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<document id="2014A60F466004F20359A215EE09DF84" ID-DOI="10.5281/zenodo.8074943" ID-ISSN="2754-9844" ID-Zenodo-Dep="8074943" ID-ZooBank="BCAD06E8-0AFE-46ED-B7FA-930983CD44C4" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1699388683108" checkinUser="felipe" docAuthor="Noyes, John Stuart" docDate="2023" docId="03BA87A7FFAFFFDBFE1FBA22A59FFA92" docLanguage="en" docName="TaxMonogNeotropHymenopt.2.1-921.pdf.imf" docOrigin="Taxonomic Monographs on Neotropical Hymenoptera (Oxford, England) 2 (11)" docSource="http://dx.doi.org/10.5281/zenodo.8074943" docStyle="DocumentStyle:DAA2ADC4E839520FEEF1E0A10B782A41.2:TaxMonogNeotropHymenopt.2022-.journal_article" docStyleId="DAA2ADC4E839520FEEF1E0A10B782A41" docStyleName="TaxMonogNeotropHymenopt.2022-.journal_article" docStyleVersion="2" docTitle="Coccidencyrtus thuryus Noyes 2023, sp.nov." docType="treatment" docVersion="3" lastPageNumber="123" masterDocId="FF83FFDFFFD5FFA7FFE2BC24A610FFC5" masterDocTitle="ENCYRTIDAE OF COSTA RICA (HYMENOPTERA: CHALCIDOIDEA), 4 Subfamily Encyrtinae: tribes Arrhenophagini, Habrolepidini, Cerapterocerini, Cheiloneurini, Trechnitini, Cercobelini, Polaszekiini, Protyndarichoidini, Gahaniellini and Syrphophagini (part), mainly primary parasitoids and hyperparasitoids of Coccoidea and Psylloidea (Hemiptera)" masterLastPageNumber="5239" masterPageNumber="5228" pageNumber="121" updateTime="1699893637884" updateUser="felipe" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="3DB2940A6B905A4CD879E30493BA3DE1">ENCYRTIDAE OF COSTA RICA (HYMENOPTERA: CHALCIDOIDEA), 4 Subfamily Encyrtinae: tribes Arrhenophagini, Habrolepidini, Cerapterocerini, Cheiloneurini, Trechnitini, Cercobelini, Polaszekiini, Protyndarichoidini, Gahaniellini and Syrphophagini (part), mainly primary parasitoids and hyperparasitoids of Coccoidea and Psylloidea (Hemiptera)</mods:title>
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<mods:namePart id="B2D30AE38A7B8D3B643CC346A83BC25A">Noyes, John Stuart</mods:namePart>
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<mods:title id="94E32FA28876A69C172432AB1BD9BE47">Taxonomic Monographs on Neotropical Hymenoptera</mods:title>
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<mods:date id="8B10B9490639538140F8CD8731F0F972">2023</mods:date>
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<treatment id="03BA87A7FFAFFFDBFE1FBA22A59FFA92" LSID="urn:lsid:plazi:treatment:03BA87A7FFAFFFDBFE1FBA22A59FFA92" httpUri="http://treatment.plazi.org/id/03BA87A7FFAFFFDBFE1FBA22A59FFA92" lastPageId="124" lastPageNumber="123" pageId="122" pageNumber="121">
<subSubSection id="C309653AFFAFFFDDFE1FBA22A555F9DA" box="[509,837,1542,1567]" pageId="122" pageNumber="121" type="nomenclature">
<paragraph id="8BAC36B1FFAFFFDDFE1FBA22A555F9DA" blockId="122.[509,837,1542,1593]" box="[509,837,1542,1567]" pageId="122" pageNumber="121">
<heading id="D0E481DDFFAFFFDDFE1FBA22A555F9DA" bold="true" box="[509,837,1542,1567]" centered="true" fontSize="10" level="2" pageId="122" pageNumber="121" reason="2">
<emphasis id="B967EAA3FFAFFFDDFE1FBA22A555F9DA" bold="true" box="[509,837,1542,1567]" pageId="122" pageNumber="121">
<taxonomicName id="4C134D32FFAFFFDDFE1FBA22A4E4F9DB" authority="Noyes, 2023" authorityName="Noyes" authorityYear="2023" box="[509,756,1542,1566]" class="Insecta" family="Encyrtidae" genus="Coccidencyrtus" kingdom="Animalia" order="Hymenoptera" pageId="122" pageNumber="121" phylum="Arthropoda" rank="species" species="thuryus" status="sp. nov.">
<emphasis id="B967EAA3FFAFFFDDFE1FBA22A4E4F9DB" bold="true" box="[509,756,1542,1566]" italics="true" pageId="122" pageNumber="121">Coccidencyrtus thuryus</emphasis>
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<taxonomicNameLabel id="A25457D8FFAFFFDDFD18BA23A555F9DA" box="[762,837,1543,1567]" pageId="122" pageNumber="121" rank="species">sp.nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C309653AFFAFFFDDFDFBBA05A538F9FC" box="[537,808,1569,1593]" pageId="122" pageNumber="121" type="description">
<paragraph id="8BAC36B1FFAFFFDDFDFBBA05A538F9FC" blockId="122.[509,837,1542,1593]" box="[537,808,1569,1593]" pageId="122" pageNumber="121">
(
<figureCitation id="13282A34FFAFFFDDFDC2BA05A4BBF9FC" box="[544,683,1569,1593]" captionStart="Figs 280-293" captionStartId="757.[181,226,1526,1550]" captionTargetBox="[182,1163,229,1507]" captionTargetId="figure-10@757.[181,1163,223,1508]" captionTargetPageId="757" captionText="Figs 280-293. Coccidencyrtus solivus, HT (280) hypopygium. Coccidencyrtus thuryus (281) head, facial view; (282) antenna; (283) apex of fore wing venation; (284) fore wing, smaller specimen; (285) fore wing; (286) mesosoma, dorsal view. Quetsira tinthrax (287) mesosoma, dorsal view; (288) antenna (a, outer aspect of clava; b, inner aspect of clava); (289) fore wing; (290) head, facial aspect; (291) apex of fore wing venation; (292) hypopygium; (293) ovipositor." pageId="122" pageNumber="121">Figs 281-286</figureCitation>
; Hab. E 47)
</paragraph>
</subSubSection>
<subSubSection id="C309653AFFAFFFDBFF57BA72A771FCC9" lastPageId="124" lastPageNumber="123" pageId="122" pageNumber="121" type="diagnosis">
<paragraph id="8BAC36B1FFAFFFDCFF57BA72A74CFDD9" blockId="122.[181,1165,1622,1700]" lastBlockId="123.[181,1165,222,540]" lastPageId="123" lastPageNumber="122" pageId="122" pageNumber="121">
DIAGNOSIS.
<emphasis id="B967EAA3FFAFFFDDFEAEBA72A788F9AB" box="[332,408,1622,1646]" italics="true" pageId="122" pageNumber="121">Female</emphasis>
(overall length about
<quantity id="4CEB9B54FFAFFFDDFD9CBA72A4E2F9AB" box="[638,754,1622,1646]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" metricValueMax="9.0" metricValueMin="6.0" pageId="122" pageNumber="121" unit="mm" value="0.75" valueMax="0.9" valueMin="0.6">0.6-0.9mm</quantity>
): scape largely dark brown; pedicel and funicle yellow, clava pale brown; all coxae white or very pale yellow, hind coxa may be partially brown; fore femur and tibia very pale yellow; mid and hind femora pale yellow with a dark brown subapical band; mid tibia pale yellow with a dark brown subbasal band; hind tibia pale yellow with a weak subbasal brown band; gaster dorsally dark brown, medially and ventrally pale orange; frontovertex about half head width; antenna (
<figureCitation id="13282A34FFAEFFDCFD38BD30A521FEE9" box="[730,817,276,300]" captionStart="Figs 280-293" captionStartId="757.[181,226,1526,1550]" captionTargetBox="[182,1163,229,1507]" captionTargetId="figure-10@757.[181,1163,223,1508]" captionTargetPageId="757" captionText="Figs 280-293. Coccidencyrtus solivus, HT (280) hypopygium. Coccidencyrtus thuryus (281) head, facial view; (282) antenna; (283) apex of fore wing venation; (284) fore wing, smaller specimen; (285) fore wing; (286) mesosoma, dorsal view. Quetsira tinthrax (287) mesosoma, dorsal view; (288) antenna (a, outer aspect of clava; b, inner aspect of clava); (289) fore wing; (290) head, facial aspect; (291) apex of fore wing venation; (292) hypopygium; (293) ovipositor." pageId="123" pageNumber="122">Fig. 282</figureCitation>
) with scape about 3-3.5X as long as broad, funicle with F1 narrower than pedicel, funicle with F1-F3 subquadrate, F4-6 larger and slightly longer than broad; linear sensilla present on F2, and F4-F6; clava very nearly as long as funicle, about 4X as long as broad; palp formula 3-2; scutellum (
<figureCitation id="13282A34FFAEFFDCFCB6BD40A5BCFEB9" box="[852,940,356,380]" captionStart="Figs 280-293" captionStartId="757.[181,226,1526,1550]" captionTargetBox="[182,1163,229,1507]" captionTargetId="figure-10@757.[181,1163,223,1508]" captionTargetPageId="757" captionText="Figs 280-293. Coccidencyrtus solivus, HT (280) hypopygium. Coccidencyrtus thuryus (281) head, facial view; (282) antenna; (283) apex of fore wing venation; (284) fore wing, smaller specimen; (285) fore wing; (286) mesosoma, dorsal view. Quetsira tinthrax (287) mesosoma, dorsal view; (288) antenna (a, outer aspect of clava; b, inner aspect of clava); (289) fore wing; (290) head, facial aspect; (291) apex of fore wing venation; (292) hypopygium; (293) ovipositor." pageId="123" pageNumber="122">Fig. 286</figureCitation>
) with 3 pairs of setae and longitudinally elongate polygonally reticulate to striate-reticulate sculpture, deeper than that on mesoscutum; mid basitarsus with 5-6 ventral pegs; mid tibial spur about as long as basitarsus; fore wing with stigmal vein slightly longer than postmarginal vein and about as long as marginal vein(
<figureCitation id="13282A34FFAEFFDCFF09BDEAA751FE22" box="[235,321,462,487]" captionStart="Figs 280-293" captionStartId="757.[181,226,1526,1550]" captionTargetBox="[182,1163,229,1507]" captionTargetId="figure-10@757.[181,1163,223,1508]" captionTargetPageId="757" captionText="Figs 280-293. Coccidencyrtus solivus, HT (280) hypopygium. Coccidencyrtus thuryus (281) head, facial view; (282) antenna; (283) apex of fore wing venation; (284) fore wing, smaller specimen; (285) fore wing; (286) mesosoma, dorsal view. Quetsira tinthrax (287) mesosoma, dorsal view; (288) antenna (a, outer aspect of clava; b, inner aspect of clava); (289) fore wing; (290) head, facial aspect; (291) apex of fore wing venation; (292) hypopygium; (293) ovipositor." pageId="123" pageNumber="122">Fig. 283</figureCitation>
); hypopygium reaching about 0.6X towards apex of gaster; ovipositor, about 3.5-4X as long as gonostylus and slightly exserted with exserted part a little shorter than mid tibial spur.
<emphasis id="B967EAA3FFAEFFDCFF57BE20A6F9FDD9" box="[181,233,516,540]" italics="true" pageId="123" pageNumber="122">Male</emphasis>
: unknown.
</paragraph>
<paragraph id="8BAC36B1FFAEFFDCFF57BE1DA293FD94" blockId="123.[181,1165,569,1687]" box="[181,1155,569,593]" pageId="123" pageNumber="122">
<materialsCitation id="3B7B3CECFFAEFFDCFF57BE1DA26EFD94" box="[181,1150,569,593]" collectionCode="CPD" pageId="123" pageNumber="122" specimenCount="1" typeStatus="holotype">
Female
<emphasis id="B967EAA3FFAEFFDCFEE4BE1DA71EFD94" bold="true" box="[262,270,569,593]" pageId="123" pageNumber="122">(</emphasis>
<typeStatus id="54A88813FFAEFFDCFEEDBE1DA778FD94" box="[271,360,569,593]" pageId="123" pageNumber="122" type="holotype">holotype</typeStatus>
<emphasis id="B967EAA3FFAEFFDCFE85BE1DA77FFD94" bold="true" box="[359,367,569,593]" pageId="123" pageNumber="122">)</emphasis>
: length, including ovipositor,
<quantity id="4CEB9B54FFAEFFDCFD40BE1DA4E9FD94" box="[674,761,569,593]" metricMagnitude="-4" metricUnit="m" metricValue="7.3" pageId="123" pageNumber="122" unit="mm" value="0.73">0.73mm</quantity>
; excluding ovipositor,
<quantity id="4CEB9B54FFAEFFDCFC3DBE1DA223FD94" box="[991,1075,569,593]" metricMagnitude="-4" metricUnit="m" metricValue="6.8" pageId="123" pageNumber="122" unit="mm" value="0.68">0.68mm</quantity>
(
<collectionCode id="ED02AE74FFAEFFDCFBA3BE1DA268FD94" box="[1089,1144,569,593]" pageId="123" pageNumber="122">CPD</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BAC36B1FFAEFFDCFF39BE70A47BFB44" blockId="123.[181,1165,569,1687]" pageId="123" pageNumber="122">
Head dark brown to black with a weak to moderate metallic lustre; occiput with a weak brassy sheen; frontovertex generally metallic dark green mixed coppery, area between posterior ocelli and occipital margin coppery; area above scrobes coppery; scrobal area, temple and gena with a weak brassy and coppery purple sheen; frontovertex with a few, inconspicuous, scattered, brown setae; gena and interantennal prominence with a few scattered, very inconspicuous, brown setae; maxillary palpus white; radicle dark brown; scape dark brown with extreme apex yellow; pedicel and funicle yellow; clava pale brown; thorax dark brown to black with a metallic lustre; pronotum with a weak coppery purple and brassy sheen; mesoscutum relatively bright metallic dark green mixed coppery near margins and in anterior half; tegula dark purple-brown with a weak brassy sheen; axilla with a weak coppery purple and brassy sheen; scutellum dorsally virtually matt, but with a very weak coppery purple sheen, extreme side and apex weakly shining and slightly brassy; mesopleuron with a moderate purple sheen; all coxae very pale yellow; fore femur and tibia very pale yellow, tarsus slightly dusky, pale orange; mid femur very pale yellow in proximal 0.5X, a broad, dark brown subapical band, extreme apex white, tibia white to very pale yellow with a narrow, subbasal dark, brown band, spur and tarsus very pale orange; hind femur with proximal 0.6X very pale yellow, extreme apex white and a pale brown, slightly diagonal, subapical band, tibia pale yellow with a diffuse, narrow, pale brown, subbasal band, tarsus slightly dusky, pale orange; wings mainly hyaline, but fore wing with a small, pale fuscous patch below the marginal and stigmal veins (
<figureCitation id="13282A34FFAEFFDCFE9BB810A7C4FB89" box="[377,468,1076,1100]" captionStart="Figs 280-293" captionStartId="757.[181,226,1526,1550]" captionTargetBox="[182,1163,229,1507]" captionTargetId="figure-10@757.[181,1163,223,1508]" captionTargetPageId="757" captionText="Figs 280-293. Coccidencyrtus solivus, HT (280) hypopygium. Coccidencyrtus thuryus (281) head, facial view; (282) antenna; (283) apex of fore wing venation; (284) fore wing, smaller specimen; (285) fore wing; (286) mesosoma, dorsal view. Quetsira tinthrax (287) mesosoma, dorsal view; (288) antenna (a, outer aspect of clava; b, inner aspect of clava); (289) fore wing; (290) head, facial aspect; (291) apex of fore wing venation; (292) hypopygium; (293) ovipositor." pageId="123" pageNumber="122">Fig. 283</figureCitation>
); propodeum dark brown with a weak purple sheen; gaster dorsally dark purple-brown on side, medially pale orange, median area a little dusky posteriorly, side and venter pale orange; gonostylus pale brown.
</paragraph>
<paragraph id="8BAC36B1FFAEFFDCFF39B8A0A5ADF9B9" blockId="123.[181,1165,569,1687]" pageId="123" pageNumber="122">
Head (
<figureCitation id="13282A34FFAEFFDCFEC3B8A0A769FB59" box="[289,377,1156,1180]" captionStart="Figs 280-293" captionStartId="757.[181,226,1526,1550]" captionTargetBox="[182,1163,229,1507]" captionTargetId="figure-10@757.[181,1163,223,1508]" captionTargetPageId="757" captionText="Figs 280-293. Coccidencyrtus solivus, HT (280) hypopygium. Coccidencyrtus thuryus (281) head, facial view; (282) antenna; (283) apex of fore wing venation; (284) fore wing, smaller specimen; (285) fore wing; (286) mesosoma, dorsal view. Quetsira tinthrax (287) mesosoma, dorsal view; (288) antenna (a, outer aspect of clava; b, inner aspect of clava); (289) fore wing; (290) head, facial aspect; (291) apex of fore wing venation; (292) hypopygium; (293) ovipositor." pageId="123" pageNumber="122">Fig. 281</figureCitation>
) about 2.3X as wide as frontovertex, in profile about 1.5X as high as deep, tangent to frontovertex forming an angle of about 100° with that of face; occipital margin more or less sharp, not carinate; no distinct oval depression adjacent to eye near occipital margin; ocelli forming an angle of about 85°; frontovertex clearly widening in front of anterior ocellus, with distinct polygonally reticulate sculpture of mesh size subequal to, or conspicuously larger than, eye facet, not shallower above top of scrobes; temple and gena with relatively shallow, longitudinally elongate, fine polygonally reticulate sculpture; top of scrobes with distinct, regular, polygonally reticulate sculpture; interantennal prominence and mouth margin with slightly shallower, polygonally reticulate sculpture; scrobes shallow, indistinctly margined, broadly ∩-shaped; interantennal prominence dorsally narrowly rounded; antenna as in
<figureCitation id="13282A34FFAEFFDCFC0EB950A256FA49" box="[1004,1094,1396,1420]" captionStart="Figs 280-293" captionStartId="757.[181,226,1526,1550]" captionTargetBox="[182,1163,229,1507]" captionTargetId="figure-10@757.[181,1163,223,1508]" captionTargetPageId="757" captionText="Figs 280-293. Coccidencyrtus solivus, HT (280) hypopygium. Coccidencyrtus thuryus (281) head, facial view; (282) antenna; (283) apex of fore wing venation; (284) fore wing, smaller specimen; (285) fore wing; (286) mesosoma, dorsal view. Quetsira tinthrax (287) mesosoma, dorsal view; (288) antenna (a, outer aspect of clava; b, inner aspect of clava); (289) fore wing; (290) head, facial aspect; (291) apex of fore wing venation; (292) hypopygium; (293) ovipositor." pageId="123" pageNumber="122">Fig. 282</figureCitation>
; scape very slightly broadened and flattened, about 3.1X as long as broad; funicle with F1-F3 relatively small, subquadrate and subequal, F4-F5 slightly larger, subequal and a little longer than broad, F6 larger and clearly longer than broad; clava moderately slender, about 4X as long as broad, about 1.5X as wide as F6, not downcurved ventrally, very nearly as long as funicle, sutures parallel, sensory area relatively small, at apex only giving it a rounded or weak obliquely truncate appearance; eye slightly overreaching occipital margin, clothed with dense, conspicuous setae, each a little longer than diameter of facet; malar sulcus present, but very shallow and indistinct; clypeal margin evenly very slightly concave. Relative measurements: HW 43.5,
<collectingRegion id="49D7F853FFAEFFDCFC13BA6EA206F9A4" box="[1009,1046,1610,1633]" country="Swaziland" name="Hhohho" pageId="123" pageNumber="122">HH</collectingRegion>
32, FV 19, POL 7.5, OOL 2.5, OCL 2, AOL 5.5, EL 25, EW 21, MS 12, SL 17, SW 5.5.
</paragraph>
<paragraph id="8BAC36B1FFAEFFDBFF39BA5BA799FE23" blockId="123.[181,1165,569,1687]" lastBlockId="124.[181,1165,222,727]" lastPageId="124" lastPageNumber="123" pageId="123" pageNumber="122">
Thorax (
<figureCitation id="13282A34FFAEFFDCFED6BA5BA79DF952" box="[308,397,1663,1687]" captionStart="Figs 280-293" captionStartId="757.[181,226,1526,1550]" captionTargetBox="[182,1163,229,1507]" captionTargetId="figure-10@757.[181,1163,223,1508]" captionTargetPageId="757" captionText="Figs 280-293. Coccidencyrtus solivus, HT (280) hypopygium. Coccidencyrtus thuryus (281) head, facial view; (282) antenna; (283) apex of fore wing venation; (284) fore wing, smaller specimen; (285) fore wing; (286) mesosoma, dorsal view. Quetsira tinthrax (287) mesosoma, dorsal view; (288) antenna (a, outer aspect of clava; b, inner aspect of clava); (289) fore wing; (290) head, facial aspect; (291) apex of fore wing venation; (292) hypopygium; (293) ovipositor." pageId="123" pageNumber="122">Fig. 286</figureCitation>
) with pronotum hardly exposed behind head, its posterior margin broadly concave, not angular medially; pronotum and mesoscutum with polygonally reticulate sculpture that is about as deep as that in ocellar area, and of similar mesh; axilla with similar sculpture to mesoscutum but clearly shallower; scutellum dorsally with very fine, slightly longitudinally elongate, polygonally reticulate sculpture that is a little deeper than sculpture on mesoscutum, extreme side and apex completely smooth and shiny; visible part of mesoscutum about 1.9X as broad as long; scutellum about 1.2X as broad as long, apex with a pair of slender, elongate setae; hind femur about 3.3X as long as broad; mid basitarsus with 5 ventral pegs; mid tibial spur about as long as basitarsus; fore wing with venation and setation as in
<figureCitation id="13282A34FFA9FFDBFCA5BDBDA5C6FE74" box="[839,982,409,433]" captionStart="Figs 280-293" captionStartId="757.[181,226,1526,1550]" captionTargetBox="[182,1163,229,1507]" captionTargetId="figure-10@757.[181,1163,223,1508]" captionTargetPageId="757" captionText="Figs 280-293. Coccidencyrtus solivus, HT (280) hypopygium. Coccidencyrtus thuryus (281) head, facial view; (282) antenna; (283) apex of fore wing venation; (284) fore wing, smaller specimen; (285) fore wing; (286) mesosoma, dorsal view. Quetsira tinthrax (287) mesosoma, dorsal view; (288) antenna (a, outer aspect of clava; b, inner aspect of clava); (289) fore wing; (290) head, facial aspect; (291) apex of fore wing venation; (292) hypopygium; (293) ovipositor." pageId="124" pageNumber="123">Figs 283, 285</figureCitation>
; propodeum (
<figureCitation id="13282A34FFA9FFDBFB81BDBDA6CCFE09" captionStart="Figs 280-293" captionStartId="757.[181,226,1526,1550]" captionTargetBox="[182,1163,229,1507]" captionTargetId="figure-10@757.[181,1163,223,1508]" captionTargetPageId="757" captionText="Figs 280-293. Coccidencyrtus solivus, HT (280) hypopygium. Coccidencyrtus thuryus (281) head, facial view; (282) antenna; (283) apex of fore wing venation; (284) fore wing, smaller specimen; (285) fore wing; (286) mesosoma, dorsal view. Quetsira tinthrax (287) mesosoma, dorsal view; (288) antenna (a, outer aspect of clava; b, inner aspect of clava); (289) fore wing; (290) head, facial aspect; (291) apex of fore wing venation; (292) hypopygium; (293) ovipositor." pageId="124" pageNumber="123">Fig. 286</figureCitation>
) medially about 0.05X as long as scutellum. Relative measurements: FWL 43, FWW 17.5; HWL 28, HWW 5.5.
</paragraph>
<paragraph id="8BAC36B1FFA9FFDBFF39BDCDA7CCFDF3" blockId="124.[181,1165,222,727]" pageId="124" pageNumber="123">Gaster with hypopygium reaching about 0.6X to apex; last tergite about 0.65X as long as mid tibia, with apex rounded; ovipositor exserted, the exserted part about 0.6X as long as mid tibial spur or about 0.2X mid tibia.</paragraph>
<paragraph id="8BAC36B1FFA9FFDBFF39BE1EA256FDA9" blockId="124.[181,1165,222,727]" pageId="124" pageNumber="123">
<typeStatus id="54A88813FFA9FFDBFF39BE1EA724FD97" box="[219,308,570,594]" pageId="124" pageNumber="123" type="paratype">Paratype</typeStatus>
. Mandible with two acute teeth and an obtuse upper tooth; palp formula 3-2; funicle with linear sensilla present on F2, F4-F6. Relative measurements: OL 99, GL 27 [MT 91].
</paragraph>
<paragraph id="8BAC36B1FFA9FFDBFF39BE4AA5AEFD12" blockId="124.[181,1165,222,727]" pageId="124" pageNumber="123">
Variation. Females vary in overall length from
<quantity id="4CEB9B54FFA9FFDBFD28BE4AA545FD43" box="[714,853,622,646]" metricMagnitude="-4" metricUnit="m" metricValue="7.1499999999999995" metricValueMax="8.5" metricValueMin="5.8" pageId="124" pageNumber="123" unit="mm" value="0.715" valueMax="0.85" valueMin="0.58">0.58-0.85mm</quantity>
, the terminal segments of the gaster may be completely brown dorsally, specimens from higher altitudes have the hind coxa partially brown externally, the fore wing varies between 2.28-2.46X (
<figureCitation id="13282A34FFA9FFDBFC62BE80A206FD79" box="[896,1046,676,700]" captionStart="Figs 280-293" captionStartId="757.[181,226,1526,1550]" captionTargetBox="[182,1163,229,1507]" captionTargetId="figure-10@757.[181,1163,223,1508]" captionTargetPageId="757" captionText="Figs 280-293. Coccidencyrtus solivus, HT (280) hypopygium. Coccidencyrtus thuryus (281) head, facial view; (282) antenna; (283) apex of fore wing venation; (284) fore wing, smaller specimen; (285) fore wing; (286) mesosoma, dorsal view. Quetsira tinthrax (287) mesosoma, dorsal view; (288) antenna (a, outer aspect of clava; b, inner aspect of clava); (289) fore wing; (290) head, facial aspect; (291) apex of fore wing venation; (292) hypopygium; (293) ovipositor." pageId="124" pageNumber="123">Figs. 284, 285</figureCitation>
) as long as broad and the ovipositor varies from about 3.5-4X as long as the gonostylus.
</paragraph>
<paragraph id="8BAC36B1FFA9FFDBFF57BED0A771FCC9" blockId="124.[181,353,756,780]" box="[181,353,756,780]" pageId="124" pageNumber="123">Male. Unknown.</paragraph>
</subSubSection>
<subSubSection id="C309653AFFA9FFDBFF57BF0DA7F9FC84" box="[181,489,809,833]" pageId="124" pageNumber="123" type="distribution">
<paragraph id="8BAC36B1FFA9FFDBFF57BF0DA7F9FC84" blockId="124.[181,489,809,833]" box="[181,489,809,833]" pageId="124" pageNumber="123">
DISTRIBUTION.
<collectingCountry id="F3047621FFA9FFDBFE91BF0DA7F5FC84" box="[371,485,809,833]" name="Costa Rica" pageId="124" pageNumber="123">Costa Rica</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C309653AFFA9FFDBFF57BF7BA76EFCB3" box="[181,382,862,887]" pageId="124" pageNumber="123" type="biology_ecology">
<paragraph id="8BAC36B1FFA9FFDBFF57BF7BA76EFCB3" blockId="124.[181,382,862,887]" box="[181,382,862,887]" pageId="124" pageNumber="123">HOSTS. Unknown.</paragraph>
</subSubSection>
<subSubSection id="C309653AFFA9FFDBFF57BFB2A728FAE5" pageId="124" pageNumber="123" type="materials_examined">
<paragraph id="8BAC36B1FFA9FFDBFF57BFB2A7BDFC6E" blockId="124.[181,1165,918,1313]" box="[181,429,918,939]" pageId="124" pageNumber="123">MATERIAL EXAMINED.</paragraph>
<paragraph id="8BAC36B1FFA9FFDBFF39BF95A728FAE5" blockId="124.[181,1165,918,1313]" pageId="124" pageNumber="123">
Type material.
<typeStatus id="54A88813FFA9FFDBFE89BF94A7AFFC03" box="[363,447,944,966]" pageId="124" pageNumber="123" type="holotype">Holotype</typeStatus>
E:
<emphasis id="B967EAA3FFA9FFDBFE08BF94A468FC03" bold="true" box="[490,632,944,966]" pageId="124" pageNumber="123">
<collectingCountry id="F3047621FFA9FFDBFE08BF94A468FC03" box="[490,632,944,966]" name="Costa Rica" pageId="124" pageNumber="123">COSTA RICA</collectingCountry>
</emphasis>
,
<emphasis id="B967EAA3FFA9FFDBFD65BF94A4E5FC00" bold="true" box="[647,757,944,965]" pageId="124" pageNumber="123">
<collectingRegion id="49D7F853FFA9FFDBFD65BF94A4E5FC00" box="[647,757,944,965]" country="Costa Rica" name="Puntarenas" pageId="124" pageNumber="123">Puntarenas</collectingRegion>
</emphasis>
, San Vito, Las Cruces, Wilson Bot. Gdns,
<geoCoordinate id="EE275076FFA9FFDBFF57BFEFA6E9FC24" box="[181,249,971,993]" degrees="8" direction="north" minutes="46" orientation="latitude" pageId="124" pageNumber="123" precision="925" value="8.766666">8°46N</geoCoordinate>
<geoCoordinate id="EE275076FFA9FFDBFF1DBFEFA742FC24" box="[255,338,971,993]" degrees="82" direction="west" minutes="57" orientation="longitude" pageId="124" pageNumber="123" precision="925" value="-82.95">82°57W</geoCoordinate>
,
<date id="FFAD1071FFA9FFDBFEBFBFEFA7DCFC24" box="[349,460,971,993]" pageId="124" pageNumber="123" value="2007-02-07" valueMax="2007-02-19" valueMin="2007-02-07">7-19.ii.2007</date>
(C. Hansson).
<typeStatus id="54A88813FFA9FFDBFDBABFEFA4A4FC25" box="[600,692,971,992]" pageId="124" pageNumber="123" type="paratype">Paratypes</typeStatus>
:
<emphasis id="B967EAA3FFA9FFDBFD5FBFEFA559FC24" bold="true" box="[701,841,971,993]" pageId="124" pageNumber="123">
<collectingCountry id="F3047621FFA9FFDBFD5FBFEFA559FC24" box="[701,841,971,993]" name="Costa Rica" pageId="124" pageNumber="123">COSTA RICA</collectingCountry>
</emphasis>
, 2E,
<emphasis id="B967EAA3FFA9FFDBFC9CBFEFA5FFFC24" bold="true" box="[894,1007,971,993]" pageId="124" pageNumber="123">
<collectingRegion id="49D7F853FFA9FFDBFC9CBFEFA5FFFC24" box="[894,1007,971,993]" country="Costa Rica" name="Guanacaste" pageId="124" pageNumber="123">Guanacaste</collectingRegion>
</emphasis>
, PN Santa
<taxonomicName id="4C134D32FFA9FFDBFBBBBFEFA75FFC3E" authority=", Bosque Humedo" authorityName="Bosque Humedo" class="Magnoliopsida" family="Rosaceae" genus="Rosa" kingdom="Plantae" order="Rosales" pageId="124" pageNumber="123" phylum="Tracheophyta" rank="genus">Rosa, Bosque Humedo</taxonomicName>
,
<geoCoordinate id="EE275076FFA9FFDBFEBABFC1A7B6FC3E" box="[344,422,997,1019]" degrees="10" direction="north" minutes="51" orientation="latitude" pageId="124" pageNumber="123" precision="925" value="10.85">10°51N</geoCoordinate>
<geoCoordinate id="EE275076FFA9FFDBFE4EBFC1A410FC3E" box="[428,512,997,1019]" degrees="85" direction="west" minutes="37" orientation="longitude" pageId="124" pageNumber="123" precision="925" value="-85.61667">85°37W</geoCoordinate>
.
<quantity id="4CEB9B54FFA9FFDBFDEBBFC2A42FFC3E" box="[521,575,998,1019]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="124" pageNumber="123" unit="m" value="300.0">300m</quantity>
,
<date id="FFAD1071FFA9FFDBFDA5BFC1A4B3FC3E" box="[583,675,997,1019]" pageId="124" pageNumber="123" value="2007-02-20">20.ii.2007</date>
(J.S. Noyes); 1E,
<collectingRegion id="49D7F853FFA9FFDBFCAEBFC2A5A7FC3E" box="[844,951,998,1019]" country="Costa Rica" name="Guanacaste" pageId="124" pageNumber="123">Guanacaste</collectingRegion>
,
<taxonomicName id="4C134D32FFA9FFDBFC22BFC2A266FC3E" authority="(ACG), 1100" authorityName=", Estac. Cacao" authorityYear="1100" baseAuthorityName="ACG" baseAuthorityYear="1100" box="[960,1142,998,1020]" class="Magnoliopsida" family="Ericaceae" genus="Vaccinium" kingdom="Plantae" order="Ericales" pageId="124" pageNumber="123" phylum="Tracheophyta" rank="species" species="cacao">
Cacao (ACG),
<quantity id="4CEB9B54FFA9FFDBFBAABFC2A266FC3E" box="[1096,1142,998,1019]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="124" pageNumber="123" unit="m" value="1100.0">1100</quantity>
</taxonomicName>
m, MT/YPT,
<date id="FFAD1071FFA9FFDBFEF0B824A787FBD3" box="[274,407,1024,1046]" pageId="124" pageNumber="123" value="1996-01-26" valueMax="1996-02-24" valueMin="1996-01-26">26.i-24.ii.1996</date>
(J.S. Noyes); 1E,
<emphasis id="B967EAA3FFA9FFDBFDDDB824A4BDFBD0" bold="true" box="[575,685,1024,1045]" pageId="124" pageNumber="123">
<collectingRegion id="49D7F853FFA9FFDBFDDDB824A4BDFBD0" box="[575,685,1024,1045]" country="Costa Rica" name="Puntarenas" pageId="124" pageNumber="123">Puntarenas</collectingRegion>
</emphasis>
, RB Carara,
<geoCoordinate id="EE275076FFA9FFDBFCC0B824A576FBD3" box="[802,870,1024,1046]" degrees="9" direction="north" minutes="46" orientation="latitude" pageId="124" pageNumber="123" precision="925" value="9.766666">9°46N</geoCoordinate>
<geoCoordinate id="EE275076FFA9FFDBFC88B824A5ADFBD3" box="[874,957,1024,1046]" degrees="84" direction="west" minutes="57" orientation="longitude" pageId="124" pageNumber="123" precision="925" value="-84.95">84°57W</geoCoordinate>
,
<quantity id="4CEB9B54FFA9FFDBFC24B824A5E1FBD0" box="[966,1009,1024,1045]" metricMagnitude="1" metricUnit="m" metricValue="4.0" pageId="124" pageNumber="123" unit="m" value="40.0">40m</quantity>
,
<date id="FFAD1071FFA9FFDBFC1BB824A273FBD3" box="[1017,1123,1024,1046]" pageId="124" pageNumber="123" value="2005-03-01" valueMax="2005-03-07" valueMin="2005-03-01">1-7.iii.2005</date>
((C. Hansson); 1E,
<collectingRegion id="49D7F853FFA9FFDBFEA6B83FA7BAFBF5" box="[324,426,1051,1072]" country="Costa Rica" name="Puntarenas" pageId="124" pageNumber="123">Puntarenas</collectingRegion>
, Buenos Aires, Est. Altamira, Send. los Gigantes, LS 572200 331700, #57880, vii-viii.2000 (
<taxonomicName id="4C134D32FFA9FFDBFED5B812A797FB8E" box="[311,391,1077,1099]" class="Insecta" family="Diaspididae" genus="Diaspis" kingdom="Animalia" order="Hemiptera" pageId="124" pageNumber="123" phylum="Arthropoda" rank="species" species="rubi">D. Rubí</taxonomicName>
); 1E,
<collectingRegion id="49D7F853FFA9FFDBFE24B812A43CFB8E" box="[454,556,1078,1099]" country="Costa Rica" name="Puntarenas" pageId="124" pageNumber="123">Puntarenas</collectingRegion>
, Est. Altamira, Send. los Gigantes, LS 331800 572100,
<quantity id="4CEB9B54FFA9FFDBFBA5B811A299FB8E" box="[1095,1161,1077,1099]" metricMagnitude="3" metricUnit="m" metricValue="1.46" pageId="124" pageNumber="123" unit="m" value="1460.0">1460m</quantity>
,
<date id="FFAD1071FFA9FFDBFF57B874A6E5FBA3" box="[181,245,1104,1126]" pageId="124" pageNumber="123" value="2002-02">ii.2002</date>
(C. Hansson,
<taxonomicName id="4C134D32FFA9FFDBFE9CB875A7DCFBA3" box="[382,460,1104,1126]" class="Insecta" family="Diaspididae" genus="Diaspis" kingdom="Animalia" order="Hemiptera" pageId="124" pageNumber="123" phylum="Arthropoda" rank="species" species="rubi">D. Rubí</taxonomicName>
); 2E,
<collectingRegion id="49D7F853FFA9FFDBFDEBB875A47FFBA3" box="[521,623,1105,1126]" country="Costa Rica" name="Puntarenas" pageId="124" pageNumber="123">Puntarenas</collectingRegion>
, San Vito. Las Cruces, Wilson Bot. Gdns,
<quantity id="4CEB9B54FFA9FFDBFBE7B874A256FBA3" box="[1029,1094,1104,1126]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="124" pageNumber="123" unit="m" value="1500.0">1500m</quantity>
,
<date id="FFAD1071FFA9FFDBFBB3B874A6EBFB44" pageId="124" pageNumber="123" value="1990-03-18" valueMax="1990-03-22" valueMin="1990-03-18">18-22. iii.1990</date>
(J.S. Noyes); 2E, same data as
<typeStatus id="54A88813FFA9FFDBFDC0B84FA464FB44" box="[546,628,1131,1153]" pageId="124" pageNumber="123" type="holotype">holotype</typeStatus>
; 1E,
<collectingRegion id="49D7F853FFA9FFDBFD45B84FA51DFB45" box="[679,781,1131,1152]" country="Costa Rica" name="Puntarenas" pageId="124" pageNumber="123">Puntarenas</collectingRegion>
, San Vito, Las Cruces, Wilson Bot. Gdns,
<geoCoordinate id="EE275076FFA9FFDBFF57B8A1A6E9FB5E" box="[181,249,1157,1179]" degrees="8" direction="north" minutes="46" orientation="latitude" pageId="124" pageNumber="123" precision="925" value="8.766666">8°46N</geoCoordinate>
<geoCoordinate id="EE275076FFA9FFDBFF1CB8A1A741FB5E" box="[254,337,1157,1179]" degrees="82" direction="west" minutes="57" orientation="longitude" pageId="124" pageNumber="123" precision="925" value="-82.95">82°57W</geoCoordinate>
,
<quantity id="4CEB9B54FFA9FFDBFEB8B8A1A7C0FB5E" box="[346,464,1157,1179]" metricMagnitude="3" metricUnit="m" metricValue="1.275" metricValueMax="1.35" metricValueMin="1.2" pageId="124" pageNumber="123" unit="m" value="1275.0" valueMax="1350.0" valueMin="1200.0">1200-1350m</quantity>
,
<date id="FFAD1071FFA9FFDBFE3BB8A1A425FB5E" box="[473,565,1157,1179]" pageId="124" pageNumber="123" value="2015-02-19">19.ii.2015</date>
(J.S. Noyes); 11E,
<collectingRegion id="49D7F853FFA9FFDBFD05B8A2A55DFB5E" box="[743,845,1158,1179]" country="Costa Rica" name="Puntarenas" pageId="124" pageNumber="123">Puntarenas</collectingRegion>
, San Vito, Est. Las Alturas,
<geoCoordinate id="EE275076FFA9FFDBFBABB8A1A29CFB5E" box="[1097,1164,1157,1179]" degrees="8" direction="north" minutes="57" orientation="latitude" pageId="124" pageNumber="123" precision="925" value="8.95">8°57N</geoCoordinate>
<geoCoordinate id="EE275076FFA9FFDBFF57B884A719FB73" box="[181,265,1184,1206]" degrees="82" direction="west" minutes="50" orientation="longitude" pageId="124" pageNumber="123" precision="925" value="-82.833336">82°50W</geoCoordinate>
,
<quantity id="4CEB9B54FFA9FFDBFEF7B884A746FB73" box="[277,342,1184,1206]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="124" pageNumber="123" unit="m" value="1500.0">1500m</quantity>
,
<date id="FFAD1071FFA9FFDBFE80B884A7AEFB73" box="[354,446,1184,1206]" pageId="124" pageNumber="123" value="2015-02-20">20.ii.2015</date>
(J.S. Noyes); 1E,
<collectingRegion id="49D7F853FFA9FFDBFD91B885A4C9FB73" box="[627,729,1185,1206]" country="Costa Rica" name="Puntarenas" pageId="124" pageNumber="123">Puntarenas</collectingRegion>
, Pen.
<taxonomicName id="4C134D32FFA9FFDBFCF6B884A529FB70" box="[788,825,1184,1205]" class="Magnoliopsida" family="Rubiaceae" genus="Osa" kingdom="Plantae" order="Gentianales" pageId="124" pageNumber="123" phylum="Tracheophyta" rank="genus">Osa</taxonomicName>
,
<quantity id="4CEB9B54FFA9FFDBFCA4B884A569FB73" box="[838,889,1184,1206]" metricMagnitude="4" metricUnit="m" metricValue="2.3" pageId="124" pageNumber="123" unit="km" value="23.0">23km</quantity>
N. Puerto Jiménez,
<quantity id="4CEB9B54FFA9FFDBFBDEB884A277FB70" box="[1084,1127,1184,1205]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="124" pageNumber="123" unit="m" value="10.0">10m</quantity>
, xi-xii.1992 (P. Hanson). 1E,
<collectingRegion id="49D7F853FFA9FFDBFE49B89FA401FB15" box="[427,529,1211,1232]" country="Costa Rica" name="Puntarenas" pageId="124" pageNumber="123">Puntarenas</collectingRegion>
, Golfito, Est. Agujas, LS 526550
<quantity id="4CEB9B54FFA9FFDBFCAFB89FA5C4FB15" box="[845,980,1211,1233]" metricMagnitude="8" metricUnit="m" metricValue="2.767503" pageId="124" pageNumber="123" unit="m" value="2.767503E8">276750, 300m</quantity>
, #57901,
<date id="FFAD1071FFA9FFDBFBD2B89FA29CFB14" box="[1072,1164,1211,1233]" pageId="124" pageNumber="123" value="2000-02-21">21.ii.2000</date>
(J. Azofeifa); 1E,
<collectingRegion id="49D7F853FFA9FFDBFE80B8F2A7D8FB2E" box="[354,456,1238,1259]" country="Costa Rica" name="Puntarenas" pageId="124" pageNumber="123">Puntarenas</collectingRegion>
, PN Corcovado, Los Patos,
<geoCoordinate id="EE275076FFA9FFDBFD30B8F2A505FB2E" box="[722,789,1238,1259]" degrees="8" direction="north" minutes="33" orientation="latitude" pageId="124" pageNumber="123" precision="925" value="8.55">8°33N</geoCoordinate>
<geoCoordinate id="EE275076FFA9FFDBFCFFB8F2A560FB2E" box="[797,880,1238,1259]" degrees="83" direction="west" minutes="30" orientation="longitude" pageId="124" pageNumber="123" precision="925" value="-83.5">83°30W</geoCoordinate>
,
<quantity id="4CEB9B54FFA9FFDBFC9EB8F2A5A3FB2E" box="[892,947,1238,1259]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="124" pageNumber="123" unit="m" value="200.0">200m</quantity>
,
<date id="FFAD1071FFA9FFDBFC5CB8F1A5EDFB2E" box="[958,1021,1237,1259]" pageId="124" pageNumber="123" value="2000-02">ii.2000</date>
(J.S. Noyes, J. Azofeifa); 5E,
<emphasis id="B967EAA3FFA9FFDBFEA2B8CBA790FAC0" bold="true" box="[320,384,1263,1285]" pageId="124" pageNumber="123">
<collectingRegion id="49D7F853FFA9FFDBFEA2B8CBA790FAC0" box="[320,384,1263,1285]" country="Costa Rica" name="Limon" pageId="124" pageNumber="123">Limón</collectingRegion>
</emphasis>
, PN Cahuita,
<quantity id="4CEB9B54FFA9FFDBFE1CB8D4A40DFAC3" box="[510,541,1264,1286]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="124" pageNumber="123" unit="m" value="5.0">5m</quantity>
,
<date id="FFAD1071FFA9FFDBFDC4B8D4A46DFAC3" box="[550,637,1264,1286]" pageId="124" pageNumber="123" value="1991-03-01">1.iii.1991</date>
(J.S. Noyes).
<typeStatus id="54A88813FFA9FFDBFD1EB8D4A540FAC3" box="[764,848,1264,1286]" pageId="124" pageNumber="123" type="holotype">Holotype</typeStatus>
in
<collectionCode id="ED02AE74FFA9FFDBFC8FB8D4A5D0FAC0" box="[877,960,1264,1285]" pageId="124" pageNumber="123">MZUCR</collectionCode>
,
<typeStatus id="54A88813FFA9FFDBFC2AB8D5A20FFAC3" box="[968,1055,1265,1286]" pageId="124" pageNumber="123" type="paratype">paratypes</typeStatus>
in
<collectionCode id="ED02AE74FFA9FFDBFBD9B8D4A29CFAC0" box="[1083,1164,1264,1285]" pageId="124" pageNumber="123">MZUCR</collectionCode>
and
<collectionCode id="ED02AE74FFA9FFDBFF3EB92FA725FAE5" box="[220,309,1291,1312]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="124" pageNumber="123" type="Museum">NHMUK</collectionCode>
.
</paragraph>
</subSubSection>
<subSubSection id="C309653AFFA9FFDBFF57B91BA59FFA92" box="[181,911,1342,1367]" pageId="124" pageNumber="123" type="discussion">
<paragraph id="8BAC36B1FFA9FFDBFF57B91BA59FFA92" blockId="124.[181,911,1342,1367]" box="[181,911,1342,1367]" pageId="124" pageNumber="123">
COMMENTS. See comments under
<taxonomicName id="4C134D32FFA9FFDBFDC9B91BA44EFA93" authorityName="Noyes" authorityYear="2023" box="[555,606,1343,1366]" class="Insecta" family="Encyrtidae" genus="Coccidencyrtus" kingdom="Animalia" order="Hymenoptera" pageId="124" pageNumber="123" phylum="Arthropoda" rank="species" species="aries">
<emphasis id="B967EAA3FFA9FFDBFDC9B91BA44EFA93" box="[555,606,1343,1366]" italics="true" pageId="124" pageNumber="123">aries</emphasis>
</taxonomicName>
(p. 91) and
<taxonomicName id="4C134D32FFA9FFDBFD3BB91AA522FA93" authorityName="Noyes" authorityYear="2023" box="[729,818,1342,1366]" class="Insecta" family="Encyrtidae" genus="Coccidencyrtus" kingdom="Animalia" order="Hymenoptera" pageId="124" pageNumber="123" phylum="Arthropoda" rank="species" species="harpales">
<emphasis id="B967EAA3FFA9FFDBFD3BB91AA522FA93" box="[729,818,1342,1366]" italics="true" pageId="124" pageNumber="123">harpales</emphasis>
</taxonomicName>
(p. 120).
</paragraph>
</subSubSection>
</treatment>
</document>