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<document id="351BD9B17C5FA2CD82487F54C1CA595D" ID-CLB-Dataset="22877" ID-DOI="10.5852/ejt.2020.629" ID-GBIF-Dataset="3722aa65-b935-4294-ac20-0d1179da9015" ID-Zenodo-Dep="3762280" ID-ZooBank="ECEBD539-6E3E-45BE-A0CB-264DF3270CC0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1587578640316" checkinUser="valdenar" docAuthor="Herbert, David G." docDate="2020" docId="03C787854625FFE94190443C3036F968" docLanguage="en" docName="ejt-629_herbert.pdf.imf" docOrigin="European Journal of Taxonomy 629" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Phialodonta introtuberculata Herbert 2020, gen. et comb. nov." docType="treatment" docVersion="7" lastPageNumber="44" masterDocId="FFFEFFFD460CFFC2400243213138FFD9" masterDocTitle="Revision of the aperturally dentate Charopidae (Gastropoda: Stylommatophora) of southern Africa - genus Afrodonta s. lat., with description of five new genera, twelve new species and one new subspecies" masterLastPageNumber="55" masterPageNumber="1" pageNumber="42" updateTime="1698835565728" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="747E34489A538F0AD58A7F7CB670303E">Revision of the aperturally dentate Charopidae (Gastropoda: Stylommatophora) of southern Africa - genus Afrodonta s. lat., with description of five new genera, twelve new species and one new subspecies</mods:title>
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<mods:namePart id="9884DE26B3D57569679B79121179589D">Herbert, David G.</mods:namePart>
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<mods:affiliation id="770A5264152D96E2783CDB6FC1037A46">Department of Natural Sciences, National Museum Wales, Cathays Park, Cardiff, CF 10 3 NP, UK. Formerly at KwaZulu-Natal Museum, P. Bag 9070, Pietermaritzburg 3201, South Africa</mods:affiliation>
<mods:nameIdentifier id="53C4160BCC767842873175F2C0D7E1F2" type="email">phasianella@gmail.com</mods:nameIdentifier>
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<mods:title id="EA4C9FB3C4D8A0D99D1E5DF8E8C3FA40">European Journal of Taxonomy</mods:title>
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<mods:date id="45CBB6B30BC60F031F5936466B971E6C">2020</mods:date>
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<treatment id="03C787854625FFE94190443C3036F968" ID-DOI="http://doi.org/10.5281/zenodo.3804750" ID-GBIF-Taxon="163732363" ID-Zenodo-Dep="3804750" LSID="urn:lsid:plazi:treatment:03C787854625FFE94190443C3036F968" httpUri="http://treatment.plazi.org/id/03C787854625FFE94190443C3036F968" lastPageId="43" lastPageNumber="44" pageId="41" pageNumber="42">
<subSubSection id="C37465184625FFEB4190443C3599F8E1" box="[402,1185,1821,1848]" pageId="41" pageNumber="42" type="nomenclature">
<paragraph id="8BD136934625FFEB4190443C3599F8E1" blockId="41.[402,1185,1821,1883]" box="[402,1185,1821,1848]" pageId="41" pageNumber="42">
<heading id="D09981FF4625FFEB4190443C3599F8E1" box="[402,1185,1821,1848]" centered="true" fontSize="11" level="2" pageId="41" pageNumber="42" reason="2">
<taxonomicName id="4C6E4D104625FFEB4190443C32FAF8E1" ID-CoL="7SP54" authority="(Connolly, 1933)" authorityName="Herbert" authorityYear="2020" baseAuthorityName="Connolly" baseAuthorityYear="1933" box="[402,962,1821,1848]" class="Gastropoda" family="Charopidae" genus="Phialodonta" kingdom="Animalia" order="Stylommatophora" pageId="41" pageNumber="42" phylum="Mollusca" rank="species" species="introtuberculata" status="gen. et comb. nov.">
<emphasis id="B91AEA814625FFEB4190443C33CBF8E1" bold="true" box="[402,755,1821,1848]" italics="true" pageId="41" pageNumber="42">Phialodonta introtuberculata</emphasis>
(
<bibRefCitation id="EFFF4B624625FFEB4301443F3282F8E1" author="Connolly M." box="[771,954,1822,1848]" pageId="41" pageNumber="42" pagination="145 - 150" refId="ref26638" refString="Connolly M. 1933. On South African Endodontidae. Part 2. Annals of the Natal Museum 7 (2): 145 - 150." type="journal article" year="1933">Connolly, 1933</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A22957FA4625FFEB43C8443F3599F8E1" box="[970,1185,1822,1848]" pageId="41" pageNumber="42" rank="species">gen. et comb. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37465184625FFEB428B44603293F882" box="[649,939,1857,1883]" pageId="41" pageNumber="42" type="description">
<paragraph id="8BD136934625FFEB428B44603293F882" blockId="41.[402,1185,1821,1883]" box="[649,939,1857,1883]" pageId="41" pageNumber="42">
<figureCitation id="13552A164625FFEB428B446033DBF882" box="[649,739,1857,1883]" captionStart="Fig" captionStartId="36.[189,232,1860,1886]" captionTargetBox="[302,1285,1199,1820]" captionTargetId="figure@36.[302,1285,1199,1820]" captionTargetPageId="36" captionText="Fig. 14. Distribution of Iterodonta gen. nov. and Phialodonta gen. nov. species. Iterodonta ammonita gen. et sp. nov. (brown triangles), Phialodonta agulhasae gen. et sp. nov. (orange square), P. atromontana gen. et sp. nov. (turquoise square), P. aviana gen. et sp. nov. (royal blue squares), P. introtuberculata (Connolly, 1933) gen. et comb. nov. (red squares), P. perfida (Burnup, 1912) gen. et comb. nov. (green square), P. rivalalea gen. et sp. nov. (pink squares). Contour at 1000 m." figureDoi="http://doi.org/10.5281/zenodo.3762308" httpUri="https://zenodo.org/record/3762308/files/figure.png" pageId="41" pageNumber="42">Figs 14</figureCitation>
,
<figureCitation id="13552A164625FFEB42ED44603233F882" box="[751,779,1857,1883]" captionStart="Fig" captionStartId="42.[189,232,1895,1921]" captionTargetBox="[190,1395,677,1852]" captionTargetId="figure@42.[189,1399,667,1855]" captionTargetPageId="42" captionText="Fig. 16. Shells of Phialodonta gen. nov. species. AD. P. introtuberculata (Connolly, 1933) gen. et comb. nov., Karkloof, KwaZulu-Natal, diameter 1.87 mm (NMSA W2438). EH. P. perfida (Burnup, 1912) gen. et comb. nov., paratype, diameter 1.84 mm (NMSA 2602/T617). IL. P. rivalalea gen. et sp. nov., holotype, diameter 1.70 mm (NMSA W3114 /T4251)." figureDoi="http://doi.org/10.5281/zenodo.3762312" httpUri="https://zenodo.org/record/3762312/files/figure.png" pageId="41" pageNumber="42">16</figureCitation>
AD, 19GH
</paragraph>
</subSubSection>
<subSubSection id="C37465184625FFEB40BF445A3316F832" pageId="41" pageNumber="42" type="reference_group">
<paragraph id="8BD136934625FFEB40BF445A3571F84F" blockId="41.[189,1097,1915,1942]" box="[189,1097,1915,1942]" pageId="41" pageNumber="42">
<treatmentCitationGroup id="AB7E11BD4625FFEB40BF445A3571F84F" box="[189,1097,1915,1942]" pageId="41" pageNumber="42">
<treatmentCitation id="0ACF10824625FFEB40BF445A33CAF84C" author="Connolly M." box="[189,754,1915,1941]" page="147" pageId="41" pageNumber="42" year="1933">
<taxonomicName id="4C6E4D104625FFEB40BF445A33CAF84C" authority="Connolly, 1933: 147" authorityName="Connolly" authorityPageNumber="147" authorityYear="1933" box="[189,754,1915,1941]" class="Gastropoda" family="Endodontidae" genus="Afrodonta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="41" pageNumber="42" phylum="Mollusca" rank="species" species="introtuberculata">
<emphasis id="B91AEA814625FFEB40BF445A30C0F84C" box="[189,504,1915,1941]" italics="true" pageId="41" pageNumber="42">Afrodonta introtuberculata</emphasis>
<bibRefCitation id="EFFF4B624625FFEB4202445A33CAF84C" author="Connolly M." box="[512,754,1915,1941]" pageId="41" pageNumber="42" pagination="145 - 150" refId="ref26638" refString="Connolly M. 1933. On South African Endodontidae. Part 2. Annals of the Natal Museum 7 (2): 145 - 150." type="journal article" year="1933">Connolly, 1933: 147</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, text-fig. 1(9), pl. 7, figs 58.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BD136934625FFEB40BF448C3316F832" blockId="41.[189,1399,1965,2027]" pageId="41" pageNumber="42">
<treatmentCitationGroup id="AB7E11BD4625FFEB40BF448C3316F832" pageId="41" pageNumber="42">
<taxonomicName id="4C6E4D104625FFEB40BF448C30C2F81E" authorityName="Connolly" authorityYear="1933" box="[189,506,1965,1991]" class="Gastropoda" family="Endodontidae" genus="Afrodonta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="41" pageNumber="42" phylum="Mollusca" rank="species" species="introtuberculata">
<emphasis id="B91AEA814625FFEB40BF448C30C2F81E" box="[189,506,1965,1991]" italics="true" pageId="41" pageNumber="42">Afrodonta introtuberculata</emphasis>
</taxonomicName>
<treatmentCitation id="0ACF10824625FFEB421E448C3236F811" author="Connolly M." box="[540,782,1965,1992]" page="255" pageId="41" pageNumber="42" year="1939">
<bibRefCitation id="EFFF4B624625FFEB421E448C3236F811" author="Connolly M." box="[540,782,1965,1992]" pageId="41" pageNumber="42" pagination="1 - 660" refId="ref26665" refString="Connolly M. 1939. A monographic survey of South African non-marine Mollusca. Annals of the South African Museum 33: 1 - 660." type="journal article" year="1939">Connolly 1939: 255</bibRefCitation>
</treatmentCitation>
, text-fig. 19(9). —
<treatmentCitation id="0ACF10824625FFEB43FA448C35F0F81E" author="Solem A." box="[1016,1224,1965,1992]" page="361" pageId="41" pageNumber="42" year="1970">
<bibRefCitation id="EFFF4B624625FFEB43FA448C35F0F81E" author="Solem A." box="[1016,1224,1965,1992]" pageId="41" pageNumber="42" pagination="342 - 364" refId="ref27176" refString="Solem A. 1970. The land snail genus Afrodonta (Mollusca: Gastropoda: Endodontidae). Annals of the Natal Museum 20: 342 - 364." type="journal article" year="1970">Solem 1970: 361</bibRefCitation>
</treatmentCitation>
. —
<treatmentCitation id="0ACF10824625FFEB44FF448C30FEF832" author="Herbert D. &amp; Kilburn D." page="247" pageId="41" pageNumber="42" year="2004">
<bibRefCitation id="EFFF4B624625FFEB44FF448C30FEF832" author="Herbert D. &amp; Kilburn D." pageId="41" pageNumber="42" refId="ref26755" refString="Herbert D. &amp; Kilburn D. 2004. Field Guide to the Land Snails and Slugs of Eastern South Africa. Natal Museum, Pietermaritzburg." type="book" year="2004">Herbert &amp; Kilburn 2004: 247</bibRefCitation>
</treatmentCitation>
, text-fig.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C37465184626FFE840BF422A35B4FDAF" pageId="42" pageNumber="43" type="diagnosis">
<paragraph id="8BD136934626FFE840BF422A300EFEFC" blockId="42.[189,310,267,293]" box="[189,310,267,293]" pageId="42" pageNumber="43">
<heading id="D09981FF4626FFE840BF422A300EFEFC" bold="true" box="[189,310,267,293]" fontSize="11" level="3" pageId="42" pageNumber="43" reason="3">
<emphasis id="B91AEA814626FFE840BF422A300EFEFC" bold="true" box="[189,310,267,293]" pageId="42" pageNumber="43">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BD136934626FFE840BF421B35B4FDAF" blockId="42.[189,1399,314,630]" pageId="42" pageNumber="43">Shell relatively large, narrowly planorboid, spire slightly sunken; protoconch initially smooth, becoming microscopically shagreened and finally with low axial undulations (diameter ±350 μm); teleoconch sculptured by relatively strong, close-set, compound axial riblets, intervals between riblets with 34 microscopic axial threads; spiral sculpture scarcely evident even in umbilicus; parietal and basocolumellar dentition lacking; palatal region with 13 axially aligned pairs of denticles set back ⅛ to ½ whorl behind outer lip, outermost pair usually visible through aperture, the others apparent only through translucent shell; lower denticle well below periphery, upper one slightly above periphery; number and position of denticle pairs somewhat variable and related to degree of development; umbilicus shallow and very wide. Shell translucent, honey-brown when fresh; diameter up to 2.0 mm.</paragraph>
</subSubSection>
<caption id="DF11661B4626FFE840BF4446324EF835" ID-DOI="http://doi.org/10.5281/zenodo.3762312" ID-Zenodo-Dep="3762312" httpUri="https://zenodo.org/record/3762312/files/figure.png" pageId="42" pageNumber="43" startId="42.[189,232,1895,1921]" targetBox="[190,1395,677,1852]" targetPageId="42">
<paragraph id="8BD136934626FFE840BF4446324EF835" blockId="42.[189,1398,1895,2028]" pageId="42" pageNumber="43">
<emphasis id="B91AEA814626FFE840BF44463025F858" bold="true" box="[189,285,1895,1921]" pageId="42" pageNumber="43">Fig. 16.</emphasis>
Shells of
<taxonomicName id="4C6E4D104626FFE8419F44463316F858" authority="Herbert, 2020" authorityName="Herbert" authorityYear="2020" box="[413,558,1895,1921]" class="Gastropoda" family="Charopidae" genus="Phialodonta" kingdom="Animalia" order="Stylommatophora" pageId="42" pageNumber="43" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="B91AEA814626FFE8419F44463316F858" box="[413,558,1895,1921]" italics="true" pageId="42" pageNumber="43">Phialodonta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22957FA4626FFE8423B44493390F85B" box="[569,680,1896,1922]" pageId="42" pageNumber="43" rank="genus">gen. nov.</taxonomicNameLabel>
species.
<emphasis id="B91AEA814626FFE8431B4446326AF858" bold="true" box="[793,850,1895,1921]" pageId="42" pageNumber="43">AD</emphasis>
.
<taxonomicName id="4C6E4D104626FFE8436644493421F85B" authority="(Connolly, 1933)" authorityName="Herbert" authorityYear="2020" baseAuthorityName="Connolly" baseAuthorityYear="1933" box="[868,1305,1895,1922]" class="Gastropoda" family="Endodontidae" genus="Afrodonta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="42" pageNumber="43" phylum="Mollusca" rank="species" species="introtuberculata" status="gen. et comb. nov.">
<emphasis id="B91AEA814626FFE843664449357BF858" box="[868,1091,1895,1921]" italics="true" pageId="42" pageNumber="43">P. introtuberculata</emphasis>
(
<bibRefCitation id="EFFF4B624626FFE8445444463429F85B" author="Connolly M." box="[1110,1297,1895,1922]" pageId="42" pageNumber="43" pagination="145 - 150" refId="ref26638" refString="Connolly M. 1933. On South African Endodontidae. Part 2. Annals of the Natal Museum 7 (2): 145 - 150." type="journal article" year="1933">Connolly, 1933</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A22957FA4626FFE845274449307AF87C" pageId="42" pageNumber="43" rank="species">gen. et comb. nov.</taxonomicNameLabel>
, Karkloof, KwaZulu-Natal, diameter 1.87 mm (NMSA W2438).
<emphasis id="B91AEA814626FFE8444844AA35BBF87C" bold="true" box="[1098,1155,1931,1957]" pageId="42" pageNumber="43">EH</emphasis>
.
<taxonomicName id="4C6E4D104626FFE8449144AD3039F811" authority="(Burnup, 1912)" authorityName="Herbert" authorityYear="2020" baseAuthorityName="Burnup" baseAuthorityYear="1912" class="Gastropoda" family="Charopidae" genus="Phialodonta" kingdom="Animalia" order="Stylommatophora" pageId="42" pageNumber="43" phylum="Mollusca" rank="species" species="perfida" status="gen. et comb. nov.">
<emphasis id="B91AEA814626FFE8449144AD343CF87C" box="[1171,1284,1931,1957]" italics="true" pageId="42" pageNumber="43">P. perfida</emphasis>
(
<bibRefCitation id="EFFF4B624626FFE8451744AA31C1F811" author="Burnup H. C." pageId="42" pageNumber="43" pagination="333 - 343" refId="ref26463" refString="Burnup H. C. 1912. On Afrodonta Melv. and Pons., with descriptions of new species. Annals of the Natal Museum 2 (3): 333 - 343." type="journal article" year="1912">Burnup, 1912</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A22957FA4626FFE8410A448F30D8F811" box="[264,480,1966,1992]" pageId="42" pageNumber="43" rank="species">gen. et comb. nov.</taxonomicNameLabel>
, paratype, diameter 1.84 mm (NMSA 2602/T617).
<emphasis id="B91AEA814626FFE84435448F355DF811" bold="true" box="[1079,1125,1966,1992]" pageId="42" pageNumber="43">IL</emphasis>
.
<taxonomicName id="4C6E4D104626FFE84471448E35C2F811" authority="Herbert, 2020" authorityName="Herbert" authorityYear="2020" box="[1139,1274,1966,1992]" class="Gastropoda" family="Charopidae" genus="Phialodonta" kingdom="Animalia" order="Stylommatophora" pageId="42" pageNumber="43" phylum="Mollusca" rank="species" species="rivalalea" status="gen. et sp. nov.">
<emphasis id="B91AEA814626FFE84471448E35C2F811" box="[1139,1274,1966,1992]" italics="true" pageId="42" pageNumber="43">P. rivalalea</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22957FA4626FFE84502448F31C9F835" pageId="42" pageNumber="43" rank="species">gen. et sp. nov.</taxonomicNameLabel>
, holotype, diameter 1.70 mm (NMSA W3114 /T4251).
</paragraph>
</caption>
<subSubSection id="C37465184627FFE940BF422A35AAFB9A" pageId="43" pageNumber="44" type="materials_examined">
<paragraph id="8BD136934627FFE940BF422A3092FEFC" blockId="43.[189,426,267,293]" box="[189,426,267,293]" pageId="43" pageNumber="44">
<heading id="D09981FF4627FFE940BF422A3092FEFC" bold="true" box="[189,426,267,293]" fontSize="11" level="3" pageId="43" pageNumber="44" reason="3">
<emphasis id="B91AEA814627FFE940BF422A3092FEFC" bold="true" box="[189,426,267,293]" pageId="43" pageNumber="44">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BD136934627FFE940E8421A3064FE8C" blockId="43.[189,1399,315,413]" box="[234,348,315,341]" pageId="43" pageNumber="44">
<emphasis id="B91AEA814627FFE940E8421A3064FE8C" bold="true" box="[234,348,315,341]" pageId="43" pageNumber="44">
<typeStatus id="54D588314627FFE940E8421A3064FE8C" box="[234,348,315,341]" pageId="43" pageNumber="44" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BD136934627FFE940BF427E3079FE44" blockId="43.[189,1399,315,413]" pageId="43" pageNumber="44">
<materialsCitation id="3B063CCE4627FFE940BF427E3079FE44" ID-GBIF-Occurrence="2605305701" collectionCode="NHMUK" collectorName="A. J. Taynton" country="South Africa" location="KwaZulu-Natal" pageId="43" pageNumber="44" specimenCode="NHMUK 1931.6.4.1." specimenCount="1" stateProvince="KwaZulu-Natal" typeStatus="holotype">
<collectingCountry id="F37976034627FFE940BF427E30ABFEA0" box="[189,403,351,377]" name="South Africa" pageId="43" pageNumber="44">SOUTH AFRICA</collectingCountry>
<collectingRegion id="49AAF8714627FFE941B4427E3349FEA0" box="[438,625,351,377]" country="South Africa" name="KwaZulu-Natal" pageId="43" pageNumber="44">KwaZulu-Natal</collectingRegion>
,
<locationDeviation id="9A632FB14627FFE94283427E3264FEA0" box="[641,860,351,377]" pageId="43" pageNumber="44">Nottingham Road</locationDeviation>
; prior to 1928;
<collectorName id="269B53454627FFE94423427E3584FEA0" box="[1057,1212,351,377]" pageId="43" pageNumber="44">A.J. Taynton</collectorName>
leg.;
<specimenCode id="DBC89EE84627FFE9450A427E3079FE44" pageId="43" pageNumber="44">NHMUK 1931.6.4.1.</specimenCode>
</materialsCitation>
</paragraph>
<paragraph id="8BD136934627FFE940E842E7305FFE39" blockId="43.[189,1399,454,552]" box="[234,359,454,480]" pageId="43" pageNumber="44">
<emphasis id="B91AEA814627FFE940E842E7305FFE39" bold="true" box="[234,359,454,480]" pageId="43" pageNumber="44">
<typeStatus id="54D588314627FFE940E842E7305FFE39" box="[234,359,454,480]" pageId="43" pageNumber="44" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BD136934627FFE940BF42CB31C7FDF1" blockId="43.[189,1399,454,552]" pageId="43" pageNumber="44">
<materialsCitation id="3B063CCE4627FFE940BF42CB31C3FDF1" ID-GBIF-Occurrence="2605305738" collectionCode="NHMUK" collectorName="A. J. Taynton" country="South Africa" location="SOUTH AFRICA" pageId="43" pageNumber="44" specimenCode="NHMUK 1937.12.30.2759 to 2761" specimenCount="3" stateProvince="KwaZulu-Natal" typeStatus="paratype">
<collectingCountry id="F37976034627FFE940BF42CB30A9FDDD" box="[189,401,490,516]" name="South Africa" pageId="43" pageNumber="44">SOUTH AFRICA</collectingCountry>
<specimenCount id="9D68FD1A4627FFE941AC42CB337DFDDD" box="[430,581,490,516]" pageId="43" pageNumber="44" type="generic">3 specimens</specimenCount>
; same collection data as for holotype;
<specimenCode id="DBC89EE84627FFE9441542CA31C3FDF1" pageId="43" pageNumber="44">NHMUK 1937.12.30.2759 to 2761</specimenCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8BD136934627FFE940E84170309EFDB2" blockId="43.[189,1399,593,1092]" box="[234,422,593,619]" pageId="43" pageNumber="44">
<emphasis id="B91AEA814627FFE940E84170309EFDB2" bold="true" box="[234,422,593,619]" pageId="43" pageNumber="44">Other material</emphasis>
</paragraph>
<paragraph id="8BD136934627FFE940BF415435AAFB9A" blockId="43.[189,1399,593,1092]" pageId="43" pageNumber="44">
<materialsCitation id="3B063CCE4627FFE940BF415430E4FD01" ID-GBIF-Occurrence="2605305705" collectedFrom="mistbelt Podocarpus forest, in leaf-litter" collectingDate="2002-06-22" collectionCode="NMSA" collectorName="D. G. Herbert" country="South Africa" county="Karkloof" elevation="1300" latitude="-29.3009" location="Holbeck" longLatPrecision="6" longitude="30.3615" municipality="Mbona Private Nat. Res." pageId="43" pageNumber="44" specimenCode="W216" specimenCount="4" stateProvince="KwaZulu-Natal">
<collectingCountry id="F37976034627FFE940BF415430ABFD56" box="[189,403,629,655]" name="South Africa" pageId="43" pageNumber="44">SOUTH AFRICA</collectingCountry>
<emphasis id="B91AEA814627FFE941B841573347FD49" bold="true" box="[442,639,630,656]" pageId="43" pageNumber="44">
<collectingRegion id="49AAF8714627FFE941B841573347FD49" box="[442,639,630,656]" country="South Africa" name="KwaZulu-Natal" pageId="43" pageNumber="44">KwaZulu-Natal</collectingRegion>
</emphasis>
<specimenCount id="9D68FD1A4627FFE942A041543202FD56" box="[674,826,629,655]" pageId="43" pageNumber="44" type="generic">4 specimens</specimenCount>
;
<collectingCounty id="62B04E1F4627FFE9434941543280FD56" box="[843,952,629,655]" pageId="43" pageNumber="44">Karkloof</collectingCounty>
,
<collectingMunicipality id="6BB5ACE94627FFE943CA415435C2FD49" box="[968,1274,629,656]" pageId="43" pageNumber="44">Mbona Private Nat. Res.</collectingMunicipality>
,
<location id="8EB160484627FFE945094154344AFD56" LSID="urn:lsid:plazi:treatment:03C787854625FFE94190443C3036F968:8EB160484627FFE945094154344AFD56" box="[1291,1394,629,655]" country="South Africa" county="Karkloof" latitude="-29.3009" longLatPrecision="6" longitude="30.3615" municipality="Mbona Private Nat. Res." name="Holbeck" pageId="43" pageNumber="44" stateProvince="KwaZulu-Natal">Holbeck</location>
;
<geoCoordinate id="EE5A50544627FFE940BF41BB307AFD6D" box="[189,322,666,692]" degrees="29.3009" direction="south" orientation="latitude" pageId="43" pageNumber="44" precision="5" value="-29.3009">29.3009° S</geoCoordinate>
,
<geoCoordinate id="EE5A50544627FFE9414D41B830EFFD6D" box="[335,471,665,692]" degrees="30.3615" direction="east" orientation="longitude" pageId="43" pageNumber="44" precision="5" value="30.3615">30.3615° E</geoCoordinate>
;
<elevation id="0043D1A04627FFE941E641BB3346FD6D" box="[484,638,666,692]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="43" pageNumber="44" unit="m" value="1300.0">
<quantity id="4C969B764627FFE941E641BB3306FD6D" box="[484,574,666,692]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="43" pageNumber="44" unit="m" value="1300.0">1300 m</quantity>
a.s.l.
</elevation>
;
<date id="FFD010534627FFE9428E41BB321FFD6D" box="[652,807,666,692]" pageId="43" pageNumber="44" value="2002-06-22">
<collectingDate id="EF94E9BB4627FFE9428E41BB321FFD6D" box="[652,807,666,692]" pageId="43" pageNumber="44" value="2002-06-22">22 Jun. 2002</collectingDate>
</date>
;
<collectorName id="269B53454627FFE9433641BB32F7FD6D" box="[820,975,666,692]" pageId="43" pageNumber="44">D.G. Herbert</collectorName>
leg.;
<collectedFrom id="3872756E4627FFE9441041BB3016FD01" pageId="43" pageNumber="44">
mistbelt
<taxonomicName id="4C6E4D104627FFE9447A41BB343EFD6D" box="[1144,1286,666,692]" class="Pinopsida" family="Podocarpaceae" genus="Podocarpus" kingdom="Plantae" order="Pinales" pageId="43" pageNumber="44" phylum="Tracheophyta" rank="genus">
<emphasis id="B91AEA814627FFE9447A41BB343EFD6D" box="[1144,1286,666,692]" italics="true" pageId="43" pageNumber="44">Podocarpus</emphasis>
</taxonomicName>
forest, in leaf-litter
</collectedFrom>
;
<collectionCode id="ED7FAE564627FFE94138419F30B5FD01" box="[314,397,702,728]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="43" pageNumber="44" type="Museum">NMSA</collectionCode>
<specimenCode id="DBC89EE84627FFE94196419F30E4FD01" box="[404,476,702,728]" collectionCode="NMSA" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="43" pageNumber="44" type="Museum">W216</specimenCode>
</materialsCitation>
<materialsCitation id="3B063CCE4627FFE941F7419F3055FCF8" ID-GBIF-Occurrence="2605305688" collectedFrom="mistbelt Podocarpus forest, in leaf-litter" collectingDate="2004-04-25" collectionCode="NMSA" collectorName="D. G. Herbert" country="South Africa" county="Karkloof" elevation="1400" latitude="-29.30119" location="Holbeck area" longLatPrecision="1" longitude="30.35899" municipality="Mbona Private Nat. Res." pageId="43" pageNumber="44" specimenCode="W1810" specimenCount="1" stateProvince="KwaZulu-Natal">
<specimenCount id="9D68FD1A4627FFE941F7419F3346FD01" box="[501,638,702,728]" pageId="43" pageNumber="44" type="generic">1 specimen</specimenCount>
;
<collectingCounty id="62B04E1F4627FFE94288419F33CFFD01" box="[650,759,702,728]" pageId="43" pageNumber="44">Karkloof</collectingCounty>
,
<collectingMunicipality id="6BB5ACE94627FFE94300419F351FFD00" box="[770,1063,702,729]" pageId="43" pageNumber="44">Mbona Private Nat. Res.</collectingMunicipality>
,
<location id="8EB160484627FFE94431419F35EBFD00" LSID="urn:lsid:plazi:treatment:03C787854625FFE94190443C3036F968:8EB160484627FFE94431419F35EBFD00" box="[1075,1235,702,729]" country="South Africa" county="Karkloof" latitude="-29.30119" longLatPrecision="1" longitude="30.35899" municipality="Mbona Private Nat. Res." name="Holbeck area" pageId="43" pageNumber="44" stateProvince="KwaZulu-Natal">Holbeck area</location>
;
<geoCoordinate id="EE5A50544627FFE944DD419F3449FD01" box="[1247,1393,702,728]" degrees="29.30119" direction="south" orientation="latitude" pageId="43" pageNumber="44" precision="1" value="-29.30119">29.30119° S</geoCoordinate>
,
<geoCoordinate id="EE5A50544627FFE940BF41C3306CFD24" box="[189,340,738,765]" degrees="30.35899" direction="east" orientation="longitude" pageId="43" pageNumber="44" precision="1" value="30.35899">30.35899° E</geoCoordinate>
;
<elevation id="0043D1A04627FFE9416341C330C4FD25" box="[353,508,738,765]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="43" pageNumber="44" unit="m" value="1400.0">
<quantity id="4C969B764627FFE9416341C33083FD24" box="[353,443,738,765]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="43" pageNumber="44" unit="m" value="1400.0">1400 m</quantity>
a.s.l.
</elevation>
;
<date id="FFD010534627FFE9420841C33390FD25" box="[522,680,738,765]" pageId="43" pageNumber="44" value="2004-04-25">
<collectingDate id="EF94E9BB4627FFE9420841C33390FD25" box="[522,680,738,765]" pageId="43" pageNumber="44" value="2004-04-25">25 Apr. 2004</collectingDate>
</date>
;
<collectorName id="269B53454627FFE942B741C33268FD25" box="[693,848,738,764]" pageId="43" pageNumber="44">D.G. Herbert</collectorName>
leg.;
<collectedFrom id="3872756E4627FFE9439141C3344AFD25" box="[915,1394,738,764]" pageId="43" pageNumber="44">
mistbelt
<taxonomicName id="4C6E4D104627FFE943F841C335B0FD25" box="[1018,1160,738,764]" class="Pinopsida" family="Podocarpaceae" genus="Podocarpus" kingdom="Plantae" order="Pinales" pageId="43" pageNumber="44" phylum="Tracheophyta" rank="genus">
<emphasis id="B91AEA814627FFE943F841C335B0FD25" box="[1018,1160,738,764]" italics="true" pageId="43" pageNumber="44">Podocarpus</emphasis>
</taxonomicName>
forest, in leaf-litter
</collectedFrom>
;
<collectionCode id="ED7FAE564627FFE940BF40263028FCF8" box="[189,272,775,801]" collectionName="South Africa, Kwa-Zulu Natal, Pietermaritzburg, Natal Museum" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="43" pageNumber="44" type="Museum">NMSA</collectionCode>
<specimenCode id="DBC89EE84627FFE9411540263055FCF8" box="[279,365,775,801]" collectionCode="NMSA" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="43" pageNumber="44" type="Museum">W1810</specimenCode>
</materialsCitation>
<materialsCitation id="3B063CCE4627FFE941874026344EFC9C" ID-GBIF-Occurrence="2605305717" collectedFrom="mistbelt Podocarpus forest, in leaf-litter" collectingDate="2004-10-24" collectionCode="NMSA" collectorName="D. G. Herbert" country="South Africa" county="Karkloof" elevation="1330" latitude="-29.30339" location="Holbeck" longLatPrecision="1" longitude="30.36525" municipality="Mbona Private Nat. Res." pageId="43" pageNumber="44" specimenCode="W2438" specimenCount="3" stateProvince="KwaZulu-Natal">
<specimenCount id="9D68FD1A4627FFE9418740263321FCF8" box="[389,537,775,801]" pageId="43" pageNumber="44" type="generic">3 specimens</specimenCount>
;
<collectingCounty id="62B04E1F4627FFE94226402633A9FCF8" box="[548,657,775,801]" pageId="43" pageNumber="44">Karkloof</collectingCounty>
,
<collectingMunicipality id="6BB5ACE94627FFE9429E40263286FCF8" box="[668,958,775,801]" pageId="43" pageNumber="44">Mbona Private Nat. Res.</collectingMunicipality>
,
<location id="8EB160484627FFE943C840263509FCF8" LSID="urn:lsid:plazi:treatment:03C787854625FFE94190443C3036F968:8EB160484627FFE943C840263509FCF8" box="[970,1073,775,801]" country="South Africa" county="Karkloof" latitude="-29.30339" longLatPrecision="1" longitude="30.36525" municipality="Mbona Private Nat. Res." name="Holbeck" pageId="43" pageNumber="44" stateProvince="KwaZulu-Natal">Holbeck</location>
;
<geoCoordinate id="EE5A50544627FFE9443F402635F7FCF8" box="[1085,1231,775,801]" degrees="29.30339" direction="south" orientation="latitude" pageId="43" pageNumber="44" precision="1" value="-29.30339">29.30339° S</geoCoordinate>
,
<geoCoordinate id="EE5A50544627FFE944DE40273449FCF8" box="[1244,1393,774,801]" degrees="30.36525" direction="east" orientation="longitude" pageId="43" pageNumber="44" precision="1" value="30.36525">30.36525° E</geoCoordinate>
;
<elevation id="0043D1A04627FFE940BF400A306DFC9C" box="[189,341,811,838]" metricMagnitude="3" metricUnit="m" metricValue="1.33" pageId="43" pageNumber="44" unit="m" value="1330.0">
<quantity id="4C969B764627FFE940BF400A302DFC9F" box="[189,277,811,838]" metricMagnitude="3" metricUnit="m" metricValue="1.33" pageId="43" pageNumber="44" unit="m" value="1330.0">1330 m</quantity>
a.s.l.
</elevation>
;
<date id="FFD010534627FFE94160400A30C4FC9C" box="[354,508,811,837]" pageId="43" pageNumber="44" value="2004-10-24">
<collectingDate id="EF94E9BB4627FFE94160400A30C4FC9C" box="[354,508,811,837]" pageId="43" pageNumber="44" value="2004-10-24">24 Oct. 2004</collectingDate>
</date>
;
<collectorName id="269B53454627FFE9420A400A339AFC9C" box="[520,674,811,837]" pageId="43" pageNumber="44">D.G. Herbert</collectorName>
leg.;
<collectedFrom id="3872756E4627FFE942E0400A3584FC9C" box="[738,1212,811,837]" pageId="43" pageNumber="44">
mistbelt
<taxonomicName id="4C6E4D104627FFE94345400A32EDFC9C" box="[839,981,811,837]" class="Pinopsida" family="Podocarpaceae" genus="Podocarpus" kingdom="Plantae" order="Pinales" pageId="43" pageNumber="44" phylum="Tracheophyta" rank="genus">
<emphasis id="B91AEA814627FFE94345400A32EDFC9C" box="[839,981,811,837]" italics="true" pageId="43" pageNumber="44">Podocarpus</emphasis>
</taxonomicName>
forest, in leaf-litter
</collectedFrom>
;
<collectionCode id="ED7FAE564627FFE944C5400A3422FC9C" box="[1223,1306,811,837]" collectionName="South Africa, Kwa-Zulu Natal, Pietermaritzburg, Natal Museum" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="43" pageNumber="44" type="Museum">NMSA</collectionCode>
<specimenCode id="DBC89EE84627FFE94522400A344EFC9C" box="[1312,1398,811,837]" collectionCode="NMSA" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="43" pageNumber="44" type="Museum">W2438</specimenCode>
</materialsCitation>
<materialsCitation id="3B063CCE4627FFE940D1406E3035FC6B" ID-GBIF-Occurrence="2605305731" collectedFrom="mistbelt Podocarpus forest, in leaf-litter" collectingDate="1998-11-23" collectionCode="NMSA" collectorName="D. G. Herbert &amp; M. Seddon &amp; P. Tattersfield" country="South Africa" elevation="1350" latitude="-29.3183" location="Leopard's Bush" longLatPrecision="6" longitude="30.258" municipality="Karkloof" pageId="43" pageNumber="44" specimenCode="V8417" specimenCount="1" stateProvince="KwaZulu-Natal">
<specimenCount id="9D68FD1A4627FFE940D1406E3067FCB0" box="[211,351,847,873]" pageId="43" pageNumber="44" type="generic">1 specimen</specimenCount>
;
<collectingMunicipality id="6BB5ACE94627FFE9416D406E30E4FCB0" box="[367,476,847,873]" pageId="43" pageNumber="44">Karkloof</collectingMunicipality>
,
<location id="8EB160484627FFE941E9406E3394FCB0" LSID="urn:lsid:plazi:treatment:03C787854625FFE94190443C3036F968:8EB160484627FFE941E9406E3394FCB0" box="[491,684,847,873]" country="South Africa" latitude="-29.3183" longLatPrecision="6" longitude="30.258" municipality="Karkloof" name="Leopard's Bush" pageId="43" pageNumber="44" stateProvince="KwaZulu-Natal">Leopards Bush</location>
;
<geoCoordinate id="EE5A50544627FFE942B9406E327CFCB0" box="[699,836,847,873]" degrees="29.3183" direction="south" orientation="latitude" pageId="43" pageNumber="44" precision="5" value="-29.3183">29.3183° S</geoCoordinate>
,
<geoCoordinate id="EE5A50544627FFE94357406E32D8FCB3" box="[853,992,847,874]" degrees="30.2580" direction="east" orientation="longitude" pageId="43" pageNumber="44" precision="5" value="30.258">30.2580° E</geoCoordinate>
; ca
<elevation id="0043D1A04627FFE94414406E358EFCB0" box="[1046,1206,847,874]" metricMagnitude="3" metricUnit="m" metricValue="1.35" pageId="43" pageNumber="44" unit="m" value="1350.0">
<quantity id="4C969B764627FFE94414406E354BFCB3" box="[1046,1139,847,874]" metricMagnitude="3" metricUnit="m" metricValue="1.35" pageId="43" pageNumber="44" unit="m" value="1350.0">1350 m</quantity>
a.s.l.
</elevation>
;
<date id="FFD010534627FFE944CA406E3449FCB0" box="[1224,1393,847,874]" pageId="43" pageNumber="44" value="1998-11-23">
<collectingDate id="EF94E9BB4627FFE944CA406E3449FCB0" box="[1224,1393,847,874]" pageId="43" pageNumber="44" value="1998-11-23">23 Nov. 1998</collectingDate>
</date>
;
<collectorName id="269B53454627FFE940BF40553067FC57" box="[189,351,884,910]" pageId="43" pageNumber="44">D.G. Herbert</collectorName>
,
<collectorName id="269B53454627FFE9416D405530CCFC57" box="[367,500,884,910]" pageId="43" pageNumber="44">M. Seddon</collectorName>
and
<collectorName id="269B53454627FFE94234405533E2FC57" box="[566,730,884,910]" pageId="43" pageNumber="44">P. Tattersfield</collectorName>
leg.;
<collectedFrom id="3872756E4627FFE943274055342BFC57" box="[805,1299,884,910]" pageId="43" pageNumber="44">
mistbelt
<taxonomicName id="4C6E4D104627FFE9439240553526FC57" box="[912,1054,884,910]" class="Pinopsida" family="Podocarpaceae" genus="Podocarpus" kingdom="Plantae" order="Pinales" pageId="43" pageNumber="44" phylum="Tracheophyta" rank="genus">
<emphasis id="B91AEA814627FFE9439240553526FC57" box="[912,1054,884,910]" italics="true" pageId="43" pageNumber="44">Podocarpus</emphasis>
</taxonomicName>
forest, in leaf-litter
</collectedFrom>
;
<collectionCode id="ED7FAE564627FFE945214055344EFC57" box="[1315,1398,884,910]" collectionName="South Africa, Kwa-Zulu Natal, Pietermaritzburg, Natal Museum" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="43" pageNumber="44" type="Museum">NMSA</collectionCode>
<specimenCode id="DBC89EE84627FFE940BF40B93035FC6B" box="[189,269,920,946]" collectionCode="NMSA" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="43" pageNumber="44" type="Museum">V8417</specimenCode>
</materialsCitation>
<materialsCitation id="3B063CCE4627FFE9412740B932E0FC0F" ID-GBIF-Occurrence="2605305623" collectedFrom="mistbelt Podocarpus forest, in leaf-litter" collectingDate="2003-12-01" collectionCode="NMSA" collectorName="A. Moussalli &amp; D. Stuart- Fox" country="South Africa" latitude="-29.3" location="Karkloof Nat. Res." longLatPrecision="736" longitude="30.23" pageId="43" pageNumber="44" specimenCode="W3789" specimenCount="3" stateProvince="KwaZulu-Natal">
<specimenCount id="9D68FD1A4627FFE9412740B93081FC6B" box="[293,441,920,946]" pageId="43" pageNumber="44" type="generic">3 specimens</specimenCount>
;
<location id="8EB160484627FFE941C640B9339CFC6A" LSID="urn:lsid:plazi:treatment:03C787854625FFE94190443C3036F968:8EB160484627FFE941C640B9339CFC6A" box="[452,676,920,947]" country="South Africa" latitude="-29.3" longLatPrecision="736" longitude="30.23" name="Karkloof Nat. Res." pageId="43" pageNumber="44" stateProvince="KwaZulu-Natal">Karkloof Nat. Res.</location>
;
<geoCoordinate id="EE5A50544627FFE942B240B9322EFC6B" box="[688,790,920,946]" degrees="29.30" direction="south" orientation="latitude" pageId="43" pageNumber="44" precision="555" value="-29.3">29.30° S</geoCoordinate>
,
<geoCoordinate id="EE5A50544627FFE9432140B932B5FC6A" box="[803,909,920,947]" degrees="30.23" direction="east" orientation="longitude" pageId="43" pageNumber="44" precision="555" value="30.23">30.23° E</geoCoordinate>
;
<date id="FFD010534627FFE9439B40B93512FC6B" box="[921,1066,920,947]" pageId="43" pageNumber="44" value="2003-12-01">
<collectingDate id="EF94E9BB4627FFE9439B40B93512FC6B" box="[921,1066,920,947]" pageId="43" pageNumber="44" value="2003-12-01">1 Dec. 2003</collectingDate>
</date>
;
<collectorName id="269B53454627FFE9443640B935F3FC6B" box="[1076,1227,920,946]" pageId="43" pageNumber="44">A. Moussalli</collectorName>
and
<collectorName id="269B53454627FFE9450140B831D3FC0E" pageId="43" pageNumber="44">D. Stuart- Fox</collectorName>
leg.;
<collectedFrom id="3872756E4627FFE94130409D322FFC0F" box="[306,791,956,982]" pageId="43" pageNumber="44">
mistbelt
<taxonomicName id="4C6E4D104627FFE94198409D3310FC0F" box="[410,552,956,982]" class="Pinopsida" family="Podocarpaceae" genus="Podocarpus" kingdom="Plantae" order="Pinales" pageId="43" pageNumber="44" phylum="Tracheophyta" rank="genus">
<emphasis id="B91AEA814627FFE94198409D3310FC0F" box="[410,552,956,982]" italics="true" pageId="43" pageNumber="44">Podocarpus</emphasis>
</taxonomicName>
forest, in leaf-litter
</collectedFrom>
;
<collectionCode id="ED7FAE564627FFE94327409D3240FC0F" box="[805,888,956,982]" collectionName="South Africa, Kwa-Zulu Natal, Pietermaritzburg, Natal Museum" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="43" pageNumber="44" type="Museum">NMSA</collectionCode>
<specimenCode id="DBC89EE84627FFE94380409D32E0FC0F" box="[898,984,956,982]" collectionCode="NMSA" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="43" pageNumber="44" type="Museum">W3789</specimenCode>
</materialsCitation>
<materialsCitation id="3B063CCE4627FFE943F4409C354FFC22" ID-GBIF-Occurrence="2605305744" collectionCode="NMSA" collectorName="A. J. Taynton" country="South Africa" elevation="1500" latitude="-29.358" location="Nottingham Road" longLatPrecision="73" longitude="29.995" pageId="43" pageNumber="44" specimenCode="A9191" specimenCount="7" stateProvince="KwaZulu-Natal">
<specimenCount id="9D68FD1A4627FFE943F4409C35B4FC0F" box="[1014,1164,956,983]" pageId="43" pageNumber="44" type="generic">7 specimens</specimenCount>
;
<location id="8EB160484627FFE94498409D344AFC0F" LSID="urn:lsid:plazi:treatment:03C787854625FFE94190443C3036F968:8EB160484627FFE94498409D344AFC0F" box="[1178,1394,956,982]" country="South Africa" latitude="-29.358" longLatPrecision="73" longitude="29.995" name="Nottingham Road" pageId="43" pageNumber="44" stateProvince="KwaZulu-Natal">Nottingham Road</location>
;
<geoCoordinate id="EE5A50544627FFE940BF40C1300AFC22" box="[189,306,992,1019]" degrees="29.358" direction="south" orientation="latitude" pageId="43" pageNumber="44" precision="55" value="-29.358">29.358° S</geoCoordinate>
,
<geoCoordinate id="EE5A50544627FFE9413C40C1308DFC22" box="[318,437,992,1019]" degrees="29.995" direction="east" orientation="longitude" pageId="43" pageNumber="44" precision="55" value="29.995">29.995° E</geoCoordinate>
; ca
<elevation id="0043D1A04627FFE941E340C13340FC22" box="[481,632,992,1019]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="43" pageNumber="44" unit="m" value="1500.0">
<quantity id="4C969B764627FFE941E340C13301FC22" box="[481,569,992,1019]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="43" pageNumber="44" unit="m" value="1500.0">1500 m</quantity>
a.s.l.
</elevation>
; pre-1928;
<collectorName id="269B53454627FFE942F940C032A8FC22" box="[763,912,993,1019]" pageId="43" pageNumber="44">A.J. Taynton</collectorName>
leg.;
<collectionCode id="ED7FAE564627FFE943CD40C0351AFC22" box="[975,1058,993,1019]" collectionName="South Africa, Kwa-Zulu Natal, Pietermaritzburg, Natal Museum" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="43" pageNumber="44" type="Museum">NMSA</collectionCode>
<specimenCode id="DBC89EE84627FFE9442540C0354FFC22" box="[1063,1143,993,1019]" collectionCode="NMSA" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="43" pageNumber="44" type="Museum">A9191</specimenCode>
</materialsCitation>
<materialsCitation id="3B063CCE4627FFE9448C40C133C9FBC6" ID-GBIF-Occurrence="2605305634" collectionCode="NMSA" collectorName="A. J. Taynton" country="South Africa" elevation="1500" latitude="-29.358" location="Nottingham Road" longLatPrecision="73" longitude="29.995" pageId="43" pageNumber="44" specimenCode="A9192" specimenCount="46" stateProvince="KwaZulu-Natal">
<specimenCount id="9D68FD1A4627FFE9448C40C13417FC22" box="[1166,1327,992,1019]" pageId="43" pageNumber="44" type="generic">46 specimens</specimenCount>
; same collection data as for preceding;
<collectionCode id="ED7FAE564627FFE94244472433A1FBC6" box="[582,665,1029,1055]" collectionName="South Africa, Kwa-Zulu Natal, Pietermaritzburg, Natal Museum" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="43" pageNumber="44" type="Museum">NMSA</collectionCode>
<specimenCode id="DBC89EE84627FFE942A3472433C9FBC6" box="[673,753,1029,1055]" collectionCode="NMSA" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="43" pageNumber="44" type="Museum">A9192</specimenCode>
</materialsCitation>
<materialsCitation id="3B063CCE4627FFE9430F4724305EFB9A" ID-GBIF-Occurrence="2605305685" collectionCode="NMSA" collectorName="A. J. Taynton" country="South Africa" elevation="1500" latitude="-29.358" location="Nottingham Road" longLatPrecision="73" longitude="29.995" pageId="43" pageNumber="44" specimenCode="A9193" specimenCount="3" stateProvince="KwaZulu-Natal">
<specimenCount id="9D68FD1A4627FFE9430F4724329BFBC6" box="[781,931,1029,1055]" pageId="43" pageNumber="44" type="generic">3 specimens</specimenCount>
; same collection data as for preceding;
<collectionCode id="ED7FAE564627FFE940BF47083028FB9A" box="[189,272,1065,1091]" collectionName="South Africa, Kwa-Zulu Natal, Pietermaritzburg, Natal Museum" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="43" pageNumber="44" type="Museum">NMSA</collectionCode>
<specimenCode id="DBC89EE84627FFE941144708305EFB9A" box="[278,358,1065,1091]" collectionCode="NMSA" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="43" pageNumber="44" type="Museum">A9193</specimenCode>
</materialsCitation>
<materialsCitation id="3B063CCE4627FFE9417D470835B6FB9A" ID-GBIF-Occurrence="2605305642" box="[383,1166,1065,1092]" collectionCode="NMSA" collectorName="A. J. Taynton" country="South Africa" elevation="1500" latitude="-29.358" location="Nottingham Road" longLatPrecision="73" longitude="29.995" pageId="43" pageNumber="44" specimenCode="A9221" specimenCount="6" stateProvince="KwaZulu-Natal">
<specimenCount id="9D68FD1A4627FFE9417D4708332BFB9A" box="[383,531,1065,1091]" pageId="43" pageNumber="44" type="generic">6 specimens</specimenCount>
; same collection data as for preceding;
<collectionCode id="ED7FAE564627FFE943E04708350DFB9A" box="[994,1077,1065,1091]" collectionName="South Africa, Kwa-Zulu Natal, Pietermaritzburg, Natal Museum" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="43" pageNumber="44" type="Museum">NMSA</collectionCode>
<specimenCode id="DBC89EE84627FFE94438470835B6FB9A" box="[1082,1166,1065,1091]" collectionCode="NMSA" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="43" pageNumber="44" type="Museum">A9221</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C37465184627FFE940BF474D35E3FA9E" pageId="43" pageNumber="44" type="distribution">
<paragraph id="8BD136934627FFE940BF474D330CFB5F" blockId="43.[189,564,1132,1158]" box="[189,564,1132,1158]" pageId="43" pageNumber="44">
<heading id="D09981FF4627FFE940BF474D330CFB5F" bold="true" box="[189,564,1132,1158]" fontSize="11" level="3" pageId="43" pageNumber="44" reason="3">
<emphasis id="B91AEA814627FFE940BF474D330CFB5F" bold="true" box="[189,564,1132,1158]" pageId="43" pageNumber="44">Distribution and conservation</emphasis>
</heading>
</paragraph>
<paragraph id="8BD136934627FFE940BF47BD35E3FA9E" blockId="43.[189,1399,1179,1351]" pageId="43" pageNumber="44">
A narrow-range endemic (
<figureCitation id="13552A164627FFE9420147BA3358FB6F" box="[515,608,1179,1206]" captionStart="Fig" captionStartId="36.[189,232,1860,1886]" captionTargetBox="[302,1285,1199,1820]" captionTargetId="figure@36.[302,1285,1199,1820]" captionTargetPageId="36" captionText="Fig. 14. Distribution of Iterodonta gen. nov. and Phialodonta gen. nov. species. Iterodonta ammonita gen. et sp. nov. (brown triangles), Phialodonta agulhasae gen. et sp. nov. (orange square), P. atromontana gen. et sp. nov. (turquoise square), P. aviana gen. et sp. nov. (royal blue squares), P. introtuberculata (Connolly, 1933) gen. et comb. nov. (red squares), P. perfida (Burnup, 1912) gen. et comb. nov. (green square), P. rivalalea gen. et sp. nov. (pink squares). Contour at 1000 m." figureDoi="http://doi.org/10.5281/zenodo.3762308" httpUri="https://zenodo.org/record/3762308/files/figure.png" pageId="43" pageNumber="44">Fig. 14</figureCitation>
), known only from the KarkloofNottingham Road area in the
<collectingRegion id="49AAF8714627FFE940BF47E1304DFB03" box="[189,373,1216,1242]" country="South Africa" name="KwaZulu-Natal" pageId="43" pageNumber="44">KwaZulu-Natal</collectingRegion>
<collectingRegion id="49AAF8714627FFE9417C47E130C8FB03" box="[382,496,1216,1242]" country="Zimbabwe" name="Midlands" pageId="43" pageNumber="44">Midlands</collectingRegion>
, at
<quantity id="4C969B764627FFE9421E47E133F8FB03" box="[540,704,1216,1243]" metricMagnitude="3" metricUnit="m" metricValue="1.4" metricValueMax="1.5" metricValueMin="1.3" pageId="43" pageNumber="44" unit="m" value="1400.0" valueMax="1500.0" valueMin="1300.0">13001500 m</quantity>
a.s.l.; in leaf-litter of southern mistbelt forest. Judging by the numbers of specimens in the original samples collected by A.J. Taynton in the
<collectingRegion id="49AAF8714627FFE944A147C53417FB27" box="[1187,1327,1252,1278]" country="United Kingdom" name="Nottingham" pageId="43" pageNumber="44">Nottingham</collectingRegion>
Road area (pre-1928), the species may be locally common or abundant. There are a number of provincial and private nature reserves in this area in which the species has been collected in recent years.
</paragraph>
</subSubSection>
<subSubSection id="C37465184627FFE940BF46513036F968" pageId="43" pageNumber="44" type="discussion">
<paragraph id="8BD136934627FFE940BF46513017FA53" blockId="43.[189,303,1392,1418]" box="[189,303,1392,1418]" pageId="43" pageNumber="44">
<heading id="D09981FF4627FFE940BF46513017FA53" bold="true" box="[189,303,1392,1418]" fontSize="11" level="3" pageId="43" pageNumber="44" reason="3">
<emphasis id="B91AEA814627FFE940BF46513017FA53" bold="true" box="[189,303,1392,1418]" pageId="43" pageNumber="44">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BD136934627FFE940BF46BE344FF9DB" blockId="43.[189,1399,1439,1538]" pageId="43" pageNumber="44">
<taxonomicName id="4C6E4D104627FFE940BF46BE332CFA60" authorityName="Herbert" authorityYear="2020" baseAuthorityName="Connolly" baseAuthorityYear="1933" box="[189,532,1439,1465]" class="Gastropoda" family="Charopidae" genus="Phialodonta" kingdom="Animalia" order="Stylommatophora" pageId="43" pageNumber="44" phylum="Mollusca" rank="species" species="introtuberculata" status="gen. et comb. nov.">
<emphasis id="B91AEA814627FFE940BF46BE332CFA60" box="[189,532,1439,1465]" italics="true" pageId="43" pageNumber="44">Phialodonta introtuberculata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22957FA4627FFE9421E468133CDFA63" box="[540,757,1439,1466]" pageId="43" pageNumber="44" rank="species">gen. et comb. nov.</taxonomicNameLabel>
is distinguished from other species in the genus by its much flatter shell and paired palatal dentition.
<taxonomicName id="4C6E4D104627FFE942D846E232D7FA04" authority="Herbert, 2020" authorityName="Herbert" authorityYear="2020" box="[730,1007,1475,1501]" class="Gastropoda" family="Endodontidae" genus="Afrodonta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="43" pageNumber="44" phylum="Mollusca" rank="species" species="geminodonta" status="sp. nov.">
<emphasis id="B91AEA814627FFE942D846E232D7FA04" box="[730,1007,1475,1501]" italics="true" pageId="43" pageNumber="44">Afrodonta geminodonta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22957FA4627FFE943F446E53568FA07" box="[1014,1104,1476,1502]" pageId="43" pageNumber="44" rank="species">sp. nov.</taxonomicNameLabel>
has similar sets of paired palatal denticles, but its spire is raised, the axial sculpture much finer and the umbilicus much narrower.
</paragraph>
<paragraph id="8BD136934627FFE940BF450B3036F968" blockId="43.[189,1399,1578,1713]" pageId="43" pageNumber="44">
Specimens somewhat resembling
<taxonomicName id="4C6E4D104627FFE94247450A3226F99D" authorityName="Herbert" authorityYear="2020" baseAuthorityName="Connolly" baseAuthorityYear="1933" box="[581,798,1578,1604]" class="Gastropoda" family="Charopidae" genus="Phialodonta" kingdom="Animalia" order="Stylommatophora" pageId="43" pageNumber="44" phylum="Mollusca" rank="species" species="introtuberculata" status="gen. et comb. nov.">
<emphasis id="B91AEA814627FFE94247450A3226F99D" box="[581,798,1578,1604]" italics="true" pageId="43" pageNumber="44">P. introtuberculata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22957FA4627FFE94321450A32CCF99C" box="[803,1012,1578,1605]" pageId="43" pageNumber="44" rank="species">gen. et comb. nov.</taxonomicNameLabel>
have been found in the Creighton area (Hlabeni Forest,
<geoCoordinate id="EE5A50544627FFE941B5456F3313F9B0" box="[439,555,1614,1641]" degrees="29.975" direction="south" orientation="latitude" pageId="43" pageNumber="44" precision="55" value="-29.975">29.975° S</geoCoordinate>
,
<geoCoordinate id="EE5A50544627FFE94234456E3395F9B0" box="[566,685,1615,1641]" degrees="29.742" direction="east" orientation="longitude" pageId="43" pageNumber="44" precision="55" value="29.742">29.742° E</geoCoordinate>
, NMSA V5219). However, although they have similar palatal dentition, their axial sculpture is noticeably finer, the umbilicus not as wide and they are milky-white in colour.
</paragraph>
</subSubSection>
</treatment>
</document>