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<document id="51F5B04FF08188DA4E382BAE7ED4609E" ID-DOI="10.11646/zootaxa.5293.1.7" ID-GBIF-Dataset="cf7c803a-60d3-471c-b8d1-34a67225a4ee" ID-ISSN="1175-5326" ID-Zenodo-Dep="7959879" ID-ZooBank="CA911076-3579-4BBF-8553-5A3D668D5C30" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="carolina" checkinTime="1684796687446" checkinUser="plazi" docAuthor="Tamamitsu, Saito &amp; Lien, Vu Van" docDate="2023" docId="038C87FEFFB6FF9BFF6FFB509918AF2D" docLanguage="en" docName="zootaxa.5293.1.7.pdf" docOrigin="Zootaxa 5293 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5293.1.7" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Bassarona teuta subsp. anh T. Saito &amp; Vu 2023, subspec. nov." docType="treatment" docVersion="1" lastPageNumber="169" masterDocId="FFB5FF86FFB0FF93FFF8FFD79C30AE48" masterDocTitle="Four new subspecies of butterflies (Lepidoptera, Papilionoidea) from Phu Quoc Island, southern Vietnam" masterLastPageNumber="170" masterPageNumber="161" pageNumber="167" updateTime="1684958038065" updateUser="carolina" zenodo-license-document="CLOSED">
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<mods:titleInfo id="178816D1C294E994A7A384D84707B9A4">
<mods:title id="DF056EAC4089C81FF6ADE6059A3972E7">Four new subspecies of butterflies (Lepidoptera, Papilionoidea) from Phu Quoc Island, southern Vietnam</mods:title>
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<mods:namePart id="D9024F7F28E8472C7AE672E800D36730">Tamamitsu, Saito</mods:namePart>
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<mods:namePart id="448805DEE9B5C4B8F89DD599C758810E">Lien, Vu Van</mods:namePart>
<mods:affiliation id="F46E896458CB996479800B5EFD029343">Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Cau Giay, Hanoi, Vietnam.</mods:affiliation>
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<mods:title id="435302FFC5B4611133B74B26AA5D4979">Zootaxa</mods:title>
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<mods:part id="61855EF3CDD073E095D77585811E4932">
<mods:date id="BEF4462C0F334ED3946AE400D1942D98">2023</mods:date>
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<mods:number id="3756CE03A1D04FA0E9696DB1300F5450">2023-05-19</mods:number>
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<mods:classification id="8D40E4D7F1D124A8524E508B0D0A1B46">journal article</mods:classification>
<mods:identifier id="D1A0332F34B8F2799D773712E8FCA65E" type="DOI">10.11646/zootaxa.5293.1.7</mods:identifier>
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<treatment id="038C87FEFFB6FF9BFF6FFB509918AF2D" ID-DOI="http://doi.org/10.5281/zenodo.7959863" ID-GBIF-Taxon="209628360" ID-Zenodo-Dep="7959863" LSID="urn:lsid:plazi:treatment:038C87FEFFB6FF9BFF6FFB509918AF2D" httpUri="http://treatment.plazi.org/id/038C87FEFFB6FF9BFF6FFB509918AF2D" lastPageId="8" lastPageNumber="169" pageId="6" pageNumber="167">
<subSubSection id="C33F6563FFB6FF95FF6FFB509EDDAAE9" box="[151,749,1158,1185]" pageId="6" pageNumber="167" type="nomenclature">
<paragraph id="8B9A36E8FFB6FF95FF6FFB509EDDAAE9" blockId="6.[151,749,1158,1221]" box="[151,749,1158,1185]" pageId="6" pageNumber="167">
<heading id="D0D28184FFB6FF95FF6FFB509EDDAAE9" bold="true" box="[151,749,1158,1185]" fontSize="11" level="1" pageId="6" pageNumber="167" reason="1">
<emphasis id="B951EAFAFFB6FF95FF6FFB509EDDAAE9" bold="true" box="[151,749,1158,1185]" pageId="6" pageNumber="167">
<taxonomicName id="4C254D6BFFB6FF95FF6FFB509E72AAE9" authority="T. Saito &amp; Vu" authorityName="T. Saito &amp; Vu" authorityYear="2023" box="[151,578,1158,1185]" class="Insecta" family="Nymphalidae" genus="Bassarona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="167" phylum="Arthropoda" rank="subSpecies" species="teuta" status="subspec. nov." subSpecies="anh">
<emphasis id="B951EAFAFFB6FF95FF6FFB509DA2AAE9" bold="true" box="[151,402,1158,1185]" italics="true" pageId="6" pageNumber="167">Bassarona teuta anh</emphasis>
T. Saito &amp; Vu
</taxonomicName>
<taxonomicNameLabel id="A2625781FFB6FF95FDB1FB509EDDAAE9" box="[585,749,1159,1185]" pageId="6" pageNumber="167" rank="subSpecies">subspec. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33F6563FFB6FF95FF6FFB7C9912A94D" pageId="6" pageNumber="167" type="description">
<paragraph id="8B9A36E8FFB6FF95FF6FFB7C9CC5AA8D" blockId="6.[151,749,1158,1221]" box="[151,245,1195,1221]" pageId="6" pageNumber="167">
<figureCitation id="131E2A6DFFB6FF95FF6FFB7C9CC5AA8D" box="[151,245,1195,1221]" captionStart="FIGURE 6" captionStartId="7.[151,250,1928,1952]" captionTargetBox="[309,1274,277,1904]" captionTargetId="figure-68@7.[309,1278,273,1904]" captionTargetPageId="7" captionText="FIGURE 6. Bassarona teuta anh T. Saito &amp; Vu ssp. nov., A–B—J HT, B—Ditto underside; C–D—PT ♀ same location as HT.; E—J Bassarona teuta gupta (de Nicéville, 1886) Kao Soi Dao (E. Thailand), F—Ditto un.; G—♀ same location, H—Ditto underside." figureDoi="http://doi.org/10.5281/zenodo.7959889" httpUri="https://zenodo.org/record/7959889/files/figure.png" pageId="6" pageNumber="167">Figure 6</figureCitation>
</paragraph>
<paragraph id="8B9A36E8FFB6FF95FF6FFB259D1BAB44" blockId="6.[151,1437,1266,1977]" box="[151,299,1266,1292]" pageId="6" pageNumber="167">
<heading id="D0D28184FFB6FF95FF6FFB259D1BAB44" bold="true" box="[151,299,1266,1292]" fontSize="11" level="1" pageId="6" pageNumber="167" reason="1">
<emphasis id="B951EAFAFFB6FF95FF6FFB259D1BAB44" bold="true" box="[151,299,1266,1292]" pageId="6" pageNumber="167">Description:</emphasis>
</heading>
</paragraph>
<paragraph id="8B9A36E8FFB6FF95FF3FFAC09D8CAB89" blockId="6.[151,1437,1266,1977]" pageId="6" pageNumber="167">
<emphasis id="B951EAFAFFB6FF95FF3FFAC09D34AB79" bold="true" box="[199,260,1303,1329]" pageId="6" pageNumber="167">Male</emphasis>
.
<emphasis id="B951EAFAFFB6FF95FEEDFAC19DA0AB78" bold="true" box="[277,400,1302,1328]" pageId="6" pageNumber="167">Upperside</emphasis>
. Both wings ground colour is black. Series of discal yellowish spots well developed and similar to the male patterns in other subspecies; this discal band slightly narrower. Blackish submarginal band on both wings obscure. Underside: both wings have dark brown ground colour with dark blackish stains at different areas; series of discal whitish spots narrower than in other subspecies but somewhat similar to that in subspecies g
<emphasis id="B951EAFAFFB6FF95FF5DFA709D34AB89" box="[165,260,1447,1473]" italics="true" pageId="6" pageNumber="167">oodrichi</emphasis>
(Distant, 1886).
</paragraph>
<paragraph id="8B9A36E8FFB6FF95FF3FFA1C99ACA819" blockId="6.[151,1437,1266,1977]" pageId="6" pageNumber="167">
<emphasis id="B951EAFAFFB6FF95FF3FFA1C9D9DABAC" bold="true" box="[199,429,1482,1509]" pageId="6" pageNumber="167">Female. Upperside.</emphasis>
Both wings ground colour is dark blackish brown. Forewing: Series of discal whitish spots extends from cell 2 to 6; whitish spots in cells 2 to 4 much smaller than elongated spots in cells 5 and 6. Submarginal black fascia includes separated black spots in cells 1b to 6. Hindwing: Series of whitish discal spots reduced to dot like spots only in cells 5-7; submarginal black fascia is divided into black arrow-shaped spots in cells 1a to 7.
</paragraph>
<paragraph id="8B9A36E8FFB6FF95FF6FF98D9EC6A8A9" blockId="6.[151,1437,1266,1977]" pageId="6" pageNumber="167">
<emphasis id="B951EAFAFFB6FF95FF6FF98D9D22A83C" bold="true" box="[151,274,1626,1652]" pageId="6" pageNumber="167">Underside</emphasis>
. Both wings ground colour is grayish brown with blurred areas of blackish hue. Forewing with series of separated whitish spots in cells 1b to 6; elongated spots in cells 5 and 6 larger; submarginal fascia obscure and blackish markings reduced to small dots. Hindwing with rather distinct but thin whitish discal band; submarginal black fascia reduced into separate small black strokes.
</paragraph>
<paragraph id="8B9A36E8FFB6FF95FF3FF93D9912A94D" blockId="6.[151,1437,1266,1977]" box="[199,1314,1770,1797]" pageId="6" pageNumber="167">
<emphasis id="B951EAFAFFB6FF95FF3FF93D9DA1A94D" bold="true" box="[199,401,1770,1797]" pageId="6" pageNumber="167">Forewing length.</emphasis>
J
<quantity id="4CDD9B0DFFB6FF95FE4CF93D9E67A94D" box="[436,599,1770,1797]" metricMagnitude="-2" metricUnit="m" metricValue="3.3849999999999993" metricValueMax="3.53" metricValueMin="3.2399999999999998" pageId="6" pageNumber="167" unit="mm" value="33.849999999999994" valueMax="35.3" valueMin="32.4">32.4–35.3 mm</quantity>
(n = 3, avg.
<quantity id="4CDD9B0DFFB6FF95FD1DF93C9F7DA94D" box="[741,845,1771,1797]" metricMagnitude="-2" metricUnit="m" metricValue="3.3899999999999997" pageId="6" pageNumber="167" unit="mm" value="33.9">33.9 mm</quantity>
);
<specimenCount id="9D23FD61FFB6FF95FC99F93C9F46A94D" box="[865,886,1771,1797]" pageId="6" pageNumber="167" type="female">♀</specimenCount>
<quantity id="4CDD9B0DFFB6FF95FC85F93D9810A94D" box="[893,1056,1770,1797]" metricMagnitude="-2" metricUnit="m" metricValue="4.25" metricValueMax="4.52" metricValueMin="3.9799999999999995" pageId="6" pageNumber="167" unit="mm" value="42.5" valueMax="45.2" valueMin="39.8">39.8–45.2 mm</quantity>
(n = 7, avg. 43.0 mm).
</paragraph>
</subSubSection>
<subSubSection id="C33F6563FFB6FF94FF3FF8D99F9EAEB1" lastPageId="7" lastPageNumber="168" pageId="6" pageNumber="167" type="diagnosis">
<paragraph id="8B9A36E8FFB6FF95FF3FF8D99D69A9F1" blockId="6.[151,1437,1266,1977]" pageId="6" pageNumber="167">
<emphasis id="B951EAFAFFB6FF95FF3FF8D99D72A960" bold="true" box="[199,322,1806,1832]" pageId="6" pageNumber="167">Diagnosis.</emphasis>
The population of males from Phu Quoc Is reveals similarity to males of other known subspecies, such as
<taxonomicName id="4C254D6BFFB6FF95FF0CF8E49D1CA904" box="[244,300,1843,1868]" class="Insecta" family="Nymphalidae" genus="Bassarona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="167" phylum="Arthropoda" rank="species" species="teuta">
<emphasis id="B951EAFAFFB6FF95FF0CF8E49D1CA904" box="[244,300,1843,1868]" italics="true" pageId="6" pageNumber="167">teuta</emphasis>
</taxonomicName>
,
<emphasis id="B951EAFAFFB6FF95FEC4F8E49D4CA904" box="[316,380,1843,1868]" italics="true" pageId="6" pageNumber="167">gupta</emphasis>
,
<emphasis id="B951EAFAFFB6FF95FE73F8E49DC8A905" box="[395,504,1843,1869]" italics="true" pageId="6" pageNumber="167">goodrichi</emphasis>
,
<emphasis id="B951EAFAFFB6FF95FDF0F8E49EB8A904" box="[520,648,1843,1868]" italics="true" pageId="6" pageNumber="167">rayana etc.</emphasis>
It is quite difficult to distinguish the features by which it is possible to separate males of a new taxon. Among such characters, we can note the blacker colour of the upperside of both wings; the absence of submarginal black fasciae on the upper surface of the wings, as well as smaller whitish spots of the disc stripe.
</paragraph>
<paragraph id="8B9A36E8FFB7FF94FF3FFF409F9EAEB1" blockId="7.[151,1437,151,249]" pageId="7" pageNumber="168">
The greatest differences from other subspecies of
<taxonomicName id="4C254D6BFFB7FF94FD14FF409F90AEF8" box="[748,928,151,176]" class="Insecta" family="Nymphalidae" genus="Bassarona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="168" phylum="Arthropoda" rank="species" species="teuta">
<emphasis id="B951EAFAFFB7FF94FD14FF409F90AEF8" box="[748,928,151,176]" italics="true" pageId="7" pageNumber="168">Bassarona teuta</emphasis>
</taxonomicName>
is demonstrated by the population of females. Dark brown, almost black ground colour of the upper surface of both wings, as well as the reduction of the number and size of whitish discal spots are the main features of the new taxon.
</paragraph>
</subSubSection>
<caption id="DF5A6660FFB7FF94FF6FF85F9D31A9A1" ID-DOI="http://doi.org/10.5281/zenodo.7959889" ID-Zenodo-Dep="7959889" httpUri="https://zenodo.org/record/7959889/files/figure.png" pageId="7" pageNumber="168" startId="7.[151,250,1928,1952]" targetBox="[309,1274,277,1904]" targetPageId="7" targetType="figure">
<paragraph id="8B9A36E8FFB7FF94FF6FF85F9D31A9A1" blockId="7.[151,1436,1928,2025]" pageId="7" pageNumber="168">
<emphasis id="B951EAFAFFB7FF94FF6FF85F9D3CA9E8" bold="true" box="[151,268,1928,1952]" pageId="7" pageNumber="168">FIGURE 6</emphasis>
.
<taxonomicName id="4C254D6BFFB7FF94FEE0F85E9E45A9E8" authority="T. Saito &amp; Vu" authorityName="T. Saito &amp; Vu" authorityYear="2023" box="[280,629,1929,1953]" class="Insecta" family="Nymphalidae" genus="Bassarona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="168" phylum="Arthropoda" rank="subSpecies" species="teuta" status="ssp. nov." subSpecies="anh">
<emphasis id="B951EAFAFFB7FF94FEE0F85E9DD7A9E9" box="[280,487,1929,1953]" italics="true" pageId="7" pageNumber="168">Bassarona teuta anh</emphasis>
T. Saito &amp; Vu
</taxonomicName>
<taxonomicNameLabel id="A2625781FFB7FF94FD83F85E9EE5A9E8" box="[635,725,1929,1952]" pageId="7" pageNumber="168" rank="subSpecies">ssp. nov.</taxonomicNameLabel>
,
<emphasis id="B951EAFAFFB7FF94FD26F85F9F3EA9E9" bold="true" box="[734,782,1928,1953]" pageId="7" pageNumber="168">A–B</emphasis>
—J HT,
<emphasis id="B951EAFAFFB7FF94FC93F85E9F4CA9E9" bold="true" box="[875,892,1929,1953]" pageId="7" pageNumber="168">B</emphasis>
—Ditto underside;
<emphasis id="B951EAFAFFB7FF94FBC6F85F98B9A9E9" bold="true" box="[1086,1161,1928,1953]" pageId="7" pageNumber="168">C–D—</emphasis>
PT ♀ same location as HT.;
<emphasis id="B951EAFAFFB7FF94FF6FF87A9CF1A98D" bold="true" box="[151,193,1965,1989]" pageId="7" pageNumber="168">E—</emphasis>
J
<taxonomicName id="4C254D6BFFB7FF94FF24F87A9F17A98C" authority="(de Niceville, 1886) Kao Soi Dao" authorityName="Kao Soi Dao" baseAuthorityName="de Niceville" baseAuthorityYear="1886" box="[220,807,1964,1989]" class="Insecta" family="Nymphalidae" genus="Bassarona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="168" phylum="Arthropoda" rank="subSpecies" species="teuta" subSpecies="gupta">
<emphasis id="B951EAFAFFB7FF94FF24F87A9DF4A98C" box="[220,452,1965,1988]" italics="true" pageId="7" pageNumber="168">Bassarona teuta gupta</emphasis>
(de Nicéville, 1886) Kao Soi Dao
</taxonomicName>
(E. Thailand),
<emphasis id="B951EAFAFFB7FF94FC3DF87A9FE4A98D" bold="true" box="[965,980,1965,1989]" pageId="7" pageNumber="168">F</emphasis>
—Ditto un.;
<emphasis id="B951EAFAFFB7FF94FBA0F87B98B5A98C" bold="true" box="[1112,1157,1964,1988]" pageId="7" pageNumber="168">G—</emphasis>
♀ same location,
<emphasis id="B951EAFAFFB7FF94FAC3F87A9958A98D" bold="true" box="[1339,1384,1965,1989]" pageId="7" pageNumber="168">H—</emphasis>
Ditto underside.
</paragraph>
</caption>
<subSubSection id="C33F6563FFB8FF9BFF3FFF409918AF2D" pageId="8" pageNumber="169" type="materials_examined">
<paragraph id="8B9A36E8FFB8FF9BFF3FFF409EE9AEB1" blockId="8.[151,1437,150,357]" pageId="8" pageNumber="169">
<emphasis id="B951EAFAFFB8FF9BFF3FFF409DD7AEF9" bold="true" box="[199,487,150,177]" pageId="8" pageNumber="169">
Type material.
<materialsCitation id="3B4D3CB5FFB8FF9BFE82FF409DEFAE9D" collectingDate="2015-07-04" collectorName="T. Saito" country="Vietnam" location="Ganh Dau Commune" municipality="Phu Quoc Is." pageId="8" pageNumber="169" specimenCount="1" specimenCount-female="1" stateProvince="Kien Giang province" typeStatus="holotype">
<typeStatus id="549E884AFFB8FF9BFE82FF409DD7AEF9" box="[378,487,151,177]" pageId="8" pageNumber="169">Holotype</typeStatus>
.
<specimenCount id="9D23FD61FFB8FF9BFE0DFF409E3AAEF9" box="[501,522,151,177]" pageId="8" pageNumber="169" type="female">♀</specimenCount>
S.
<collectingCountry id="F3327678FFB8FF9BFDD4FF409EBEAEF9" box="[556,654,151,177]" name="Vietnam" pageId="8" pageNumber="169">Vietnam</collectingCountry>
,
<collectingRegion id="49E1F80AFFB8FF9BFD60FF409FB5AEF9" box="[664,901,151,177]" country="Vietnam" name="Kien Giang" pageId="8" pageNumber="169">Kien Giang province</collectingRegion>
,
<collectingMunicipality id="6BFEAC92FFB8FF9BFC77FF409810AEF9" box="[911,1056,151,177]" pageId="8" pageNumber="169">Phu Quoc Is.</collectingMunicipality>
,
<location id="8EFA6033FFB8FF9BFBD4FF40992CAEF9" LSID="urn:lsid:plazi:treatment:038C87FEFFB6FF9BFF6FFB509918AF2D:8EFA6033FFB8FF9BFBD4FF40992CAEF9" box="[1068,1308,151,177]" country="Vietnam" municipality="Phu Quoc Is." name="Ganh Dau Commune" pageId="8" pageNumber="169" stateProvince="Kien Giang province">Ganh Dau Commune</location>
,
<location id="8EFA6033FFB8FF9BFADEFF409CF6AE9D" LSID="urn:lsid:plazi:treatment:038C87FEFFB6FF9BFF6FFB509918AF2D:8EFA6033FFB8FF9BFADEFF409CF6AE9D" country="Vietnam" municipality="Phu Quoc Is." name="Suoi Ganh Dau" pageId="8" pageNumber="169" stateProvince="Kien Giang province">Suoi Ganh Dau</location>
<date id="FF9B1028FFB8FF9BFF37FF6D9D7AAE9D" box="[207,330,186,213]" pageId="8" pageNumber="169" value="2015-07-04">
<collectingDate id="EFDFE9C0FFB8FF9BFF37FF6D9D7AAE9D" box="[207,330,186,213]" pageId="8" pageNumber="169" value="2015-07-04">4.VII.2015</collectingDate>
</date>
<collectorName id="26D0533EFFB8FF9BFEABFF6C9D9DAE9D" box="[339,429,187,213]" pageId="8" pageNumber="169">T. Saito</collectorName>
leg.
</materialsCitation>
</emphasis>
<emphasis id="B951EAFAFFB8FF9BFE10FF6C9E5AAE9D" bold="true" box="[488,618,187,213]" pageId="8" pageNumber="169">
<typeStatus id="549E884AFFB8FF9BFE10FF6C9E54AE9D" box="[488,612,187,213]" pageId="8" pageNumber="169">Paratypes</typeStatus>
:
</emphasis>
3J
<specimenCount id="9D23FD61FFB8FF9BFD6CFF6C9E8AAE9D" box="[660,698,187,213]" pageId="8" pageNumber="169" type="female">6♀</specimenCount>
, same location as
<typeStatus id="549E884AFFB8FF9BFC69FF6C9FCCAE9D" box="[913,1020,187,213]" pageId="8" pageNumber="169">Holotype</typeStatus>
, 3J
<specimenCount id="9D23FD61FFB8FF9BFBCEFF6C9869AE9D" box="[1078,1113,187,213]" pageId="8" pageNumber="169" type="female">3♀</specimenCount>
<date id="FF9B1028FFB8FF9BFB9BFF6D98EEAE9D" box="[1123,1246,186,213]" pageId="8" pageNumber="169" value="2015-07-04">4.VII.2015</date>
T. Saito leg.,
<specimenCount id="9D23FD61FFB8FF9BFA84FF6C99ADAE9D" box="[1404,1437,187,213]" pageId="8" pageNumber="169" type="female">2♀</specimenCount>
<date id="FF9B1028FFB8FF9BFF6FFF099D22AEB1" box="[151,274,222,249]" pageId="8" pageNumber="169" value="2015-07-03">3.VII.2015</date>
T. Saito leg.,
<specimenCount id="9D23FD61FFB8FF9BFE55FF089DF9AEB1" box="[429,457,223,249]" pageId="8" pageNumber="169" type="female">1♀</specimenCount>
<date id="FF9B1028FFB8FF9BFE28FF099E7CAEB1" box="[464,588,222,249]" pageId="8" pageNumber="169" value="2015-07-05">5.VII.2015</date>
T. Saito leg.
</paragraph>
<paragraph id="8B9A36E8FFB8FF9BFF3FFED4993DAF55" blockId="8.[151,1437,150,357]" box="[199,1293,259,285]" pageId="8" pageNumber="169">
<emphasis id="B951EAFAFFB8FF9BFF3FFED49D7EAF55" bold="true" box="[199,334,259,285]" pageId="8" pageNumber="169">Etymology.</emphasis>
The subspecific name is dedicated to Mrs. Anh, who is the wife of the second author.
</paragraph>
<paragraph id="8B9A36E8FFB8FF9BFF3FFEF19918AF2D" blockId="8.[151,1437,150,357]" pageId="8" pageNumber="169">
<emphasis id="B951EAFAFFB8FF9BFF3FFEF19D60AF08" bold="true" box="[199,336,294,320]" pageId="8" pageNumber="169">Depositary.</emphasis>
The
<typeStatus id="549E884AFFB8FF9BFE73FEF09DC8AF09" box="[395,504,295,321]" pageId="8" pageNumber="169">holotypes</typeStatus>
of four subspecies are deposited in the Research Institute of Evolution Biology in
<collectingRegion id="49E1F80AFFB8FF9BFF6FFE9C9CD1AF2D" box="[151,225,331,357]" country="Japan" name="Tokyo" pageId="8" pageNumber="169">Tokyo</collectingRegion>
,
<collectingCountry id="F3327678FFB8FF9BFF14FE9C9D1FAF2D" box="[236,303,331,357]" name="Japan" pageId="8" pageNumber="169">Japan</collectingCountry>
. The
<typeStatus id="549E884AFFB8FF9BFE94FE9C9DE9AF2D" box="[364,473,331,357]" pageId="8" pageNumber="169">Paratypes</typeStatus>
are deposited in the
<collectingCountry id="F3327678FFB8FF9BFD39FE9C9F10AF2D" box="[705,800,331,357]" name="Vietnam" pageId="8" pageNumber="169">Vietnam</collectingCountry>
National Museum of Nature,
<collectingRegion id="49E1F80AFFB8FF9BFB88FE9C9887AF2D" box="[1136,1207,331,357]" country="Vietnam" name="Ha Noi" pageId="8" pageNumber="169">Hanoi</collectingRegion>
,
<collectingCountry id="F3327678FFB8FF9BFB3AFE9C9915AF2D" box="[1218,1317,331,357]" name="Vietnam" pageId="8" pageNumber="169">Vietnam</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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