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<document id="C60BAEED9415D84389CAF7C9BA5A06D2" ID-CLB-Dataset="22877" ID-DOI="10.5852/ejt.2020.629" ID-GBIF-Dataset="3722aa65-b935-4294-ac20-0d1179da9015" ID-Zenodo-Dep="3762280" ID-ZooBank="ECEBD539-6E3E-45BE-A0CB-264DF3270CC0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1587578640316" checkinUser="valdenar" docAuthor="Herbert, David G." docDate="2020" docId="03C78785462EFFE6423C46C7344FFE39" docLanguage="en" docName="ejt-629_herbert.pdf.imf" docOrigin="European Journal of Taxonomy 629" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Iterodonta ammonita Herbert 2020, gen. et sp. nov." docType="treatment" docUuid="A33983DE-4995-4FB7-A54D-83F7E4F897D8" docUuidSource="ZooBank" docVersion="7" lastPageNumber="37" masterDocId="FFFEFFFD460CFFC2400243213138FFD9" masterDocTitle="Revision of the aperturally dentate Charopidae (Gastropoda: Stylommatophora) of southern Africa - genus Afrodonta s. lat., with description of five new genera, twelve new species and one new subspecies" masterLastPageNumber="55" masterPageNumber="1" pageNumber="35" updateTime="1698835565728" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="0137B237B124A7DF2F4FB4B7405D0368">Revision of the aperturally dentate Charopidae (Gastropoda: Stylommatophora) of southern Africa - genus Afrodonta s. lat., with description of five new genera, twelve new species and one new subspecies</mods:title>
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<mods:name id="DF6A988E8BF6C69BA0755BF7157760C7" type="personal">
<mods:role id="0B33BE2F0572109B8BAB5C8F64EB58A1">
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<mods:namePart id="C96C16AC724407B3320D1E7EE0A96936">Herbert, David G.</mods:namePart>
<mods:nameIdentifier id="CF9E8065EB984D7687FF52357BF5E975" type="LSID">urn:lsid:zoobank.org:author:0C09EE45-6198-482E-857A-EF690C2A016F</mods:nameIdentifier>
<mods:affiliation id="94961B59EF7EB4A38F2227ABA0948398">Department of Natural Sciences, National Museum Wales, Cathays Park, Cardiff, CF 10 3 NP, UK. Formerly at KwaZulu-Natal Museum, P. Bag 9070, Pietermaritzburg 3201, South Africa</mods:affiliation>
<mods:nameIdentifier id="83EECC42C07051670927CBAE03DC3700" type="email">phasianella@gmail.com</mods:nameIdentifier>
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<mods:title id="129A9D8806F04DE25BEAEC9FB7BE71F3">European Journal of Taxonomy</mods:title>
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<mods:part id="3F0737E6B07583B21A726D01FD00AD81">
<mods:date id="DCE7AB4392A7F6B9E4AFD84D239DFFED">2020</mods:date>
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<mods:number id="4154020CE9EB6035F5BAE55E0A6861D6">2020-04-17</mods:number>
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<treatment id="03C78785462EFFE6423C46C7344FFE39" ID-DOI="http://doi.org/10.5281/zenodo.3804766" ID-GBIF-Taxon="163732356" ID-Zenodo-Dep="3804766" LSID="urn:lsid:zoobank.org:act:A33983DE-4995-4FB7-A54D-83F7E4F897D8" httpUri="http://treatment.plazi.org/id/03C78785462EFFE6423C46C7344FFE39" lastPageId="36" lastPageNumber="37" pageId="34" pageNumber="35">
<subSubSection id="C3746518462EFFE0423C46C735FBF9FF" pageId="34" pageNumber="35" type="nomenclature">
<paragraph id="8BD13693462EFFE0423C46C732CEF9D9" blockId="34.[368,1219,1510,1610]" box="[574,1014,1510,1537]" pageId="34" pageNumber="35">
<heading id="D09981FF462EFFE0423C46C732CEF9D9" box="[574,1014,1510,1537]" centered="true" fontSize="11" level="2" pageId="34" pageNumber="35" reason="2">
<taxonomicName id="4C6E4D10462EFFE0423C46C73206F9D9" ID-CoL="7RRJ3" authority="Herbert, 2020" authorityName="Herbert" authorityYear="2020" box="[574,830,1510,1537]" class="Gastropoda" family="Charopidae" genus="Iterodonta" kingdom="Animalia" order="Stylommatophora" pageId="34" pageNumber="35" phylum="Mollusca" rank="species" species="ammonita" status="gen. et sp. nov.">
<emphasis id="B91AEA81462EFFE0423C46C73206F9D9" bold="true" box="[574,830,1510,1537]" italics="true" pageId="34" pageNumber="35">Iterodonta ammonita</emphasis>
</taxonomicName>
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</heading>
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urn:lsid:zoobank.org:act:
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</uri>
</paragraph>
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<subSubSection id="C3746518462EFFE042A4451132B5F993" box="[678,909,1584,1610]" pageId="34" pageNumber="35" type="description">
<paragraph id="8BD13693462EFFE042A4451132B5F993" blockId="34.[368,1219,1510,1610]" box="[678,909,1584,1610]" pageId="34" pageNumber="35">
<figureCitation id="13552A16462EFFE042A4451133C7F993" box="[678,767,1584,1610]" captionStart="Fig" captionStartId="35.[189,232,2001,2027]" captionTargetBox="[195,1390,1564,1953]" captionTargetId="figure@35.[189,1398,1559,1960]" captionTargetPageId="35" captionText="Fig. 13. Iterodonta ammonita gen. et sp. nov., holotype, diameter 1.45 mm (NMSA W5826/T4258)." figureDoi="http://doi.org/10.5281/zenodo.3762306" httpUri="https://zenodo.org/record/3762306/files/figure.png" pageId="34" pageNumber="35">Figs 13</figureCitation>
,
<figureCitation id="13552A16462EFFE0430E45113214F993" box="[780,812,1584,1610]" captionStart="Fig" captionStartId="36.[189,232,1860,1886]" captionTargetBox="[302,1285,1199,1820]" captionTargetId="figure@36.[302,1285,1199,1820]" captionTargetPageId="36" captionText="Fig. 14. Distribution of Iterodonta gen. nov. and Phialodonta gen. nov. species. Iterodonta ammonita gen. et sp. nov. (brown triangles), Phialodonta agulhasae gen. et sp. nov. (orange square), P. atromontana gen. et sp. nov. (turquoise square), P. aviana gen. et sp. nov. (royal blue squares), P. introtuberculata (Connolly, 1933) gen. et comb. nov. (red squares), P. perfida (Burnup, 1912) gen. et comb. nov. (green square), P. rivalalea gen. et sp. nov. (pink squares). Contour at 1000 m." figureDoi="http://doi.org/10.5281/zenodo.3762308" httpUri="https://zenodo.org/record/3762308/files/figure.png" pageId="34" pageNumber="35">14</figureCitation>
,
<figureCitation id="13552A16462EFFE0433A4511326BF993" box="[824,851,1584,1610]" captionStart="Fig" captionStartId="47.[189,232,1792,1818]" captionTargetBox="[193,1394,556,1748]" captionTargetId="figure@47.[189,1398,554,1751]" captionTargetPageId="47" captionText="Fig. 19. Iterodonta gen. nov. and Phialodonta gen. nov. species, detail of protoconch morphology and teleoconch microsculpture. AB. Iterodonta ammonita gen. et sp. nov., paratype (NMSA P1017/T4259). CD. Phialodonta atromontana gen. et sp. nov., paratype (NMSA P1016/T4255). EF. P. aviana gen. et sp. nov., paratype (NMSA W3524/T4247). GH. P. introtuberculata gen. et comb. nov. (NMSA W216). IJ. P. perfida gen. et comb. nov., paratype (NMSA 2602/T617). KL. P. rivalalea gen. et sp. nov., paratype (NMSA V4734/T4250). Scale bars for protoconchs = 100 μm; scale bars for microsculpture = 50 μm." figureDoi="http://doi.org/10.5281/zenodo.3762314" httpUri="https://zenodo.org/record/3762314/files/figure.png" pageId="34" pageNumber="35">19</figureCitation>
AB
</paragraph>
</subSubSection>
<subSubSection id="C3746518462EFFE040BF45523533F8A1" pageId="34" pageNumber="35" type="diagnosis">
<paragraph id="8BD13693462EFFE040BF4552300EF954" blockId="34.[189,310,1651,1677]" box="[189,310,1651,1677]" pageId="34" pageNumber="35">
<heading id="D09981FF462EFFE040BF4552300EF954" bold="true" box="[189,310,1651,1677]" fontSize="11" level="3" pageId="34" pageNumber="35" reason="3">
<emphasis id="B91AEA81462EFFE040BF4552300EF954" bold="true" box="[189,310,1651,1677]" pageId="34" pageNumber="35">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BD13693462EFFE040BF45853533F8A1" blockId="34.[189,1398,1700,1912]" pageId="34" pageNumber="35">
Shell small, symmetrically biconcave; protoconch initially with traces of close-set spiral threads, later portion with spiral threads and close-set axial riblets producing a fine, reticulate sculpture; teleoconch sculpture of close-set, compound axial riblets with finer, uneven intermediary axial threads; aperture lacking parietal and columellar dentition; palatal region with 13 broad, prosocline, axial ridge-like calluses set back from lip, visible through translucent shell; umbilicus very wide and shallow. Shell translucent, straw-brown to pale honey-brown; diameter up to
<quantity id="4C969B76462EFFE04399447C353FF8A1" box="[923,1031,1885,1912]" metricMagnitude="-3" metricUnit="m" metricValue="1.68" pageId="34" pageNumber="35" unit="mm" value="1.68">1.68 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C3746518462EFFE040BF44803591F832" box="[189,1193,1953,2028]" pageId="34" pageNumber="35" type="etymology">
<paragraph id="8BD13693462EFFE040BF4480307CF862" blockId="34.[189,324,1953,1979]" box="[189,324,1953,1979]" pageId="34" pageNumber="35">
<heading id="D09981FF462EFFE040BF4480307CF862" bold="true" box="[189,324,1953,1979]" fontSize="11" level="3" pageId="34" pageNumber="35" reason="3">
<emphasis id="B91AEA81462EFFE040BF4480307CF862" bold="true" box="[189,324,1953,1979]" pageId="34" pageNumber="35">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BD13693462EFFE040BF44F03591F832" blockId="34.[189,1193,2001,2028]" box="[189,1193,2001,2028]" pageId="34" pageNumber="35">From ammonite (Ammonoidea); with reference to the symmetrically biconcave shell.</paragraph>
</subSubSection>
<subSubSection id="C3746518462FFFE140BF422A3029FD55" pageId="35" pageNumber="36" type="materials_examined">
<paragraph id="8BD13693462FFFE140BF422A3092FEFC" blockId="35.[189,426,267,293]" box="[189,426,267,293]" pageId="35" pageNumber="36">
<heading id="D09981FF462FFFE140BF422A3092FEFC" bold="true" box="[189,426,267,293]" fontSize="11" level="3" pageId="35" pageNumber="36" reason="3">
<emphasis id="B91AEA81462FFFE140BF422A3092FEFC" bold="true" box="[189,426,267,293]" pageId="35" pageNumber="36">Material examined</emphasis>
</heading>
</paragraph>
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<emphasis id="B91AEA81462FFFE140E8421B3064FE8D" bold="true" box="[234,348,314,340]" pageId="35" pageNumber="36">
<typeStatus id="54D58831462FFFE140E8421B3064FE8D" box="[234,348,314,340]" pageId="35" pageNumber="36" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BD13693462FFFE140BF427C353AFE67" blockId="35.[189,1398,314,446]" pageId="35" pageNumber="36">
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,
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,
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(Oudebosch);
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,
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;
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a.s.l.
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;
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<collectingDate id="EF94E9BB462FFFE1415C42A130C6FE43" box="[350,510,384,410]" pageId="35" pageNumber="36" value="2007-10-11">11 Oct. 2007</collectingDate>
</date>
;
<collectorName id="269B5345462FFFE1420C42A13393FE43" box="[526,683,384,410]" pageId="35" pageNumber="36">D.G. Herbert</collectorName>
and
<collectorName id="269B5345462FFFE142E842A13251FE43" box="[746,873,384,410]" pageId="35" pageNumber="36">L.S. Davis</collectorName>
leg.;
<collectedFrom id="3872756E462FFFE143B042A13003FE64" pageId="35" pageNumber="36">afrotemperate forest, in leaf-litter and under logs</collectedFrom>
; diameter
<quantity id="4C969B76462FFFE141B742823319FE67" box="[437,545,419,446]" metricMagnitude="-3" metricUnit="m" metricValue="1.45" pageId="35" pageNumber="36" unit="mm" value="1.45">1.45 mm</quantity>
, height
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;
<collectionCode id="ED7FAE56462FFFE142F742823270FE64" box="[757,840,419,445]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="35" pageNumber="36" type="Museum">NMSA</collectionCode>
<specimenCode id="DBC89EE8462FFFE1434D4282353AFE67" box="[847,1026,419,446]" pageId="35" pageNumber="36">W5826/T4258.</specimenCode>
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</paragraph>
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<typeStatus id="54D58831462FFFE140E842C4305FFE26" box="[234,359,485,511]" pageId="35" pageNumber="36" type="paratype">Paratypes</typeStatus>
</emphasis>
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<collectingRegion id="49AAF871462FFFE141A84129332CFDF8" box="[426,532,519,546]" pageId="35" pageNumber="36">W. Cape</collectingRegion>
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<specimenCount id="9D68FD1A462FFFE1422E41293387FDFB" box="[556,703,520,546]" pageId="35" pageNumber="36" type="generic">3 specimens</specimenCount>
; same collection data as for holotype;
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<specimenCode id="DBC89EE8462FFFE144D54126344EFDFB" box="[1239,1398,519,546]" pageId="35" pageNumber="36">P1017/T4259</specimenCode>
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<materialsCitation id="3B063CCE462FFFE140D1410A3035FD55" ID-GBIF-Occurrence="2605305713" collectingDate="2007-10-13" collectionCode="NMSA" collectorName="D. G. Herbert &amp; L. S. Davis" country="South Africa" elevation="345" latitude="-33.98994" location="Grootvadersbosch Nat. Res." longLatPrecision="1" longitude="20.81658" pageId="35" pageNumber="36" specimenCode="W5899/ T4256" specimenCount="1" stateProvince="Western Cape" typeStatus="paratype">
<specimenCount id="9D68FD1A462FFFE140D1410A3058FD9C" box="[211,352,555,581]" pageId="35" pageNumber="36" type="generic">1 specimen</specimenCount>
;
<location id="8EB16048462FFFE14172410A33FDFD9C" LSID="urn:lsid:plazi:treatment:03C78785462EFFE6423C46C7344FFE39:8EB16048462FFFE14172410A33FDFD9C" box="[368,709,555,581]" country="South Africa" latitude="-33.98994" longLatPrecision="1" longitude="20.81658" name="Grootvadersbosch Nat. Res." pageId="35" pageNumber="36" stateProvince="Western Cape">Grootvadersbosch Nat. Res.</location>
;
<geoCoordinate id="EE5A5054462FFFE142D7410A3255FD9C" box="[725,877,555,581]" degrees="33.98994" direction="south" orientation="latitude" pageId="35" pageNumber="36" precision="1" value="-33.98994">33.98994° S</geoCoordinate>
,
<geoCoordinate id="EE5A5054462FFFE1437C410A3520FD9C" box="[894,1048,555,582]" degrees="20.81658" direction="east" orientation="longitude" pageId="35" pageNumber="36" precision="1" value="20.81658">20.81658° E</geoCoordinate>
;
<elevation id="0043D1A0462FFFE1442B410A3584FD9C" box="[1065,1212,555,582]" metricMagnitude="2" metricUnit="m" metricValue="3.45" pageId="35" pageNumber="36" unit="m" value="345.0">
<quantity id="4C969B76462FFFE1442B410A354FFD9C" box="[1065,1143,555,582]" metricMagnitude="2" metricUnit="m" metricValue="3.45" pageId="35" pageNumber="36" unit="m" value="345.0">345 m</quantity>
a.s.l.
</elevation>
;
<date id="FFD01053462FFFE144CF410A3449FD9C" box="[1229,1393,555,581]" pageId="35" pageNumber="36" value="2007-10-13">
<collectingDate id="EF94E9BB462FFFE144CF410A3449FD9C" box="[1229,1393,555,581]" pageId="35" pageNumber="36" value="2007-10-13">13 Oct. 2007</collectingDate>
</date>
;
<collectorName id="269B5345462FFFE140BF416F3062FDB1" box="[189,346,590,616]" pageId="35" pageNumber="36">D.G. Herbert</collectorName>
and
<collectorName id="269B5345462FFFE1419A416F332FFDB1" box="[408,535,590,616]" pageId="35" pageNumber="36">L.S. Davis</collectorName>
leg.; afrotemperate forest, in leaf-litter and under logs;
<collectionCode id="ED7FAE56462FFFE144B9416F3436FDB1" box="[1211,1294,590,616]" collectionName="South Africa, Kwa-Zulu Natal, Pietermaritzburg, Natal Museum" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="35" pageNumber="36" type="Museum">NMSA</collectionCode>
<specimenCode id="DBC89EE8462FFFE1451A416F3035FD55" pageId="35" pageNumber="36">W5899/ T4256</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3746518462FFFE140BF4193345DFB1C" pageId="35" pageNumber="36" type="description">
<paragraph id="8BD13693462FFFE140BF41933076FD15" blockId="35.[189,334,690,716]" box="[189,334,690,716]" pageId="35" pageNumber="36">
<heading id="D09981FF462FFFE140BF41933076FD15" bold="true" box="[189,334,690,716]" fontSize="11" level="3" pageId="35" pageNumber="36" reason="3">
<emphasis id="B91AEA81462FFFE140BF41933076FD15" bold="true" box="[189,334,690,716]" pageId="35" pageNumber="36">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BD13693462FFFE140BF41C0345DFB1C" blockId="35.[189,1399,737,1221]" pageId="35" pageNumber="36">
Shell small, diameter up to
<quantity id="4C969B76462FFFE141F141C03364FD25" box="[499,604,737,764]" metricMagnitude="-3" metricUnit="m" metricValue="1.68" pageId="35" pageNumber="36" unit="mm" value="1.68">1.68 mm</quantity>
, H/D ratio ±0.48; symmetrically biconcave, whorls tightly coiled, but not conspicuously deep; spire sunken, but not deeply so; last adult whorl not descendant; suture strongly indented and apical and basal portions of whorls strongly convex, less so at mid-whorl; periphery evenly convex. Protoconch comprising apical cap plus approx. 1.25 whorls; diameter ±330 μm; initially with traces of close-set spiral threads, later portion with spiral threads and close-set axial riblets producing a fine, reticulate sculpture. Teleoconch of up to 3.5 whorls; sculptured by distinct, close-set compound, orthocline axial riblets with finer, uneven intermediary axial threads; intervals between riblets approx. twice riblet width at whorl periphery; spiral sculpture relatively distinct, comprising microscopic spiral threads, most obvious in intervals between riblets; threads coarsest below suture and in umbilicus, but present throughout. Umbilicus very wide and relatively shallow (
<figureCitation id="13552A16462FFFE143A5473F3528FBE1" box="[935,1040,1054,1080]" captionStart="Fig" captionStartId="35.[189,232,2001,2027]" captionTargetBox="[195,1390,1564,1953]" captionTargetId="figure@35.[189,1398,1559,1960]" captionTargetPageId="35" captionText="Fig. 13. Iterodonta ammonita gen. et sp. nov., holotype, diameter 1.45 mm (NMSA W5826/T4258)." figureDoi="http://doi.org/10.5281/zenodo.3762306" httpUri="https://zenodo.org/record/3762306/files/figure.png" pageId="35" pageNumber="36">Fig. 13B</figureCitation>
). Aperture narrow, more or less symmetrically lunate, with apical limit rounded; parietal and columellar dentition lacking; palatal region with 13 broad, prosocline, axial, ridge-like calluses set back ⅛–½ whorl from lip (position variable), visible through translucent shell (
<figureCitation id="13552A16462FFFE1424F47A93383FB7B" box="[589,699,1160,1186]" captionStart="Fig" captionStartId="35.[189,232,2001,2027]" captionTargetBox="[195,1390,1564,1953]" captionTargetId="figure@35.[189,1398,1559,1960]" captionTargetPageId="35" captionText="Fig. 13. Iterodonta ammonita gen. et sp. nov., holotype, diameter 1.45 mm (NMSA W5826/T4258)." figureDoi="http://doi.org/10.5281/zenodo.3762306" httpUri="https://zenodo.org/record/3762306/files/figure.png" pageId="35" pageNumber="36">Fig. 13D</figureCitation>
); calluses slightly curved, one usually well developed, the others in the process of resorption or deposition. Shell translucent, straw-brown to pale honey-brown.
</paragraph>
</subSubSection>
<subSubSection id="C3746518462FFFE140BF47CD343CFA18" pageId="35" pageNumber="36" type="distribution">
<paragraph id="8BD13693462FFFE140BF47CD330CFADF" blockId="35.[189,564,1260,1286]" box="[189,564,1260,1286]" pageId="35" pageNumber="36">
<heading id="D09981FF462FFFE140BF47CD330CFADF" bold="true" box="[189,564,1260,1286]" fontSize="11" level="3" pageId="35" pageNumber="36" reason="3">
<emphasis id="B91AEA81462FFFE140BF47CD330CFADF" bold="true" box="[189,564,1260,1286]" pageId="35" pageNumber="36">Distribution and conservation</emphasis>
</heading>
</paragraph>
<paragraph id="8BD13693462FFFE140BF463A343CFA18" blockId="35.[189,1399,1307,1474]" pageId="35" pageNumber="36">
A narrow-range endemic (
<figureCitation id="13552A16462FFFE141F8463A336BFAEC" box="[506,595,1307,1333]" captionStart="Fig" captionStartId="36.[189,232,1860,1886]" captionTargetBox="[302,1285,1199,1820]" captionTargetId="figure@36.[302,1285,1199,1820]" captionTargetPageId="36" captionText="Fig. 14. Distribution of Iterodonta gen. nov. and Phialodonta gen. nov. species. Iterodonta ammonita gen. et sp. nov. (brown triangles), Phialodonta agulhasae gen. et sp. nov. (orange square), P. atromontana gen. et sp. nov. (turquoise square), P. aviana gen. et sp. nov. (royal blue squares), P. introtuberculata (Connolly, 1933) gen. et comb. nov. (red squares), P. perfida (Burnup, 1912) gen. et comb. nov. (green square), P. rivalalea gen. et sp. nov. (pink squares). Contour at 1000 m." figureDoi="http://doi.org/10.5281/zenodo.3762308" httpUri="https://zenodo.org/record/3762308/files/figure.png" pageId="35" pageNumber="36">Fig. 14</figureCitation>
), known only from the south-facing slopes of the Langeberge and Riviersonderendberge, W.
<collectingRegion id="49AAF871462FFFE141FC461F3305FA81" box="[510,573,1342,1368]" country="South Africa" name="Western Cape" pageId="35" pageNumber="36">Cape</collectingRegion>
, at
<quantity id="4C969B76462FFFE1426C461F33C0FA80" box="[622,760,1342,1369]" metricMagnitude="2" metricUnit="m" metricValue="3.75" metricValueMax="4.5" metricValueMin="3.0" pageId="35" pageNumber="36" unit="m" value="375.0" valueMax="450.0" valueMin="300.0">300450 m</quantity>
a.s.l.; in leaf-litter of southern afrotemperate forest. Grootvadersbosch Nature Reserve is a formally protected area and the indigenous forest of the Langeberge and Riviersonderendberge are generally well managed, with additional formally protected areas that should be surveyed in the hope of finding additional extant colonies of this species.
</paragraph>
</subSubSection>
<caption id="DF11661B462FFFE140BF44F03469F835" ID-DOI="http://doi.org/10.5281/zenodo.3762306" ID-Zenodo-Dep="3762306" box="[189,1361,2001,2028]" httpUri="https://zenodo.org/record/3762306/files/figure.png" pageId="35" pageNumber="36" startId="35.[189,232,2001,2027]" targetBox="[195,1390,1564,1953]" targetPageId="35">
<paragraph id="8BD13693462FFFE140BF44F03469F835" blockId="35.[189,1361,2001,2028]" box="[189,1361,2001,2028]" pageId="35" pageNumber="36">
<emphasis id="B91AEA81462FFFE140BF44F03021F832" bold="true" box="[189,281,2001,2027]" pageId="35" pageNumber="36">Fig. 13.</emphasis>
<taxonomicName id="4C6E4D10462FFFE1412244F0332FF832" authority="Herbert, 2020" authorityName="Herbert" authorityYear="2020" box="[288,535,2001,2027]" class="Gastropoda" family="Charopidae" genus="Iterodonta" kingdom="Animalia" order="Stylommatophora" pageId="35" pageNumber="36" phylum="Mollusca" rank="species" species="ammonita" status="gen. et sp. nov.">
<emphasis id="B91AEA81462FFFE1412244F0332FF832" box="[288,535,2001,2027]" italics="true" pageId="35" pageNumber="36">Iterodonta ammonita</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22957FA462FFFE1421D44F333EAF835" box="[543,722,2002,2028]" pageId="35" pageNumber="36" rank="species">gen. et sp. nov.</taxonomicNameLabel>
, holotype, diameter 1.45 mm (NMSA W5826/T4258).
</paragraph>
</caption>
<subSubSection id="C37465184628FFE640BF422A344FFE39" pageId="36" pageNumber="37" type="discussion">
<paragraph id="8BD136934628FFE640BF422A3017FEFC" blockId="36.[189,303,267,293]" box="[189,303,267,293]" pageId="36" pageNumber="37">
<heading id="D09981FF4628FFE640BF422A3017FEFC" bold="true" box="[189,303,267,293]" fontSize="11" level="3" pageId="36" pageNumber="37" reason="3">
<emphasis id="B91AEA814628FFE640BF422A3017FEFC" bold="true" box="[189,303,267,293]" pageId="36" pageNumber="37">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BD136934628FFE640BF4218344FFE39" blockId="36.[189,1399,313,481]" pageId="36" pageNumber="37">
<taxonomicName id="4C6E4D104628FFE640BF4218308CFE8A" authority="Herbert, 2020" authorityName="Herbert" authorityYear="2020" box="[189,436,313,339]" class="Gastropoda" family="Charopidae" genus="Iterodonta" kingdom="Animalia" order="Stylommatophora" pageId="36" pageNumber="37" phylum="Mollusca" rank="species" species="ammonita" status="gen. et sp. nov.">
<emphasis id="B91AEA814628FFE640BF4218308CFE8A" box="[189,436,313,339]" italics="true" pageId="36" pageNumber="37">Iterodonta ammonita</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22957FA4628FFE641B9421B3353FE8D" box="[443,619,314,340]" pageId="36" pageNumber="37" rank="species">gen. et sp. nov.</taxonomicNameLabel>
is highly distinctive amongst the southern African charopid fauna. The only other African species with similar palatal dentition is
<taxonomicName id="4C6E4D104628FFE643D9427C344FFEAE" authority="Connolly, 1925" authorityName="Connolly" authorityYear="1925" box="[987,1399,348,375]" class="Gastropoda" family="Endodontidae" genus="Endodonta" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="36" pageNumber="37" phylum="Mollusca" rank="species" species="kempi">
<emphasis id="B91AEA814628FFE643D9427C3597FEAE" box="[987,1199,349,375]" italics="true" pageId="36" pageNumber="37">Endodonta kempi</emphasis>
<bibRefCitation id="EFFF4B624628FFE644BE427D344FFEAE" author="Connolly M." box="[1212,1399,348,375]" pageId="36" pageNumber="37" pagination="105 - 220" refId="ref26598" refString="Connolly M. 1925. The non-marine Mollusca of Portuguese East Africa. Transactions of the Royal Society of South Africa 12 (3): 105 - 220. https: // doi. org / 10.1080 / 00359192409519300" type="journal article" year="1925">Connolly, 1925</bibRefCitation>
</taxonomicName>
recorded from
<collectingCountry id="F37976034628FFE6417242A130F8FE43" box="[368,448,384,410]" name="Kenya" pageId="36" pageNumber="37">Kenya</collectingCountry>
,
<collectingCountry id="F37976034628FFE641CF42A1331FFE43" box="[461,551,384,410]" name="Malawi" pageId="36" pageNumber="37">Malaŵi</collectingCountry>
and
<collectingCountry id="F37976034628FFE6426642A13386FE43" box="[612,702,384,410]" name="Zambia" pageId="36" pageNumber="37">Zambia</collectingCountry>
(
<bibRefCitation id="EFFF4B624628FFE642CD42A13245FE43" author="Bruggen A. C. van" box="[719,893,384,410]" pageId="36" pageNumber="37" pagination="1 - 17" refId="ref26318" refString="Bruggen A. C. van. 1988. A record of the genus Cerastua (Mollusca, Gastropoda Pulmonata: Enidae) from Zambia, with a preliminary list of the terrestrial molluscs of that country. Proceedings of the Koninklijke Nederlandse Akademie van Wetenschappen 91 (1): 1 - 17." type="journal article" year="1988">Bruggen 1988</bibRefCitation>
,
<bibRefCitation id="EFFF4B624628FFE6438942A132F2FE43" author="Bruggen A. C. van" box="[907,970,384,410]" pageId="36" pageNumber="37" pagination="17 - 37" refId="ref26373" refString="Bruggen A. C. van. 2007. Terrestrial molluscs recorded from Mt. Mulanje, Malawi. Nyala 24: 17 - 37." type="journal article" year="2007">2007</bibRefCitation>
), but in that species the spire is not sunken, the umbilicus is narrower and the protoconch lacks distinctive sculpture.
<bibRefCitation id="EFFF4B624628FFE644714282342AFE64" author="Solem A." box="[1139,1298,419,445]" pageId="36" pageNumber="37" pagination="342 - 364" refId="ref27176" refString="Solem A. 1970. The land snail genus Afrodonta (Mollusca: Gastropoda: Endodontidae). Annals of the Natal Museum 20: 342 - 364." type="journal article" year="1970">Solem (1970)</bibRefCitation>
referred the latter to
<taxonomicName id="4C6E4D104628FFE6414A42E73086FE39" authorityName="Melvill &amp; Ponsonby" authorityYear="1908" box="[328,446,454,480]" class="Gastropoda" family="Endodontidae" genus="Afrodonta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="36" pageNumber="37" phylum="Mollusca" rank="genus">
<emphasis id="B91AEA814628FFE6414A42E73086FE39" box="[328,446,454,480]" italics="true" pageId="36" pageNumber="37">Afrodonta</emphasis>
</taxonomicName>
, but this seems improbable and its true relationships require further investigation.
</paragraph>
</subSubSection>
</treatment>
</document>