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<document id="9C507E6C249E9D282C3D44EAE8B715D9" ID-CLB-Dataset="25163" ID-DOI="10.11646/zootaxa.4688.3.3" ID-GBIF-Dataset="a9af11dc-d705-48ed-96fb-3b323accc19e" ID-ISSN="1175-5326" ID-Zenodo-Dep="3516791" ID-ZooBank="B8F6F528-F616-4EA1-8CA7-FA2FA89A34B9" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1571832732088" checkinUser="plazi" docAuthor="Thandar, Ahmed S." docDate="2019" docId="038C87E5BF63FF9CFF50F9607A6160A4" docLanguage="en" docName="zootaxa.4688.3.3.pdf" docOrigin="Zootaxa 4688 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Stolus canescens" docType="treatment" docVersion="9" lastPageNumber="368" masterDocId="FFB5FF9DBF67FF9BFFC7FF8C7E106073" masterDocTitle="On some lesser known sea cucumbers in the Natural History Museum, UK (Echinodermata: Holothuroidea)" masterLastPageNumber="381" masterPageNumber="361" pageNumber="365" updateTime="1698760707621" updateUser="ExternalLinkService">
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<mods:title id="4BE0350DADD38B89D1D6E7A7F0F05B5A">On some lesser known sea cucumbers in the Natural History Museum, UK (Echinodermata: Holothuroidea)</mods:title>
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<treatment id="038C87E5BF63FF9CFF50F9607A6160A4" ID-DOI="http://doi.org/10.5281/zenodo.5687796" ID-GBIF-Taxon="159812122" ID-Zenodo-Dep="5687796" LSID="urn:lsid:plazi:treatment:038C87E5BF63FF9CFF50F9607A6160A4" httpUri="http://treatment.plazi.org/id/038C87E5BF63FF9CFF50F9607A6160A4" lastPageId="7" lastPageNumber="368" pageId="4" pageNumber="365">
<subSubSection id="C33F6578BF63FF9FFF50F9607C356775" box="[151,549,1772,1799]" pageId="4" pageNumber="365" type="nomenclature">
<paragraph id="8B9A36F3BF63FF9FFF50F9607C356775" blockId="4.[151,549,1772,1835]" box="[151,549,1772,1799]" pageId="4" pageNumber="365">
<heading id="D0D2819FBF63FF9FFF50F9607C356775" bold="true" box="[151,549,1772,1799]" fontSize="11" level="1" pageId="4" pageNumber="365" reason="1">
<taxonomicName id="4C254D70BF63FF9FFF50F9607C356775" ID-CoL="52TKX" authority="(Semper, 1867)" baseAuthorityName="Semper" baseAuthorityYear="1867" box="[151,549,1772,1799]" class="Holothuroidea" family="Phyllophoridae" genus="Stolus" kingdom="Animalia" order="Dendrochirotida" pageId="4" pageNumber="365" phylum="Echinodermata" rank="species" species="canescens">
<emphasis id="B951EAE1BF63FF9FFF50F9607C356775" bold="true" box="[151,549,1772,1799]" pageId="4" pageNumber="365">
<emphasis id="B951EAE1BF63FF9FFF50F9607F746774" bold="true" box="[151,356,1772,1799]" italics="true" pageId="4" pageNumber="365">Stolus canescens</emphasis>
(
<bibRefCitation id="EFB44B02BF63FF9FFEB4F9607C0D6775" author="Semper, C." box="[371,541,1772,1798]" pageId="4" pageNumber="365" refId="ref12610" refString="Semper, C. (1867) Holothurien. Reisen im Archipel der Philippen. Holothurien. 2. Wissenschaftliche Resultate. Wiesbaden, Leipzig, 288 pp. https: // doi. org / 10.5962 / bhl. title. 11687" type="book" year="1867">Semper, 1867</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<paragraph id="8B9A36F3BF63FF9FFF50F89C7F226758" blockId="4.[151,549,1772,1835]" box="[151,306,1808,1835]" pageId="4" pageNumber="365">
<figureCitation id="131E2A76BF63FF9FFF50F89C7F106758" box="[151,256,1808,1835]" captionStart="FIGURE 2" captionStartId="5.[151,250,1932,1957]" captionTargetBox="[174,1401,166,1907]" captionTargetId="figure@5.[174,1414,166,1912]" captionTargetPageId="5" captionText="FIGURE 2. Stolus canescens Semper, 1868. Admirality Is. NHMUK 1892.1.14.27. A. Specimen entire; B. Part of calcareous ring; C. Introvert ossicles; D. Rods from tentacle branches; E. Ossicles from tentacle stalk; F. Buttons and plates from body wall; G. Tube feet tables. (All ossicles drawn to same scale). MDIR = mid-dorsal interradial plate." figureDoi="http://doi.org/10.5281/zenodo.3516795" httpUri="https://zenodo.org/record/3516795/files/figure.png" pageId="4" pageNumber="365">Figures 2</figureCitation>
&amp;
<figureCitation id="131E2A76BF63FF9FFEE3F89D7F226758" box="[292,306,1809,1835]" captionStart="FIGURE 3" captionStartId="6.[151,250,878,903]" captionTargetBox="[218,1369,166,858]" captionTargetId="figure@6.[218,1369,166,858]" captionTargetPageId="6" captionText="FIGURE 3. Scanning electron micrograph of ossicles of Stolus canescens Semper, 1868.Admirality Is. NHMUK 1892.1.14.27. A. Tentacle ossicles; B. Tube feet tables; C. Tube foot end-plate; D. Body wall ossicles." figureDoi="http://doi.org/10.5281/zenodo.3516797" httpUri="https://zenodo.org/record/3516797/files/figure.png" pageId="4" pageNumber="365">3</figureCitation>
</paragraph>
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<subSubSection id="C33F6578BF63FF9FFF50F8D97AD367FF" pageId="4" pageNumber="365" type="reference_group">
<paragraph id="8B9A36F3BF63FF9FFF50F8D97A2D671E" blockId="4.[151,1219,1876,1932]" box="[151,1085,1876,1901]" pageId="4" pageNumber="365">
<treatmentCitationGroup id="AB3511DDBF63FF9FFF50F8D97A2D671E" box="[151,1085,1876,1901]" pageId="4" pageNumber="365">
<taxonomicName id="4C254D70BF63FF9FFF50F8D97C22671E" authority="Semper, 1867: 48" authorityName="Semper" authorityPageNumber="48" authorityYear="1867" box="[151,562,1876,1901]" class="Holothuroidea" family="Cucumariidae" genus="Cucumaria" kingdom="Animalia" order="Dendrochirotida" pageId="4" pageNumber="365" phylum="Echinodermata" rank="species" species="canescens">
<emphasis id="B951EAE1BF63FF9FFF50F8D97F67671E" box="[151,375,1877,1901]" italics="true" pageId="4" pageNumber="365">Cucumaria canescens</emphasis>
<treatmentCitation id="0A8410E2BF63FF9FFEBAF8D97C22671E" author="Semper, C." box="[381,562,1876,1901]" page="48" pageId="4" pageNumber="365" year="1867">
<bibRefCitation id="EFB44B02BF63FF9FFEBAF8D97C22671E" author="Semper, C." box="[381,562,1876,1901]" pageId="4" pageNumber="365" refId="ref12610" refString="Semper, C. (1867) Holothurien. Reisen im Archipel der Philippen. Holothurien. 2. Wissenschaftliche Resultate. Wiesbaden, Leipzig, 288 pp. https: // doi. org / 10.5962 / bhl. title. 11687" type="book" year="1867">Semper, 1867: 48</bibRefCitation>
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, pl. 13, fig. 6; pl. 14, figs. 3, 9, 10; pl. 15, figs. 23.
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</paragraph>
<paragraph id="8B9A36F3BF63FF9FFF50F8FF7AD367FF" blockId="4.[151,1219,1876,1932]" box="[151,1219,1907,1932]" pageId="4" pageNumber="365">
<treatmentCitationGroup id="AB3511DDBF63FF9FFF50F8FF7AD367FF" box="[151,1219,1907,1932]" pageId="4" pageNumber="365">
<taxonomicName id="4C254D70BF63FF9FFF50F8FF7C4167F8" ID-CoL="52TKX" authority="Clark &amp; Rowe, 1971: 182" authorityName="Clark &amp; Rowe" authorityPageNumber="182" authorityYear="1971" box="[151,593,1907,1932]" class="Holothuroidea" family="Phyllophoridae" genus="Stolus" kingdom="Animalia" order="Dendrochirotida" pageId="4" pageNumber="365" phylum="Echinodermata" rank="species" species="canescens">
<emphasis id="B951EAE1BF63FF9FFF50F8FF7F5267F8" box="[151,322,1907,1931]" italics="true" pageId="4" pageNumber="365">Stolus canescens</emphasis>
<treatmentCitation id="0A8410E2BF63FF9FFE8FF8FF7C4167F8" author="Clark, H. L. &amp; Rowe, F. W. E." box="[328,593,1907,1932]" page="182" pageId="4" pageNumber="365" year="1971">
<bibRefCitation id="EFB44B02BF63FF9FFE8FF8FF7C4167F8" author="Clark, H. L. &amp; Rowe, F. W. E." box="[328,593,1907,1932]" pageId="4" pageNumber="365" pagination="171 - 209" refId="ref11100" refString="Clark, H. L. &amp; Rowe, F. W. E. (1971) Monograph of shallow-water Indo-West Pacific echinoderms. British Museum, London, 39 pp. [pp. 171 - 209]" type="book chapter" year="1971">Clark &amp; Rowe, 1971: 182</bibRefCitation>
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</taxonomicName>
, pl. 29, fig. 7;
<treatmentCitation id="0A8410E2BF63FF9FFD22F8FF7DF167F8" author="Liao, Y. &amp; Clark, A. M." box="[741,993,1907,1932]" page="497" pageId="4" pageNumber="365" year="1995">
<bibRefCitation id="EFB44B02BF63FF9FFD22F8FF7DF167F8" author="Liao, Y. &amp; Clark, A. M." box="[741,993,1907,1932]" pageId="4" pageNumber="365" pagination="422 - 613" refId="ref11759" refString="Liao, Y. &amp; Clark, A. M. (1995) The echinoderms of southern China. Science Press, Beijing, 422 - 613." type="book chapter" year="1995">Liao &amp; Clark, 1995: 497</bibRefCitation>
</treatmentCitation>
, fig. 301, pl. 13, fig.6.
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</paragraph>
</subSubSection>
<subSubSection id="C33F6578BF63FF9FFF50F8357F52678B" pageId="4" pageNumber="365" type="materials_examined">
<paragraph id="8B9A36F3BF63FF9FFF50F8357F52678B" blockId="4.[151,1437,1977,2040]" pageId="4" pageNumber="365">
<emphasis id="B951EAE1BF63FF9FFF50F8357F6A67A7" bold="true" box="[151,378,1977,2004]" pageId="4" pageNumber="365">Material examined</emphasis>
.
<materialsCitation id="3B4D3CAEBF63FF9FFE4FF8367F52678B" ID-GBIF-Occurrence="2429072688" collectionCode="NHMUK" latitude="-13.416667" longLatPrecision="1291" longitude="126.0" pageId="4" pageNumber="365" specimenCode="NHMUK 1892.1" specimenCount="1">
<specimenCode id="DB839E88BF63FF9FFE4FF8367C5967A7" box="[392,585,1978,2004]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="4" pageNumber="365" type="Museum">NHMUK 1892.1</specimenCode>
.14.27,
<taxonomicName id="4C254D70BF63FF9FFD5DF8367D4967A7" authority="Bank" authorityName="Bank" box="[666,857,1978,2004]" class="Holothuroidea" family="Holothuriidae" genus="Holothuria" kingdom="Animalia" order="Aspidochirotida" pageId="4" pageNumber="365" phylum="Echinodermata" rank="genus">
<emphasis id="B951EAE1BF63FF9FFD5DF8367D0667A7" box="[666,790,1978,2004]" italics="true" pageId="4" pageNumber="365">Holothuria</emphasis>
Bank
</taxonomicName>
(
<geoCoordinate id="EE115034BF63FF9FFCAEF8367DD867A7" box="[873,968,1977,2004]" degrees="13" direction="south" minutes="25" orientation="latitude" pageId="4" pageNumber="365" precision="925" value="-13.416667">13 25 S</geoCoordinate>
,
<geoCoordinate id="EE115034BF63FF9FFC12F8367A2767A7" box="[981,1079,1978,2004]" degrees="126" direction="east" minutes="0" orientation="longitude" pageId="4" pageNumber="365" precision="925" value="126.0">126 0 E</geoCoordinate>
), presented by Lords of the Admiralty,
<specimenCount id="9D23FD7ABF63FF9FFF33F8527F52678B" box="[244,322,2014,2040]" pageId="4" pageNumber="365" type="generic">1 spec.</specimenCount>
</materialsCitation>
</paragraph>
</subSubSection>
<caption id="DF5A667BBF62FF9EFF50F8007A52679E" ID-DOI="http://doi.org/10.5281/zenodo.3516795" ID-Zenodo-Dep="3516795" httpUri="https://zenodo.org/record/3516795/files/figure.png" pageId="5" pageNumber="366" startId="5.[151,250,1932,1957]" targetBox="[174,1401,166,1907]" targetPageId="5">
<paragraph id="8B9A36F3BF62FF9EFF50F8007A52679E" blockId="5.[151,1436,1932,2029]" pageId="5" pageNumber="366">
<emphasis id="B951EAE1BF62FF9EFF50F8007F0467D6" bold="true" box="[151,276,1932,1957]" pageId="5" pageNumber="366">FIGURE 2.</emphasis>
<taxonomicName id="4C254D70BF62FF9EFEDCF8017C4E67D6" authority="Semper, 1868" authorityName="Semper" authorityYear="1868" box="[283,606,1932,1957]" class="Holothuroidea" family="Phyllophoridae" genus="Stolus" kingdom="Animalia" order="Dendrochirotida" pageId="5" pageNumber="366" phylum="Echinodermata" rank="species" species="canescens">
<emphasis id="B951EAE1BF62FF9EFEDCF8017FD767D7" box="[283,455,1933,1957]" italics="true" pageId="5" pageNumber="366">Stolus canescens</emphasis>
Semper, 1868
</taxonomicName>
. Admirality Is. NHMUK 1892.1.14.27. A. Specimen entire; B. Part of calcareous ring; C. Introvert ossicles; D. Rods from tentacle branches; E. Ossicles from tentacle stalk; F. Buttons and plates from body wall; G. Tube feet tables. (All ossicles drawn to same scale). MDIR = mid-dorsal interradial plate.
</paragraph>
</caption>
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<paragraph id="8B9A36F3BF61FF9DFF50FCE27A1F63D8" blockId="6.[151,1437,878,939]" pageId="6" pageNumber="367">
<emphasis id="B951EAE1BF61FF9DFF50FCE27F0263F4" bold="true" box="[151,274,878,903]" pageId="6" pageNumber="367">FIGURE 3.</emphasis>
Scanning electron micrograph of ossicles of
<taxonomicName id="4C254D70BF61FF9DFD12FCE27A0263F4" authority="Semper, 1868" authorityName="Semper" authorityYear="1868" box="[725,1042,878,903]" class="Holothuroidea" family="Phyllophoridae" genus="Stolus" kingdom="Animalia" order="Dendrochirotida" pageId="6" pageNumber="367" phylum="Echinodermata" rank="species" species="canescens">
<emphasis id="B951EAE1BF61FF9DFD12FCE27D6F63F5" box="[725,895,878,902]" italics="true" pageId="6" pageNumber="367">Stolus canescens</emphasis>
Semper, 1868
</taxonomicName>
.Admirality Is. NHMUK 1892.1.14.27. A. Tentacle ossicles; B. Tube feet tables; C. Tube foot end-plate; D. Body wall ossicles.
</paragraph>
</caption>
<subSubSection id="C33F6578BF61FF9DFF00FC547B8D66B0" pageId="6" pageNumber="367" type="description">
<paragraph id="8B9A36F3BF61FF9DFF00FC547CA46544" blockId="6.[151,1437,984,2019]" pageId="6" pageNumber="367">
<emphasis id="B951EAE1BF61FF9DFF00FC547F416380" bold="true" box="[199,337,984,1011]" pageId="6" pageNumber="367">Description</emphasis>
. Specimen fusiform, pentagonal in cross section, U-shaped with an attenuated posterior end; length
<quantity id="4CDD9B16BF61FF9DFF50FC707EF96464" box="[151,233,1020,1047]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" pageId="6" pageNumber="367" unit="mm" value="35.0">35 mm</quantity>
, breadth in mid-body
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. Skin rigid, white in alcohol. Tube feet in rows only in ambulacra, ventral feet longer than dorsal, elongated, in double rows mid-ventrally but reducing to single rows anteriorly and posteriorly. Dorsal tube feet only in single rows per ambulacrum, with one or two also scattered in interambulacra dorso-laterally. Mouth and anus terminal. Anal teeth present but cannot be demonstrated with ease. Calcareous ring complex, radial and interradial plates asymmetrical and fused; radial plates deeply incised, i.e. bifurcating before the posterior margin of the interradial plates; interradial plates with triangular anterior edge; both radial and interradial plates with incipient fragmentation, posterior processes also fragmented. Polian vesicle single and saccate. Stone canal single, short, straight, embedded in dorsal mesentery; madreporite small, oval. Retractor muscles well developed, attached to anterior tip of radial plates of calcareous ring.
</paragraph>
<paragraph id="8B9A36F3BF61FF9DFF00FACD7B8D66B0" blockId="6.[151,1437,984,2019]" pageId="6" pageNumber="367">
Ossicles of body wall circular to oval table-like buttons with undulating margin (
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, average
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), perforated with 48 holes and 912 large knobs, with a half ring on one side and a spire on opposite side; spire swollen at base by the characteristic double knobs, usually ending in a single blunt point or usually a tripartite spire with a bi-nodular base bearing a central projection; some buttons without spire and/or half ring. Tube feet tables with curved disc usually perforated by a single hole at each end; spire two-pillared, with or without a horizontal bar, pillars fused for most of their length, ending in a single blunt point. Length of disc
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(average
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); height of spire ca.
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. Tentacle stalk contains three
<typeStatus id="549E8851BF61FF9DFD3FF9957D236640" box="[760,819,1561,1587]" pageId="6" pageNumber="367">types</typeStatus>
of ossicles: large rods (average length
<quantity id="4CDD9B16BF61FF9DFB25F9957B2B6640" box="[1250,1339,1561,1587]" metricMagnitude="-1" metricUnit="m" metricValue="1.3699999999999999" pageId="6" pageNumber="367" unit="mm" value="137.0">137 µm</quantity>
), minute rods (average length
<quantity id="4CDD9B16BF61FF9DFEB8F9B07FD86624" box="[383,456,1596,1623]" metricMagnitude="-2" metricUnit="m" metricValue="8.6" pageId="6" pageNumber="367" unit="mm" value="86.0">86 µm</quantity>
) and rosettes (average
<quantity id="4CDD9B16BF61FF9DFD01F9B17D016624" box="[710,785,1597,1623]" metricMagnitude="-2" metricUnit="m" metricValue="2.7" pageId="6" pageNumber="367" unit="mm" value="27.0">27 µm</quantity>
). Large rods of stalk perforated at ends with short marginal projections; minute rods smooth, rosettes of the closed
<typeStatus id="549E8851BF61FF9DFCCAF9ED7D516608" box="[781,833,1633,1659]" pageId="6" pageNumber="367">type</typeStatus>
. Tentacle branches with both medium to minute rods (
<quantity id="4CDD9B16BF61FF9DFF58F9087F0666EC" box="[159,278,1668,1695]" metricMagnitude="-2" metricUnit="m" metricValue="4.1" metricValueMax="5.6" metricValueMin="2.6" pageId="6" pageNumber="367" unit="mm" value="41.0" valueMax="56.0" valueMin="26.0">2656 µm</quantity>
, average
<quantity id="4CDD9B16BF61FF9DFE45F9097FDF66EC" box="[386,463,1669,1695]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="6" pageNumber="367" unit="mm" value="40.0">40 µm</quantity>
), medium-sized rods delicate, may be perforated at ends; slender rods minute, curved, usually perforated at ends, sometimes branched, branches with or without perforations. Introvert devoid of ossicles.
</paragraph>
</subSubSection>
<subSubSection id="C33F6578BF61FF9DFF00F9407F1F677F" pageId="6" pageNumber="367" type="distribution">
<paragraph id="8B9A36F3BF61FF9DFF00F9407F1F677F" blockId="6.[151,1437,984,2019]" pageId="6" pageNumber="367">
<emphasis id="B951EAE1BF61FF9DFF00F9407F496694" bold="true" box="[199,345,1740,1767]" pageId="6" pageNumber="367">Distribution</emphasis>
.
<collectingCountry id="F3327663BF61FF9DFEACF9417FA16694" box="[363,433,1741,1767]" name="China" pageId="6" pageNumber="367">China</collectingCountry>
,
<collectingCountry id="F3327663BF61FF9DFE78F9417C2D6694" box="[447,573,1741,1767]" name="Philippines" pageId="6" pageNumber="367">Philippines</collectingCountry>
and East Indies (
<bibRefCitation id="EFB44B02BF61FF9DFCCBF9417DFE6694" author="Liao, Y. &amp; Clark, A. M." box="[780,1006,1740,1767]" pageId="6" pageNumber="367" pagination="422 - 613" refId="ref11759" refString="Liao, Y. &amp; Clark, A. M. (1995) The echinoderms of southern China. Science Press, Beijing, 422 - 613." type="book chapter" year="1995">Liao &amp; Clark 1995</bibRefCitation>
) and now
<taxonomicName id="4C254D70BF61FF9DFBA9F9417B3D6694" authority="Bank" authorityName="Bank" box="[1134,1325,1741,1767]" class="Holothuroidea" family="Holothuriidae" genus="Holothuria" kingdom="Animalia" order="Aspidochirotida" pageId="6" pageNumber="367" phylum="Echinodermata" rank="genus">Holothuria Bank</taxonomicName>
(Western Australia).
</paragraph>
</subSubSection>
<subSubSection id="C33F6578BF61FF9CFF00F8997A6160A4" lastPageId="7" lastPageNumber="368" pageId="6" pageNumber="367" type="discussion">
<paragraph id="8B9A36F3BF61FF9DFF00F8997C496790" blockId="6.[151,1437,984,2019]" pageId="6" pageNumber="367">
<emphasis id="B951EAE1BF61FF9DFF00F8997F24675C" bold="true" box="[199,308,1813,1839]" pageId="6" pageNumber="367">Remarks</emphasis>
. This is a well-known West-Pacific form originally described by Semper (1868) from
<collectingRegion id="49E1F811BF61FF9DFB3AF8997B54675C" box="[1277,1348,1813,1839]" country="Philippines" name="Bohol" pageId="6" pageNumber="367">Bohol</collectingRegion>
,
<collectingCountry id="F3327663BF61FF9DFA97F8997EC66720" name="Philippines" pageId="6" pageNumber="367">Philippines</collectingCountry>
. It is well illustrated by Semper and more recently by
<bibRefCitation id="EFB44B02BF61FF9DFC83F8B57A206720" author="Liao, Y. &amp; Clark, A. M." box="[836,1072,1848,1875]" pageId="6" pageNumber="367" pagination="422 - 613" refId="ref11759" refString="Liao, Y. &amp; Clark, A. M. (1995) The echinoderms of southern China. Science Press, Beijing, 422 - 613." type="book chapter" year="1995">Liao &amp; Clark (1995)</bibRefCitation>
and its distribution recorded by
<bibRefCitation id="EFB44B02BF61FF9DFF50F8D17F9C6704" author="Clark, H. L. &amp; Rowe, F. W. E." box="[151,396,1885,1911]" pageId="6" pageNumber="367" pagination="171 - 209" refId="ref11100" refString="Clark, H. L. &amp; Rowe, F. W. E. (1971) Monograph of shallow-water Indo-West Pacific echinoderms. British Museum, London, 39 pp. [pp. 171 - 209]" type="book chapter" year="1971">Clark &amp; Rowe (1971)</bibRefCitation>
and
<bibRefCitation id="EFB44B02BF61FF9DFE04F8D17CAD6704" author="Rowe, F. W. E. &amp; Gates, J." box="[451,701,1884,1911]" pageId="6" pageNumber="367" pagination="1 - 510" refId="ref12560" refString="Rowe, F. W. E. &amp; Gates, J. (1995) Echinodermata. In: Wells, A. (Ed.), Zoological Catalogue of Australia. Vol. 33. CSIRO, Melbourne, pp. i-xiii + 1 - 510." type="book chapter" year="1995">Rowe &amp; Gates (1995)</bibRefCitation>
. However, judging from the literature it has not been encountered frequently. Its most characteristic feature are the large knobs on the buttons, a varying number of marginal knobs and usually a tripartite spire with a bi-nodular base bearing a central projection.
<bibRefCitation id="EFB44B02BF61FF9DFBE4F8297AC367CC" author="Thandar, A. S." box="[1059,1235,1956,1983]" pageId="6" pageNumber="367" pagination="115 - 126" refId="ref12798" refString="Thandar, A. S. (2005) Two new species of Stolus Selenka (Echinodermata: Holothuroidea: Dendrochirotida: Phyllophoridae) from off the east coast of South Africa, with a revised key to the genus. African Zoology, 40 (1), 115 - 126." type="journal article" year="2005">Thandar (2005)</bibRefCitation>
provides a useful key to the species of
<taxonomicName id="4C254D70BF61FF9DFE44F8457FD76790" authorityName="Selenka" authorityYear="1867" box="[387,455,1993,2019]" class="Holothuroidea" family="Phyllophoridae" genus="Stolus" kingdom="Animalia" order="Dendrochirotida" pageId="6" pageNumber="367" phylum="Echinodermata" rank="genus">
<emphasis id="B951EAE1BF61FF9DFE44F8457FD76790" box="[387,455,1993,2019]" italics="true" pageId="6" pageNumber="367">Stolus</emphasis>
</taxonomicName>
then known.
</paragraph>
<paragraph id="8B9A36F3BF60FF9CFF00FF157A6160A4" blockId="7.[151,1436,153,215]" pageId="7" pageNumber="368">
The above Admiralty Island record is the first record of the species with locality data in the NHMUK online catalogue, last updated
<date id="FF9B1033BF60FF9CFE59FF317C3060A4" box="[414,544,189,215]" pageId="7" pageNumber="368" value="2017-03-02">03.02.2017</date>
. The existing record is without locality or other data.
</paragraph>
</subSubSection>
</treatment>
</document>