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<document id="A69403A8F2DEA1A0F83E73AC52F558E6" ID-CLB-Dataset="32506" ID-DOI="10.11646/zootaxa.4304.1.1" ID-GBIF-Dataset="3b9bd2c4-7dee-46a3-a36d-80077b88a78b" ID-ISSN="1175-5326" ID-Zenodo-Dep="842289" ID-ZooBank="3D557C94-0783-4C39-80C3-9C321DA94800" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1502687522877" checkinUser="plazi" docAuthor="Moore, Kirrily M., Alderslade, Philip &amp; Miller, Karen J." docDate="2017" docId="039B87ED3EF0FF0AFF4BE7187F0DDFA2" docLanguage="en" docName="zootaxa.4304.1.1.pdf" docOrigin="Zootaxa 4304 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Victorgorgia nyahae Moore &amp; Alderslade &amp; Miller 2017, sp. nov." docType="treatment" docUuid="401F301E-C5CD-43D0-83F4-F663CB47ECD2" docUuidSource="ZooBank" docVersion="10" lastPageNumber="187" masterDocId="FFA2FF953E41FFB0FFDCE4287D2BDE2A" masterDocTitle="A taxonomic revision of Anthothela (Octocorallia: Scleraxonia: Anthothelidae) and related genera, with the addition of new taxa, using morphological and molecular data" masterLastPageNumber="212" masterPageNumber="1" pageNumber="178" updateTime="1698466576984" updateUser="ExternalLinkService">
<mods:mods id="D3D21F12B5970E45F31568B3677168E4" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="A59D01D6A7E0DB65D9C335BA9A34F285">
<mods:title id="332F2178F4D4F04DF4F63A95F120F319">A taxonomic revision of Anthothela (Octocorallia: Scleraxonia: Anthothelidae) and related genera, with the addition of new taxa, using morphological and molecular data</mods:title>
</mods:titleInfo>
<mods:name id="D71DE00EC46CEAD0490EDAA4A6DD9945" type="personal">
<mods:role id="BBCEF72D2762E25AADE5533BE5E508EA">
<mods:roleTerm id="3451D1169CE8D822FFCCEEBEA39232AC">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="886FDC011FD0B39F435AC88BFF206D54">Moore, Kirrily M.</mods:namePart>
</mods:name>
<mods:name id="FB2893CC62A6E9AD845418AD82333AFA" type="personal">
<mods:role id="C70F7D92907E9DF030483170F2CE2937">
<mods:roleTerm id="083277FA1E7EA74E6DA5B44BC0D7A8D5">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0F8C924EF85461A7786A06CF7B900EBC">Alderslade, Philip</mods:namePart>
</mods:name>
<mods:name id="6821138F9838A0175DB7BB7425A0AD11" type="personal">
<mods:role id="EA4B7A2E691B645F87D4757BB09718C6">
<mods:roleTerm id="95D50414281B39586C8ACA5162FF3D10">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8FE7BE21D87C2F7C57C9C6106E75D37F">Miller, Karen J.</mods:namePart>
</mods:name>
<mods:typeOfResource id="8DEB3152439883EFBD040414854E34B6">text</mods:typeOfResource>
<mods:relatedItem id="2ADE832DECA2EB90B1CEBFCBD1CB3AF4" type="host">
<mods:titleInfo id="20C9462DD62254FB3482DD254AD91CF3">
<mods:title id="244246437581B58EAA0FAB5404E5BCC9">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="2C3A54B0B7221ED28E3E0FC6C746E3DB">
<mods:date id="447875C603BEEAD4A9F8DA7AE13449FD">2017</mods:date>
<mods:detail id="A585DA53CE18475A436D402BF363135A" type="volume">
<mods:number id="B0ECBDCB4E3DC904A41ED9E425AAA5C3">4304</mods:number>
</mods:detail>
<mods:detail id="9D09996EB5D8F8AE012A15040A545374" type="issue">
<mods:number id="0FD51C035757FEBC4A66B1B943005E74">1</mods:number>
</mods:detail>
<mods:extent id="A461FCE3EFEEA919A51BC97FD52D451F" unit="page">
<mods:start id="5AF7106782AA0A135F72DA35FEDD1D8A">1</mods:start>
<mods:end id="4008FB01F927D763F9E57DB845E702B0">212</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="2A6A7F8A0C9981F97C4FC9396EE13BBD">journal article</mods:classification>
<mods:identifier id="4D83AC90D03EB876AC5BCAA302BFD2D6" type="CLB-Dataset">32506</mods:identifier>
<mods:identifier id="80671BA99466A75CE3703DCE0C91C629" type="DOI">10.11646/zootaxa.4304.1.1</mods:identifier>
<mods:identifier id="870E0D1DBFA9D35532745E64C2F6DB34" type="GBIF-Dataset">3b9bd2c4-7dee-46a3-a36d-80077b88a78b</mods:identifier>
<mods:identifier id="5F18696AA9F5DB3039DFAE97E330618B" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="8DD199707B90C8659D0BA6239DF7DF84" type="Zenodo-Dep">842289</mods:identifier>
<mods:identifier id="B169B54F9034D6403FBF803B186FB895" type="ZooBank">3D557C94-0783-4C39-80C3-9C321DA94800</mods:identifier>
</mods:mods>
<treatment id="039B87ED3EF0FF0AFF4BE7187F0DDFA2" ID-DOI="http://doi.org/10.5281/zenodo.6015397" ID-GBIF-Taxon="132631029" ID-Zenodo-Dep="6015397" LSID="urn:lsid:zoobank.org:act:401F301E-C5CD-43D0-83F4-F663CB47ECD2" httpUri="http://treatment.plazi.org/id/039B87ED3EF0FF0AFF4BE7187F0DDFA2" lastPageId="186" lastPageNumber="187" pageId="177" pageNumber="178">
<subSubSection id="C32865703EF0FF01FF4BE7187C61DDA5" pageId="177" pageNumber="178" type="nomenclature">
<paragraph id="8B8D36FB3EF0FF01FF4BE7187E7CDD60" blockId="177.[151,902,816,911]" box="[151,855,816,842]" pageId="177" pageNumber="178">
<heading id="D0C581973EF0FF01FF4BE7187E7CDD60" bold="true" box="[151,855,816,842]" fontSize="11" level="1" pageId="177" pageNumber="178" reason="1">
<emphasis id="B946EAE93EF0FF01FF4BE7187E7CDD60" bold="true" box="[151,855,816,842]" pageId="177" pageNumber="178">
<taxonomicName id="4C324D783EF0FF01FF4BE7187CA5DD60" authority="Moore &amp; Alderslade &amp; Miller, 2017" authorityName="Moore &amp; Alderslade &amp; Miller" authorityYear="2017" box="[151,398,816,842]" class="Anthozoa" family="Anthothelidae" genus="Victorgorgia" kingdom="Animalia" order="Alcyonacea" pageId="177" pageNumber="178" phylum="Cnidaria" rank="species" species="nyahae" status="sp. nov.">
<emphasis id="B946EAE93EF0FF01FF4BE7187CA5DD60" bold="true" box="[151,398,816,842]" italics="true" pageId="177" pageNumber="178">Victorgorgia nyahae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27557923EF0FF01FE49E7187CD9DD60" box="[405,498,816,842]" pageId="177" pageNumber="178" rank="species">sp. nov.</taxonomicNameLabel>
Moore, Alderslade &amp; Miller
</emphasis>
</heading>
</paragraph>
<paragraph id="8B8D36FB3EF0FF01FF4BE77B7C61DDA5" blockId="177.[151,902,816,911]" pageId="177" pageNumber="178">
http://zoobank.org/401F301E-C5CD-43D0-83F4-F663CB47ECD2 (
<figureCitation id="13092A7E3EF0FF01FF43E75E7C69DDA5" box="[159,322,886,911]" captionStart-0="FIGURE 139" captionStart-1="FIGURE 140" captionStart-2="FIGURE 141" captionStart-3="FIGURE 142" captionStart-4="FIGURE 143" captionStart-5="FIGURE 144" captionStart-6="FIGURE 145" captionStart-7="FIGURE 146" captionStartId-0="178.[151,250,1908,1930]" captionStartId-1="179.[151,250,1550,1572]" captionStartId-2="180.[151,250,1992,2014]" captionStartId-3="181.[151,250,1518,1540]" captionStartId-4="182.[151,250,1080,1102]" captionStartId-5="183.[151,250,2010,2032]" captionStartId-6="184.[151,250,1759,1781]" captionStartId-7="185.[151,250,1976,1998]" captionTargetBox-0="[151,1435,193,1880]" captionTargetBox-1="[170,1432,193,1517]" captionTargetBox-2="[165,1428,193,1956]" captionTargetBox-3="[165,1423,193,1486]" captionTargetBox-4="[151,1433,445,1042]" captionTargetBox-5="[153,1435,193,1977]" captionTargetBox-6="[162,1434,193,1721]" captionTargetBox-7="[182,1398,193,1939]" captionTargetId-0="figure@178.[151,1435,193,1886]" captionTargetId-1="figure@179.[151,1435,193,1529]" captionTargetId-2="figure@180.[165,1428,193,1970]" captionTargetId-3="figure@181.[165,1428,193,1497]" captionTargetId-4="figure@182.[151,1435,445,1059]" captionTargetId-5="figure@183.[151,1435,193,1989]" captionTargetId-6="figure@184.[151,1435,193,1738]" captionTargetId-7="figure@185.[173,1415,193,1950]" captionTargetPageId-0="178" captionTargetPageId-1="179" captionTargetPageId-2="180" captionTargetPageId-3="181" captionTargetPageId-4="182" captionTargetPageId-5="183" captionTargetPageId-6="184" captionTargetPageId-7="185" captionText-0="FIGURE 139. Ƒictorgorgia nyahae n. sp., holotype: A. Colony; B – C. Branch tip; D. Polyp. (A – B. Courtesy of Karen Gowlett- Holmes, CSIRO O &amp; A)." captionText-1="FIGURE 140. Ƒictorgorgia nyahae n. sp., holotype: A. Polyp head; B. Cross-section of medulla; C. Decalcified cross-section of medulla." captionText-2="FIGURE 141. Ƒictorgorgia nyahae n. sp., holotype, sclerites: Point (a. spear tips)." captionText-3="FIGURE 142. Ƒictorgorgia nyahae n. sp., holotype: A. Pinnule, tentacular and point sclerites in situ; B. Tentacle rachis sclerites." captionText-4="FIGURE 143. Ƒictorgorgia nyahae n. sp., holotype, sclerites: Tentacular and pinnule (a. flat rods with jagged edges)." captionText-5="FIGURE 144. Ƒictorgorgia nyahae n. sp., holotype, sclerites: Calyx (a. thorny tip)." captionText-6="FIGURE 145. Ƒictorgorgia nyahae n. sp., holotype, sclerites: Cortex." captionText-7="FIGURE 146. Ƒictorgorgia nyahae n. sp., holotype, sclerites: Medulla." httpUri-0="https://zenodo.org/record/842567/files/figure.png" httpUri-1="https://zenodo.org/record/842569/files/figure.png" httpUri-2="https://zenodo.org/record/842571/files/figure.png" httpUri-3="https://zenodo.org/record/842573/files/figure.png" httpUri-4="https://zenodo.org/record/842575/files/figure.png" httpUri-5="https://zenodo.org/record/842577/files/figure.png" httpUri-6="https://zenodo.org/record/842579/files/figure.png" httpUri-7="https://zenodo.org/record/842581/files/figure.png" pageId="177" pageNumber="178">Figs. 139–146</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C32865703EF0FF01FF4BE79E7DFDDAAF" pageId="177" pageNumber="178" type="materials_examined">
<paragraph id="8B8D36FB3EF0FF01FF4BE79E7EE1DA32" blockId="177.[151,1437,950,2037]" pageId="177" pageNumber="178">
<materialsCitation id="3B5A3CA63EF0FF01FF4BE79E7C9CDA33" ID-GBIF-Occurrence="1586203850" collectionCode="TMAG, CSIRO" collectorName="RV Southern Surveyor" country="Australia" latitude="-43.934" location="Material" municipality="Southern Surveyor" pageId="177" pageNumber="178" specimenCount="1" stateProvince="Tasman" typeStatus="holotype">
<emphasis id="B946EAE93EF0FF01FF4BE79E7FFFDDE5" bold="true" box="[151,724,950,975]" pageId="177" pageNumber="178">
<location id="8EED60203EF0FF01FF4BE79E7DD4DDE5" LSID="urn:lsid:plazi:treatment:039B87ED3EF0FF0AFF4BE7187F0DDFA2:8EED60203EF0FF01FF4BE79E7DD4DDE5" box="[151,255,950,975]" country="Australia" latitude="-43.934" municipality="Southern Surveyor" name="Material" pageId="177" pageNumber="178" stateProvince="Tasman">Material</location>
examined.
<typeStatus id="548988593EF0FF01FE4AE79E7F2CDDE5" box="[406,519,950,975]" pageId="177" pageNumber="178" type="holotype">Holotype</typeStatus>
:
<collectionCode id="ED23AE3E3EF0FF01FDC7E79E7F59DDE5" box="[539,626,950,975]" country="Australia" httpUri="http://grbio.org/cool/88vw-h26m" name="Tasmanian Museum and Art Gallery" pageId="177" pageNumber="178">TMAG</collectionCode>
K3988,
</emphasis>
Cascade Seamount, Huon Commonwealth Marine Reserve (CMR), SW
<collectingRegion id="49F6F8193EF0FF01FEEEE7F37CA2DDD8" box="[306,393,987,1010]" country="New Zealand" name="Tasman" pageId="177" pageNumber="178">Tasman</collectingRegion>
Sea,
<collectingCountry id="F325766B3EF0FF01FE0DE7F27F10DDD9" box="[465,571,986,1011]" name="Australia" pageId="177" pageNumber="178">Australia</collectingCountry>
,
<collectionCode id="ED23AE3E3EF0FF01FD97E7F27FB5DDD8" box="[587,670,986,1010]" country="Australia" httpUri="http://grbio.org/cool/0k2v-y6wz" name="Australian National Fish Collection" pageId="177" pageNumber="178">CSIRO</collectionCode>
<collectorName id="26C7532D3EF0FF01FD76E7F37E9ADDD8" box="[682,945,986,1011]" pageId="177" pageNumber="178">
RV
<collectingMunicipality id="6BE9AC813EF0FF01FD07E7F27E9ADDD8" box="[731,945,986,1011]" pageId="177" pageNumber="178">
<emphasis id="B946EAE93EF0FF01FD07E7F27E9ADDD8" box="[731,945,986,1011]" italics="true" pageId="177" pageNumber="178">Southern Surveyor</emphasis>
</collectingMunicipality>
</collectorName>
, stn. 75, sample 32, (SS
<date id="FF8C103B3EF0FF01FB0AE7F27805DDD9" box="[1238,1326,986,1011]" pageId="177" pageNumber="178" value="02-07-20">200702</date>
/075-032) 43.92–
<geoCoordinate id="EE06503C3EF0FF01FF34E0287C65DA32" box="[232,334,1024,1048]" direction="south" orientation="latitude" pageId="177" pageNumber="178" precision="55" value="-43.934">43.934°S</geoCoordinate>
, 150.463
</materialsCitation>
–150.479°E, depth
<quantity id="4CCA9B1E3EF0FF01FD51E0287E25DA32" box="[653,782,1024,1049]" metricMagnitude="2" metricUnit="m" metricValue="6.25" metricValueMax="6.6" metricValueMin="5.9" pageId="177" pageNumber="178" unit="m" value="625.0" valueMax="660.0" valueMin="590.0">590–660 m</quantity>
,
<date id="FF8C103B3EF0FF01FCC6E0287EECDA32" box="[794,967,1020,1049]" pageId="177" pageNumber="178" value="2007-04-10">10th April 2007</date>
.
</paragraph>
<paragraph id="8B8D36FB3EF0FF01FF1BE00B7DFDDAAF" blockId="177.[151,1437,950,2037]" pageId="177" pageNumber="178">
<materialsCitation id="3B5A3CA63EF0FF01FF1BE00B7DFDDAAF" ID-GBIF-Occurrence="1586203847" collectingDate="2007-04-10" collectionCode="TMAG, CSIRO" collectorName="RV Southern Surveyor" country="Australia" elevation="625" latitude="-43.915" location="Other" longLatPrecision="404" longitude="150.46" municipality="Southern Surveyor" pageId="177" pageNumber="178" specimenCount="1" stateProvince="Tasman">
<emphasis id="B946EAE93EF0FF01FF1BE00B7F6CDA16" bold="true" box="[199,583,1059,1084]" pageId="177" pageNumber="178">
<location id="8EED60203EF0FF01FF1BE00B7C24DA16" LSID="urn:lsid:plazi:treatment:039B87ED3EF0FF0AFF4BE7187F0DDFA2:8EED60203EF0FF01FF1BE00B7C24DA16" box="[199,271,1059,1084]" country="Australia" latitude="-43.915" longLatPrecision="404" longitude="150.46" municipality="Southern Surveyor" name="Other" pageId="177" pageNumber="178" stateProvince="Tasman">Other</location>
material:
<collectionCode id="ED23AE3E3EF0FF01FE4DE00B7CC3DA16" box="[401,488,1059,1084]" country="Australia" httpUri="http://grbio.org/cool/88vw-h26m" name="Tasmanian Museum and Art Gallery" pageId="177" pageNumber="178">TMAG</collectionCode>
K3989,
</emphasis>
Cascade Seamount, Huon CMR, SW
<collectingRegion id="49F6F8193EF0FF01FBD4E00C7974DA11" box="[1032,1119,1060,1083]" country="New Zealand" name="Tasman" pageId="177" pageNumber="178">Tasman</collectingRegion>
Sea,
<collectingCountry id="F325766B3EF0FF01FB7FE00B7827DA16" box="[1187,1292,1059,1084]" name="Australia" pageId="177" pageNumber="178">Australia</collectingCountry>
,
<collectionCode id="ED23AE3E3EF0FF01FAC7E00C7846DA16" box="[1307,1389,1060,1084]" country="Australia" httpUri="http://grbio.org/cool/0k2v-y6wz" name="Australian National Fish Collection" pageId="177" pageNumber="178">CSIRO</collectionCode>
<collectorName id="26C7532D3EF0FF01FAA4E00C7C42DA4B" pageId="177" pageNumber="178">
RV
<collectingMunicipality id="6BE9AC813EF0FF01FF4BE0617C42DA4B" box="[151,361,1097,1122]" pageId="177" pageNumber="178">
<emphasis id="B946EAE93EF0FF01FF4BE0617C42DA4B" box="[151,361,1097,1122]" italics="true" pageId="177" pageNumber="178">Southern Surveyor</emphasis>
</collectingMunicipality>
</collectorName>
, stn. 77, sample 0 0 3, (SS
<date id="FF8C103B3EF0FF01FD53E0627FC2DA48" box="[655,745,1098,1122]" pageId="177" pageNumber="178" value="02-07-20">200702</date>
/077-003),
<geoCoordinate id="EE06503C3EF0FF01FCB9E0617EE4DA48" box="[869,975,1097,1122]" direction="south" orientation="latitude" pageId="177" pageNumber="178" precision="55" value="-43.915">43.915°S</geoCoordinate>
,
<geoCoordinate id="EE06503C3EF0FF01FC01E0617963DA48" box="[989,1096,1097,1122]" direction="east" orientation="longitude" pageId="177" pageNumber="178" precision="555" value="150.46">150.46°E</geoCoordinate>
, depth
<elevation id="001FD1C83EF0FF01FB42E061780ADA4B" box="[1182,1313,1097,1122]" metricMagnitude="2" metricUnit="m" metricValue="6.25" metricValueMax="6.6" metricValueMin="5.9" pageId="177" pageNumber="178" unit="m" value="625.0" valueMax="660.0" valueMin="590.0">
<quantity id="4CCA9B1E3EF0FF01FB42E061780ADA4B" box="[1182,1313,1097,1122]" metricMagnitude="2" metricUnit="m" metricValue="6.25" metricValueMax="6.6" metricValueMin="5.9" pageId="177" pageNumber="178" unit="m" value="625.0" valueMax="660.0" valueMin="590.0">590–660 m</quantity>
</elevation>
,
<collectingDate id="EFC8E9D33EF0FF01FAF3E0627DF8DAAF" pageId="177" pageNumber="178" value="2007-04-10">
<date id="FF8C103B3EF0FF01FAF3E0627DF8DAAF" pageId="177" pageNumber="178" value="2007-04-10">10th April 2007</date>
</collectingDate>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C32865703EF0FF08FF1BE0A77CB3D9C0" lastPageId="184" lastPageNumber="185" pageId="177" pageNumber="178" type="description">
<paragraph id="8B8D36FB3EF0FF01FF1BE0A77C70DA82" blockId="177.[151,1437,950,2037]" box="[199,347,1167,1192]" pageId="177" pageNumber="178">
<emphasis id="B946EAE93EF0FF01FF1BE0A77C70DA82" bold="true" box="[199,347,1167,1192]" pageId="177" pageNumber="178">Description:</emphasis>
</paragraph>
<paragraph id="8B8D36FB3EF0FF01FF1BE09B7E4BDBE8" blockId="177.[151,1437,950,2037]" pageId="177" pageNumber="178">
<emphasis id="B946EAE93EF0FF01FF1BE09B7C43DAE6" bold="true" box="[199,360,1203,1228]" pageId="177" pageNumber="178">Colony form:</emphasis>
The
<typeStatus id="548988593EF0FF01FE7EE09B7F2FDAE6" box="[418,516,1203,1228]" pageId="177" pageNumber="178" type="holotype">holotype</typeStatus>
is a complete colony, now broken into three fragments (
<figureCitation id="13092A7E3EF0FF01FBA0E09B79F6DAE1" box="[1148,1245,1203,1228]" captionStart="FIGURE 139" captionStartId="178.[151,250,1908,1930]" captionTargetBox="[151,1435,193,1880]" captionTargetId="figure@178.[151,1435,193,1886]" captionTargetPageId="178" captionText="FIGURE 139. Ƒictorgorgia nyahae n. sp., holotype: A. Colony; B – C. Branch tip; D. Polyp. (A – B. Courtesy of Karen Gowlett- Holmes, CSIRO O &amp; A)." httpUri="https://zenodo.org/record/842567/files/figure.png" pageId="177" pageNumber="178">Fig. 139</figureCitation>
A). When whole, the colony had three main branches emanating from a small holdfast, encrusting a piece of coral rubble. The largest of these branches is approximately
<quantity id="4CCA9B1E3EF0FF01FDF8E0D27F58DB3B" box="[548,627,1274,1298]" metricMagnitude="-2" metricUnit="m" metricValue="2.9" pageId="177" pageNumber="178" unit="mm" value="29.0">29 mm</quantity>
in length with a diameter ranging between
<quantity id="4CCA9B1E3EF0FF01FB87E0D279CFDB3B" box="[1115,1252,1274,1298]" metricMagnitude="-3" metricUnit="m" metricValue="2.35" metricValueMax="2.9" metricValueMin="1.8" pageId="177" pageNumber="178" unit="mm" value="2.35" valueMax="2.9" valueMin="1.8">1.8–2.9 mm</quantity>
, and has a small group of polyps branching off close to the base. The other colony fragments are approximately
<quantity id="4CCA9B1E3EF0FF01FB3AE134784ADB1E" box="[1254,1377,1308,1333]" metricMagnitude="-2" metricUnit="m" metricValue="2.7" metricValueMax="2.9" metricValueMin="2.5" pageId="177" pageNumber="178" unit="mm" value="27.0" valueMax="29.0" valueMin="25.0">25–29 mm</quantity>
long with slightly narrower diameters (
<quantity id="4CCA9B1E3EF0FF01FDC7E1687F8CDB72" box="[539,679,1344,1368]" metricMagnitude="-3" metricUnit="m" metricValue="1.95" metricValueMax="2.1" metricValueMin="1.8" pageId="177" pageNumber="178" unit="mm" value="1.95" valueMax="2.1" valueMin="1.8">1.8–2.1 mm</quantity>
) and no secondary branching. The branches are occasionally bent and twisted, and often not circular in cross-section being more likely to be irregularly narrowed and distorted. No anastomoses are present. The colony holdfast is an encrusting membrane with a few scattered polyps. The colony is in good condition—preservation was directly into 70% alcohol.
</paragraph>
<paragraph id="8B8D36FB3EF0FF01FF1BE1E47E40D865" blockId="177.[151,1437,950,2037]" pageId="177" pageNumber="178">
Calyces and polyps are distributed along and all around the branches. They are particularly crowded towards the branch tip and so form terminal polyp bunches (
<figureCitation id="13092A7E3EF0FF01FD29E1C77E73D822" box="[757,856,1519,1544]" captionStart="FIGURE 139" captionStartId="178.[151,250,1908,1930]" captionTargetBox="[151,1435,193,1880]" captionTargetId="figure@178.[151,1435,193,1886]" captionTargetPageId="178" captionText="FIGURE 139. Ƒictorgorgia nyahae n. sp., holotype: A. Colony; B – C. Branch tip; D. Polyp. (A – B. Courtesy of Karen Gowlett- Holmes, CSIRO O &amp; A)." httpUri="https://zenodo.org/record/842567/files/figure.png" pageId="177" pageNumber="178">Fig. 139</figureCitation>
B, C). All calyces and polyps project substantially from the colony, mostly at right angles to the branches, with large, prominent calyces and rounded polyp heads. There is little space between calyces and only a few are isolated.
</paragraph>
<paragraph id="8B8D36FB3EF0FF01FF1BE2717DDAD892" blockId="177.[151,1437,950,2037]" pageId="177" pageNumber="178">
<emphasis id="B946EAE93EF0FF01FF1BE2717C0FD858" bold="true" box="[199,292,1625,1650]" pageId="177" pageNumber="178">Colour:</emphasis>
The photograph taken soon after collection shows the polyps and the distal half of the calyces are lilac, the retracted tentacles are purple and the rest of the colony is light cream. The colony is now light beige in alcohol.
</paragraph>
<paragraph id="8B8D36FB3EF0FF01FF1BE2EB7974D9DF" blockId="177.[151,1437,950,2037]" pageId="177" pageNumber="178">
<emphasis id="B946EAE93EF0FF01FF1BE2EB7C9DD8F6" bold="true" box="[199,438,1731,1756]" pageId="177" pageNumber="178">Calyces and polyps:</emphasis>
The straight-sided calyces are large relative to the branch diameter, and range from
<quantity id="4CCA9B1E3EF0FF01FAB7E2EB7DC5D8D4" metricMagnitude="-3" metricUnit="m" metricValue="3.15" metricValueMax="3.9" metricValueMin="2.4" pageId="177" pageNumber="178" unit="mm" value="3.15" valueMax="3.9" valueMin="2.4">2.4– 3.9 mm</quantity>
tall and
<quantity id="4CCA9B1E3EF0FF01FE8DE2CE7CF2D8D4" box="[337,473,1766,1791]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="2.6" metricValueMin="1.4" pageId="177" pageNumber="178" unit="mm" value="2.0" valueMax="2.6" valueMin="1.4">1.4–2.6 mm</quantity>
wide. There are no polyps with the head fully retracted into the calyx although some are slightly retracted such that the polyp head is sitting on the lip of the calyx. Occasionally a partially retracted polyp head causes the calyx lip to flare out while the polyp head flattens (
<figureCitation id="13092A7E3EF0FF01FC0CE3047905D96F" box="[976,1070,1836,1861]" captionStart="FIGURE 139" captionStartId="178.[151,250,1908,1930]" captionTargetBox="[151,1435,193,1880]" captionTargetId="figure@178.[151,1435,193,1886]" captionTargetPageId="178" captionText="FIGURE 139. Ƒictorgorgia nyahae n. sp., holotype: A. Colony; B – C. Branch tip; D. Polyp. (A – B. Courtesy of Karen Gowlett- Holmes, CSIRO O &amp; A)." httpUri="https://zenodo.org/record/842567/files/figure.png" pageId="177" pageNumber="178">Fig. 139</figureCitation>
Ca) making the whole polyp and calyx quite short and squat, however the majority of the polyps are exsert. Exsert polyps extended between
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above the lip of the calyx, and the polyp heads have diameters of approximately
<quantity id="4CCA9B1E3EF0FF01FB92E35B79E5D9A1" box="[1102,1230,1907,1932]" metricMagnitude="-3" metricUnit="m" metricValue="2.375" metricValueMax="2.75" metricValueMin="2.0" pageId="177" pageNumber="178" unit="mm" value="2.375" valueMax="2.75" valueMin="2.0">2–2.75 mm</quantity>
(
<figureCitation id="13092A7E3EF0FF01FB00E35B7816D9A1" box="[1244,1341,1907,1932]" captionStart="FIGURE 139" captionStartId="178.[151,250,1908,1930]" captionTargetBox="[151,1435,193,1880]" captionTargetId="figure@178.[151,1435,193,1886]" captionTargetPageId="178" captionText="FIGURE 139. Ƒictorgorgia nyahae n. sp., holotype: A. Colony; B – C. Branch tip; D. Polyp. (A – B. Courtesy of Karen Gowlett- Holmes, CSIRO O &amp; A)." httpUri="https://zenodo.org/record/842567/files/figure.png" pageId="177" pageNumber="178">Fig. 139</figureCitation>
D). Both polyps and calyces are covered in a dense layer of crowded sclerites. Most polyps have the tentacles tightly folded over the polyp mouth so the polyp heads are rounded mounds with eight segments (
<figureCitation id="13092A7E3EF0FF01FB49E39179D7D9F8" box="[1173,1276,1977,2002]" captionStart="FIGURE 140" captionStartId="179.[151,250,1550,1572]" captionTargetBox="[170,1432,193,1517]" captionTargetId="figure@179.[151,1435,193,1529]" captionTargetPageId="179" captionText="FIGURE 140. Ƒictorgorgia nyahae n. sp., holotype: A. Polyp head; B. Cross-section of medulla; C. Decalcified cross-section of medulla." httpUri="https://zenodo.org/record/842569/files/figure.png" pageId="177" pageNumber="178">Fig. 140</figureCitation>
A). There are approximately 8–10 pinnules arranged in a single row along each side of the tentacles.
</paragraph>
<caption id="DF4D66733EF3FF02FF4BE35C7CA0D983" httpUri="https://zenodo.org/record/842567/files/figure.png" pageId="178" pageNumber="179" targetBox="[151,1435,193,1880]" targetPageId="178">
<paragraph id="8B8D36FB3EF3FF02FF4BE35C7CA0D983" blockId="178.[151,1436,1908,1961]" pageId="178" pageNumber="179">
<emphasis id="B946EAE93EF3FF02FF4BE35C7C07D9A0" bold="true" box="[151,300,1908,1930]" pageId="178" pageNumber="179">FIGURE 139.</emphasis>
<taxonomicName id="4C324D783EF3FF02FEEDE35D7CD4D9A0" authority="Moore &amp; Alderslade &amp; Miller, 2017" authorityName="Moore &amp; Alderslade &amp; Miller" authorityYear="2017" box="[305,511,1908,1930]" class="Anthozoa" family="Anthothelidae" genus="Fictorgorgia" kingdom="Animalia" order="Alcyonacea" pageId="178" pageNumber="179" phylum="Cnidaria" rank="species" species="nyahae" status="sp. nov.">
<emphasis id="B946EAE93EF3FF02FEEDE35D7CD4D9A0" box="[305,511,1908,1930]" italics="true" pageId="178" pageNumber="179">Ƒictorgorgia nyahae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27557923EF3FF02FDD9E35C7F15D9A0" box="[517,574,1908,1930]" pageId="178" pageNumber="179" rank="species">
<emphasis id="B946EAE93EF3FF02FDD9E35C7F15D9A0" bold="true" box="[517,574,1908,1930]" pageId="178" pageNumber="179">n. sp.</emphasis>
</taxonomicNameLabel>
, holotype: A. Colony; B–C. Branch tip; D. Polyp. (A–B. Courtesy of Karen Gowlett- Holmes, CSIRO O&amp;A).
</paragraph>
</caption>
<caption id="DF4D66733EF2FF03FF4BE2267C20D869" httpUri="https://zenodo.org/record/842569/files/figure.png" pageId="179" pageNumber="180" targetBox="[170,1432,193,1517]" targetPageId="179">
<paragraph id="8B8D36FB3EF2FF03FF4BE2267C20D869" blockId="179.[151,1436,1550,1603]" pageId="179" pageNumber="180">
<emphasis id="B946EAE93EF2FF03FF4BE2267C07D80E" bold="true" box="[151,300,1550,1572]" pageId="179" pageNumber="180">FIGURE 140.</emphasis>
<taxonomicName id="4C324D783EF2FF03FEEFE2277F29D80E" authority="Moore &amp; Alderslade &amp; Miller, 2017" authorityName="Moore &amp; Alderslade &amp; Miller" authorityYear="2017" box="[307,514,1550,1572]" class="Anthozoa" family="Anthothelidae" genus="Fictorgorgia" kingdom="Animalia" order="Alcyonacea" pageId="179" pageNumber="180" phylum="Cnidaria" rank="species" species="nyahae" status="sp. nov.">
<emphasis id="B946EAE93EF2FF03FEEFE2277F29D80E" box="[307,514,1550,1572]" italics="true" pageId="179" pageNumber="180">Ƒictorgorgia nyahae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27557923EF2FF03FDD5E2267F69D80E" box="[521,578,1550,1572]" pageId="179" pageNumber="180" rank="species">
<emphasis id="B946EAE93EF2FF03FDD5E2267F69D80E" bold="true" box="[521,578,1550,1572]" pageId="179" pageNumber="180">n. sp.</emphasis>
</taxonomicNameLabel>
, holotype: A. Polyp head; B. Cross-section of medulla; C. Decalcified cross-section of medulla.
</paragraph>
</caption>
<paragraph id="8B8D36FB3EF2FF03FF1BE2427FA1D9ED" blockId="179.[151,1437,1642,1991]" pageId="179" pageNumber="180">
<emphasis id="B946EAE93EF2FF03FF1BE2427CEAD8A9" bold="true" box="[199,449,1642,1667]" pageId="179" pageNumber="180">Medulla and Cortex:</emphasis>
The branches of the scleraxonian colony are composed of a central medulla, made up of tightly packed longitudinal sclerites, surrounded by a thin cortex. The cortex and medulla are separated by a crowded series of longitudinal canals which anastomose to form an encircling boundary space allowing the cortex to be easily separated from the medulla. At the breakage point of the largest piece of colony, it can be seen that the medulla is extensively penetrated by longitudinal canals (
<figureCitation id="13092A7E3EF2FF03FCF0E2D27EA4D939" box="[812,911,1786,1811]" captionStart="FIGURE 140" captionStartId="179.[151,250,1550,1572]" captionTargetBox="[170,1432,193,1517]" captionTargetId="figure@179.[151,1435,193,1529]" captionTargetPageId="179" captionText="FIGURE 140. Ƒictorgorgia nyahae n. sp., holotype: A. Polyp head; B. Cross-section of medulla; C. Decalcified cross-section of medulla." httpUri="https://zenodo.org/record/842569/files/figure.png" pageId="179" pageNumber="180">Fig. 140</figureCitation>
B, C). Many are large and encircled by a thin transparent layer of mesogloea, as mentioned by
<bibRefCitation id="EFA34B0A3EF2FF03FD6BE336791CD91D" author="Lopez-Gonzalez" box="[695,1079,1822,1847]" pageId="179" pageNumber="180" refString="Lopez-Gonzalez, P. J. &amp; Briand, P. (2002) A new scleraxonian genus from Josephine Bank, north-eastern Atlantic (Cnidaria, Octocorallia). Hydrobiologia, 482, 97 - 105. http: // dx. doi. org / 10.1023 / A: 1021256230344" type="journal article" year="2002">López-González and Briand, 2002</bibRefCitation>
for
<taxonomicName id="4C324D783EF2FF03FBB8E33779D3D91D" authorityName="Lopez-Gonzalez &amp; Briand" authorityYear="2002" box="[1124,1272,1822,1847]" class="Anthozoa" family="Anthothelidae" genus="Fictorgorgia" kingdom="Animalia" order="Alcyonacea" pageId="179" pageNumber="180" phylum="Cnidaria" rank="species" species="josephinae">
<emphasis id="B946EAE93EF2FF03FBB8E33779D3D91D" box="[1124,1272,1822,1847]" italics="true" pageId="179" pageNumber="180">V. josephinae</emphasis>
</taxonomicName>
. However, not all the canals have this layer and many are small and indistinct. For polyps which are arranged along and perpendicular to the branches, gastric cavities terminate at the medulla. For polyps which form the polyp clumps at the branch tip, their body cavity can extend internally down the branch for a short distance, eventually merging into the coelenteric canals in the central medulla.
</paragraph>
<caption id="DF4D66733EF5FF04FF4BE3E07EF5D9F4" box="[151,990,1992,2014]" httpUri="https://zenodo.org/record/842571/files/figure.png" pageId="180" pageNumber="181" targetBox="[165,1428,193,1956]" targetPageId="180">
<paragraph id="8B8D36FB3EF5FF04FF4BE3E07EF5D9F4" blockId="180.[151,990,1992,2014]" box="[151,990,1992,2014]" pageId="180" pageNumber="181">
<emphasis id="B946EAE93EF5FF04FF4BE3E07C07D9F4" bold="true" box="[151,300,1992,2014]" pageId="180" pageNumber="181">FIGURE 141.</emphasis>
<taxonomicName id="4C324D783EF5FF04FEEEE3E17F2AD9F4" authority="Moore &amp; Alderslade &amp; Miller, 2017" authorityName="Moore &amp; Alderslade &amp; Miller" authorityYear="2017" box="[306,513,1992,2014]" class="Anthozoa" family="Anthothelidae" genus="Fictorgorgia" kingdom="Animalia" order="Alcyonacea" pageId="180" pageNumber="181" phylum="Cnidaria" rank="species" species="nyahae" status="sp. nov.">
<emphasis id="B946EAE93EF5FF04FEEEE3E17F2AD9F4" box="[306,513,1992,2014]" italics="true" pageId="180" pageNumber="181">Ƒictorgorgia nyahae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27557923EF5FF04FDDBE3E07F6BD9F4" box="[519,576,1992,2014]" pageId="180" pageNumber="181" rank="species">
<emphasis id="B946EAE93EF5FF04FDDBE3E07F6BD9F4" bold="true" box="[519,576,1992,2014]" pageId="180" pageNumber="181">n. sp.</emphasis>
</taxonomicNameLabel>
, holotype, sclerites: Point (a. spear tips).
</paragraph>
</caption>
<caption id="DF4D66733EF4FF05FF4BE1C67DDAD808" httpUri="https://zenodo.org/record/842573/files/figure.png" pageId="181" pageNumber="182" targetBox="[165,1423,193,1486]" targetPageId="181">
<paragraph id="8B8D36FB3EF4FF05FF4BE1C67DDAD808" blockId="181.[151,1435,1518,1570]" pageId="181" pageNumber="182">
<emphasis id="B946EAE93EF4FF05FF4BE1C67C1BD82E" bold="true" box="[151,304,1518,1540]" pageId="181" pageNumber="182">FIGURE 142.</emphasis>
<taxonomicName id="4C324D783EF4FF05FEE6E1C77F26D82E" authority="Moore &amp; Alderslade &amp; Miller, 2017" authorityName="Moore &amp; Alderslade &amp; Miller" authorityYear="2017" box="[314,525,1518,1540]" class="Anthozoa" family="Anthothelidae" genus="Fictorgorgia" kingdom="Animalia" order="Alcyonacea" pageId="181" pageNumber="182" phylum="Cnidaria" rank="species" species="nyahae" status="sp. nov.">
<emphasis id="B946EAE93EF4FF05FEE6E1C77F26D82E" box="[314,525,1518,1540]" italics="true" pageId="181" pageNumber="182">Ƒictorgorgia nyahae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27557923EF4FF05FDCBE1C67F7ED82E" box="[535,597,1518,1540]" pageId="181" pageNumber="182" rank="species">
<emphasis id="B946EAE93EF4FF05FDCBE1C67F7ED82E" bold="true" box="[535,597,1518,1540]" pageId="181" pageNumber="182">n. sp.</emphasis>
</taxonomicNameLabel>
, holotype: A. Pinnule, tentacular and point sclerites in situ; B. Tentacle rachis sclerites.
</paragraph>
</caption>
<paragraph id="8B8D36FB3EF4FF05FF1BE2627CC2D9C5" blockId="181.[151,1437,1610,2031]" pageId="181" pageNumber="182">
<emphasis id="B946EAE93EF4FF05FF1BE2627C1ED849" bold="true" box="[199,309,1610,1635]" pageId="181" pageNumber="182">Sclerites:</emphasis>
The polyps and calyces are strongly protected with a dense covering of sclerites, predominately tuberculate sticks and spindles. On the polyp head, sclerites are arranged as a collaret and eight spiky points, with the sclerites bunched and crowded (
<figureCitation id="13092A7E3EF4FF05FDF6E2BA7FBDD880" box="[554,662,1682,1707]" captionStart="FIGURE 139" captionStartId="178.[151,250,1908,1930]" captionTargetBox="[151,1435,193,1880]" captionTargetId="figure@178.[151,1435,193,1886]" captionTargetPageId="178" captionText="FIGURE 139. Ƒictorgorgia nyahae n. sp., holotype: A. Colony; B – C. Branch tip; D. Polyp. (A – B. Courtesy of Karen Gowlett- Holmes, CSIRO O &amp; A)." httpUri="https://zenodo.org/record/842567/files/figure.png" pageId="181" pageNumber="182">Figs. 139</figureCitation>
D; 140A). Around the head, approximately 10–15 transverse rows of sclerites form the collaret. Sclerites then angle
<emphasis id="B946EAE93EF4FF05FD67E29F7E13D8E5" box="[699,824,1718,1743]" italics="true" pageId="181" pageNumber="182">en chevron</emphasis>
up into the points, and eventually are longitudinally arranged at the tip of the points where the tentacle rachis begins. The collaret and point sclerites are mostly composed of tuberculate or warty sticks and spindles that are slightly curved or straight (
<figureCitation id="13092A7E3EF4FF05FBA7E2D679F6D93D" box="[1147,1245,1790,1815]" captionStart="FIGURE 141" captionStartId="180.[151,250,1992,2014]" captionTargetBox="[165,1428,193,1956]" captionTargetId="figure@180.[165,1428,193,1970]" captionTargetPageId="180" captionText="FIGURE 141. Ƒictorgorgia nyahae n. sp., holotype, sclerites: Point (a. spear tips)." httpUri="https://zenodo.org/record/842571/files/figure.png" pageId="181" pageNumber="182">Fig. 141</figureCitation>
). The points also have protruding, often large, warty spindles and clubs which are arranged with their modified spear-tip sticking out from the polyp head at the top of the points (
<figureCitation id="13092A7E3EF4FF05FD46E36E7E2CD975" box="[666,775,1862,1887]" captionStart="FIGURE 140" captionStartId="179.[151,250,1550,1572]" captionTargetBox="[170,1432,193,1517]" captionTargetId="figure@179.[151,1435,193,1529]" captionTargetPageId="179" captionText="FIGURE 140. Ƒictorgorgia nyahae n. sp., holotype: A. Polyp head; B. Cross-section of medulla; C. Decalcified cross-section of medulla." httpUri="https://zenodo.org/record/842569/files/figure.png" pageId="181" pageNumber="182">Figs. 140</figureCitation>
A; 141a). These sclerites give the polyp head a distinctive spiky appearance. Simple sclerites from the points and collaret range from
<quantity id="4CCA9B1E3EF4FF05FBDEE3427983D9A8" box="[1026,1192,1898,1923]" metricMagnitude="-4" metricUnit="m" metricValue="4.85" metricValueMax="6.5" metricValueMin="3.2" pageId="181" pageNumber="182" unit="mm" value="0.485" valueMax="0.65" valueMin="0.32">0.32–0.65 mm</quantity>
long while the more complex sclerites usually range from
<quantity id="4CCA9B1E3EF4FF05FD8AE3A67FC8D98C" box="[598,739,1934,1959]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="181" pageNumber="182" unit="mm" value="0.6" valueMax="0.7" valueMin="0.5">0.5–0.7 mm</quantity>
long with occasional longer sclerites up to
<quantity id="4CCA9B1E3EF4FF05FB28E3A7784AD98C" box="[1268,1377,1935,1959]" metricMagnitude="-4" metricUnit="m" metricValue="9.1" pageId="181" pageNumber="182" unit="mm" value="0.91">0.91 mm</quantity>
. The sclerites from the polyp neck are simple tuberculate sticks and spindles, arranged obliquely, becoming transverse in the slightly contracted polyps.
</paragraph>
<paragraph id="8B8D36FB3EF7FF06FF1BE4BF7E95DFA2" blockId="182.[151,1437,151,392]" pageId="182" pageNumber="183">
Sclerites continue longitudinally along the aboral side of the tentacles from the points (
<figureCitation id="13092A7E3EF7FF06FB14E4BF7807DE9A" box="[1224,1324,151,176]" captionStart="FIGURE 142" captionStartId="181.[151,250,1518,1540]" captionTargetBox="[165,1423,193,1486]" captionTargetId="figure@181.[165,1428,193,1497]" captionTargetPageId="181" captionText="FIGURE 142. Ƒictorgorgia nyahae n. sp., holotype: A. Pinnule, tentacular and point sclerites in situ; B. Tentacle rachis sclerites." httpUri="https://zenodo.org/record/842573/files/figure.png" pageId="182" pageNumber="183">Fig. 142</figureCitation>
A). These sclerites are sharply pointed thorn clubs with a tuberculate handle and a bent, spiny or thorny tip (
<figureCitation id="13092A7E3EF7FF06FB3BE4947862DEFE" box="[1255,1353,188,213]" captionStart="FIGURE 142" captionStartId="181.[151,250,1518,1540]" captionTargetBox="[165,1423,193,1486]" captionTargetId="figure@181.[165,1428,193,1497]" captionTargetPageId="181" captionText="FIGURE 142. Ƒictorgorgia nyahae n. sp., holotype: A. Pinnule, tentacular and point sclerites in situ; B. Tentacle rachis sclerites." httpUri="https://zenodo.org/record/842573/files/figure.png" pageId="182" pageNumber="183">Fig. 142</figureCitation>
B). The thorny tip of the sclerites is distad in the tentacle and the bent tips project up from the tentacle rachis making it very bristly. The sclerites decrease in size distally along the tentacle rachis, grading from approximately 0.5 to
<quantity id="4CCA9B1E3EF7FF06FAEEE52C78B2DF36" box="[1330,1433,260,284]" metricMagnitude="-4" metricUnit="m" metricValue="1.7999999999999998" pageId="182" pageNumber="183" unit="mm" value="0.18">0.18 mm</quantity>
. Amongst these thorn clubs are
<taxonomicName id="4C324D783EF7FF06FE2EE50F7F41DF6A" authorityName="Lopez-Gonzalez &amp; Briand" authorityYear="2002" box="[498,618,295,320]" class="Anthozoa" family="Anthothelidae" genus="Fictorgorgia" kingdom="Animalia" order="Alcyonacea" pageId="182" pageNumber="183" phylum="Cnidaria" rank="species" species="josephinae">josephinae</taxonomicName>
clubs (
<figureCitation id="13092A7E3EF7FF06FD60E50F7E30DF6A" box="[700,795,295,320]" captionStart="FIGURE 143" captionStartId="182.[151,250,1080,1102]" captionTargetBox="[151,1433,445,1042]" captionTargetId="figure@182.[151,1435,445,1059]" captionTargetPageId="182" captionText="FIGURE 143. Ƒictorgorgia nyahae n. sp., holotype, sclerites: Tentacular and pinnule (a. flat rods with jagged edges)." httpUri="https://zenodo.org/record/842575/files/figure.png" pageId="182" pageNumber="183">Fig. 143</figureCitation>
) with a long, mostly smooth handle and a rounded spiny, bent tip. These tend to occur along the sides of the tentacles and extend into the pinnules while the thorn clubs are mostly restricted to the middle proximal ridge of the rachis (
<figureCitation id="13092A7E3EF7FF06FCE6E5477EB0DFA2" box="[826,923,367,392]" captionStart="FIGURE 142" captionStartId="181.[151,250,1518,1540]" captionTargetBox="[165,1423,193,1486]" captionTargetId="figure@181.[165,1428,193,1497]" captionTargetPageId="181" captionText="FIGURE 142. Ƒictorgorgia nyahae n. sp., holotype: A. Pinnule, tentacular and point sclerites in situ; B. Tentacle rachis sclerites." httpUri="https://zenodo.org/record/842573/files/figure.png" pageId="182" pageNumber="183">Fig. 142</figureCitation>
A).
</paragraph>
<caption id="DF4D66733EF7FF06FF4BE010786ADA64" box="[151,1345,1080,1102]" httpUri="https://zenodo.org/record/842575/files/figure.png" pageId="182" pageNumber="183" targetBox="[151,1433,445,1042]" targetPageId="182">
<paragraph id="8B8D36FB3EF7FF06FF4BE010786ADA64" blockId="182.[151,1345,1080,1102]" box="[151,1345,1080,1102]" pageId="182" pageNumber="183">
<emphasis id="B946EAE93EF7FF06FF4BE0107C07DA64" bold="true" box="[151,300,1080,1102]" pageId="182" pageNumber="183">FIGURE 143.</emphasis>
<taxonomicName id="4C324D783EF7FF06FEEEE0117F2ADA64" authority="Moore &amp; Alderslade &amp; Miller, 2017" authorityName="Moore &amp; Alderslade &amp; Miller" authorityYear="2017" box="[306,513,1080,1102]" class="Anthozoa" family="Anthothelidae" genus="Fictorgorgia" kingdom="Animalia" order="Alcyonacea" pageId="182" pageNumber="183" phylum="Cnidaria" rank="species" species="nyahae" status="sp. nov.">
<emphasis id="B946EAE93EF7FF06FEEEE0117F2ADA64" box="[306,513,1080,1102]" italics="true" pageId="182" pageNumber="183">Ƒictorgorgia nyahae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27557923EF7FF06FDDBE0107F6BDA64" box="[519,576,1080,1102]" pageId="182" pageNumber="183" rank="species">
<emphasis id="B946EAE93EF7FF06FDDBE0107F6BDA64" bold="true" box="[519,576,1080,1102]" pageId="182" pageNumber="183">n. sp.</emphasis>
</taxonomicNameLabel>
, holotype, sclerites: Tentacular and pinnule (a. flat rods with jagged edges).
</paragraph>
</caption>
<paragraph id="8B8D36FB3EF7FF06FF1BE05D791CDB68" blockId="182.[151,1437,1141,1994]" pageId="182" pageNumber="183">
Sclerites,
<quantity id="4CCA9B1E3EF7FF06FEE5E05E7CF9DAA7" box="[313,466,1142,1166]" metricMagnitude="-4" metricUnit="m" metricValue="1.85" metricValueMax="2.7" metricValueMin="1.0" pageId="182" pageNumber="183" unit="mm" value="0.185" valueMax="0.27" valueMin="0.1">0.1–0.27 mm</quantity>
long, are crowded longitudinally in the pinnules, reaching approximately half way down. They are mostly sparsely tuberculate, flat rods, straight clubs and
<taxonomicName id="4C324D783EF7FF06FC3FE0B17970DA98" authorityName="Lopez-Gonzalez &amp; Briand" authorityYear="2002" box="[995,1115,1177,1202]" class="Anthozoa" family="Anthothelidae" genus="Fictorgorgia" kingdom="Animalia" order="Alcyonacea" pageId="182" pageNumber="183" phylum="Cnidaria" rank="species" species="josephinae">josephinae</taxonomicName>
clubs with the clubbed tips pointing distad (
<figureCitation id="13092A7E3EF7FF06FE93E0957C99DAFC" box="[335,434,1213,1238]" captionStart="FIGURE 143" captionStartId="182.[151,250,1080,1102]" captionTargetBox="[151,1433,445,1042]" captionTargetId="figure@182.[151,1435,445,1059]" captionTargetPageId="182" captionText="FIGURE 143. Ƒictorgorgia nyahae n. sp., holotype, sclerites: Tentacular and pinnule (a. flat rods with jagged edges)." httpUri="https://zenodo.org/record/842575/files/figure.png" pageId="182" pageNumber="183">Fig. 143</figureCitation>
). Some of the larger sclerites have a reasonably well-developed clubbed tip, blurring the distinction between pinnule and tentacle rachis sclerites. There are also very small, flat rods with jagged edges (
<figureCitation id="13092A7E3EF7FF06FAB3E0C97D94DB34" captionStart="FIGURE 143" captionStartId="182.[151,250,1080,1102]" captionTargetBox="[151,1433,445,1042]" captionTargetId="figure@182.[151,1435,445,1059]" captionTargetPageId="182" captionText="FIGURE 143. Ƒictorgorgia nyahae n. sp., holotype, sclerites: Tentacular and pinnule (a. flat rods with jagged edges)." httpUri="https://zenodo.org/record/842575/files/figure.png" pageId="182" pageNumber="183">Fig. 143</figureCitation>
a)—these appear to crowd around the distal end of the tentacle and can appear to be arranged transversely although this may simply be an artefact of contraction of the tentacles (
<figureCitation id="13092A7E3EF7FF06FC68E101793FDB68" box="[948,1044,1321,1346]" captionStart="FIGURE 142" captionStartId="181.[151,250,1518,1540]" captionTargetBox="[165,1423,193,1486]" captionTargetId="figure@181.[165,1428,193,1497]" captionTargetPageId="181" captionText="FIGURE 142. Ƒictorgorgia nyahae n. sp., holotype: A. Pinnule, tentacular and point sclerites in situ; B. Tentacle rachis sclerites." httpUri="https://zenodo.org/record/842573/files/figure.png" pageId="182" pageNumber="183">Fig. 142</figureCitation>
A).
</paragraph>
<paragraph id="8B8D36FB3EF7FF06FF1BE1657903D814" blockId="182.[151,1437,1141,1994]" pageId="182" pageNumber="183">
Calyces are covered in tightly packed sclerites, almost all sticks and spindles with tubercles varying from simple to complex branched warts (
<figureCitation id="13092A7E3EF7FF06FDF1E1597FBADBA0" box="[557,657,1393,1418]" captionStart="FIGURE 144" captionStartId="183.[151,250,2010,2032]" captionTargetBox="[153,1435,193,1977]" captionTargetId="figure@183.[151,1435,193,1989]" captionTargetPageId="183" captionText="FIGURE 144. Ƒictorgorgia nyahae n. sp., holotype, sclerites: Calyx (a. thorny tip)." httpUri="https://zenodo.org/record/842577/files/figure.png" pageId="182" pageNumber="183">Fig. 144</figureCitation>
). Sclerites can range from
<quantity id="4CCA9B1E3EF7FF06FC1FE15A794CDBA3" box="[963,1127,1394,1418]" metricMagnitude="-4" metricUnit="m" metricValue="5.05" metricValueMax="8.8" metricValueMin="1.3" pageId="182" pageNumber="183" unit="mm" value="0.505" valueMax="0.88" valueMin="0.13">0.13–0.88 mm</quantity>
long but most are between approximately
<quantity id="4CCA9B1E3EF7FF06FE9CE1BD7CCCDB87" box="[320,487,1429,1454]" metricMagnitude="-4" metricUnit="m" metricValue="5.700000000000001" metricValueMax="7.800000000000001" metricValueMin="3.5999999999999996" pageId="182" pageNumber="183" unit="mm" value="0.57" valueMax="0.78" valueMin="0.36">0.36–0.78 mm</quantity>
. Bulky spindles with complex warts can have a spiny tip (
<figureCitation id="13092A7E3EF7FF06FBA0E1BD79F7DB84" box="[1148,1244,1429,1454]" captionStart="FIGURE 144" captionStartId="183.[151,250,2010,2032]" captionTargetBox="[153,1435,193,1977]" captionTargetId="figure@183.[151,1435,193,1989]" captionTargetPageId="183" captionText="FIGURE 144. Ƒictorgorgia nyahae n. sp., holotype, sclerites: Calyx (a. thorny tip)." httpUri="https://zenodo.org/record/842577/files/figure.png" pageId="182" pageNumber="183">Fig. 144</figureCitation>
a) although these are not as developed as those present in the points and tentacles. Most sclerites are positioned longitudinally on the calyx, and tend to group into columns so the calyx has eight vaguely defined longitudinal ridges ending at the lip as eight indistinct mounds. The warty spindles are all orientated with the thorny tip projecting out from the calyx, sometimes giving the calyx a slightly prickly appearance particularly near the lip.
</paragraph>
<paragraph id="8B8D36FB3EF7FF06FF1BE2627FA9D848" blockId="182.[151,1437,1141,1994]" box="[199,642,1609,1634]" pageId="182" pageNumber="183">No sclerites were found in the pharynx.</paragraph>
<paragraph id="8B8D36FB3EF7FF06FF1BE2457F3AD8E7" blockId="182.[151,1437,1141,1994]" pageId="182" pageNumber="183">
The thin cortex of the colony is composed of a dense layer of longitudinally arranged sticks and spindles from
<quantity id="4CCA9B1E3EF7FF06FF4BE2BA7C16D883" box="[151,317,1682,1706]" metricMagnitude="-4" metricUnit="m" metricValue="4.699999999999999" metricValueMax="7.1" metricValueMin="2.3" pageId="182" pageNumber="183" unit="mm" value="0.47" valueMax="0.71" valueMin="0.23">0.23–0.71 mm</quantity>
long (
<figureCitation id="13092A7E3EF7FF06FE57E2B97CDAD880" box="[395,497,1681,1706]" captionStart="FIGURE 145" captionStartId="184.[151,250,1759,1781]" captionTargetBox="[162,1434,193,1721]" captionTargetId="figure@184.[151,1435,193,1738]" captionTargetPageId="184" captionText="FIGURE 145. Ƒictorgorgia nyahae n. sp., holotype, sclerites: Cortex." httpUri="https://zenodo.org/record/842579/files/figure.png" pageId="182" pageNumber="183">Fig. 145</figureCitation>
). Most sclerites have mainly simple tubercles but there are some spindles with a dense covering of complex warts.
</paragraph>
<paragraph id="8B8D36FB3EF7FF06FF1BE2F1784FD98C" blockId="182.[151,1437,1141,1994]" pageId="182" pageNumber="183">
The medulla is composed of tightly packed longitudinal sclerites—mostly tuberculate and warty sticks and spindles (
<figureCitation id="13092A7E3EF7FF06FED8E2D57C43D93C" box="[260,360,1789,1814]" captionStart="FIGURE 146" captionStartId="185.[151,250,1976,1998]" captionTargetBox="[182,1398,193,1939]" captionTargetId="figure@185.[173,1415,193,1950]" captionTargetPageId="185" captionText="FIGURE 146. Ƒictorgorgia nyahae n. sp., holotype, sclerites: Medulla." httpUri="https://zenodo.org/record/842581/files/figure.png" pageId="182" pageNumber="183">Fig. 146</figureCitation>
). Occasionally there are large sticks and spindles with only sparse warts, often with branches, forks and fused areas. Bulky spindles with complex warts are also present but only infrequently. Most sclerites are from
<quantity id="4CCA9B1E3EF7FF06FF09E36D7C53D977" box="[213,376,1861,1886]" metricMagnitude="-4" metricUnit="m" metricValue="5.3" metricValueMax="7.6" metricValueMin="3.0" pageId="182" pageNumber="183" unit="mm" value="0.53" valueMax="0.76" valueMin="0.3">0.30–0.76 mm</quantity>
long although longer sclerites up to
<quantity id="4CCA9B1E3EF7FF06FCC9E36E7E7DD977" box="[789,854,1862,1886]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="182" pageNumber="183" unit="mm" value="1.0">1 mm</quantity>
were observed. It was difficult to ensure these long sclerites remained undamaged during sampling so the prevalence of these cannot be estimated. Similar to the calyx and surface, small, mostly smooth spindles, from
<quantity id="4CCA9B1E3EF7FF06FD18E3A67E4CD98F" box="[708,871,1934,1958]" metricMagnitude="-4" metricUnit="m" metricValue="1.6999999999999997" metricValueMax="2.2" metricValueMin="1.2" pageId="182" pageNumber="183" unit="mm" value="0.16999999999999998" valueMax="0.22" valueMin="0.12">0.12–0.22 mm</quantity>
in length, occur amongst the bigger sclerites.
</paragraph>
<paragraph id="8B8D36FB3EF7FF06FF1BE3997E72D9E0" blockId="182.[151,1437,1141,1994]" box="[199,857,1969,1994]" pageId="182" pageNumber="183">Sclerites are uniformly transparent under transmitted light.</paragraph>
<caption id="DF4D66733EF6FF07FF4BE3F27EC3D9DA" box="[151,1000,2010,2032]" httpUri="https://zenodo.org/record/842577/files/figure.png" pageId="183" pageNumber="184" targetBox="[153,1435,193,1977]" targetPageId="183">
<paragraph id="8B8D36FB3EF6FF07FF4BE3F27EC3D9DA" blockId="183.[151,1000,2010,2032]" box="[151,1000,2010,2032]" pageId="183" pageNumber="184">
<emphasis id="B946EAE93EF6FF07FF4BE3F27C07D9DA" bold="true" box="[151,300,2010,2032]" pageId="183" pageNumber="184">FIGURE 144.</emphasis>
<taxonomicName id="4C324D783EF6FF07FEEEE3F37F2AD9DA" authority="Moore &amp; Alderslade &amp; Miller, 2017" authorityName="Moore &amp; Alderslade &amp; Miller" authorityYear="2017" box="[306,513,2010,2032]" class="Anthozoa" family="Anthothelidae" genus="Fictorgorgia" kingdom="Animalia" order="Alcyonacea" pageId="183" pageNumber="184" phylum="Cnidaria" rank="species" species="nyahae" status="sp. nov.">
<emphasis id="B946EAE93EF6FF07FEEEE3F37F2AD9DA" box="[306,513,2010,2032]" italics="true" pageId="183" pageNumber="184">Ƒictorgorgia nyahae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27557923EF6FF07FDDBE3F27F6BD9DA" box="[519,576,2010,2032]" pageId="183" pageNumber="184" rank="species">
<emphasis id="B946EAE93EF6FF07FDDBE3F27F6BD9DA" bold="true" box="[519,576,2010,2032]" pageId="183" pageNumber="184">n. sp.</emphasis>
</taxonomicNameLabel>
, holotype, sclerites: Calyx (a. thorny tip).
</paragraph>
</caption>
<caption id="DF4D66733EF9FF08FF4BE2F77E75D8DF" box="[151,862,1759,1782]" httpUri="https://zenodo.org/record/842579/files/figure.png" pageId="184" pageNumber="185" targetBox="[162,1434,193,1721]" targetPageId="184">
<paragraph id="8B8D36FB3EF9FF08FF4BE2F77E75D8DF" blockId="184.[151,862,1759,1782]" box="[151,862,1759,1782]" pageId="184" pageNumber="185">
<emphasis id="B946EAE93EF9FF08FF4BE2F77C07D8DF" bold="true" box="[151,300,1759,1781]" pageId="184" pageNumber="185">FIGURE 145.</emphasis>
<taxonomicName id="4C324D783EF9FF08FEEEE2C97F2AD8DF" authority="Moore &amp; Alderslade &amp; Miller, 2017" authorityName="Moore &amp; Alderslade &amp; Miller" authorityYear="2017" box="[306,513,1759,1782]" class="Anthozoa" family="Anthothelidae" genus="Fictorgorgia" kingdom="Animalia" order="Alcyonacea" pageId="184" pageNumber="185" phylum="Cnidaria" rank="species" species="nyahae" status="sp. nov.">
<emphasis id="B946EAE93EF9FF08FEEEE2C97F2AD8DF" box="[306,513,1759,1782]" italics="true" pageId="184" pageNumber="185">Ƒictorgorgia nyahae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27557923EF9FF08FDDBE2F77F6BD8DF" box="[519,576,1759,1781]" pageId="184" pageNumber="185" rank="species">
<emphasis id="B946EAE93EF9FF08FDDBE2F77F6BD8DF" bold="true" box="[519,576,1759,1781]" pageId="184" pageNumber="185">n. sp.</emphasis>
</taxonomicNameLabel>
, holotype, sclerites: Cortex.
</paragraph>
</caption>
<paragraph id="8B8D36FB3EF9FF08FF1BE3347CB3D9C0" blockId="184.[151,1437,1820,2026]" pageId="184" pageNumber="185">
<emphasis id="B946EAE93EF9FF08FF1BE3347C7BD91F" bold="true" box="[199,336,1820,1845]" pageId="184" pageNumber="185">Variability:</emphasis>
The specimen TMAG K3989 is a small fragment closely resembling the
<typeStatus id="548988593EF9FF08FB57E33479DAD91F" box="[1163,1265,1820,1845]" pageId="184" pageNumber="185" type="holotype">holotype</typeStatus>
. It is a piece of a narrow branch with polyps which have a similarly prickly appearance, obvious large, straight-sided calyces and polyps which are mostly extended. No polyps are fully retracted. The sclerites are similar to those in the
<typeStatus id="548988593EF9FF08FAEFE34D78B2D954" box="[1331,1433,1893,1918]" pageId="184" pageNumber="185" type="holotype">holotype</typeStatus>
. Unfortunately there is no live photograph of this colony. However, there is a significant genetic difference between the sequences from this specimen when compared with those from the
<typeStatus id="548988593EF9FF08FC76E3857927D9EC" box="[938,1036,1965,1990]" pageId="184" pageNumber="185" type="holotype">holotype</typeStatus>
(see discussion below) thus it is not included as a
<typeStatus id="548988593EF9FF08FEECE3FA7CBFD9C0" box="[304,404,2002,2026]" pageId="184" pageNumber="185" type="paratype">paratype</typeStatus>
.
</paragraph>
</subSubSection>
<caption id="DF4D66733EF8FF09FF4BE3907E45D9E4" box="[151,878,1976,1998]" httpUri="https://zenodo.org/record/842581/files/figure.png" pageId="185" pageNumber="186" targetBox="[182,1398,193,1939]" targetPageId="185">
<paragraph id="8B8D36FB3EF8FF09FF4BE3907E45D9E4" blockId="185.[151,878,1976,1998]" box="[151,878,1976,1998]" pageId="185" pageNumber="186">
<emphasis id="B946EAE93EF8FF09FF4BE3907C07D9E4" bold="true" box="[151,300,1976,1998]" pageId="185" pageNumber="186">FIGURE 146.</emphasis>
<taxonomicName id="4C324D783EF8FF09FEEEE3917F2AD9E4" authority="Moore &amp; Alderslade &amp; Miller, 2017" authorityName="Moore &amp; Alderslade &amp; Miller" authorityYear="2017" box="[306,513,1976,1998]" class="Anthozoa" family="Anthothelidae" genus="Fictorgorgia" kingdom="Animalia" order="Alcyonacea" pageId="185" pageNumber="186" phylum="Cnidaria" rank="species" species="nyahae" status="sp. nov.">
<emphasis id="B946EAE93EF8FF09FEEEE3917F2AD9E4" box="[306,513,1976,1998]" italics="true" pageId="185" pageNumber="186">Ƒictorgorgia nyahae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27557923EF8FF09FDDBE3907F6BD9E4" box="[519,576,1976,1998]" pageId="185" pageNumber="186" rank="species">
<emphasis id="B946EAE93EF8FF09FDDBE3907F6BD9E4" bold="true" box="[519,576,1976,1998]" pageId="185" pageNumber="186">n. sp.</emphasis>
</taxonomicNameLabel>
, holotype, sclerites: Medulla.
</paragraph>
</caption>
<subSubSection id="C32865703EFBFF0AFF1BE4BF7CF7DEFE" pageId="186" pageNumber="187" type="distribution">
<paragraph id="8B8D36FB3EFBFF0AFF1BE4BF7FF8DE85" blockId="186.[151,1436,151,392]" box="[199,723,151,176]" pageId="186" pageNumber="187">
<emphasis id="B946EAE93EFBFF0AFF1BE4BF7C49DE9A" bold="true" box="[199,354,151,176]" pageId="186" pageNumber="187">Distribution:</emphasis>
Southern Tasmanian seamounts.
</paragraph>
<paragraph id="8B8D36FB3EFBFF0AFF1BE4947CF7DEFE" blockId="186.[151,1436,151,392]" box="[199,476,188,213]" pageId="186" pageNumber="187">
<emphasis id="B946EAE93EFBFF0AFF1BE4947C32DEFF" bold="true" box="[199,281,188,213]" pageId="186" pageNumber="187">Depth:</emphasis>
590–660 metres.
</paragraph>
</subSubSection>
<subSubSection id="C32865703EFBFF0AFF1BE4F77E69DF6A" pageId="186" pageNumber="187" type="discussion">
<paragraph id="8B8D36FB3EFBFF0AFF1BE4F77E69DF6A" blockId="186.[151,1436,151,392]" pageId="186" pageNumber="187">
<emphasis id="B946EAE93EFBFF0AFF1BE4F77C16DED2" bold="true" box="[199,317,223,248]" pageId="186" pageNumber="187">Remarks:</emphasis>
<taxonomicName id="4C324D783EFBFF0AFE98E4C97C9BDED2" authority="Moore &amp; Alderslade &amp; Miller, 2017" authorityName="Moore &amp; Alderslade &amp; Miller" authorityYear="2017" box="[324,432,223,248]" class="Anthozoa" family="Anthothelidae" genus="Fictorgorgia" kingdom="Animalia" order="Alcyonacea" pageId="186" pageNumber="187" phylum="Cnidaria" rank="species" species="nyahae" status="sp. nov.">
<emphasis id="B946EAE93EFBFF0AFE98E4C97C9BDED2" box="[324,432,223,248]" italics="true" pageId="186" pageNumber="187">V. nyahae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27557923EFBFF0AFE6BE4F77CDDDED2" box="[439,502,223,248]" pageId="186" pageNumber="187" rank="species">
<emphasis id="B946EAE93EFBFF0AFE6BE4F77CDDDED2" bold="true" box="[439,502,223,248]" pageId="186" pageNumber="187">n. sp.</emphasis>
</taxonomicNameLabel>
is differentiated from other
<taxonomicName id="4C324D783EFBFF0AFCEDE4C97E95DED2" authorityName="Lopez-Gonzalez &amp; Briand" authorityYear="2002" box="[817,958,225,248]" class="Anthozoa" family="Anthothelidae" genus="Victorgorgia" kingdom="Animalia" order="Alcyonacea" pageId="186" pageNumber="187" phylum="Cnidaria" rank="genus">
<emphasis id="B946EAE93EFBFF0AFCEDE4C97E95DED2" box="[817,958,225,248]" italics="true" pageId="186" pageNumber="187">Victorgorgia</emphasis>
</taxonomicName>
species by the presence of sharply pointed thorn clubs and spear-tipped spindles in the tentacles, points and top of the calyx. See molecular section for details of the divergent sequences obtained from the two specimens.
</paragraph>
</subSubSection>
<subSubSection id="C32865703EFBFF0AFF1BE5647F0DDFA2" pageId="186" pageNumber="187" type="etymology">
<paragraph id="8B8D36FB3EFBFF0AFF1BE5647F0DDFA2" blockId="186.[151,1436,151,392]" pageId="186" pageNumber="187">
<emphasis id="B946EAE93EFBFF0AFF1BE5647C7ADF4F" bold="true" box="[199,337,332,357]" pageId="186" pageNumber="187">Etymology:</emphasis>
The species is named after the first author’s daughter, Nyah, as a small recompense for a childhood overshadowed by a PhD.
</paragraph>
</subSubSection>
</treatment>
</document>
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