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<mods:title id="65036FDDB0E26F1125FBAEA14EB64536">Four new species and two new records of Nola Leach, 1815 from China (Lepidoptera: Nolidae: Nolinae)</mods:title>
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<mods:namePart id="F9E550E91F386577184C6BC66B18B4C0">Hu, Yan-Qing</mods:namePart>
<mods:affiliation id="6B4C01B641C364BCFB8933B4CCE9EFC4">Lepidoptera Research Group, School of Life Science and Engineering, Southwest University of Science and Technology, Mianyang, 621010, China.</mods:affiliation>
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<treatment id="038C87FFA32D3C02F38BFA05FF2E22B0" ID-DOI="http://doi.org/10.5281/zenodo.12733295" ID-GBIF-Taxon="235080675" ID-Zenodo-Dep="12733295" LSID="urn:lsid:plazi:treatment:038C87FFA32D3C02F38BFA05FF2E22B0" httpUri="http://treatment.plazi.org/id/038C87FFA32D3C02F38BFA05FF2E22B0" lastPageId="5" lastPageNumber="499" pageId="3" pageNumber="497">
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<paragraph id="8B9A36E9A32D3C04F38BFA05FDBE26C7" blockId="3.[151,613,1471,1534]" box="[151,613,1471,1498]" pageId="3" pageNumber="497">
<heading id="D0D28185A32D3C04F38BFA05FDBE26C7" bold="true" box="[151,613,1471,1498]" fontSize="11" level="1" pageId="3" pageNumber="497" reason="1">
<emphasis id="B951EAFBA32D3C04F38BFA05FDBE26C7" bold="true" box="[151,613,1471,1498]" pageId="3" pageNumber="497">
<taxonomicName id="4C254D6AA32D3C04F38BFA05FDDB26C7" ID-CoL="CGP9X" authority="Yu &amp; Hu" authorityName="Yu &amp; Hu" authorityYear="2024" box="[151,512,1471,1498]" class="Insecta" family="Nolidae" genus="Nola" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="497" phylum="Arthropoda" rank="species" species="hutiaoxiaensis" status="sp. nov.">
<emphasis id="B951EAFBA32D3C04F38BFA05FE5226C7" bold="true" box="[151,393,1471,1498]" italics="true" pageId="3" pageNumber="497">Nola hutiaoxiaensis</emphasis>
Yu &amp; Hu
</taxonomicName>
<taxonomicNameLabel id="A2625780A32D3C04F114FA7AFDBE26C7" box="[520,613,1472,1498]" pageId="3" pageNumber="497" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="C33F6562A32D3C04F38BFA5EFEF526E3" box="[151,302,1508,1534]" pageId="3" pageNumber="497" type="description">
<paragraph id="8B9A36E9A32D3C04F38BFA5EFEF526E3" blockId="3.[151,613,1471,1534]" box="[151,302,1508,1534]" pageId="3" pageNumber="497">
(
<figureCitation id="131E2A6CA32D3C04F383FA5EFEFE26E3" box="[159,293,1508,1534]" captionStart="FIGURES" captionStartId="4.[151,266,1824,1848]" captionTargetBox="[160,1422,190,1786]" captionTargetId="figure-18@4.[151,1436,181,1800]" captionTargetPageId="4" captionText="FIGURES. 1121. Nola spp., adult habitus and genitalia. 1113. Nola mojiangensis sp. nov. 11. Male habitus (holotype); 12, 13. Male genitalia (holotype). 1416. Nola hutiaoxiaensis sp. nov. 14. Male habitus (holotype); 15, 16. Male genitalia (holotype). 17, 18. Nola ochrolopha. 17. Female habitus; 18. Female genitalia. 1921. Nola meilingchani. 19. Male habitus; 20, 21. Male genitalia." figureDoi="http://doi.org/10.5281/zenodo.12733233" httpUri="https://zenodo.org/record/12733233/files/figure.png" pageId="3" pageNumber="497">Figs. 1416</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C33F6562A32D3C04F38BF996FD302577" pageId="3" pageNumber="497" type="materials_examined">
<paragraph id="8B9A36E9A32D3C04F38BF996FD302577" blockId="3.[151,1437,1580,2038]" pageId="3" pageNumber="497">
<emphasis id="B951EAFBA32D3C04F38BF996FE9D255B" bold="true" box="[151,326,1580,1606]" pageId="3" pageNumber="497">Type material.</emphasis>
<materialsCitation id="3B4D3CB4A32D3C04F251F996FA46255A" ID-GBIF-Occurrence="4905299311" box="[333,1437,1580,1607]" collectingDate="2004-04-26" collectingDateMax="2004-04-28" collectingDateMin="2004-04-26" collectorName="Min Wang." location="Shangri-la" municipality="Hutiaoxia Scenic Area" pageId="3" pageNumber="497" specimenCount="1" stateProvince="Yunnan" typeStatus="holotype">
<typeStatus id="549E884BA32D3C04F251F996FE61255B" box="[333,442,1580,1606]" pageId="3" pageNumber="497" type="holotype">Holotype</typeStatus>
: male,
<collectingMunicipality id="6BFEAC93A32D3C04F116F996FCD1255B" box="[522,778,1580,1606]" pageId="3" pageNumber="497">Hutiaoxia Scenic Area</collectingMunicipality>
,
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,
<collectingRegion id="49E1F80BA32D3C04F085F997FBEF255A" box="[921,1076,1581,1607]" country="China" name="Yunnan" pageId="3" pageNumber="497">Yunnan Prov.</collectingRegion>
,
<date id="FF9B1029A32D3C04F723F996FB3A255B" box="[1087,1249,1580,1606]" pageId="3" pageNumber="497" value="2004-04-26" valueMax="2004-04-28" valueMin="2004-04-26">
<collectingDate id="EFDFE9C1A32D3C04F723F996FB3A255B" box="[1087,1249,1580,1606]" pageId="3" pageNumber="497" value="2004-04-26" valueMax="2004-04-28" valueMin="2004-04-26">2628.iv.2004</collectingDate>
</date>
, leg.
<collectorName id="26D0533FA32D3C04F603F996FA46255A" box="[1311,1437,1580,1607]" pageId="3" pageNumber="497">Min Wang.</collectorName>
</materialsCitation>
<materialsCitation id="3B4D3CB4A32D3C04F38BF9EBFD3C2577" ID-GBIF-Occurrence="4905299304" box="[151,743,1616,1643]" collectingDate="2004-04-26" collectingDateMax="2004-04-28" collectingDateMin="2004-04-26" collectorName="Min Wang." location="Shangri-la" municipality="Hutiaoxia Scenic Area" pageId="3" pageNumber="497" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="paratype">
<typeStatus id="549E884BA32D3C04F38BF9EBFF262576" box="[151,253,1617,1643]" pageId="3" pageNumber="497" type="paratype">Paratype</typeStatus>
:
<specimenCount id="9D23FD60A32D3C04F214F9EAFE8D2577" box="[264,342,1616,1642]" count="1" pageId="3" pageNumber="497" type="male">1 male</specimenCount>
, with the same data as the holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33F6562A32D3C04F3DBF9CEFC0F2402" pageId="3" pageNumber="497" type="diagnosis">
<paragraph id="8B9A36E9A32D3C04F3DBF9CEFC0F2402" blockId="3.[151,1437,1580,2038]" pageId="3" pageNumber="497">
<emphasis id="B951EAFBA32D3C04F3DBF9CEFE992593" bold="true" box="[199,322,1652,1678]" pageId="3" pageNumber="497">Diagnosis.</emphasis>
Externally, the new species can be distinguished with other
<taxonomicName id="4C254D6AA32D3C04F0EEF9CEFB592593" box="[1010,1154,1652,1678]" pageId="3" pageNumber="497">
<emphasis id="B951EAFBA32D3C04F0EEF9CEFBF32593" box="[1010,1064,1652,1678]" italics="true" pageId="3" pageNumber="497">Nola</emphasis>
species
</taxonomicName>
by the remarkable black median fascia on the forewings. In the male genitalia, the new species resembles
<taxonomicName id="4C254D6AA32D3C04F735F922FE9425CB" authority="Laszlo, Ronkay &amp; Ronkay, 2014" authorityName="Laszlo, Ronkay &amp; Ronkay" authorityYear="2014" class="Insecta" family="Nolidae" genus="Nola" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="497" phylum="Arthropoda" rank="species" species="yelagarciai">
<emphasis id="B951EAFBA32D3C04F735F922FB3F25AF" box="[1065,1252,1688,1714]" italics="true" pageId="3" pageNumber="497">Nola yelagarciai</emphasis>
<bibRefCitation id="EFB44B18A32D3C04F7F0F922FE9425CB" author="Laszlo, G. M. &amp; Ronkay, G. &amp; Ronkay, L." pageId="3" pageNumber="497" pagination="201 - 228" refId="ref3668" refString="Laszlo, G. M., Ronkay, G. &amp; Ronkay, L. (2014) Taxonomic studies on the genus Nola Leach, 1815 with description of 33 new species from Asia (Lepidoptera, Noctuoidea, Nolidae, Nolini). Fibigeriana Supplement, 2, 201 - 228." type="journal article" year="2014">László, Ronkay &amp; Ronkay, 2014</bibRefCitation>
</taxonomicName>
by the apically bilobate valva and the short arms, but the difference features are found as follows: the new species has a S-shaped harpe and a short aedeagus with a long cornutus, while
<taxonomicName id="4C254D6AA32D3C04F749F95BFB2825E7" authorityName="Laszlo, Ronkay &amp; Ronkay" authorityYear="2014" box="[1109,1267,1760,1786]" class="Insecta" family="Nolidae" genus="Nola" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="497" phylum="Arthropoda" rank="species" species="yelagarciai">
<emphasis id="B951EAFBA32D3C04F749F95BFB2825E7" box="[1109,1267,1760,1786]" italics="true" pageId="3" pageNumber="497">N. yelagarciai</emphasis>
</taxonomicName>
has an apically slender and basally wide harpe and a long aedeagus with a short cornutus.
</paragraph>
</subSubSection>
<subSubSection id="C33F6562A32D3C02F3DBF892FE79225D" lastPageId="5" lastPageNumber="499" pageId="3" pageNumber="497" type="description">
<paragraph id="8B9A36E9A32D3C04F3DBF892FF0024EB" blockId="3.[151,1437,1580,2038]" pageId="3" pageNumber="497">
<emphasis id="B951EAFBA32D3C04F3DBF892FE82245F" bold="true" box="[199,345,1832,1858]" pageId="3" pageNumber="497">Description.</emphasis>
Adult (
<figureCitation id="131E2A6CA32D3C04F2AEF892FDD3245F" box="[434,520,1832,1858]" captionStart="FIGURES" captionStartId="4.[151,266,1824,1848]" captionTargetBox="[160,1422,190,1786]" captionTargetId="figure-18@4.[151,1436,181,1800]" captionTargetPageId="4" captionText="FIGURES. 1121. Nola spp., adult habitus and genitalia. 1113. Nola mojiangensis sp. nov. 11. Male habitus (holotype); 12, 13. Male genitalia (holotype). 1416. Nola hutiaoxiaensis sp. nov. 14. Male habitus (holotype); 15, 16. Male genitalia (holotype). 17, 18. Nola ochrolopha. 17. Female habitus; 18. Female genitalia. 1921. Nola meilingchani. 19. Male habitus; 20, 21. Male genitalia." figureDoi="http://doi.org/10.5281/zenodo.12733233" httpUri="https://zenodo.org/record/12733233/files/figure.png" pageId="3" pageNumber="497">Fig. 14</figureCitation>
). Wingspan
<quantity id="4CDD9B0CA32D3C04F18BF892FCCD245E" box="[663,790,1832,1859]" metricMagnitude="-2" metricUnit="m" metricValue="1.75" metricValueMax="1.8" metricValueMin="1.7" pageId="3" pageNumber="497" unit="mm" value="17.5" valueMax="18.0" valueMin="17.0">1718 mm</quantity>
. Head and collar white; labial palpus white with brown. Thorax and abdomen grayish brown. Forewing ground color grayish brown, a black median fascia between the antemedial line and the medial line; the basal line undistinguished; the antemedial line black, arc-shaped; the medial line black, zigzagged; the postmedial line brown, wavy; the subterminal line brown, arc-shaped, close to outer margin; the terminal line faint and brown; cilia brown. Hind wing ground color grayish white; discal spot pale black.
</paragraph>
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<paragraph id="8B9A36E9A32A3C03F38BF89AFE8D24B9" blockId="4.[151,1436,1824,1956]" pageId="4" pageNumber="498">
<emphasis id="B951EAFBA32A3C03F38BF89AFE872425" bold="true" box="[151,348,1824,1848]" pageId="4" pageNumber="498">FIGURES. 1121.</emphasis>
<taxonomicName id="4C254D6AA32A3C03F279F89AFE172425" box="[357,460,1824,1848]" class="Insecta" family="Nolidae" genus="Nola" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="498" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B951EAFBA32A3C03F279F89AFE4D2425" box="[357,406,1824,1848]" italics="true" pageId="4" pageNumber="498">Nola</emphasis>
spp.
</taxonomicName>
, adult habitus and genitalia. 1113.
<taxonomicName id="4C254D6AA32A3C03F050F89AFBD72425" authorityName="Yu &amp; Hu" authorityYear="2024" box="[844,1036,1824,1848]" class="Insecta" family="Nolidae" genus="Nola" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="498" phylum="Arthropoda" rank="species" species="mojiangensis" status="sp. nov.">
<emphasis id="B951EAFBA32A3C03F050F89AFBD72425" box="[844,1036,1824,1848]" italics="true" pageId="4" pageNumber="498">Nola mojiangensis</emphasis>
</taxonomicName>
<emphasis id="B951EAFBA32A3C03F70AF89AFBB22425" bold="true" box="[1046,1129,1824,1848]" pageId="4" pageNumber="498">
<taxonomicNameLabel id="A2625780A32A3C03F70AF89AFBB22425" box="[1046,1129,1824,1848]" pageId="4" pageNumber="498" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
11. Male habitus (holotype); 12, 13. Male genitalia (holotype). 1416.
<taxonomicName id="4C254D6AA32A3C03F153F8FEFCC72441" authorityName="Yu &amp; Hu" authorityYear="2024" box="[591,796,1860,1884]" class="Insecta" family="Nolidae" genus="Nola" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="498" phylum="Arthropoda" rank="species" species="hutiaoxiaensis" status="sp. nov.">
<emphasis id="B951EAFBA32A3C03F153F8FEFCC72441" box="[591,796,1860,1884]" italics="true" pageId="4" pageNumber="498">Nola hutiaoxiaensis</emphasis>
</taxonomicName>
<emphasis id="B951EAFBA32A3C03F03AF8FEFCA22441" bold="true" box="[806,889,1860,1884]" pageId="4" pageNumber="498">
<taxonomicNameLabel id="A2625780A32A3C03F03AF8FEFCA22441" box="[806,889,1860,1884]" pageId="4" pageNumber="498" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
14. Male habitus (holotype); 15, 16. Male genitalia (holotype). 17, 18.
<emphasis id="B951EAFBA32A3C03F244F8D2FDD1249D" box="[344,522,1896,1920]" italics="true" pageId="4" pageNumber="498">
<taxonomicName id="4C254D6AA32A3C03F244F8D2FDDD249D" baseAuthorityName="Hampson" baseAuthorityYear="1911" box="[344,518,1896,1920]" class="Insecta" family="Nolidae" genus="Nola" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="498" phylum="Arthropoda" rank="species" species="ochrolopha">Nola ochrolopha</taxonomicName>
.
</emphasis>
17. Female habitus; 18. Female genitalia. 1921.
<taxonomicName id="4C254D6AA32A3C03F71CF8D2FB60249D" authorityName="Laszlo, Ronkay &amp; Ronkay" authorityYear="2014" box="[1024,1211,1896,1920]" class="Insecta" family="Nolidae" genus="Nola" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="498" phylum="Arthropoda" rank="species" species="meilingchani">
<emphasis id="B951EAFBA32A3C03F71CF8D2FB60249D" box="[1024,1211,1896,1920]" italics="true" pageId="4" pageNumber="498">Nola meilingchani</emphasis>
</taxonomicName>
. 19. Male habitus; 20, 21. Male genitalia.
</paragraph>
</caption>
<paragraph id="8B9A36E9A32B3C02F3DBFF2CFC652201" blockId="5.[151,1437,150,429]" pageId="5" pageNumber="499">
<emphasis id="B951EAFBA32B3C02F3DBFF2CFEAF23AD" bold="true" box="[199,372,150,176]" pageId="5" pageNumber="499">Male genitalia</emphasis>
(
<figureCitation id="131E2A6CA32B3C02F299FF2CFDC923AD" box="[389,530,150,177]" captionStart="FIGURES" captionStartId="4.[151,266,1824,1848]" captionTargetBox="[160,1422,190,1786]" captionTargetId="figure-18@4.[151,1436,181,1800]" captionTargetPageId="4" captionText="FIGURES. 1121. Nola spp., adult habitus and genitalia. 1113. Nola mojiangensis sp. nov. 11. Male habitus (holotype); 12, 13. Male genitalia (holotype). 1416. Nola hutiaoxiaensis sp. nov. 14. Male habitus (holotype); 15, 16. Male genitalia (holotype). 17, 18. Nola ochrolopha. 17. Female habitus; 18. Female genitalia. 1921. Nola meilingchani. 19. Male habitus; 20, 21. Male genitalia." figureDoi="http://doi.org/10.5281/zenodo.12733233" httpUri="https://zenodo.org/record/12733233/files/figure.png" pageId="5" pageNumber="499">Figs. 15, 16</figureCitation>
). Uncus degenerated; tegumen medium size; valva bilobate apically, the dorsal arm longer than the ventral arm; the dorsal arm sclerotized at the costal margin, rounded at the apex; the ventral arm sclerotized at the ventral margin; harpe S-shaped, pointed at the apex; sacculus medium size; saccus V-shaped. Aedeagus short, with an anteriorly curved spine; vesica without cornuti.
</paragraph>
<paragraph id="8B9A36E9A32B3C02F3DBFE9CFE79225D" blockId="5.[151,1437,150,429]" box="[199,418,294,320]" pageId="5" pageNumber="499">
<emphasis id="B951EAFBA32B3C02F3DBFE9CFEFE225D" bold="true" box="[199,293,294,320]" pageId="5" pageNumber="499">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C33F6562A32B3C02F3DBFEF0FDBA2279" box="[199,609,330,356]" pageId="5" pageNumber="499" type="distribution">
<paragraph id="8B9A36E9A32B3C02F3DBFEF0FDBA2279" blockId="5.[151,1437,150,429]" box="[199,609,330,356]" pageId="5" pageNumber="499">
<emphasis id="B951EAFBA32B3C02F3DBFEF0FEBA2279" bold="true" box="[199,353,330,356]" pageId="5" pageNumber="499">Distribution.</emphasis>
<collectingCountry id="F3327679A32B3C02F27BFEF0FE712279" box="[359,426,330,356]" name="China" pageId="5" pageNumber="499">China</collectingCountry>
(Prov.
<collectingRegion id="49E1F80BA32B3C02F2E7FEF0FD8D2279" box="[507,598,330,356]" country="China" name="Yunnan" pageId="5" pageNumber="499">Yunnan</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="C33F6562A32B3C02F3DBFED4FF2E22B0" pageId="5" pageNumber="499" type="etymology">
<paragraph id="8B9A36E9A32B3C02F3DBFED4FF2E22B0" blockId="5.[151,1437,150,429]" pageId="5" pageNumber="499">
<emphasis id="B951EAFBA32B3C02F3DBFED4FE952295" bold="true" box="[199,334,366,392]" pageId="5" pageNumber="499">Etymology.</emphasis>
The species name is derived from the name of the type-locality, the Hutiaoxia Scenic Area, Prov.
<collectingRegion id="49E1F80BA32B3C02F38BFE29FF2B22B0" box="[151,240,403,429]" country="China" name="Yunnan" pageId="5" pageNumber="499">Yunnan</collectingRegion>
.
</paragraph>
</subSubSection>
</treatment>
</document>