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<mods:title id="39CE3257F4ED747BF9F8AC6F1CEEE422">On some lesser known sea cucumbers in the Natural History Museum, UK (Echinodermata: Holothuroidea)</mods:title>
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<taxonomicName id="4C254D70BF6DFF91FF50FF6D7C706088" ID-CoL="3JWLV" authority="(Semper, 1867)" baseAuthorityName="Semper" baseAuthorityYear="1867" box="[151,608,225,252]" class="Holothuroidea" family="Sclerodactylidae" genus="Havelockia" kingdom="Animalia" order="Dendrochirotida" pageId="10" pageNumber="371" phylum="Echinodermata" rank="species" species="versicolor">
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<emphasis id="B951EAE1BF6DFF91FF50FF6D7F8E608F" bold="true" box="[151,414,225,252]" italics="true" pageId="10" pageNumber="371">Havelockia versicolor</emphasis>
(
<bibRefCitation id="EFB44B02BF6DFF91FE6AFF6D7C476088" author="Semper, C." box="[429,599,225,251]" pageId="10" pageNumber="371" refId="ref12610" refString="Semper, C. (1867) Holothurien. Reisen im Archipel der Philippen. Holothurien. 2. Wissenschaftliche Resultate. Wiesbaden, Leipzig, 288 pp. https: // doi. org / 10.5962 / bhl. title. 11687" type="book" year="1867">Semper, 1867</bibRefCitation>
)
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<figureCitation id="131E2A76BF6DFF91FF50FE897EE5616C" box="[151,245,261,287]" captionStart="FIGURE 6" captionStartId="11.[151,250,1919,1944]" captionTargetBox="[151,1424,204,1839]" captionTargetId="figure@11.[151,1436,204,1839]" captionTargetPageId="11" captionText="FIGURE 6. Havelockia versicolor Semper, 1868. NHMUK 1981.2.6.76. Hong Kong. A. Specimen entire; B. Part of calcareous ring; C. &amp; G. Tables of dorsal body wall; D. Tube foot end-plate; E. Tube feet ossicles, F. Rosette-like rods from tentacles. (All line drawings to same scale); MDIR = mid-dorsal interradial plate." figureDoi="http://doi.org/10.5281/zenodo.3516803" httpUri="https://zenodo.org/record/3516803/files/figure.png" pageId="10" pageNumber="371">Figure 6</figureCitation>
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<paragraph id="8B9A36F3BF6DFF91FF50FEC67A3F6111" blockId="10.[151,1437,329,2019]" box="[151,1071,329,354]" pageId="10" pageNumber="371">
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<taxonomicName id="4C254D70BF6DFF91FF50FEC67C356111" authority="Semper, 1867: 49" authorityName="Semper" authorityPageNumber="49" authorityYear="1867" box="[151,549,329,354]" class="Holothuroidea" family="Cucumariidae" genus="Cucumaria" kingdom="Animalia" order="Dendrochirotida" pageId="10" pageNumber="371" phylum="Echinodermata" rank="species" species="versiclor">
<emphasis id="B951EAE1BF6DFF91FF50FEC67F7A6111" box="[151,362,330,354]" italics="true" pageId="10" pageNumber="371">Cucumaria versiclor</emphasis>
<treatmentCitation id="0A8410E2BF6DFF91FEB7FEC67C356111" author="Semper, C." box="[368,549,329,354]" page="49" pageId="10" pageNumber="371" year="1867">
<bibRefCitation id="EFB44B02BF6DFF91FEB7FEC67C356111" author="Semper, C." box="[368,549,329,354]" pageId="10" pageNumber="371" refId="ref12610" refString="Semper, C. (1867) Holothurien. Reisen im Archipel der Philippen. Holothurien. 2. Wissenschaftliche Resultate. Wiesbaden, Leipzig, 288 pp. https: // doi. org / 10.5962 / bhl. title. 11687" type="book" year="1867">Semper, 1867: 49</bibRefCitation>
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, pl. XII, fig 11, pl. XIV, fig. 8; Utinomi, 1955: 273.
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<paragraph id="8B9A36F3BF6DFF91FF50FEE47B3A61F2" blockId="10.[151,1437,329,2019]" box="[151,1322,360,385]" pageId="10" pageNumber="371">
<treatmentCitationGroup id="AB3511DDBF6DFF91FF50FEE47B3A61F2" box="[151,1322,360,385]" pageId="10" pageNumber="371">
<taxonomicName id="4C254D70BF6DFF91FF50FEE47FEE61F3" ID-CoL="329KM" authority="Ludwig, 1875: 93" authorityName="Ludwig" authorityPageNumber="93" authorityYear="1875" box="[151,510,360,385]" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="10" pageNumber="371" phylum="Echinodermata" rank="species" species="mirabilis">
<emphasis id="B951EAE1BF6DFF91FF50FEE47F5461F3" box="[151,324,360,384]" italics="true" pageId="10" pageNumber="371">Thyone mirabilis</emphasis>
<treatmentCitation id="0A8410E2BF6DFF91FE8DFEE47FEE61F3" author="Ludwig, H." box="[330,510,360,385]" page="93" pageId="10" pageNumber="371" year="1875">
<bibRefCitation id="EFB44B02BF6DFF91FE8DFEE47FEE61F3" author="Ludwig, H." box="[330,510,360,385]" pageId="10" pageNumber="371" pagination="77 - 120" refId="ref11788" refString="Ludwig, H. (1875) Beitrage zur Kenntnis der Holothurien. Arbeiten aus dem Zoologisch. Zootomischen Institut in Wurzburg, 2, 77 - 120, pls. 6 + 7." type="journal article" year="1875">Ludwig, 1875: 93</bibRefCitation>
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pl. 6, fig. 18;
<treatmentCitation id="0A8410E2BF6DFF91FD49FEE57D4A61F2" author="Lampert, K." box="[654,858,360,385]" page="162" pageId="10" pageNumber="371" year="1885">
<bibRefCitation id="EFB44B02BF6DFF91FD49FEE57D4A61F2" author="Lampert, K." box="[654,858,360,385]" pageId="10" pageNumber="371" pagination="1 - 312" refId="ref11710" refString="Lampert, K. (1885) Die Seewalzen (Holothuroidea). In: Semper, C. (Ed.), Reisen im Archipel der Philippinen, Wiesbaden, (2) 4 (3), pp. 1 - 312, 1 pl." type="journal article" year="1885">Lampert, 1885: 162</bibRefCitation>
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;;
<treatmentCitation id="0A8410E2BF6DFF91FCACFEE47A0661F3" author="Sluiter, C." box="[875,1046,360,385]" page="93" pageId="10" pageNumber="371" year="1901">
<bibRefCitation id="EFB44B02BF6DFF91FCACFEE47A0661F3" author="Sluiter, C." box="[875,1046,360,385]" pageId="10" pageNumber="371" pagination="1 - 28" refId="ref12653" refString="Sluiter, C. P (1901) Neue Holothurien aus der Tief-See des Indischen Archipels gesammelt durch die Siboga-Expedition. Tijdschrift der Nederlandsche Dierkundige Vereeniging, 7 (1), 1 - 28." type="journal article" year="1901">Sluiter, 1901: 93</bibRefCitation>
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;
<treatmentCitation id="0A8410E2BF6DFF91FBE6FEE47AD561F2" author="Kalk, M." box="[1057,1221,360,385]" page="216" pageId="10" pageNumber="371" year="1958">
<bibRefCitation id="EFB44B02BF6DFF91FBE6FEE47AD561F2" author="Kalk, M." box="[1057,1221,360,385]" pageId="10" pageNumber="371" pagination="189 - 242" refId="ref11625" refString="Kalk, M. (1958) The fauna of the intertidal rocks at Inhaca Island, Delagoa Bay. Annals of the Natal Museum, 14, 189 - 242." type="journal article" year="1958">Kalk, 1958: 216</bibRefCitation>
</treatmentCitation>
(passim).
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</paragraph>
<paragraph id="8B9A36F3BF6DFF91FF50FE0B7AD261EC" blockId="10.[151,1437,329,2019]" box="[151,1218,390,415]" pageId="10" pageNumber="371">
Non
<taxonomicName id="4C254D70BF6DFF91FF0EFE0B7C3261EC" ID-CoL="329KM" authority="Theel, 1886: 138" authorityName="Theel" authorityPageNumber="138" authorityYear="1886" box="[201,546,390,415]" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="10" pageNumber="371" phylum="Echinodermata" rank="species" species="mirabilis">
<emphasis id="B951EAE1BF6DFF91FF0EFE0B7F6661EC" box="[201,374,391,415]" italics="true" pageId="10" pageNumber="371">Thyone mirabilis</emphasis>
Théel, 1886:138
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[(=
<taxonomicName id="4C254D70BF6DFF91FD89FE0B7CC761EC" ID-CoL="56RPH" authorityName="Rowe in Rowe &amp; Gates" authorityYear="1995" box="[590,727,391,415]" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="10" pageNumber="371" phylum="Echinodermata" rank="species" species="theeli">
<emphasis id="B951EAE1BF6DFF91FD89FE0B7CC761EC" box="[590,727,391,415]" italics="true" pageId="10" pageNumber="371">Thyone theeli</emphasis>
</taxonomicName>
(jun. sec. homonym of
<taxonomicName id="4C254D70BF6DFF91FC0FFE047AA861EC" ID-CoL="3JWLV" authority="(Semper)" authorityName="Semper" baseAuthorityName="Semper" baseAuthorityYear="1867" box="[968,1208,391,415]" class="Holothuroidea" family="Sclerodactylidae" genus="Havelockia" kingdom="Animalia" order="Dendrochirotida" pageId="10" pageNumber="371" phylum="Echinodermata" rank="species" species="versicolor">
<emphasis id="B951EAE1BF6DFF91FC0FFE047A5E61EC" box="[968,1102,391,415]" italics="true" pageId="10" pageNumber="371">H. versicolor</emphasis>
(Semper)
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].
</paragraph>
<paragraph id="8B9A36F3BF6DFF91FF50FE2A7FEA61CD" blockId="10.[151,1437,329,2019]" box="[151,506,421,446]" pageId="10" pageNumber="371">
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<taxonomicName id="4C254D70BF6DFF91FF50FE2A7FE761CD" ID-CoL="329KM" authority="? Bell, 1884: 149" authorityName="Bell" authorityPageNumber="149" authorityYear="1884" box="[151,503,421,446]" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="10" pageNumber="371" phylum="Echinodermata" rank="species" species="mirabilis">
<emphasis id="B951EAE1BF6DFF91FF50FE2A7F4161CD" box="[151,337,422,446]" italics="true" pageId="10" pageNumber="371">Thyone mirabilis?</emphasis>
<treatmentCitation id="0A8410E2BF6DFF91FE90FE297FE761CD" author="Bell, J. F." box="[343,503,421,446]" page="149" pageId="10" pageNumber="371" year="1884">
<bibRefCitation id="EFB44B02BF6DFF91FE90FE297FE761CD" author="Bell, J. F." box="[343,503,421,446]" pageId="10" pageNumber="371" pagination="1881 - 1882" refId="ref10895" refString="Bell, J. F. (1884) Echinodermata: In: Coppinger, R. W. (Ed.), Report on the zoological collections made in the Indo-Pacific Ocean during the voyage of HMS. ' Alert', 1881 - 1882. Printed by order of the Trustees, London, pp. 117 - 177 + 509 - 512." type="book chapter" year="1884">Bell, 1884: 149</bibRefCitation>
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.
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</paragraph>
<paragraph id="8B9A36F3BF6DFF91FF50FE487C2461AF" blockId="10.[151,1437,329,2019]" box="[151,564,452,476]" pageId="10" pageNumber="371">
<treatmentCitationGroup id="AB3511DDBF6DFF91FF50FE487C2461AF" box="[151,564,452,476]" pageId="10" pageNumber="371">
<emphasis id="B951EAE1BF6DFF91FF50FE487F7461AF" box="[151,356,452,476]" italics="true" pageId="10" pageNumber="371">
<taxonomicName id="4C254D70BF6DFF91FF50FE487EF261AF" ID-CoL="7Q4FF" authorityName="Oken" authorityYear="1815" box="[151,226,452,476]" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="10" pageNumber="371" phylum="Echinodermata" rank="genus">Thyone</taxonomicName>
(?) calcarea
</emphasis>
<treatmentCitation id="0A8410E2BF6DFF91FEADFE497C2061AF" author="Pearson, J." box="[362,560,452,476]" page="194" pageId="10" pageNumber="371" year="1903">
<bibRefCitation id="EFB44B02BF6DFF91FEADFE497C2061AF" author="Pearson, J." box="[362,560,452,476]" pageId="10" pageNumber="371" pagination="181 - 208" refId="ref12504" refString="Pearson, J. (1903) Report on the Holothuroidea collected by Prof. Herdman, at Ceylon, in 1902. Report to the Government of Ceylon on the Pearl Oyster Fisheries of the Gulf of Manaar, 1 (Supplement 5), 181 - 208, pls. 1 - 3." type="journal article" year="1903">Pearson, 1903: 194</bibRefCitation>
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.
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</paragraph>
<paragraph id="8B9A36F3BF6DFF91FF50FE6F7B7D6188" blockId="10.[151,1437,329,2019]" box="[151,1389,482,507]" pageId="10" pageNumber="371">
<treatmentCitationGroup id="AB3511DDBF6DFF91FF50FE6F7B7D6188" box="[151,1389,482,507]" pageId="10" pageNumber="371">
<taxonomicName id="4C254D70BF6DFF91FF50FE6F7C2D6188" ID-CoL="3JWL8" authority="Pearson, 1903: 197" authorityName="Pearson" authorityPageNumber="197" authorityYear="1903" box="[151,573,483,507]" class="Holothuroidea" family="Sclerodactylidae" genus="Havelockia" kingdom="Animalia" order="Dendrochirotida" pageId="10" pageNumber="371" phylum="Echinodermata" rank="species" species="herdmani">
<emphasis id="B951EAE1BF6DFF91FF50FE6F7F616188" box="[151,369,483,507]" italics="true" pageId="10" pageNumber="371">Havelockia herdmani</emphasis>
<treatmentCitation id="0A8410E2BF6DFF91FEB0FE6F7C2D6188" author="Pearson, J." box="[375,573,483,507]" page="197" pageId="10" pageNumber="371" year="1903">
<bibRefCitation id="EFB44B02BF6DFF91FEB0FE6F7C2D6188" author="Pearson, J." box="[375,573,483,507]" pageId="10" pageNumber="371" pagination="181 - 208" refId="ref12504" refString="Pearson, J. (1903) Report on the Holothuroidea collected by Prof. Herdman, at Ceylon, in 1902. Report to the Government of Ceylon on the Pearl Oyster Fisheries of the Gulf of Manaar, 1 (Supplement 5), 181 - 208, pls. 1 - 3." type="journal article" year="1903">Pearson, 1903: 197</bibRefCitation>
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;
<treatmentCitation id="0A8410E2BF6DFF91FD8FFE6E7D606188" author="Koehler, R. &amp; Vaney, C." box="[584,880,482,507]" page="25" pageId="10" pageNumber="371" year="1908">
<bibRefCitation id="EFB44B02BF6DFF91FD8FFE6E7D606188" author="Koehler, R. &amp; Vaney, C." box="[584,880,482,507]" pageId="10" pageNumber="371" refId="ref11657" refString="Koehler, R. &amp; Vaney, C. (1908) Holothuries recuielles par L' INVESTIGATOR dans l'Ocean Indien. II. Les Holothuries Littorales. Echinodermata of the Indian Museum. Printed by order of the trustees of the Indian Museum, Calcutta, 53 pp., 3 pls." type="book" year="1908">Koehler and Vaney, 1908: 25</bibRefCitation>
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;
<treatmentCitation id="0A8410E2BF6DFF91FCBCFE6E7A546188" author="Panning, A." box="[891,1092,482,507]" page="466" pageId="10" pageNumber="371" year="1949">
<bibRefCitation id="EFB44B02BF6DFF91FCBCFE6E7A546188" author="Panning, A." box="[891,1092,482,507]" pageId="10" pageNumber="371" pagination="404 - 470" refId="ref12386" refString="Panning, A. (1949) Versuch einer Neuordnung der Familie Cucumariidae (Holothurioidea, Dendrochirota). Zoologische Jahrbucher Abtheilung fur Systematik, Geographie und Biologie der Thiere, 78, 404 - 470." type="journal article" year="1949">Panning, 1949: 466</bibRefCitation>
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;
<treatmentCitation id="0A8410E2BF6DFF91FB89FE6E7B796188" author="Clark, H. L. &amp; Rowe, F. W. E." box="[1102,1385,482,507]" page="180" pageId="10" pageNumber="371" year="1971">
<bibRefCitation id="EFB44B02BF6DFF91FB89FE6E7B796188" author="Clark, H. L. &amp; Rowe, F. W. E." box="[1102,1385,482,507]" pageId="10" pageNumber="371" pagination="171 - 209" refId="ref11100" refString="Clark, H. L. &amp; Rowe, F. W. E. (1971) Monograph of shallow-water Indo-West Pacific echinoderms. British Museum, London, 39 pp. [pp. 171 - 209]" type="book chapter" year="1971">Clark and Rowe, 1971: 180</bibRefCitation>
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.
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</paragraph>
<paragraph id="8B9A36F3BF6DFF91FF50FD8E7C0F6269" blockId="10.[151,1437,329,2019]" box="[151,543,513,538]" pageId="10" pageNumber="371">
<treatmentCitationGroup id="AB3511DDBF6DFF91FF50FD8E7C0F6269" box="[151,543,513,538]" pageId="10" pageNumber="371">
<taxonomicName id="4C254D70BF6DFF91FF50FD8E7C0B6269" authority="Ekman, 1918: 35" authorityName="Ekman" authorityPageNumber="35" authorityYear="1918" box="[151,539,513,538]" class="Holothuroidea" family="Cucumariidae" genus="Cucumaria" kingdom="Animalia" order="Dendrochirotida" pageId="10" pageNumber="371" phylum="Echinodermata" rank="species" species="areolata">
<emphasis id="B951EAE1BF6DFF91FF50FD8E7F756269" box="[151,357,514,538]" italics="true" pageId="10" pageNumber="371">Cucumaria areolata</emphasis>
<treatmentCitation id="0A8410E2BF6DFF91FEACFD8D7C0B6269" author="Ekman, S." box="[363,539,513,538]" page="35" pageId="10" pageNumber="371" year="1918">
<bibRefCitation id="EFB44B02BF6DFF91FEACFD8D7C0B6269" author="Ekman, S." box="[363,539,513,538]" pageId="10" pageNumber="371" pagination="1 - 70" refId="ref11172" refString="Ekman, S. (1918) Holothurioidea. Results of Dr. E. Mjoberg's Schwedish Scientific Expedition to Australia 1910 - 1913 part 19. Kunglinga Svenska Vetenskapsakadmiens Handlingar, 38 (6) &amp; 58 (6), 1 - 70., 5 pls." type="book chapter" year="1918">Ekman, 1918: 35</bibRefCitation>
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B9A36F3BF6DFF91FF50FDAC7A3B624A" blockId="10.[151,1437,329,2019]" box="[151,1067,544,569]" pageId="10" pageNumber="371">
<treatmentCitationGroup id="AB3511DDBF6DFF91FF50FDAC7A3B624A" box="[151,1067,544,569]" pageId="10" pageNumber="371">
<taxonomicName id="4C254D70BF6DFF91FF50FDAC7C70624A" ID-CoL="f0ec5ace-c858-4e29-86a0-81a97938d060" authority="H. L. Clark, 1938: 459" authorityName="H. L. Clark" authorityPageNumber="459" authorityYear="1938" box="[151,608,544,569]" class="Holothuroidea" family="Sclerodactylidae" genus="Pentathyone" higherTaxonomySource="CoL" kingdom="Animalia" order="Dendrochirotida" pageId="10" pageNumber="371" phylum="Echinodermata" rank="species" species="mirabilis">
<emphasis id="B951EAE1BF6DFF91FF50FDAC7F68624B" box="[151,376,544,568]" italics="true" pageId="10" pageNumber="371">Pentathyone mirabilis</emphasis>
H.L.
<treatmentCitation id="0A8410E2BF6DFF91FE74FDAC7C70624A" author="Clark, H. L." box="[435,608,544,569]" page="459" pageId="10" pageNumber="371" year="1938">
<bibRefCitation id="EFB44B02BF6DFF91FE74FDAC7C70624A" author="Clark, H. L." box="[435,608,544,569]" pageId="10" pageNumber="371" pagination="1 - 596" refId="ref11053" refString="Clark, H. L. (1938) Echinoderms from Australia. An account of collections made in 1929 and 1932. Memoir of the Museum of Comparative Zoology, Harvard University, 55, 1 - 596, 64 figs., 28 pls." type="book chapter" year="1938">Clark, 1938: 459</bibRefCitation>
</treatmentCitation>
</taxonomicName>
, pl. 6, fig. 3;
<treatmentCitation id="0A8410E2BF6DFF91FD20FDAC7DBF624A" author="Panning, A." box="[743,943,544,569]" page="459" pageId="10" pageNumber="371" year="1949">
<bibRefCitation id="EFB44B02BF6DFF91FD20FDAC7DBF624A" author="Panning, A." box="[743,943,544,569]" pageId="10" pageNumber="371" pagination="404 - 470" refId="ref12386" refString="Panning, A. (1949) Versuch einer Neuordnung der Familie Cucumariidae (Holothurioidea, Dendrochirota). Zoologische Jahrbucher Abtheilung fur Systematik, Geographie und Biologie der Thiere, 78, 404 - 470." type="journal article" year="1949">Panning, 1949: 459</bibRefCitation>
</treatmentCitation>
, text-fig 55.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B9A36F3BF6DFF91FF50FDB37C466224" blockId="10.[151,1437,329,2019]" box="[151,598,574,599]" pageId="10" pageNumber="371">
<treatmentCitationGroup id="AB3511DDBF6DFF91FF50FDB37C466224" box="[151,598,574,599]" pageId="10" pageNumber="371">
<taxonomicName id="4C254D70BF6DFF91FF50FDB37C426224" authority="Panning, 1949: 460" authorityName="Panning" authorityPageNumber="460" authorityYear="1949" box="[151,594,574,599]" class="Holothuroidea" family="Sclerodactylidae" genus="Pentathyone" higherTaxonomySource="CoL" kingdom="Animalia" order="Dendrochirotida" pageId="10" pageNumber="371" phylum="Echinodermata" rank="species" species="versicolor">
<emphasis id="B951EAE1BF6DFF91FF50FDB37F936224" box="[151,387,575,599]" italics="true" pageId="10" pageNumber="371">Pentathyone versicolor</emphasis>
<treatmentCitation id="0A8410E2BF6DFF91FE4EFDB27C426224" author="Panning, A." box="[393,594,574,599]" page="460" pageId="10" pageNumber="371" year="1949">
<bibRefCitation id="EFB44B02BF6DFF91FE4EFDB27C426224" author="Panning, A." box="[393,594,574,599]" pageId="10" pageNumber="371" pagination="404 - 470" refId="ref12386" refString="Panning, A. (1949) Versuch einer Neuordnung der Familie Cucumariidae (Holothurioidea, Dendrochirota). Zoologische Jahrbucher Abtheilung fur Systematik, Geographie und Biologie der Thiere, 78, 404 - 470." type="journal article" year="1949">Panning, 1949: 460</bibRefCitation>
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B9A36F3BF6DFF91FF50FDD27FEA6205" blockId="10.[151,1437,329,2019]" box="[151,506,605,630]" pageId="10" pageNumber="371">
<treatmentCitationGroup id="AB3511DDBF6DFF91FF50FDD27FEA6205" box="[151,506,605,630]" pageId="10" pageNumber="371">
<taxonomicName id="4C254D70BF6DFF91FF50FDD27FE56205" authority="James, 1969: 60" authorityName="James" authorityPageNumber="60" authorityYear="1969" box="[151,501,605,630]" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="10" pageNumber="371" phylum="Echinodermata" rank="species" species="herdmani">
<emphasis id="B951EAE1BF6DFF91FF50FDD27F586205" box="[151,328,606,630]" italics="true" pageId="10" pageNumber="371">Thyone herdmani</emphasis>
<treatmentCitation id="0A8410E2BF6DFF91FE88FDD27FE56205" author="James, D. B." box="[335,501,605,630]" page="60" pageId="10" pageNumber="371" year="1969">
<bibRefCitation id="EFB44B02BF6DFF91FE88FDD27FE56205" author="James, D. B." box="[335,501,605,630]" pageId="10" pageNumber="371" pagination="51 - 62" refId="ref11523" refString="James, D. B. (1969) Catalogue of echinoderms in the reference collection of the Central Marine Fisheries Research Institute. Bulletin of the Centre for Marine Fisheries Research Institute, 7, 51 - 62." type="journal article" year="1969">James, 1969: 60</bibRefCitation>
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B9A36F3BF6DFF91FF50FDF07C8162E7" blockId="10.[151,1437,329,2019]" box="[151,657,636,661]" pageId="10" pageNumber="371">
<treatmentCitationGroup id="AB3511DDBF6DFF91FF50FDF07C8162E7" box="[151,657,636,661]" pageId="10" pageNumber="371">
<taxonomicName id="4C254D70BF6DFF91FF50FDF07C9D62E7" ID-CoL="6LFMP" authority="Clark and Rowe, 1971: 180" authorityName="Clark and Rowe" authorityPageNumber="180" authorityYear="1971" box="[151,653,636,661]" class="Holothuroidea" family="Sclerodactylidae" genus="Havelockia" kingdom="Animalia" order="Dendrochirotida" pageId="10" pageNumber="371" phylum="Echinodermata" rank="species" species="mirabilis">
<emphasis id="B951EAE1BF6DFF91FF50FDF07F7D62E7" box="[151,365,636,660]" italics="true" pageId="10" pageNumber="371">Havelockia mirabilis</emphasis>
<treatmentCitation id="0A8410E2BF6DFF91FEB4FDF07C9D62E7" author="Clark, H. L. &amp; Rowe, F. W. E." box="[371,653,636,661]" page="180" pageId="10" pageNumber="371" year="1971">
<bibRefCitation id="EFB44B02BF6DFF91FEB4FDF07C9D62E7" author="Clark, H. L. &amp; Rowe, F. W. E." box="[371,653,636,661]" pageId="10" pageNumber="371" pagination="171 - 209" refId="ref11100" refString="Clark, H. L. &amp; Rowe, F. W. E. (1971) Monograph of shallow-water Indo-West Pacific echinoderms. British Museum, London, 39 pp. [pp. 171 - 209]" type="book chapter" year="1971">Clark and Rowe, 1971: 180</bibRefCitation>
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B9A36F3BF6DFF91FF50FD177CC462A1" blockId="10.[151,1437,329,2019]" pageId="10" pageNumber="371">
<treatmentCitationGroup id="AB3511DDBF6DFF91FF50FD177B8D62C0" box="[151,1437,666,691]" pageId="10" pageNumber="371">
<taxonomicName id="4C254D70BF6DFF91FF50FD177C8F62C0" ID-CoL="3JWLV" authority="Clark and Rowe, 1971: 180" authorityName="Clark and Rowe" authorityPageNumber="180" authorityYear="1971" box="[151,671,666,691]" class="Holothuroidea" family="Sclerodactylidae" genus="Havelockia" kingdom="Animalia" order="Dendrochirotida" pageId="10" pageNumber="371" phylum="Echinodermata" rank="species" species="versicolor">
<emphasis id="B951EAE1BF6DFF91FF50FD177F6962C0" box="[151,377,667,691]" italics="true" pageId="10" pageNumber="371">Havelockia versicolor</emphasis>
<treatmentCitation id="0A8410E2BF6DFF91FE46FD167C8F62C0" author="Clark, H. L. &amp; Rowe, F. W. E." box="[385,671,666,691]" page="180" pageId="10" pageNumber="371" year="1971">
<bibRefCitation id="EFB44B02BF6DFF91FE46FD167C8F62C0" author="Clark, H. L. &amp; Rowe, F. W. E." box="[385,671,666,691]" pageId="10" pageNumber="371" pagination="171 - 209" refId="ref11100" refString="Clark, H. L. &amp; Rowe, F. W. E. (1971) Monograph of shallow-water Indo-West Pacific echinoderms. British Museum, London, 39 pp. [pp. 171 - 209]" type="book chapter" year="1971">Clark and Rowe, 1971: 180</bibRefCitation>
</treatmentCitation>
</taxonomicName>
, text-fig 91b, 92h, pl. 29, fig. 13;
<treatmentCitation id="0A8410E2BF6DFF91FBC7FD177AB962C0" author="James, D. B." box="[1024,1193,666,691]" page="55" pageId="10" pageNumber="371" year="1976">
<bibRefCitation id="EFB44B02BF6DFF91FBC7FD177AB962C0" author="James, D. B." box="[1024,1193,666,691]" pageId="10" pageNumber="371" pagination="55 - 61" refId="ref11563" refString="James, D. B. (1976) Studies on Indian echinoderms. 6. Redescription of two little known holothurians with a note on an early juvenile of Holothuria scabra Jaeger from the Indian seas. Journal of the Marine Biological Association of India, 18, 55 - 61, 1 text fig., 1 text pl." type="journal article" year="1976">James, 1976: 55</bibRefCitation>
</treatmentCitation>
, text-fig 1af;
<bibRefCitation id="EFB44B02BF6DFF91FA84FD167B8D62C0" author="Thandar, A. S." box="[1347,1437,666,691]" pageId="10" pageNumber="371" pagination="290 - 304" refId="ref12747" refString="Thandar, A. S. (1989) The sclerodactylid holothurians of southern Africa with the erection of one newsubfamily and two new genera. South African Journal of Zoology, 24 (4), 290 - 304. https: // doi. org / 10.1080 / 02541858.1989.11448167" type="journal article" year="1989">Thandar,</bibRefCitation>
</treatmentCitationGroup>
1989: 294;
<bibRefCitation id="EFB44B02BF6DFF91FEFCFD357C5762A1" author="Liao, Y. &amp; Clark, A. M." box="[315,583,697,722]" pageId="10" pageNumber="371" pagination="422 - 613" refId="ref11759" refString="Liao, Y. &amp; Clark, A. M. (1995) The echinoderms of southern China. Science Press, Beijing, 422 - 613." type="book chapter" year="1995">Liao and Clark, 1995: 484</bibRefCitation>
, text fig. 292.
</paragraph>
</subSubSection>
<subSubSection id="C33F6578BF6DFF91FF50FD577C4D6368" pageId="10" pageNumber="371" type="description">
<paragraph id="8B9A36F3BF6DFF91FF50FD577C4D6368" blockId="10.[151,1437,329,2019]" pageId="10" pageNumber="371">
<emphasis id="B951EAE1BF6DFF91FF50FD577F9D6285" bold="true" box="[151,397,731,758]" pageId="10" pageNumber="371">Name-bearing types.</emphasis>
<materialsCitation id="3B4D3CAEBF6DFF91FE54FD517C496368" ID-GBIF-Occurrence="2429072694" collectionCode="NHMUK" collectorName="Hamburg Museum E" pageId="10" pageNumber="371" specimenCode="NHMUK 1884.3" specimenCount="2" typeStatus="lectotype">
<specimenCode id="DB839E88BF6DFF91FE54FD517C426285" box="[403,594,732,759]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="10" pageNumber="371" type="Museum">NHMUK 1884.3</specimenCode>
.8.32 (
<typeStatus id="549E8851BF6DFF91FD51FD507D136284" box="[662,771,732,759]" pageId="10" pageNumber="371" type="lectotype">lectotype</typeStatus>
);
<collectorName id="26D05325BF6DFF91FCD2FD507DEE6285" box="[789,1022,732,759]" pageId="10" pageNumber="371">Hamburg Museum E</collectorName>
2873 (
<typeStatus id="549E8851BF6DFF91FB84FD507AFC6284" box="[1091,1260,732,759]" pageId="10" pageNumber="371" type="paralectotype">paralectotypes</typeStatus>
); (type designations by Arumugam 2012, unpublished)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33F6578BF6DFF91FF00FCA87F7F63D9" pageId="10" pageNumber="371" type="materials_examined">
<paragraph id="8B9A36F3BF6DFF91FF00FCA87C55634D" blockId="10.[151,1437,329,2019]" box="[199,581,803,830]" pageId="10" pageNumber="371">
<materialsCitation id="3B4D3CAEBF6DFF91FF00FCA87C51634D" ID-GBIF-Occurrence="2429072696" box="[199,577,803,830]" country="Philippines" location="Bohol" pageId="10" pageNumber="371" specimenCount="1" stateProvince="Bohol" typeStatus="holotype">
<emphasis id="B951EAE1BF6DFF91FF00FCA87F76634D" bold="true" box="[199,358,803,830]" pageId="10" pageNumber="371">
<typeStatus id="549E8851BF6DFF91FF00FCA87F12634D" box="[199,258,804,830]" pageId="10" pageNumber="371">Type</typeStatus>
locality.
</emphasis>
<collectingRegion id="49E1F811BF6DFF91FEABFCA87FA3634D" box="[364,435,804,830]" country="Philippines" name="Bohol" pageId="10" pageNumber="371">Bohol</collectingRegion>
,
<collectingCountry id="F3327663BF6DFF91FE78FCA87C51634D" box="[447,577,804,830]" name="Philippines" pageId="10" pageNumber="371">Philippines</collectingCountry>
</materialsCitation>
.
</paragraph>
<paragraph id="8B9A36F3BF6DFF91FF00FCCB7A5D6310" blockId="10.[151,1437,329,2019]" box="[199,1101,839,867]" pageId="10" pageNumber="371">
<emphasis id="B951EAE1BF6DFF91FF00FCCB7FA16311" bold="true" box="[199,433,839,866]" pageId="10" pageNumber="371">Material examined.</emphasis>
<materialsCitation id="3B4D3CAEBF6DFF91FE70FCC57A5D6310" ID-GBIF-Occurrence="2429072692" box="[439,1101,840,867]" collectionCode="NHMUK" country="Hong Kong" location="Ninepins" pageId="10" pageNumber="371" specimenCount="2">
<collectionCode id="ED34AE36BF6DFF91FE70FCC57C316310" box="[439,545,841,867]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="10" pageNumber="371" type="Museum">NHMUK</collectionCode>
1981.2.6.76-77,
<location id="8EFA6028BF6DFF91FD26FCC47D566311" LSID="urn:lsid:plazi:treatment:038C87E5BF6DFF97FF50FF6D7F2265B0:8EFA6028BF6DFF91FD26FCC47D566311" box="[737,838,840,866]" country="Hong Kong" name="Ninepins" pageId="10" pageNumber="371">Ninepins</location>
ls,
<collectingCountry id="F3327663BF6DFF91FCAAFCC57DE46310" box="[877,1012,841,867]" name="Hong Kong" pageId="10" pageNumber="371">Hong Kong</collectingCountry>
,
<specimenCount id="9D23FD7ABF6DFF91FC38FCC47A5D6310" box="[1023,1101,840,867]" pageId="10" pageNumber="371" type="generic">2 spec.</specimenCount>
</materialsCitation>
</paragraph>
<paragraph id="8B9A36F3BF6DFF91FF00FCE17A2163F4" blockId="10.[151,1437,329,2019]" box="[199,1073,876,903]" pageId="10" pageNumber="371">
<materialsCitation id="3B4D3CAEBF6DFF91FF00FCE17A3E63F4" ID-GBIF-Occurrence="2429072693" box="[199,1070,876,903]" collectionCode="NHMUK" collectorName="Hamburg Museum E" pageId="10" pageNumber="371" specimenCode="NHMUK 1884.3" specimenCount="2" typeStatus="lectotype">
<specimenCode id="DB839E88BF6DFF91FF00FCE17F9763F5" box="[199,391,876,903]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="10" pageNumber="371" type="Museum">NHMUK 1884.3</specimenCode>
.8.32 (
<typeStatus id="549E8851BF6DFF91FE0BFCE07C2963F4" box="[460,569,876,903]" pageId="10" pageNumber="371" type="lectotype">lectotype</typeStatus>
);
<collectorName id="26D05325BF6DFF91FD8BFCE07D2663F5" box="[588,822,876,903]" pageId="10" pageNumber="371">Hamburg Museum E</collectorName>
2873 (
<typeStatus id="549E8851BF6DFF91FCBAFCE07A3663F4" box="[893,1062,876,903]" pageId="10" pageNumber="371" type="paralectotype">paralectotypes</typeStatus>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8B9A36F3BF6DFF91FF00FC037F7F63D9" blockId="10.[151,1437,329,2019]" box="[199,367,911,938]" pageId="10" pageNumber="371">
<emphasis id="B951EAE1BF6DFF91FF00FC037F3A63D9" bold="true" box="[199,298,911,938]" pageId="10" pageNumber="371">Habitat.</emphasis>
Coral
</paragraph>
</subSubSection>
<subSubSection id="C33F6578BF6DFF91FF00FC3F7A3066EC" pageId="10" pageNumber="371" type="description">
<paragraph id="8B9A36F3BF6DFF91FF00FC3F7C066561" blockId="10.[151,1437,329,2019]" pageId="10" pageNumber="371">
<emphasis id="B951EAE1BF6DFF91FF00FC3F7F4963BD" bold="true" box="[199,345,947,974]" pageId="10" pageNumber="371">Description.</emphasis>
Specimens cylindrical, barrel-shaped. Colour greyish, darker in ambulacra. Larger specimen eviscerated but calcareous ring intact, length of larger specimen
<quantity id="4CDD9B16BF6DFF91FC8BFC5B7D8F6380" box="[844,927,983,1011]" metricMagnitude="-2" metricUnit="m" metricValue="5.1" pageId="10" pageNumber="371" unit="mm" value="51.0">51 mm</quantity>
, breadth in mid-body
<quantity id="4CDD9B16BF6DFF91FB59FC547AE16380" box="[1182,1265,984,1011]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="10" pageNumber="371" unit="mm" value="11.0">11 mm</quantity>
; smaller specimen
<quantity id="4CDD9B16BF6DFF91FF08FC707F0F6464" box="[207,287,1020,1047]" metricMagnitude="-2" metricUnit="m" metricValue="3.6" pageId="10" pageNumber="371" unit="mm" value="36.0">36 mm</quantity>
in length,
<quantity id="4CDD9B16BF6DFF91FE5EFC707FF96464" box="[409,489,1020,1047]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="10" pageNumber="371" unit="mm" value="12.0">12 mm</quantity>
in breadth in mid-body. Tentacles 10, mid-ventral two reduced. Tube feet scattered, retractile, more numerous in ambulacra, thick with suckers of more or less same diameter but appearing larger when tube feet retracted. Ventral tube feet more developed than dorsal; dorsal often papilliform and fewer. No anal teeth. Genital papilla prominent, situated dorsally between extended introvert and body wall. Calcareous ring clearly of the sclerodactylid
<typeStatus id="549E8851BF6DFF91FEADFB017F8D64D4" box="[362,413,1165,1191]" pageId="10" pageNumber="371">type</typeStatus>
, with radial and interradial plates fused and unfragmented. Processes of radial plates not obviously fragmented except for one or two of which do show some incipient subdivision or fragmentation. Radial plates not obviously bifid anteriorly. Stone canal free, short. Madreporite sub-spherical, and brain coral-like. Polian vesicle single, elongated, sac-like.
</paragraph>
<paragraph id="8B9A36F3BF6DFF91FF00FA907A3066EC" blockId="10.[151,1437,329,2019]" pageId="10" pageNumber="371">
Ossicles of body wall as two-pillared tables with an oblong, oval to irregular disc (
<quantity id="4CDD9B16BF6DFF91FBAFFA907ACF6544" box="[1128,1247,1308,1335]" metricMagnitude="-2" metricUnit="m" metricValue="5.5" metricValueMax="8.1" metricValueMin="2.9" pageId="10" pageNumber="371" unit="mm" value="55.0" valueMax="81.0" valueMin="29.0">2981 µm</quantity>
, average
<quantity id="4CDD9B16BF6DFF91FA8DFA977B846544" box="[1354,1428,1307,1335]" metricMagnitude="-2" metricUnit="m" metricValue="5.1" pageId="10" pageNumber="371" unit="mm" value="51.0">51 µm</quantity>
) with smooth margins, perforated by usually four large central and approximately 10 marginal holes smaller than the central holes. Spire low or of moderate height (ca.
<quantity id="4CDD9B16BF6DFF91FD33FAE87D50650C" box="[756,832,1380,1407]" metricMagnitude="-2" metricUnit="m" metricValue="2.4" pageId="10" pageNumber="371" unit="mm" value="24.0">24 µm</quantity>
), sometimes elongated, always of two pillars with one or two cross-bars, terminating in a few ill-defined teeth. Tube feet deposits comprise arched tables (
<quantity id="4CDD9B16BF6DFF91FAD6FA047B8765D0" box="[1297,1431,1416,1443]" metricMagnitude="-2" metricUnit="m" metricValue="6.45" metricValueMax="10.0" metricValueMin="2.9" pageId="10" pageNumber="371" unit="mm" value="64.5" valueMax="100.0" valueMin="29.0">29100 µm</quantity>
, average
<quantity id="4CDD9B16BF6DFF91FF31FA207F5365B4" box="[246,323,1452,1479]" metricMagnitude="-2" metricUnit="m" metricValue="6.3" pageId="10" pageNumber="371" unit="mm" value="63.0">63 µm</quantity>
); end plates (ca.
<quantity id="4CDD9B16BF6DFF91FDC6FA207C4965B4" box="[513,601,1452,1479]" metricMagnitude="-1" metricUnit="m" metricValue="1.93" pageId="10" pageNumber="371" unit="mm" value="193.0">193 µm</quantity>
in diam.), well developed with minute medial holes and much larger ones outside these in two or three concentric circles. Introvert deposits only common at the base of tentacles comprising, in the larger individual, only tables with rather deformed and partially corroded disc and tall spire comprising two pillars usually separated by one or two cross-bars and terminating in 13 teeth. Rosettes absent in larger specimen but smaller specimen endowed with both well-developed tables and rosettes with the former composed of tall, usually completely or terminally fused pillars ending in 13 teeth. Tentacle stalks without ossicles, but branches comprise rods and finely branched rosette-like rods (
<quantity id="4CDD9B16BF6DFF91FD22F9087D4D66EC" box="[741,861,1668,1695]" metricMagnitude="-2" metricUnit="m" metricValue="3.65" metricValueMax="4.9" metricValueMin="2.4" pageId="10" pageNumber="371" unit="mm" value="36.5" valueMax="49.0" valueMin="24.0">2449 µm</quantity>
; average
<quantity id="4CDD9B16BF6DFF91FC00F9087A0466EC" box="[967,1044,1668,1695]" metricMagnitude="-2" metricUnit="m" metricValue="3.6" pageId="10" pageNumber="371" unit="mm" value="36.0">36 µm</quantity>
).
</paragraph>
</subSubSection>
<subSubSection id="C33F6578BF6DFF91FF00F92B7C8166B1" box="[199,657,1703,1730]" pageId="10" pageNumber="371" type="distribution">
<paragraph id="8B9A36F3BF6DFF91FF00F92B7C8166B1" blockId="10.[151,1437,329,2019]" box="[199,657,1703,1730]" pageId="10" pageNumber="371">
<emphasis id="B951EAE1BF6DFF91FF00F92B7F7166B1" bold="true" box="[199,353,1703,1730]" pageId="10" pageNumber="371">Distribution.</emphasis>
Central Indo-West Pacific.
</paragraph>
</subSubSection>
<subSubSection id="C33F6578BF6DFF97FF00F9407B1863F0" lastPageId="12" lastPageNumber="373" pageId="10" pageNumber="371" type="discussion">
<paragraph id="8B9A36F3BF6DFF97FF00F9407A6862D8" blockId="10.[151,1437,329,2019]" lastBlockId="12.[151,1437,153,1475]" lastPageId="12" lastPageNumber="373" pageId="10" pageNumber="371">
<emphasis id="B951EAE1BF6DFF91FF00F9407F246695" bold="true" box="[199,308,1740,1766]" pageId="10" pageNumber="371">Remarks</emphasis>
. Judging from the number of synonyms it is obvious that this is a very variable Indo-West Pacific species, extending from the Indo-Malayan region, to
<collectingCountry id="F3327663BF6DFF91FD21F97C7D6E6779" box="[742,894,1776,1802]" name="Mozambique" pageId="10" pageNumber="371">Mozambique</collectingCountry>
. It was only in 1976 that James synonymised the different names, and it is not known that he examined all the species he synonymised. However, this synonymy, although too sweeping, has been accepted by subsequent workers, except
<bibRefCitation id="EFB44B02BF6DFF91FC3DF8B57A816721" author="Rowe, F. W. E. &amp; Gates, J." box="[1018,1169,1847,1875]" pageId="10" pageNumber="371" pagination="1 - 510" refId="ref12560" refString="Rowe, F. W. E. &amp; Gates, J. (1995) Echinodermata. In: Wells, A. (Ed.), Zoological Catalogue of Australia. Vol. 33. CSIRO, Melbourne, pp. i-xiii + 1 - 510." type="book chapter" year="1995">Rowe (1995)</bibRefCitation>
who elevated Théelss (1886) to a nomen nov. (
<taxonomicName id="4C254D70BF6DFF91FE69F8D07C566705" authorityName="Rowe in Rowe &amp; Gates" authorityYear="1995" box="[430,582,1884,1910]" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="10" pageNumber="371" phylum="Echinodermata" rank="species" species="theeli">
<emphasis id="B951EAE1BF6DFF91FE69F8D07C566705" box="[430,582,1884,1910]" italics="true" pageId="10" pageNumber="371">Thyone theeli</emphasis>
</taxonomicName>
as a junior secondary homonym of
<taxonomicName id="4C254D70BF6DFF91FC1CF8D07AC46705" authorityName="Clark and Rowe" authorityYear="1971" box="[987,1236,1884,1910]" class="Holothuroidea" family="Sclerodactylidae" genus="Havelockia" kingdom="Animalia" order="Dendrochirotida" pageId="10" pageNumber="371" phylum="Echinodermata" rank="species" species="versicolor">
<emphasis id="B951EAE1BF6DFF91FC1CF8D07AC46705" box="[987,1236,1884,1910]" italics="true" pageId="10" pageNumber="371">Havelockia versicolor</emphasis>
</taxonomicName>
), after examining Théelss material in the NHMUK. Likewise, all the synonyms established by
<bibRefCitation id="EFB44B02BF6DFF91FC3CF80D7A8067E8" author="James, D. B." box="[1019,1168,1920,1947]" pageId="10" pageNumber="371" pagination="55 - 61" refId="ref11563" refString="James, D. B. (1976) Studies on Indian echinoderms. 6. Redescription of two little known holothurians with a note on an early juvenile of Holothuria scabra Jaeger from the Indian seas. Journal of the Marine Biological Association of India, 18, 55 - 61, 1 text fig., 1 text pl." type="journal article" year="1976">James (1976)</bibRefCitation>
need to be examined or re-examined. It is here noted that the NHMUK material from
<collectingCountry id="F3327663BF6DFF91FC86F8297DD367CC" box="[833,963,1957,1983]" name="Hong Kong" pageId="10" pageNumber="371">Hong Kong</collectingCountry>
described above, differs a little from other geographic variants (see that described below from
<collectingCountry id="F3327663BF6DFF91FD1AF8447D5A6790" box="[733,842,1992,2019]" name="Australia" pageId="10" pageNumber="371">Australia</collectingCountry>
).
<bibRefCitation id="EFB44B02BF6DFF91FC9BF8457DE36790" author="James, D. B." box="[860,1011,1992,2019]" pageId="10" pageNumber="371" pagination="55 - 61" refId="ref11563" refString="James, D. B. (1976) Studies on Indian echinoderms. 6. Redescription of two little known holothurians with a note on an early juvenile of Holothuria scabra Jaeger from the Indian seas. Journal of the Marine Biological Association of India, 18, 55 - 61, 1 text fig., 1 text pl." type="journal article" year="1976">James (1976)</bibRefCitation>
records only rosettes in the introvert and tentacles but no tables but Arumugam (2012), in a yet unpublished work, who examined some materials from both the NHMUK and the Hamburg Museum, reports both tables and rosettes in the introvert and only rosettes in the tentacles and this was confirmed by the writer. In the specimens here studied, introvert deposits were only found at the base of the tentacles, which in the larger specimen represented tables only whereas in the smaller specimen, both tables and rosettes. Arumugam (2012), on my instructions proposed designation of the NHMUK specimen (1884.3.8.32) as
<typeStatus id="549E8851BF6BFF97FE9CFEC17FD26114" box="[347,450,333,359]" pageId="12" pageNumber="373" type="lectotype">lectotype</typeStatus>
and the Hamburg Museum specimens (E2873 as
<typeStatus id="549E8851BF6BFF97FBC0FEC17ABE611B" box="[1031,1198,333,360]" pageId="12" pageNumber="373" type="paralectotype">paralectotypes</typeStatus>
). These designations are here published for the first time. The tables with their large multilocular disc and a tall spire composed of two fused or partially fused pillars terminating in 13 points, closely resemble those illustrated by Arumugam for the
<typeStatus id="549E8851BF6BFF97FF50FE357F1161A0" box="[151,257,441,467]" pageId="12" pageNumber="373" type="lectotype">lectotype</typeStatus>
. whereas the tentacle deposits match those illustrated by
<bibRefCitation id="EFB44B02BF6BFF97FC4DFE357A3161A7" author="James, D. B." box="[906,1057,441,468]" pageId="12" pageNumber="373" pagination="55 - 61" refId="ref11563" refString="James, D. B. (1976) Studies on Indian echinoderms. 6. Redescription of two little known holothurians with a note on an early juvenile of Holothuria scabra Jaeger from the Indian seas. Journal of the Marine Biological Association of India, 18, 55 - 61, 1 text fig., 1 text pl." type="journal article" year="1976">James (1976)</bibRefCitation>
as well as by Arumugam (2012). The calcareous ring of the current material differs slightly from that known for the species, but is clearly of the sclerodactylid type but the plates are distinctly fused and the processes are much shorter. In addition, in the current material, as opposed to that illustrated by
<bibRefCitation id="EFB44B02BF6BFF97FDA9FDA97D6A624C" author="Clark, H. L. &amp; Rowe, F. W. E." box="[622,890,549,575]" pageId="12" pageNumber="373" pagination="171 - 209" refId="ref11100" refString="Clark, H. L. &amp; Rowe, F. W. E. (1971) Monograph of shallow-water Indo-West Pacific echinoderms. British Museum, London, 39 pp. [pp. 171 - 209]" type="book chapter" year="1971">Clark and Rowe (1971)</bibRefCitation>
, the processes are not obviously fragmented and the radial plates likewise not obviously bifid anteriorly. These differences are here judged not valid to separate the current material since in size, colour, form and the general body wall and tube feet deposits it corresponds remarkably well with
<taxonomicName id="4C254D70BF6BFF97FEF9FD1E7FC362D8" authorityName="Clark and Rowe" authorityYear="1971" box="[318,467,657,683]" class="Holothuroidea" family="Sclerodactylidae" genus="Havelockia" kingdom="Animalia" order="Dendrochirotida" pageId="12" pageNumber="373" phylum="Echinodermata" rank="species" species="versicolor">
<emphasis id="B951EAE1BF6BFF97FEF9FD1E7FC362D8" box="[318,467,657,683]" italics="true" pageId="12" pageNumber="373">H. versicolor</emphasis>
</taxonomicName>
and its synonyms as well as the
<typeStatus id="549E8851BF6BFF97FC87FD1D7DB762D8" box="[832,935,657,683]" pageId="12" pageNumber="373" type="lectotype">lectotype</typeStatus>
here re-examined.
</paragraph>
<caption id="DF5A667BBF6CFF90FF50F8F37D2C6793" ID-DOI="http://doi.org/10.5281/zenodo.3516803" ID-Zenodo-Dep="3516803" httpUri="https://zenodo.org/record/3516803/files/figure.png" pageId="11" pageNumber="372" startId="11.[151,250,1919,1944]" targetBox="[151,1424,204,1839]" targetPageId="11">
<paragraph id="8B9A36F3BF6CFF90FF50F8F37D2C6793" blockId="11.[151,1437,1919,2016]" pageId="11" pageNumber="372">
<emphasis id="B951EAE1BF6CFF90FF50F8F37F0267EB" bold="true" box="[151,274,1919,1944]" pageId="11" pageNumber="372">FIGURE 6.</emphasis>
<taxonomicName id="4C254D70BF6CFF90FEDFF80C7C9B67EB" authority="Semper, 1868" authorityName="Semper" authorityYear="1868" box="[280,651,1919,1944]" class="Holothuroidea" family="Sclerodactylidae" genus="Havelockia" kingdom="Animalia" order="Dendrochirotida" pageId="11" pageNumber="372" phylum="Echinodermata" rank="species" species="versicolor">
<emphasis id="B951EAE1BF6CFF90FEDFF80C7FE867EB" box="[280,504,1920,1944]" italics="true" pageId="11" pageNumber="372">Havelockia versicolor</emphasis>
Semper, 1868
</taxonomicName>
. NHMUK 1981.2.6.76. Hong Kong. A. Specimen entire; B. Part of calcareous ring; C. &amp; G. Tables of dorsal body wall; D. Tube foot end-plate; E. Tube feet ossicles, F. Rosette-like rods from tentacles. (All line drawings to same scale); MDIR = mid-dorsal interradial plate.
</paragraph>
</caption>
<paragraph id="8B9A36F3BF6BFF97FF00FD387F4C6348" blockId="12.[151,1437,153,1475]" pageId="12" pageNumber="373">
<emphasis id="B951EAE1BF6BFF97FF00FD387FEB62BC" bold="true" box="[199,507,692,719]" pageId="12" pageNumber="373">Other material examined.</emphasis>
The NHMUK material also contains four other specimens referable to this species, two from Cape D Aquilar (
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,
<geoCoordinate id="EE115034BF6BFF97FDB9FD557D026280" box="[638,786,729,755]" degrees="22" direction="north" minutes="12" orientation="latitude" pageId="12" pageNumber="373" precision="15" seconds="27" value="22.2075">22°1227” N</geoCoordinate>
), south-east
<collectingCountry id="F3327663BF6BFF97FC65FD557A396280" box="[930,1065,729,755]" name="Hong Kong" pageId="12" pageNumber="373">Hong Kong</collectingCountry>
, and another two from the Great Barrier Reef. For comparison with the
<collectingCountry id="F3327663BF6BFF97FD92FD717CCA6364" box="[597,730,765,791]" name="Hong Kong" pageId="12" pageNumber="373">Hong Kong</collectingCountry>
specimens described above the latter specimens are also here briefly described.
</paragraph>
<paragraph id="8B9A36F3BF6BFF97FF00FCC97A61632C" blockId="12.[151,1437,153,1475]" box="[199,1137,836,863]" pageId="12" pageNumber="373">
1974.12.3.38, GBR, off
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coast,
<geoCoordinate id="EE115034BF6BFF97FD6DFCC97D3B632C" box="[682,811,837,863]" degrees="16" direction="south" minutes="40" orientation="latitude" pageId="12" pageNumber="373" precision="15" seconds="4" value="-16.667778">16º 40 4”S</geoCoordinate>
,
<geoCoordinate id="EE115034BF6BFF97FCFFFCC87DC0632C" box="[824,976,836,863]" degrees="145" direction="east" minutes="59" orientation="longitude" pageId="12" pageNumber="373" precision="15" seconds="1" value="145.98361">145º 59 1” E</geoCoordinate>
,
<quantity id="4CDD9B16BF6BFF97FC1BFCC97A07632C" box="[988,1047,837,863]" metricMagnitude="1" metricUnit="m" metricValue="2.4" pageId="12" pageNumber="373" unit="m" value="24.0">24 m</quantity>
,
<specimenCount id="9D23FD7ABF6BFF97FBE4FCC97A61632C" box="[1059,1137,837,863]" pageId="12" pageNumber="373" type="generic">1 spec.</specimenCount>
</paragraph>
<paragraph id="8B9A36F3BF6BFF97FF00FCE57B1863F0" blockId="12.[151,1437,153,1475]" box="[199,1288,872,899]" pageId="12" pageNumber="373">
1974.12.3.39, off
<collectingRegion id="49E1F811BF6BFF97FE48FCE57C0363F0" box="[399,531,873,899]" country="Australia" name="Queensland" pageId="12" pageNumber="373">Queensland</collectingRegion>
cost,
<geoCoordinate id="EE115034BF6BFF97FD91FCE57CF063F0" box="[598,736,873,899]" degrees="14" direction="south" minutes="24" orientation="latitude" pageId="12" pageNumber="373" precision="15" seconds="9" value="-14.402499">14° 24 9” S</geoCoordinate>
,
<geoCoordinate id="EE115034BF6BFF97FD2AFCE57D9863F0" box="[749,904,872,899]" degrees="144" direction="east" minutes="50" orientation="longitude" pageId="12" pageNumber="373" precision="15" seconds="1" value="144.8336">144° 50 1” E</geoCoordinate>
,
<quantity id="4CDD9B16BF6BFF97FC53FCE57DDF63F0" box="[916,975,873,899]" metricMagnitude="1" metricUnit="m" metricValue="1.6" pageId="12" pageNumber="373" unit="m" value="16.0">16 m</quantity>
, GBR exp., P. Gibb,
<specimenCount id="9D23FD7ABF6BFF97FB7EFCE57B1863F0" box="[1209,1288,873,899]" pageId="12" pageNumber="373" type="generic">1 spec.</specimenCount>
</paragraph>
</subSubSection>
<subSubSection id="C33F6578BF6BFF97FF00FC007F736540" pageId="12" pageNumber="373" type="description">
<paragraph id="8B9A36F3BF6BFF97FF00FC007F1C6444" blockId="12.[151,1437,153,1475]" pageId="12" pageNumber="373">
<emphasis id="B951EAE1BF6BFF97FF00FC007F4163D4" bold="true" box="[199,337,908,935]" pageId="12" pageNumber="373">Description</emphasis>
. Specimen (1974.12.3.38) cream-coloured, cucumber-shaped, larger one
<quantity id="4CDD9B16BF6BFF97FB42FC017AC363D4" box="[1157,1235,909,935]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" pageId="12" pageNumber="373" unit="mm" value="16.0">16 mm</quantity>
in length, breadth in mid-body
<quantity id="4CDD9B16BF6BFF97FEEEFC3D7F7E63B8" box="[297,366,945,971]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="12" pageNumber="373" unit="mm" value="7.0">7 mm</quantity>
. Mouth and anus terminal, the latter with poorly calcified (more or less papilliform) anal teeth. Tentacles retracted. Tube feet long and scattered, mostly in ambulacra but some also in interambulacra. Specimen not dissected and hence introvert, the number of tentacles and their deposits and other associated structures not examined.
</paragraph>
<paragraph id="8B9A36F3BF6BFF97FF00FBCD7F736540" blockId="12.[151,1437,153,1475]" pageId="12" pageNumber="373">
Specimen (1974.12.3.39) well preserved. Colour cream to white, with some podia darker than others, perhaps implying that live colouration lost. Length approximately
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, breadth in mid-body
<quantity id="4CDD9B16BF6BFF97FBB1FBE87AAB640C" box="[1142,1211,1124,1151]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="12" pageNumber="373" unit="mm" value="5.0">5 mm</quantity>
. Tube feet scattered but more numerous in ambulacra, both dorsally and ventrally, all appreciably long with suckers of same diameter. Anal teeth reduced. Specimen lacks calcareous ring and associated structures. Ossicles of body wall comprise twopillared tables with smooth disc bearing 810 holes. Ossicles of tube feet tables with arched disc bearing four holes and a spire ending in 23 teeth. End plates with large holes on periphery of plate and smaller holes concentrated in the central region.
</paragraph>
</subSubSection>
<subSubSection id="C33F6578BF6BFF97FF00FAB17F2265B0" pageId="12" pageNumber="373" type="discussion">
<paragraph id="8B9A36F3BF6BFF97FF00FAB17F2265B0" blockId="12.[151,1437,153,1475]" pageId="12" pageNumber="373">
<emphasis id="B951EAE1BF6BFF97FF00FAB17F2B6524" bold="true" box="[199,315,1341,1367]" pageId="12" pageNumber="373">Remarks.</emphasis>
From the form of body-wall ossicles and tube feet end plates, there is no doubt that these specimens also represent
<taxonomicName id="4C254D70BF6BFF97FEF9FAED7C296508" authorityName="Semper" authorityYear="1868" box="[318,569,1377,1403]" class="Holothuroidea" family="Sclerodactylidae" genus="Havelockia" kingdom="Animalia" order="Dendrochirotida" pageId="12" pageNumber="373" phylum="Echinodermata" rank="species" species="versicolor">
<emphasis id="B951EAE1BF6BFF97FEF9FAED7C296508" box="[318,569,1377,1403]" italics="true" pageId="12" pageNumber="373">Havelockia versicolor</emphasis>
</taxonomicName>
. The four specimens described above add substantially to the single extant record in the NHMUK online catalogue, based on a specimen from
<collectingRegion id="49E1F811BF6BFF97FC4DFA097DC165EC" box="[906,977,1413,1439]" country="Philippines" name="Bohol" pageId="12" pageNumber="373">Bohol</collectingRegion>
,
<collectingCountry id="F3327663BF6BFF97FC1BFA097A4D65EC" box="[988,1117,1413,1439]" name="Philippines" pageId="12" pageNumber="373">Philippines</collectingCountry>
, which is perhaps part of the
<typeStatus id="549E8851BF6BFF97FF50FA257ED765B0" box="[151,199,1449,1475]" pageId="12" pageNumber="373">type</typeStatus>
material.
</paragraph>
</subSubSection>
</treatment>
</document>