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<document id="47D2F6F7C980E628ECDEDB2F6FC65225" ID-DOI="10.5281/zenodo.274579" ID-GBIF-Dataset="eed612c2-ba22-4d53-ae53-9eaff3fc8706" ID-ISSN="1175-5326" ID-Zenodo-Dep="274579" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460000228395" checkinUser="plazi" docAuthor="E, José Mondaca" docDate="2008" docId="03C787A4FF9B322512ECF9ADFBEEF94C" docLanguage="en" docName="zt01922p032.pdf" docOrigin="Zootaxa 1922" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Apterodorcus Arrow" docType="treatment" docVersion="8" lastPageNumber="24" masterDocId="FFFEFFDCFF993226127BFFF8FF83FFBD" masterDocTitle="Revision of the genus Apterodorcus Arrow (Coleoptera: Lucanidae: Lucaninae) of southern South America" masterLastPageNumber="32" masterPageNumber="21" pageNumber="23" updateTime="1698233134854" updateUser="plazi">
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<mods:title id="9C52489B2F39B282131BC2C7428C105F">Revision of the genus Apterodorcus Arrow (Coleoptera: Lucanidae: Lucaninae) of southern South America</mods:title>
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<mods:namePart id="1B26C3EDA6B1FD38F149DF910918CA4D">E, José Mondaca</mods:namePart>
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<mods:date id="EF5C51BB6A233EC9B0AC2F2AB30933FA">2008</mods:date>
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<treatment id="03C787A4FF9B322512ECF9ADFBEEF94C" ID-DOI="http://doi.org/10.5281/zenodo.6227637" ID-GBIF-Taxon="119380866" ID-Zenodo-Dep="6227637" LSID="urn:lsid:plazi:treatment:03C787A4FF9B322512ECF9ADFBEEF94C" httpUri="http://treatment.plazi.org/id/03C787A4FF9B322512ECF9ADFBEEF94C" lastPageId="3" lastPageNumber="24" pageId="2" pageNumber="23">
<subSubSection id="C3746539FF9B322412ECF9ADFE68F9D2" box="[151,491,1621,1647]" pageId="2" pageNumber="23" type="nomenclature">
<paragraph id="8BD136B2FF9B322412ECF9ADFE68F9D2" blockId="2.[151,491,1621,1647]" box="[151,491,1621,1647]" pageId="2" pageNumber="23">
<heading id="D09981DEFF9B322412ECF9ADFE68F9D2" bold="true" box="[151,491,1621,1647]" fontSize="11" level="1" pageId="2" pageNumber="23" reason="1">
<emphasis id="B91AEAA0FF9B322412ECF9ADFE68F9D2" bold="true" box="[151,491,1621,1647]" pageId="2" pageNumber="23">
Genus
<taxonomicName id="4C6E4D31FF9B32241295F9ADFE68F9D2" ID-CoL="ZCW" authority="Arrow" authorityName="Arrow" box="[238,491,1621,1647]" class="Insecta" family="Lucanidae" genus="Apterodorcus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="B91AEAA0FF9B32241295F9ADFE12F9D2" bold="true" box="[238,401,1621,1647]" italics="true" pageId="2" pageNumber="23">Apterodorcus</emphasis>
Arrow
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3746539FF9B322412ECF958FB75F90A" box="[151,1270,1696,1719]" pageId="2" pageNumber="23" type="reference_group">
<paragraph id="8BD136B2FF9B322412ECF958FB75F90A" blockId="2.[151,1270,1696,1719]" box="[151,1270,1696,1719]" pageId="2" pageNumber="23">
<treatmentCitationGroup id="AB7E119CFF9B322412ECF958FB75F90A" box="[151,1270,1696,1719]" pageId="2" pageNumber="23">
<taxonomicName id="4C6E4D31FF9B322412ECF958FE64F90A" ID-CoL="ZCW" authority="Arrow 1943: 139" authorityName="Arrow" authorityPageNumber="139" authorityYear="1943" box="[151,487,1696,1719]" class="Insecta" family="Lucanidae" genus="Apterodorcus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="B91AEAA0FF9B322412ECF958FEA5F90A" box="[151,294,1696,1719]" italics="true" pageId="2" pageNumber="23">Apterodorcus</emphasis>
<treatmentCitation id="0ACF10A3FF9B32241357F958FE64F90A" author="Arrow" box="[300,487,1696,1719]" page="139" pageId="2" pageNumber="23" year="1943">
<bibRefCitation id="EFFF4B43FF9B32241357F958FE31F90A" author="Arrow" box="[300,434,1696,1719]" pageId="2" pageNumber="32" refString="Arrow, G. J. (1943) On the genera and nomenclature of the lucanid Coleoptera, and descriptions of a few new species. Proceedings of the Royal Entomological Society of London (B), 12, 133 - 143." type="journal article" year="1943">Arrow 1943</bibRefCitation>
: 139
</treatmentCitation>
</taxonomicName>
.
<typeStatus id="54D58810FF9B3224138AF958FDA5F90A" box="[497,550,1696,1719]" pageId="2" pageNumber="23">Type</typeStatus>
species:
<taxonomicName id="4C6E4D31FF9B322410F3F958FC02F90A" ID-CoL="6QP45" authority="Hope" authorityName="Hope" box="[648,897,1696,1719]" class="Insecta" family="Lucanidae" genus="Lucanus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="23" phylum="Arthropoda" rank="species" species="bacchus">
<emphasis id="B91AEAA0FF9B322410F3F958FCC2F90A" box="[648,833,1696,1719]" italics="true" pageId="2" pageNumber="23">Lucanus bacchus</emphasis>
Hope
</taxonomicName>
in
<bibRefCitation id="EFFF4B43FF9B322411D8F958FBDBF90A" author="Westwood" box="[931,1112,1696,1719]" pageId="2" pageNumber="32" refString="Westwood, J. O. (1845) A catalogue of the Lucanoid Coleoptera in the collection of the Rev. F. W. Hope, M. A., F. R. S., &amp; c., President of the Entomological Society of London, together with descriptions of the new species therein contained. J. C. Bridgewater, London. 31 pp." type="book" year="1845">Westwood, 1845</bibRefCitation>
, by monotypy.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3746539FF9B322512ECF91FFC6FFB9F" lastPageId="3" lastPageNumber="24" pageId="2" pageNumber="23" type="description">
<paragraph id="8BD136B2FF9B322512ECF91FFC6FFB9F" blockId="2.[151,1436,1767,2034]" lastBlockId="3.[151,1437,152,1778]" lastPageId="3" lastPageNumber="24" pageId="2" pageNumber="23">
<emphasis id="B91AEAA0FF9B322412ECF91FFEABF8BC" bold="true" box="[151,296,1767,1793]" pageId="2" pageNumber="23">Description</emphasis>
. Scarabaeoidea:
<taxonomicName id="4C6E4D31FF9B32241394F910FDEEF8BF" box="[495,621,1768,1794]" class="Insecta" family="Lucanidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="23" phylum="Arthropoda" rank="family">Lucanidae</taxonomicName>
:
<taxonomicName id="4C6E4D31FF9B32241003F910FD75F8BF" box="[632,758,1768,1794]" class="Insecta" family="Lucanidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="23" phylum="Arthropoda" rank="subFamily" subFamily="Lucaninae">Lucaninae</taxonomicName>
: Sclerostomini.
<emphasis id="B91AEAA0FF9B322411C9F91FFB96F8BC" bold="true" box="[946,1045,1767,1793]" pageId="2" pageNumber="23">Length:</emphasis>
19.235.4 mm.
<emphasis id="B91AEAA0FF9B322416AEF91FFAADF8BC" bold="true" box="[1237,1326,1767,1793]" pageId="2" pageNumber="23">Width:</emphasis>
9.014.5 mm.
<emphasis id="B91AEAA0FF9B322412AEF8F7FE9FF894" bold="true" box="[213,284,1807,1833]" pageId="2" pageNumber="23">Color</emphasis>
: Black, surface alutaceous, weakly shiny or opaque, lacking iridescence.
<emphasis id="B91AEAA0FF9B322416EFF8F7FA87F894" bold="true" box="[1172,1284,1807,1833]" pageId="2" pageNumber="23">Vestiture</emphasis>
: Apparently glabrous dorsally; minute scales visibly only at
<date id="FFD01072FF9B322410B3F8C0FD79F8EF" box="[712,762,1848,1874]" pageId="2" pageNumber="23" value="1940-10">40X</date>
near elytral apex; scales arising centrally in elytral punctures, form branched; branches tapering to acute apex (
<figureCitation id="13552A37FF9B32241144F898FC14F8C7" box="[831,919,1888,1914]" captionStart="FIGURE 12" captionStartId="10.[151,255,1594,1618]" captionTargetBox="[472,1115,714,1569]" captionTargetId="figure@10.[472,1115,714,1570]" captionTargetPageId="10" captionText="FIGURE 12. Female A. bacchus feeding at sap flow on Nothofagus near Chillán, VIII Region." httpUri="https://zenodo.org/record/274584/files/figure.png" pageId="2" pageNumber="23">
Fig.
<quantity id="4C969B57FF9B32241101F898FC14F8C7" box="[890,919,1888,1914]" metricMagnitude="-1" metricUnit="m" metricValue="3.048" pageId="2" pageNumber="23" unit="in" value="12.0">12</quantity>
</figureCitation>
in
<bibRefCitation id="EFFF4B43FF9B322411BDF898FB06F8C7" author="Holloway" box="[966,1157,1888,1914]" pageId="2" pageNumber="32" refString="Holloway, B. A. (1997) Elytral surface structures as indicators of relationships in stag beetles, with special reference to the New Zealand species (Coleoptera: Lucanidae). New Zealand Journal of Zoology, 24, 47 - 64." type="journal article" year="1997">Holloway 1997</bibRefCitation>
).
<emphasis id="B91AEAA0FF9B322416E6F8A7FB69F8C4" bold="true" box="[1181,1258,1887,1913]" pageId="2" pageNumber="23">Head:</emphasis>
Eyes partially divided by ocular canthus. Frons more or less flat, weakly excavated anteromedially, excavation parabolic; lacking anteromedial tubercle, sparsely to moderately punctate; punctures fine to moderate, becoming larger and more dense anteriorly and laterally, punctation markedly stronger and denser in female. Suprantennal gibbosity weakly developed. Both sexes with strongly developed, lobelike temporal process. Labrum setose, projecting anteriorly, appearing trilobed or rounded; median lobe subtuberculate in males. Male majors with mandibles usually as long as head, curved abruptly before apex, apex bifurcate with teeth acute; internally near middle with 23 obtuse teeth; dorsally with tooth near base. Male minors with mandibles similar to male major in dentition but everywhere smaller. Male laciniae lacking hook. Female mandibles complex, left mandible with 4 teeth (dorsal, apical, ventral, internal), right mandible with 3 teeth (dorsal, apical, ventral); dorsal surface flat or with external dorsal carina more or less produced. Female laciniae with sclerotized hook. Antennae with 10 antennomeres, geniculate, with asymmetrical, immobile club of 3 antennomeres; club mostly tomentose distally, glabrous on stem.
<emphasis id="B91AEAA0FF9A322510BBFE2FFCCAFE4C" bold="true" box="[704,841,471,497]" pageId="3" pageNumber="24">Pronotum:</emphasis>
Form broadly rounded externally or subparallel, weakly angulate in basal third, width at base narrower than elytra, broader than elytra anteriorly. Disc convex, not flattened medially. Anterior angle obtuse, sides; margin straight from angulation to posterior angle. Margins beaded (bead obsolete anteromedially), shining, not reflexed. Surface punctate; punctures fine to moderate on disc, becoming large and coalescing near margins.
<emphasis id="B91AEAA0FF9A32251113FD8FFC6CFD2C" bold="true" box="[872,1007,631,657]" pageId="3" pageNumber="24">Scutellum:</emphasis>
Form parabolic or triangular. Size reduced or not, occasionally weakly depressed below level of elytra.
<emphasis id="B91AEAA0FF9A322511BAFD67FB92FD04" bold="true" box="[961,1041,671,697]" pageId="3" pageNumber="24">Elytra</emphasis>
: Form convex, somewhat shorter in males (as long as head and pronotum combined), somewhat fused (but separable) at suture. Surface lacking obvious scales, weakly striate, punctate; punctures fine on disc, becoming larger and coalescing laterally and basally.
<emphasis id="B91AEAA0FF9A3225128CFCEFFEC5FC8C" bold="true" box="[247,326,791,817]" pageId="3" pageNumber="24">Wings</emphasis>
: Brachypterous in both sexes; wing form narrow, elongate, reaching approximately to middle of elytron.
<emphasis id="B91AEAA0FF9A32251362FCC7FEDDFCE4" bold="true" box="[281,350,831,857]" pageId="3" pageNumber="24">Legs:</emphasis>
Protibiae with 46 large external teeth. Mesotibiae and metatibiae with large external tooth, 02 smaller teeth proximally. Onychium (= empodium) with brush-like, multiple setae.
<emphasis id="B91AEAA0FF9A322516A8FC9FFA1FFC3C" bold="true" box="[1235,1436,871,897]" pageId="3" pageNumber="24">Male Genitalia:</emphasis>
Form symmetrical. Male internal sac permanently everted, sclerotized, strap-like, parallel-sided to subapex; apex triangular, not expanded distally. Parameres simple, curved. Basal piece membranous posteroventrally, dorsally. Genital capsule simple (lacking additional lobes present in some sclerostomines).
<emphasis id="B91AEAA0FF9A322516C5FC27FA17FC44" bold="true" box="[1214,1428,991,1017]" pageId="3" pageNumber="24">Female Genitalia</emphasis>
: Similar to other sclerostomines. Illustrated by
<bibRefCitation id="EFFF4B43FF9A322510CEFBF0FC6BFB9F" author="Weinreich" box="[693,1000,1032,1058]" pageId="3" pageNumber="32" refString="Weinreich, E. (1960) Revision sudamerikanischer Lucanidae (Ins. Col.), II. Die Gattungen Charagmophorus, Metadorcus, Scortizus, Apterodorcus, Beneshius, Sclerostomus und Pycnosiphorus. Senckenbergiana Biologica, 41, 41 - 95." type="journal article" year="1960">Weinreich (1960: Fig. 68)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3746539FF9A322512BDFBD7FA16FA87" pageId="3" pageNumber="24" type="diagnosis">
<paragraph id="8BD136B2FF9A322512BDFBD7FA16FA87" blockId="3.[151,1437,152,1778]" pageId="3" pageNumber="24">
<emphasis id="B91AEAA0FF9A322512BDFBD7FEC5FBF4" bold="true" box="[198,326,1071,1097]" pageId="3" pageNumber="24">Diagnosis.</emphasis>
This genus is distinguished from other New World lucanine genera by the following combination of characters: antennae with club of 3 antennomeres, with club mostly tomentose; onychium of last tarsal segment brush-like with more than 10 setae (not 24 setae as in all other New World sclerostomines); head with strong, lobe-like temporal processes in both sexes; pronotum and elytra lacking obvious vestiture.
<taxonomicName id="4C6E4D31FF9A322512ECFB28FEB7FB54" box="[151,308,1232,1257]" class="Insecta" family="Lucanidae" genus="Apterodorcus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="24" phylum="Arthropoda" rank="genus">
<emphasis id="B91AEAA0FF9A322512ECFB28FEB7FB54" box="[151,308,1232,1257]" italics="true" pageId="3" pageNumber="24">Apterodorcus</emphasis>
</taxonomicName>
species can be immediately distinguished from other southern South American sclerostomines by their larger size, and black coloration without visible scales (
<taxonomicName id="4C6E4D31FF9A322511F7FB00FBAAFAAC" box="[908,1065,1272,1297]" class="Insecta" family="Lucanidae" genus="Apterodorcus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="24" phylum="Arthropoda" rank="genus">
<emphasis id="B91AEAA0FF9A322511F7FB00FBAAFAAC" box="[908,1065,1272,1297]" italics="true" pageId="3" pageNumber="24">Apterodorcus</emphasis>
</taxonomicName>
species lack a pattern of scales on the pronotum and/or elytra commonly found in other genera of southern South American sclerostomines).
</paragraph>
</subSubSection>
<subSubSection id="C3746539FF9A322512BDFABFFB82FADF" box="[198,1025,1351,1378]" pageId="3" pageNumber="24" type="distribution">
<paragraph id="8BD136B2FF9A322512BDFABFFB82FADF" blockId="3.[151,1437,152,1778]" box="[198,1025,1351,1378]" pageId="3" pageNumber="24">
<emphasis id="B91AEAA0FF9A322512BDFABFFEDCFADC" bold="true" box="[198,351,1351,1377]" pageId="3" pageNumber="24">Distribution</emphasis>
(
<figureCitation id="13552A37FF9A32251314FAB0FE37FADF" box="[367,436,1352,1378]" captionStart="FIGURE 1" captionStartId="5.[151,255,1831,1855]" captionTargetBox="[415,1171,197,1807]" captionTargetId="figure@5.[415,1171,194,1807]" captionTargetPageId="5" captionText="FIGURE 1. Distribution of Apterodorcus species: A. bacchus (circles), and A. tristis (triangles). Regions of Chile indicated by Roman numeral, provinces of Argentina by postal code (Q = Neuquén, U = Chubut, R = Río Negro)." httpUri="https://zenodo.org/record/274580/files/figure.png" pageId="3" pageNumber="24">Fig. 1</figureCitation>
).
<collectingCountry id="F3797622FF9A322513B6FAB0FDC2FADF" box="[461,577,1352,1378]" name="Argentina" pageId="3" pageNumber="24">Argentina</collectingCountry>
and
<collectingCountry id="F3797622FF9A32251001FAB0FD3EFADF" box="[634,701,1352,1378]" name="Chile" pageId="3" pageNumber="24">Chile</collectingCountry>
, see species account below.
</paragraph>
</subSubSection>
<subSubSection id="C3746539FF9A322512BDFA97FBEEF94C" pageId="3" pageNumber="24" type="discussion">
<paragraph id="8BD136B2FF9A322512BDFA97FB59F9BF" blockId="3.[151,1437,152,1778]" pageId="3" pageNumber="24">
<emphasis id="B91AEAA0FF9A322512BDFA97FEBCFA34" bold="true" box="[198,319,1391,1417]" pageId="3" pageNumber="24">Remarks.</emphasis>
This genus is more similar to species of the
<collectingCountry id="F3797622FF9A32251129FA88FC73FA37" box="[850,1008,1392,1418]" name="New Zealand" pageId="3" pageNumber="24">New Zealand</collectingCountry>
genus
<taxonomicName id="4C6E4D31FF9A3225163EFA88FB47FA34" box="[1093,1220,1392,1417]" class="Insecta" family="Lucanidae" genus="Geodorcus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="24" phylum="Arthropoda" rank="genus">
<emphasis id="B91AEAA0FF9A3225163EFA88FB47FA34" box="[1093,1220,1392,1417]" italics="true" pageId="3" pageNumber="24">Geodorcus</emphasis>
</taxonomicName>
than to species of other New World genera. Microscopically, it shares with
<taxonomicName id="4C6E4D31FF9A3225112FFA60FC50FA0C" box="[852,979,1432,1457]" class="Insecta" family="Lucanidae" genus="Geodorcus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="24" phylum="Arthropoda" rank="genus">
<emphasis id="B91AEAA0FF9A3225112FFA60FC50FA0C" box="[852,979,1432,1457]" italics="true" pageId="3" pageNumber="24">Geodorcus</emphasis>
</taxonomicName>
similar elytral vestiture consisting of branched scales (
<bibRefCitation id="EFFF4B43FF9A32251311FA38FDA8FA67" author="Holloway" box="[362,555,1472,1498]" pageId="3" pageNumber="32" refString="Holloway, B. A. (1997) Elytral surface structures as indicators of relationships in stag beetles, with special reference to the New Zealand species (Coleoptera: Lucanidae). New Zealand Journal of Zoology, 24, 47 - 64." type="journal article" year="1997">Holloway 1997</bibRefCitation>
). The onychium between the claws is multisetose in
<taxonomicName id="4C6E4D31FF9A322516BCFA38FAE7FA64" box="[1223,1380,1472,1497]" class="Insecta" family="Lucanidae" genus="Apterodorcus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="24" phylum="Arthropoda" rank="genus">
<emphasis id="B91AEAA0FF9A322516BCFA38FAE7FA64" box="[1223,1380,1472,1497]" italics="true" pageId="3" pageNumber="24">Apterodorcus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6E4D31FF9A322512ECFA10FE94F9BC" box="[151,279,1512,1537]" class="Insecta" family="Lucanidae" genus="Geodorcus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="24" phylum="Arthropoda" rank="genus">
<emphasis id="B91AEAA0FF9A322512ECFA10FE94F9BC" box="[151,279,1512,1537]" italics="true" pageId="3" pageNumber="24">Geodorcus</emphasis>
</taxonomicName>
species, while in other New World sclerostomines it generally has only 24 setae.
</paragraph>
<paragraph id="8BD136B2FF9A322512BDF9E8FBEEF94C" blockId="3.[151,1437,152,1778]" pageId="3" pageNumber="24">
<taxonomicName id="4C6E4D31FF9A322512BDF9E8FEE0F994" box="[198,355,1552,1577]" class="Insecta" family="Lucanidae" genus="Apterodorcus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="24" phylum="Arthropoda" rank="genus">
<emphasis id="B91AEAA0FF9A322512BDF9E8FEE0F994" box="[198,355,1552,1577]" italics="true" pageId="3" pageNumber="24">Apterodorcus</emphasis>
</taxonomicName>
has been treated as monotypic with the single species
<taxonomicName id="4C6E4D31FF9A3225118FF9E8FBF4F994" box="[1012,1143,1552,1577]" class="Insecta" family="Lucanidae" genus="Lucanus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="24" phylum="Arthropoda" rank="species" species="bacchus">
<emphasis id="B91AEAA0FF9A3225118FF9E8FBF4F994" box="[1012,1143,1552,1577]" italics="true" pageId="3" pageNumber="24">A. bacchus</emphasis>
</taxonomicName>
, however the specimens from the coastal mountains south of Santiago exhibit several important differences and are not conspecific with
<taxonomicName id="4C6E4D31FF9A322512A9F998FED1F9C4" box="[210,338,1632,1657]" class="Insecta" family="Lucanidae" genus="Lucanus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="24" phylum="Arthropoda" rank="species" species="bacchus">
<emphasis id="B91AEAA0FF9A322512A9F998FED1F9C4" box="[210,338,1632,1657]" italics="true" pageId="3" pageNumber="24">A. bacchus</emphasis>
</taxonomicName>
. The genitalia of the two taxa do not appear to differ significantly, but they are of the simplest form found in Sclerostomini, so this may not be surprising. The second species was described by Deyrolle in
<bibRefCitation id="EFFF4B43FF9A322512ECF948FEA8F977" author="Parry" box="[151,299,1712,1738]" pageId="3" pageNumber="32" refString="Parry, F. J. S. (1870) A revised catalogue of the lucanoid Coleoptera; with remarks on the nomenclature, and descriptions of new species. Transactions of the Entomological Society of London, 1870, 53 - 118." type="journal article" year="1870">Parry (1870)</bibRefCitation>
as
<taxonomicName id="4C6E4D31FF9A32251328F948FDBAF974" box="[339,569,1712,1737]" class="Insecta" family="Lucanidae" genus="Sclerostomus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="24" phylum="Arthropoda" rank="species" species="tristis">
<emphasis id="B91AEAA0FF9A32251328F948FDBAF974" box="[339,569,1712,1737]" italics="true" pageId="3" pageNumber="24">Sclerostomus tristis</emphasis>
</taxonomicName>
, but probably due to its restricted range the species remained unknown to subsequent authors and it was eventually mistakenly synonymized with
<emphasis id="B91AEAA0FF9A3225119FF920FBEEF94C" box="[996,1133,1752,1777]" italics="true" pageId="3" pageNumber="24">
<taxonomicName id="4C6E4D31FF9A3225119FF920FBEAF94C" box="[996,1129,1752,1777]" class="Insecta" family="Lucanidae" genus="Lucanus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="24" phylum="Arthropoda" rank="species" species="bacchus">A. bacchus</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>