion id="49A16FE7FFC53C7097ECFD913740FDFD" box="[387,600,518,544]" country="Ecuador" name="Los Rios" pageId="7" pageNumber="72">Los Rios Province</collectingRegion>
[originally reported incorrectly as Pichinchi Province], 
<location id="8EBAF7DEFFC53C7092B4FD91346AFD9E" LSID="urn:lsid:plazi:treatment:03CC1013FFC43C7E9458F9233699F9A3:8EBAF7DEFFC53C7092B4FD91346AFD9E" country="Ecuador" latitude="0.588" longLatPrecision="78" longitude="-79.362" name="Rio Palenque Science Center" pageId="7" pageNumber="72" stateProvince="Los Rios Province">Rio Palenque Science Center</location>
; [
<geoCoordinate id="EE51C7C2FFC53C7097EBFDBE34ECFD9E" box="[388,500,553,580]" degrees="0.588" direction="north" orientation="latitude" pageId="7" pageNumber="72" precision="55" value="0.588">0.588° N</geoCoordinate>
, 
<geoCoordinate id="EE51C7C2FFC53C70946FFDBE379CFD9E" box="[512,644,553,579]" degrees="79.362" direction="west" orientation="longitude" pageId="7" pageNumber="72" precision="55" value="-79.362">79.362° W</geoCoordinate>
]; 
<date id="FFDB87C5FFC53C7094F7FDBE3621FD9E" box="[664,825,553,579]" pageId="7" pageNumber="72" value="1977-03-21">
<collectingDate id="EF9F7E2DFFC53C7094F7FDBE3621FD9E" box="[664,825,553,579]" pageId="7" pageNumber="72" value="1977-03-21">21 Mar. 1977</collectingDate>
</date>
; 
<collectorName id="2690C4D3FFC53C70952AFDBE36DFFD9E" box="[837,967,553,579]" pageId="7" pageNumber="72">J. Reiskind</collectorName>
leg.; 
<specimenCode id="DBC3097EFFC53C709266FDBE31A5FD9E" box="[1033,1213,553,579]" collectionCode="VMNH" country="USA" httpUri="http://grbio.org/cool/zxpe-c0as" name="Virginia Museum of Natural History" pageId="7" pageNumber="72">VMNH110810</specimenCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BDAA105FFC53C709685FDFC31F9FD75" blockId="7.[189,1249,619,680]" pageId="7" pageNumber="72">
<materialsCitation id="3B0DAB58FFC53C709685FDFC31C5FD75" ID-GBIF-Occurrence="4167732306" collectingDate="1977-03-21" collectionCode="VMNH" collectorName="J. Reiskind" country="Ecuador" latitude="0.588" location="ECUADOR" longLatPrecision="78" longitude="-79.362" pageId="7" pageNumber="72" specimenCode="VMNH110811, VMNH110812" specimenCount="2" specimenCount-female="2" stateProvince="Los Rios" typeStatus="paratype">
<emphasis id="B9117D17FFC53C709685FDFC347FFD58" bold="true" box="[234,359,619,645]" pageId="7" pageNumber="72">
<typeStatus id="54DE1FA7FFC53C709685FDFC347FFD58" box="[234,359,619,645]" pageId="7" pageNumber="72" type="paratype">Paratypes</typeStatus>
</emphasis>
<collectingCountry id="F372E195FFC53C7096D2FD193453FD75" box="[189,331,654,680]" name="Ecuador" pageId="7" pageNumber="72">ECUADOR</collectingCountry>
• 
<specimenCount id="9D636A8CFFC53C70970BFD1934BFFD75" box="[356,423,654,680]" pageId="7" pageNumber="72" type="female">2 ♀♀</specimenCount>
; same collection data as for holotype; 
<specimenCode id="DBC3097EFFC53C709504FD193105FD75" box="[875,1053,654,680]" collectionCode="VMNH" country="USA" httpUri="http://grbio.org/cool/zxpe-c0as" name="Virginia Museum of Natural History" pageId="7" pageNumber="72">VMNH110811</specimenCode>
, 
<specimenCode id="DBC3097EFFC53C709247FD1931C5FD75" box="[1064,1245,654,680]" collectionCode="VMNH" country="USA" httpUri="http://grbio.org/cool/zxpe-c0as" name="Virginia Museum of Natural History" pageId="7" pageNumber="72">VMNH110812</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DF1AF18DFFC53C7096D2F9613470F839" ID-DOI="http://doi.org/10.5281/zenodo.8205506" ID-Zenodo-Dep="8205506" httpUri="https://zenodo.org/record/8205506/files/figure.png" pageId="7" pageNumber="72" startId="7.[189,232,1782,1808]" targetBox="[189,1398,812,1742]" targetPageId="7">
<paragraph id="8BDAA105FFC53C7096D2F9613470F839" blockId="7.[189,1399,1782,2020]" pageId="7" pageNumber="72">
<emphasis id="B9117D17FFC53C7096D2F961340AF8CD" bold="true" box="[189,274,1782,1808]" pageId="7" pageNumber="72">Fig. 3.</emphasis>
<taxonomicName id="4C65DA86FFC53C70974EF9613795F8CC" authority="Hoffman, 2009" authorityName="Hoffman" authorityYear="2009" box="[289,653,1782,1809]" class="Diplopoda" family="Chelodesmidae" genus="Dibolostethus" kingdom="Animalia" order="Polydesmida" pageId="7" pageNumber="72" phylum="Arthropoda" rank="genus">
<emphasis id="B9117D17FFC53C70974EF96134DCF8CD" box="[289,452,1782,1808]" italics="true" pageId="7" pageNumber="72">Dibolostethus</emphasis>
<bibRefCitation id="EFF4DCF4FFC53C7097BCF9613795F8CC" author="Hoffman" box="[467,653,1782,1809]" firstAuthor="Hoffman" pageId="7" pageNumber="72" pagination="149 - 154" refId="ref9561" refString="Hoffman R. L. 2009. Chelodesmid studies. XXVI. A new genus and tribe of chelodesmid millipeds from Ecuador (Polydesmida: Chelodesmidae). International Journal of Myriapodology 2: 149 - 154. https: // doi. org / 10.1163 / 187525409 X 12577705044584" type="journal article" year="2009">Hoffman, 2009</bibRefCitation>
</taxonomicName>
somatic characters continued. 
<emphasis id="B9117D17FFC53C70927BF961314BF8CC" bold="true" box="[1044,1107,1782,1809]" pageId="7" pageNumber="72">A, E</emphasis>
. 
<taxonomicName id="4C65DA86FFC53C709207F961346FF8E9" authority="Hoffman, 2009" authorityName="Hoffman" authorityYear="2009" class="Diplopoda" family="Chelodesmidae" genus="Dibolostethus" kingdom="Animalia" order="Polydesmida" pageId="7" pageNumber="72" phylum="Arthropoda" rank="species" species="sicarius">
<emphasis id="B9117D17FFC53C709207F961306FF8CD" box="[1128,1399,1782,1808]" italics="true" pageId="7" pageNumber="72">Dibolostethus sicarius</emphasis>
<bibRefCitation id="EFF4DCF4FFC53C7096D2F88D346FF8E9" author="Hoffman" box="[189,375,1818,1844]" firstAuthor="Hoffman" pageId="7" pageNumber="72" pagination="149 - 154" refId="ref9561" refString="Hoffman R. L. 2009. Chelodesmid studies. XXVI. A new genus and tribe of chelodesmid millipeds from Ecuador (Polydesmida: Chelodesmidae). International Journal of Myriapodology 2: 149 - 154. https: // doi. org / 10.1163 / 187525409 X 12577705044584" type="journal article" year="2009">Hoffman, 2009</bibRefCitation>
</taxonomicName>
, holotype, ♂ (VMNH110810). 
<emphasis id="B9117D17FFC53C709496F88D3628F8E9" bold="true" box="[761,816,1818,1844]" pageId="7" pageNumber="72">B–D</emphasis>
. 
<taxonomicName id="4C65DA86FFC53C70952CF88D3416F88A" authority="Means, Bouzan, Martinez-Torres &amp; Ivanov" authorityName="Means, Bouzan, Martinez-Torres &amp; Ivanov" authorityYear="2023" class="Diplopoda" family="Chelodesmidae" genus="Dibolostethus" kingdom="Animalia" order="Polydesmida" pageId="7" pageNumber="72" phylum="Arthropoda" rank="species" species="kattani" status="sp. nov.">
<emphasis id="B9117D17FFC53C70952CF88D36A4F8E9" box="[835,956,1818,1844]" italics="true" pageId="7" pageNumber="72">D. kattani</emphasis>
Means, Bouzan, Martínez-Torres &amp; Ivanov
</taxonomicName>
<taxonomicNameLabel id="A222C06CFFC53C709779F8AA346BF88A" box="[278,371,1853,1879]" pageId="7" pageNumber="72" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♂ (ICN-MD-1317-1). 
<emphasis id="B9117D17FFC53C709492F8AA360AF88A" bold="true" box="[765,786,1853,1879]" pageId="7" pageNumber="72">A</emphasis>
. Coxa 2, mesal view, red arrow = gonopore. 
<emphasis id="B9117D17FFC53C70934CF8AA302EF88A" bold="true" box="[1315,1334,1853,1879]" pageId="7" pageNumber="72">B</emphasis>
. Left leg 4, posterior view, red arrow = tibial ventro-apical projection. 
<emphasis id="B9117D17FFC53C7095BBF8F736F1F8A7" bold="true" box="[980,1001,1888,1914]" pageId="7" pageNumber="72">C</emphasis>
. Left leg of 5 
<superScript id="7C100C4DFFC53C7092FAF8C831BAF8B0" attach="left" box="[1173,1186,1887,1901]" fontSize="6" pageId="7" pageNumber="72">th</superScript>
leg pair showing incrassate femur, red arrow = tibial ventro-apical projection. 
<emphasis id="B9117D17FFC53C7095FCF81436BFF840" bold="true" box="[915,935,1923,1949]" pageId="7" pageNumber="72">D</emphasis>
. Body ring 5, lateral view, red circle = anterior pair of sternal projections. 
<emphasis id="B9117D17FFC53C70940AF830376FF81C" bold="true" box="[613,631,1959,1985]" pageId="7" pageNumber="72">E</emphasis>
. Prefemur 5, red arrow = basal pore. Abbreviations: Cx = coxa; Pf = prefemur.
</paragraph>
</caption>
<subSubSection id="C37FF28EFFCA3C7E96D2FE9F31E1F9D2" lastPageId="9" lastPageNumber="74" pageId="8" pageNumber="73" type="description">
<paragraph id="8BDAA105FFCA3C7F96D2FE9F3456FEFF" blockId="8.[189,334,264,290]" box="[189,334,264,290]" pageId="8" pageNumber="73">
<heading id="D0921669FFCA3C7F96D2FE9F3456FEFF" bold="true" box="[189,334,264,290]" fontSize="11" level="3" pageId="8" pageNumber="73" reason="3">
<emphasis id="B9117D17FFCA3C7F96D2FE9F3456FEFF" bold="true" box="[189,334,264,290]" pageId="8" pageNumber="73">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BDAA105FFCA3C7F9685FEA03772FE8C" blockId="8.[186,1400,311,549]" box="[234,618,311,337]" pageId="8" pageNumber="73">
<emphasis id="B9117D17FFCA3C7F9685FEA03432FE8C" bold="true" box="[234,298,311,337]" pageId="8" pageNumber="73">Male</emphasis>
(
<typeStatus id="54DE1FA7FFCA3C7F9756FEA034BCFE8C" box="[313,420,311,337]" pageId="8" pageNumber="73" type="holotype">holotype</typeStatus>
, VMNH110810)
</paragraph>
<paragraph id="8BDAA105FFCA3C7F96D2FECD31DFFDF9" blockId="8.[186,1400,311,549]" pageId="8" pageNumber="73">
With characteristics typical for the genus. Coloration (preserved in 70% isopropanol): body rings light brown to caramel; antennae and head light yellow; legs light yellow; telson reddish, with posterior margin (= spinnerets region) yellowish. Head: epicranial macrosetae 2+2, interantennal macrosetae 1+1, frontal macrosetae 2+2. Sternites: 4 
<superScript id="7C100C4DFFCA3C7F9437FE54377DFE0C" attach="left" box="[600,613,451,465]" fontSize="6" pageId="8" pageNumber="73">th</superScript>
body ring with two thin, blade-like acute projections, tips separated throughout, nearly translucent (
<figureCitation id="135EBD80FFCA3C7F944DFE703787FDDC" box="[546,671,487,513]" captionStart="Fig" captionStartId="8.[189,232,1928,1954]" captionTargetBox="[190,1352,1052,1886]" captionTargetId="figure-271@8.[189,1398,1044,1886]" captionTargetPageId="8" captionText="Fig. 4. Body ring 4, sternal projections. A–B. Dibolostethus sicarius Hoffman, 2009, holotype, ♂ (VMNH110810). C–D. D. inopinatus Means, Bouzan &amp; Ivanov sp. nov., holotype, ♂ (VMNH110813). A, C. Lateral views. B, D. Posterolateral views. Abbreviation: Cx = coxae." figureDoi="http://doi.org/10.5281/zenodo.8205508" httpUri="https://zenodo.org/record/8205508/files/figure.png" pageId="8" pageNumber="73">Fig. 4A–B</figureCitation>
). Gonopod aperture on 7 
<superScript id="7C100C4DFFCA3C7F95D9FE7136DBFE29" attach="left" box="[950,963,486,500]" fontSize="6" pageId="8" pageNumber="73">th</superScript>
body ring oval, rim notably projecting above sternite (
<quantity id="4C9D0CE0FFCA3C7F9704FD9D34C1FDF9" box="[363,473,522,549]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" pageId="8" pageNumber="73" unit="mm" value="0.35">0.35 mm</quantity>
), with a deep U-shaped concavity (
<quantity id="4C9D0CE0FFCA3C7F9500FD9D36D3FDF9" box="[879,971,522,548]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="8" pageNumber="73" unit="mm" value="0.2">0.2 mm</quantity>
) on posterior margin.
</paragraph>
<paragraph id="8BDAA105FFCA3C7F96D2FDDB31DAFCCB" blockId="8.[186,1399,587,790]" pageId="8" pageNumber="73">
<smallCapsWord id="8D3C37D9FFCA3C7F96D2FDDB342CFDB9" baselines="607,607" box="[189,308,588,614]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Gonopods" pageId="8" pageNumber="73">GONOPODS</smallCapsWord>
(
<figureCitation id="135EBD80FFCA3C7F9729FDDB3496FDBB" box="[326,398,587,614]" captionStart="Fig" captionStartId="9.[189,232,970,996]" captionTargetBox="[189,1397,293,928]" captionTargetId="figure-255@9.[189,1398,287,928]" captionTargetPageId="9" captionText="Fig. 5. Dibolostethus sicarius Hoffman, 2009, holotype, ♂ (VMNH110810), left gonopod. A. Lateral view. B. Anterior view. C. Mesal view. Red circles = cingulum. Dashed lines = prostatic groove. Abbreviations: PfP = prefemoral process; S = solenomere." figureDoi="http://doi.org/10.5281/zenodo.8205510" httpUri="https://zenodo.org/record/8205510/files/figure.png" pageId="8" pageNumber="73">Fig. 5</figureCitation>
). Gonocoxae rounded, equivalent to about three quarters the length of the telopodite; dorsum of left and right gonocoxa with 2 and 3 (possible aberration) macrosetae, respectively. Solenomere long, slender, unbranched, apex falciform and partially enveloped by prefemoral process (
<figureCitation id="135EBD80FFCA3C7F9293FD05305DFD70" box="[1276,1349,658,685]" captionStart="Fig" captionStartId="9.[189,232,970,996]" captionTargetBox="[189,1397,293,928]" captionTargetId="figure-255@9.[189,1398,287,928]" captionTargetPageId="9" captionText="Fig. 5. Dibolostethus sicarius Hoffman, 2009, holotype, ♂ (VMNH110810), left gonopod. A. Lateral view. B. Anterior view. C. Mesal view. Red circles = cingulum. Dashed lines = prostatic groove. Abbreviations: PfP = prefemoral process; S = solenomere." figureDoi="http://doi.org/10.5281/zenodo.8205510" httpUri="https://zenodo.org/record/8205510/files/figure.png" pageId="8" pageNumber="73">Fig. 5</figureCitation>
, S). Prefemoral process: long (similar in length to solenomere), width subequal to solenomere, widening at halfway point with a medial knob before curving laterad into a cup-like structure; apex broad, with a series of lamellate flanges resembling a hammerhead shark in lateral view (
<figureCitation id="135EBD80FFCA3C7F925DFD6B3160FCCB" box="[1074,1144,763,790]" captionStart="Fig" captionStartId="9.[189,232,970,996]" captionTargetBox="[189,1397,293,928]" captionTargetId="figure-255@9.[189,1398,287,928]" captionTargetPageId="9" captionText="Fig. 5. Dibolostethus sicarius Hoffman, 2009, holotype, ♂ (VMNH110810), left gonopod. A. Lateral view. B. Anterior view. C. Mesal view. Red circles = cingulum. Dashed lines = prostatic groove. Abbreviations: PfP = prefemoral process; S = solenomere." figureDoi="http://doi.org/10.5281/zenodo.8205510" httpUri="https://zenodo.org/record/8205510/files/figure.png" pageId="8" pageNumber="73">Fig. 5</figureCitation>
, PfP).
</paragraph>
<paragraph id="8BDAA105FFCA3C7F96D2FCA934F8FC39" blockId="8.[189,1399,828,996]" pageId="8" pageNumber="73">
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<smallCapsWord id="8D3C37D9FFCA3C7F9768FCD434A8FC8B" baselines="849" box="[263,432,835,854]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="measurements" pageId="8" pageNumber="73">MEASUREMENTS</smallCapsWord>
(mm). Total length 24.8, width of 10 
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body segment 1.98. Antennomere lengths (1–7): 0.38, 0.75, 0.85, 0.85, 0.85, 0.85, 0.25. Collum: length 0.5, width 1.4. Podomere lengths of 10 
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body ring (1–7): 0.28, 0.55, 0.9, 0.34, 0.53, 0.95, 0.09. Gonopod aperture: length 0.50, width 0.78. Gonopod: length 0.99, width 0.86. Gonocoxae: length 0.63, width 0.36. Telopodite: length 0.98, width 0.31. Telson length 0.78.
</paragraph>
<caption id="DF1AF18DFFCA3C7F96D2F81F3132F834" ID-DOI="http://doi.org/10.5281/zenodo.8205508" ID-Zenodo-Dep="8205508" httpUri="https://zenodo.org/record/8205508/files/figure.png" pageId="8" pageNumber="73" startId="8.[189,232,1928,1954]" targetBox="[190,1352,1052,1886]" targetPageId="8">
<paragraph id="8BDAA105FFCA3C7F96D2F81F3132F834" blockId="8.[189,1399,1928,2025]" pageId="8" pageNumber="73">
<emphasis id="B9117D17FFCA3C7F96D2F81F3408F87F" bold="true" box="[189,272,1928,1954]" pageId="8" pageNumber="73">Fig. 4.</emphasis>
Body ring 4, sternal projections. 
<emphasis id="B9117D17FFCA3C7F94D8F81F37F7F87F" bold="true" box="[695,751,1928,1954]" pageId="8" pageNumber="73">A–B</emphasis>
. 
<taxonomicName id="4C65DA86FFCA3C7F956DF81F31CEF87F" authority="Hoffman, 2009" authorityName="Hoffman" authorityYear="2009" box="[770,1238,1928,1954]" class="Diplopoda" family="Chelodesmidae" genus="Dibolostethus" kingdom="Animalia" order="Polydesmida" pageId="8" pageNumber="73" phylum="Arthropoda" rank="species" species="sicarius">
<emphasis id="B9117D17FFCA3C7F956DF81F3117F87F" box="[770,1039,1928,1954]" italics="true" pageId="8" pageNumber="73">Dibolostethus sicarius</emphasis>
<bibRefCitation id="EFF4DCF4FFCA3C7F9274F81F31CEF87F" author="Hoffman" box="[1051,1238,1928,1954]" firstAuthor="Hoffman" pageId="8" pageNumber="73" pagination="149 - 154" refId="ref9561" refString="Hoffman R. L. 2009. Chelodesmid studies. XXVI. A new genus and tribe of chelodesmid millipeds from Ecuador (Polydesmida: Chelodesmidae). International Journal of Myriapodology 2: 149 - 154. https: // doi. org / 10.1163 / 187525409 X 12577705044584" type="journal article" year="2009">Hoffman, 2009</bibRefCitation>
</taxonomicName>
, holotype, ♂ (VMNH110810). 
<emphasis id="B9117D17FFCA3C7F97E0F83C34D0F818" bold="true" box="[399,456,1963,1989]" pageId="8" pageNumber="73">C–D</emphasis>
. 
<taxonomicName id="4C65DA86FFCA3C7F97B8F83B36B1F81B" authority="Means, Bouzan &amp; Ivanov" authorityName="Means, Bouzan &amp; Ivanov" authorityYear="2023" box="[471,937,1963,1990]" class="Diplopoda" family="Chelodesmidae" genus="Dibolostethus" kingdom="Animalia" order="Polydesmida" pageId="8" pageNumber="73" phylum="Arthropoda" rank="species" species="inopinatus" status="sp. nov.">
<emphasis id="B9117D17FFCA3C7F97B8F83B376EF818" box="[471,630,1964,1989]" italics="true" pageId="8" pageNumber="73">D. inopinatus</emphasis>
Means, Bouzan &amp; Ivanov
</taxonomicName>
<taxonomicNameLabel id="A222C06CFFCA3C7F95DFF83B3113F81B" box="[944,1035,1964,1990]" pageId="8" pageNumber="73" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♂ (VMNH110813). 
<emphasis id="B9117D17FFCA3C7F96D2F85935EEF835" bold="true" box="[189,246,1998,2024]" pageId="8" pageNumber="73">A, C</emphasis>
. Lateral views. 
<emphasis id="B9117D17FFCA3C7F97DDF85934F1F835" bold="true" box="[434,489,1998,2024]" pageId="8" pageNumber="73">B, D</emphasis>
. Posterolateral views. Abbreviation: Cx = coxae.
</paragraph>
</caption>
<caption id="DF1AF18DFFCB3C7E96D2FC5D3671FBF6" ID-DOI="http://doi.org/10.5281/zenodo.8205510" ID-Zenodo-Dep="8205510" httpUri="https://zenodo.org/record/8205510/files/figure.png" pageId="9" pageNumber="74" startId="9.[189,232,970,996]" targetBox="[189,1397,293,928]" targetPageId="9">
<paragraph id="8BDAA105FFCB3C7E96D2FC5D3671FBF6" blockId="9.[189,1399,970,1068]" pageId="9" pageNumber="74">
<emphasis id="B9117D17FFCB3C7E96D2FC5D3415FC38" bold="true" box="[189,269,970,997]" pageId="9" pageNumber="74">Fig. 5.</emphasis>
<taxonomicName id="4C65DA86FFCB3C7E9777FC5C37F9FC38" authority="Hoffman, 2009" authorityName="Hoffman" authorityYear="2009" box="[280,737,971,997]" class="Diplopoda" family="Chelodesmidae" genus="Dibolostethus" kingdom="Animalia" order="Polydesmida" pageId="9" pageNumber="74" phylum="Arthropoda" rank="species" species="sicarius">
<emphasis id="B9117D17FFCB3C7E9777FC5C3739FC38" box="[280,545,971,997]" italics="true" pageId="9" pageNumber="74">Dibolostethus sicarius</emphasis>
<bibRefCitation id="EFF4DCF4FFCB3C7E9446FC5C37F9FC38" author="Hoffman" box="[553,737,971,997]" firstAuthor="Hoffman" pageId="9" pageNumber="74" pagination="149 - 154" refId="ref9561" refString="Hoffman R. L. 2009. Chelodesmid studies. XXVI. A new genus and tribe of chelodesmid millipeds from Ecuador (Polydesmida: Chelodesmidae). International Journal of Myriapodology 2: 149 - 154. https: // doi. org / 10.1163 / 187525409 X 12577705044584" type="journal article" year="2009">Hoffman, 2009</bibRefCitation>
</taxonomicName>
, holotype, ♂ (VMNH110810), left gonopod. 
<emphasis id="B9117D17FFCB3C7E936FFC5D300CFC39" bold="true" box="[1280,1300,970,996]" pageId="9" pageNumber="74">A</emphasis>
. Lateral view. 
<emphasis id="B9117D17FFCB3C7E9767FC793404FBD5" bold="true" box="[264,284,1006,1032]" pageId="9" pageNumber="74">B</emphasis>
. Anterior view. 
<emphasis id="B9117D17FFCB3C7E9785FC7934E7FBD5" bold="true" box="[490,511,1006,1032]" pageId="9" pageNumber="74">C</emphasis>
. Mesal view. Red circles = cingulum. Dashed lines = prostatic groove. Abbreviations: PfP = prefemoral process; S = solenomere.
</paragraph>
</caption>
<paragraph id="8BDAA105FFCB3C7E9685FBE1366AFAC0" blockId="9.[189,1399,1142,1309]" pageId="9" pageNumber="74">
<emphasis id="B9117D17FFCB3C7E9685FBE1345DFB4D" bold="true" box="[234,325,1142,1168]" pageId="9" pageNumber="74">Female</emphasis>
(
<typeStatus id="54DE1FA7FFCB3C7E973AFBE134A7FB4D" box="[341,447,1142,1168]" pageId="9" pageNumber="74" type="paratype">paratype</typeStatus>
, VMNH110811) Coloration (preserved in 70% isopropanol) as in male. Body larger and more robust than male. Similar to male in general body outline and somatic characters, except sternite of 4 
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body ring without pair of acute projections, 4 
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, 5 
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, and 6 
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podosterna elevated to a lesser degree, 4 
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and 5 
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pair of legs not modified, and tibia lacking acute ventro-apical projections.
</paragraph>
<paragraph id="8BDAA105FFCB3C7E96D2FAD231E1F9D2" blockId="9.[188,1401,1347,1551]" pageId="9" pageNumber="74">
<smallCapsWord id="8D3C37D9FFCB3C7E96D2FAD235E6FA80" baselines="1368,1368" box="[189,254,1349,1375]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Body" pageId="9" pageNumber="74">BODY</smallCapsWord>
<smallCapsWord id="8D3C37D9FFCB3C7E9768FADD34A8FA80" baselines="1368" box="[263,432,1354,1373]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="measurements" pageId="9" pageNumber="74">MEASUREMENTS</smallCapsWord>
(mm). Total length 30.95, width of 10 
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body segment 2.6. Antennomere lengths (1–7): 0.38, 0.9, 0.93, 0.88, 0.85, 0.83, 0.23. Collum: length 1.0, width 1.7. Podomere lengths of 10 
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body ring (1–7): 0.3, 0.53, 0.9, 0.38, 0.45, 0.98, 0.09. Vulvar aperture width 0.56. Posterior margin of vulvar rim: well sclerotized, U-shaped; width 0.13. Vulvae: small, unmodified, setose (
<figureCitation id="135EBD80FFCB3C7E92A3FA39300BFA15" box="[1228,1299,1454,1480]" captionStart="Fig" captionStartId="13.[189,232,1928,1954]" captionTargetBox="[190,1397,1511,1885]" captionTargetId="figure-343@13.[189,1398,1505,1885]" captionTargetPageId="13" captionText="Fig. 8. Dibolostethus sicarius Hoffman, 2009, paratype, ♀ (VMNH110811), vulvae. A. Left vulva, lateroventral view, with operculum at base. B. Vulvae in situ, held appressed to 2nd leg pair coxae with valve openings facing each other. C. Left vulva, lateral view." figureDoi="http://doi.org/10.5281/zenodo.8205516" httpUri="https://zenodo.org/record/8205516/files/figure.png" pageId="9" pageNumber="74">Fig. 8</figureCitation>
); length 0.45, width 0.29. Valves symmetrical. External valve: lateral length 0.5, lateral height 0.44; dorsal length 0.39, dorsal width at midpoint 0.14. Operculum: length 0.06, width 0.21. Telson length 1.15.
</paragraph>
</subSubSection>
<subSubSection id="C37FF28EFFCB3C7E96D2F9A23699F9A3" box="[189,897,1589,1663]" pageId="9" pageNumber="74" type="distribution">
<paragraph id="8BDAA105FFCB3C7E96D2F9A2344EF992" blockId="9.[189,342,1589,1615]" box="[189,342,1589,1615]" pageId="9" pageNumber="74">
<heading id="D0921669FFCB3C7E96D2F9A2344EF992" bold="true" box="[189,342,1589,1615]" fontSize="11" level="3" pageId="9" pageNumber="74" reason="3">
<emphasis id="B9117D17FFCB3C7E96D2F9A2344EF992" bold="true" box="[189,342,1589,1615]" pageId="9" pageNumber="74">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BDAA105FFCB3C7E96D2F9F33699F9A3" blockId="9.[189,897,1636,1663]" box="[189,897,1636,1663]" pageId="9" pageNumber="74">
Herein restricted to the 
<collectingRegion id="49A16FE7FFCB3C7E97BDF9F237B5F9A3" box="[466,685,1636,1663]" country="Ecuador" name="Los Rios" pageId="9" pageNumber="74">Los Rios Province</collectingRegion>
, 
<collectingCountry id="F372E195FFCB3C7E94D6F9F33603F9A3" box="[697,795,1636,1662]" name="Ecuador" pageId="9" pageNumber="74">Ecuador</collectingCountry>
(
<figureCitation id="135EBD80FFCB3C7E9544F9F3366BF9A3" box="[811,883,1636,1662]" captionStart="Fig" captionStartId="14.[188,231,1892,1918]" captionTargetBox="[243,1357,497,1838]" captionTargetId="figure-72@14.[212,1380,484,1853]" captionTargetPageId="14" captionText="Fig. 9. Distribution of Chelodesmidae spp. in the Tropical Andes Biodiversity Hotspot, by country. Note: the type locality of Dibolostethus sicarius Hoffman, 2009 is included for completeness although it falls just outside of the boundaries of the Tropical Andes Biodiversity Hotspot. For information on these taxa see Supp. file 1." figureDoi="http://doi.org/10.5281/zenodo.8205518" httpUri="https://zenodo.org/record/8205518/files/figure.png" pageId="9" pageNumber="74">Fig. 9</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>