447 lines
57 KiB
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447 lines
57 KiB
XML
<document id="1A9DFB86273E1741C5F6BF5F48CF95A5" ID-DOI="10.5281/zenodo.5741842" ID-ISSN="1536-9307" ID-Zenodo-Dep="5741842" ID-ZooBank="DCAC2354-0168-4A66-AC36-87F1BB19EAA2" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638266231204" checkinUser="felipe" docAuthor="Kovařík, František, Lowe, Graeme, Stockmann, Mark & Šťáhlavský, František" docDate="2020" docId="038C87D0FFD55040FC32FA5FFB7E2DA5" docLanguage="en" docName="Euscorpius.325.1-140.pdf" docOrigin="Euscorpius 325" docStyle="DocumentStyle:5E96F6379CE20B0E8E4F9E7DF8E1C0DF.2:Euscorpius.2020-.journal_article" docStyleId="5E96F6379CE20B0E8E4F9E7DF8E1C0DF" docStyleName="Euscorpius.2020-.journal_article" docStyleVersion="2" docTitle="Scorpiops birulai Kovařík & Lowe & Stockmann & Šťáhlavský 2020, sp. n." docType="treatment" docUuid="FA2B49A2-6B88-4428-A1B0-DB6B75033D5B" docUuidSource="ZooBank" docVersion="3" lastPageNumber="48" masterDocId="FFB5FFA8FFF95071FFDFFFEDFFD72E46" masterDocTitle="Revision of genus-group taxa in the family Scorpiopidae Kraepelin, 1905, with description of 15 new species (Arachnida Scorpiones)" masterLastPageNumber="140" masterPageNumber="1" pageNumber="43" updateTime="1699436806083" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="3E124F59A568FB4027A5BA8F3B86B0E0">Revision of genus-group taxa in the family Scorpiopidae Kraepelin, 1905, with description of 15 new species (Arachnida Scorpiones)</mods:title>
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<taxonomicName id="4C254D45FFD5505DFC32FA5FFB6A2B8B" authority="Kovařík & Lowe & Stockmann & Šťáhlavský, 2020" authorityName="Kovařík & Lowe & Stockmann & Šťáhlavský" authorityYear="2020" box="[1005,1213,1458,1485]" class="Arachnida" family="Euscorpiidae" genus="Scorpiops" kingdom="Animalia" order="Scorpiones" pageId="44" pageNumber="43" phylum="Arthropoda" rank="species" species="birulai" status="sp. nov.">Scorpiops birulai</taxonomicName>
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(
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<figureCitation id="131E2A43FFD5505DFCE2FA39FC682BAA" box="[829,959,1492,1516]" captionStart="Figures 149–163" captionStartId="27.[116,194,1896,1918]" captionTargetBox="[114,1470,267,1796]" captionTargetId="figure-109@27.[113,1474,253,1836]" captionTargetPageId="27" captionText="Figures 149–163: Whole hemispermatophores of 15 scorpiopids. Convex (external) aspect of hemispermatophores of: Scorpiops oligotrichus (149), S. citadelle (150), S. sherwoodae sp. n. (151), S. montanus (152), S. pakseensis sp. n. (153), S. leptochirus (154), S. phatoensis sp. n. (155), S. schumacheri (156), S. wongpromi (157), S. kautti sp. n. (158), S. birulai sp. n. (159), S. anthracinus (160), S. thaomischi (161), S. dii sp. n. (162) and S. problematicus (163). Specimen numbers indicated. Scale bars: 2 mm (149–154, 157–161, 163), 1 mm (155–156, 162)." figureDoi="http://doi.org/10.5281/zenodo.5741869" httpUri="https://zenodo.org/record/5741869/files/figure.png" pageId="44" pageNumber="43">Figures 159</figureCitation>
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<figureCitation id="131E2A43FFD5505DFC14FA39FC222BAA" box="[971,1013,1492,1516]" captionStart="Figures 170–185" captionStartId="29.[113,190,1905,1927]" captionTargetBox="[121,1449,242,1879]" captionTargetId="figure-121@29.[113,1474,223,1901]" captionTargetPageId="29" captionText="Figures 170–185: Hemispermatophore capsules of 16 scorpiopids. Convex aspect of capsules of: Scorpiops oligotrichus (170), S. citadelle (171), S. sherwoodae sp. n. (172), S. montanus (173), S. pakseensis sp. n. (174), S. leptochirus (175), S. phatoensis sp. n. (176), S. schumacheri (177), S. wongpromi (178), S. kautti sp. n. (179), S. krabiensis sp. n. (180), S. birulai sp. n. (181), S. anthracinus (182), S. thaomischi (183), S. dii sp. n. (184) and S. problematicus (185). Specimen number of S. krabiensis 1829, of others as indicated in Figs. 149–163. Scale bars: 400 μm (170–175, 178–185), 200 μm (176–177)." figureDoi="http://doi.org/10.5281/zenodo.5741875" httpUri="https://zenodo.org/record/5741875/files/figure.png" pageId="44" pageNumber="43">181</figureCitation>
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<figureCitation id="131E2A43FFD5505DFBDFFA39FBFD2BAA" box="[1024,1066,1492,1516]" captionStart="Figure 186" captionStartId="30.[116,184,1815,1837]" captionTargetBox="[124,1456,261,1796]" captionTargetId="figure-14@30.[113,1474,234,1815]" captionTargetPageId="30" captionText="Figure 186: Dentition of crown-like structures (cls) and distal posterior lobe (dpl) apex of 16 scorpiopids. Species are grouped according to four major clades resolved by DNA analysis (Šťáhlavský et al., in press). To highlight inconsistency of the trichobothrial classification with molecular phylogeny, previously described species are listed under genera (in quotes) where they were originally placed according to trichobothriotaxic characters of Soleglad & Fet (2001). Cited ranges of cls denticle counts for each species represent data from both left and right hemispermatophores extracted from all available specimens: N =1, except for “Alloscorpiops” wongpromi (N = 2), “A.” anthracinus (N = 2), Scorpiops thaomischi (N = 2), and “Euscorpiops” problematicus (N = 3). Data with N> 1 reveal intraspecific stability of cls dentition patterns, and intraspecific variability of dpl denticulation patterns. Right insets: convex (upper inset) and concave (lower inset) views of whole hemispermatophore of Scorpiops kautti sp. n. Anatomical structures and measurements are indicated. Scale bar: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5741877" httpUri="https://zenodo.org/record/5741877/files/figure.png" pageId="44" pageNumber="43">186</figureCitation>
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<figureCitation id="131E2A43FFD5505DFBEAFA39FB412BAA" box="[1077,1174,1492,1516]" captionStart-0="Figures 331–336" captionStart-1="Figures 337–346" captionStart-2="Figures 347–358" captionStartId-0="45.[115,192,1979,2001]" captionStartId-1="46.[116,193,1965,1987]" captionStartId-2="47.[116,193,1948,1970]" captionTargetBox-0="[119,1470,262,1938]" captionTargetBox-1="[122,1465,230,1951]" captionTargetBox-2="[113,1474,238,1923]" captionTargetId-0="figure-9@45.[113,1474,249,1943]" captionTargetId-1="figure-14@46.[113,1474,227,1958]" captionTargetId-2="figure-8@47.[113,1474,238,1923]" captionTargetPageId-0="45" captionTargetPageId-1="46" captionTargetPageId-2="47" captionText-0="Figures 331–336: Scorpiops birulai sp. n., male holotype. Figures 331–332. dorsal (331) and ventral (332) views. Figures 333–336. Telson lateral (333), and metasoma and telson lateral (334), ventral (335), and dorsal (336) views. Scale bars: 10 mm (331–332, 334–336)." captionText-1="Figures 337–346: Scorpiops birulai sp. n., male holotype, pedipalp segments. Chela dorsal (337), external (338) and ventral (339) views. Patella dorsal (340), external (341) and ventral (342) views. Femur and trochanter dorsal (343), and ventral (344) views. Fixed (345) and movable (346) finger dentition. Trichobothrial pattern is indicated by white circles (337a–342a)." captionText-2="Figures 347–358: Figures 347–352. Scorpiops birulai sp. n., male holotype, carapace and tergites I–IV (347), coxosternal area and sternites (348), left legs I–IV, retrolateral aspect (349–352 respectively). Figures 353–358. Scorpiops sherwoodae sp. n., male holotype, carapace and tergites I–IV (353), coxosternal area and sternites (354), left legs I–IV, retrolateral aspect (355–358 respectively)." figureDoi-0="http://doi.org/10.5281/zenodo.5741895" figureDoi-1="http://doi.org/10.5281/zenodo.5741897" figureDoi-2="http://doi.org/10.5281/zenodo.5741899" httpUri-0="https://zenodo.org/record/5741895/files/figure.png" httpUri-1="https://zenodo.org/record/5741897/files/figure.png" httpUri-2="https://zenodo.org/record/5741899/files/figure.png" pageId="44" pageNumber="43">331–352</figureCitation>
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<figureCitation id="131E2A43FFD5505DFB7FFA39FB1D2BAA" box="[1184,1226,1492,1516]" captionStart="Figures 359–362" captionStartId="48.[116,193,1992,2014]" captionTargetBox="[113,1474,232,1980]" captionTargetId="figure-14@48.[113,1474,229,1983]" captionTargetPageId="48" captionText="Figures 359–362. Thailand, type localities of Scorpiops birulai sp. n. (359), S. kautti sp. n. (360), S. krabiensis sp. n. (361), and S. phatoensis sp. n. (362)." figureDoi="http://doi.org/10.5281/zenodo.5741901" httpUri="https://zenodo.org/record/5741901/files/figure.png" pageId="44" pageNumber="43">359</figureCitation>
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<figureCitation id="131E2A43FFD5505DFB0AFA39FAD72BAA" box="[1237,1280,1492,1516]" captionStart="Figure 799" captionStartId="128.[117,185,1143,1165]" captionTargetBox="[113,1474,231,1335]" captionTargetId="figure-14@128.[113,1474,231,1337]" captionTargetPageId="128" captionText="Figure 799. Geographic distribution of the family Scorpiopidae. The map plots locality" figureDoi="http://doi.org/10.5281/zenodo.5742043" httpUri="https://zenodo.org/record/5742043/files/figure.png" pageId="44" pageNumber="43">799</figureCitation>
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,
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<figureCitation id="131E2A43FFD5505DFAD4FA39FAE22BAA" box="[1291,1333,1492,1516]" captionStart="Figures 800–807" captionStartId="131.[115,192,1873,1895]" captionTargetBox="[113,1474,249,1836]" captionTargetId="figure-8@131.[113,1474,249,1836]" captionTargetPageId="131" captionText="Figures 800–807: Male karyotypes of scorpiopid species based on postpachytene. Figure 800. Scorpiops anthracinus (2n=105, 51II+III). Figure 801. Scorpiops birulai sp. n. (2n=99, 96II+III). Figure 802. Scorpiops citadelle (2n=104, 52II). Figure 803. Scorpiops dii sp. n. (2n=109, 53II+III). Figure 804. Scorpiops kautti sp. n. (2n=87, 42II+III). Figure 805. Scorpiops krabiensis sp. n. (2n=81, 39II+III). Figure 806. Scorpiops leptochirus (2n=96, 48II). Figure 807. Scorpiops montanus (2n=70, 35II). Insets show the location of 18S rDNA (red signal) on the same chromosome as in the karyogram (ppach) and additionally on chromosomes during pachytene (pach) or metaphase II (met II). Abbreviation of postpachytene configuration: II – bivalent, III – trivalent. Scale bar: 10 μm." figureDoi="http://doi.org/10.5281/zenodo.5742045" httpUri="https://zenodo.org/record/5742045/files/figure.png" pageId="44" pageNumber="43">801</figureCitation>
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<tableCitation id="C6A7037DFFD5505DFA9FFA39FA4C2BAA" box="[1344,1435,1492,1516]" captionStart="Table 3" captionStartId="44.[117,173,1000,1021]" captionText="Table 3. Comparative measurements of adults of Scorpiops birulai sp. n., S. ciki sp. n., and S. dii sp. n. Abbreviations: length (L), width (W, in carapace it corresponds to posterior width), depth (D)." pageId="44" pageNumber="43">Tables 3</tableCitation>
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<location id="8EFA601DFFD5505DFC42F999FC0B28CA" LSID="urn:lsid:plazi:treatment:038C87D0FFD55040FC32FA5FFB7E2DA5:8EFA601DFFD5505DFC42F999FC0B28CA" box="[925,988,1652,1676]" country="Thailand" latitude="9.829299" longLatPrecision="1" longitude="98.77315" name="Phato" pageId="44" pageNumber="43" stateProvince="Chumphon">Phato</location>
|
||
,
|
||
<geoCoordinate id="EE115001FFD5505DFC38F999FBA128CA" box="[999,1142,1652,1676]" degrees="9.8292990" direction="north" orientation="latitude" pageId="44" pageNumber="43" precision="1" value="9.829299">9.8292990°N</geoCoordinate>
|
||
<geoCoordinate id="EE115001FFD5505DFBA2F999FACD28CA" box="[1149,1306,1652,1676]" degrees="98.7731480" direction="east" orientation="longitude" pageId="44" pageNumber="43" precision="1" value="98.77315">98.7731480°E</geoCoordinate>
|
||
</materialsCitation>
|
||
;
|
||
<collectionCode id="ED34AE03FFD5505DFAFAF999FABF28CA" box="[1317,1384,1652,1676]" pageId="44" pageNumber="43">FKCP</collectionCode>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9A36C6FFD5505DFCEFF958FCAF290A" blockId="44.[816,1474,1716,1868]" pageId="44" pageNumber="43">
|
||
<materialsCitation id="3B4D3C9BFFD5505DFCEFF958FCE8294A" collectingDate="2019-10-28" collectionCode="TYPE, MATERIAL" country="Thailand" elevation="222" latitude="9.829299" location="Phato" longLatPrecision="1" longitude="98.77315" pageId="44" pageNumber="43" specimenCount="1" stateProvince="Chumphon">
|
||
<collectionCode id="ED34AE03FFD5505DFCEFF958FC35288A" box="[816,994,1716,1740]" pageId="44" pageNumber="43">TYPE MATERIAL</collectionCode>
|
||
.
|
||
<emphasis id="B951EAD4FFD5505DFC26F959FBB6288A" bold="true" box="[1017,1121,1716,1740]" pageId="44" pageNumber="43">
|
||
<collectingCountry id="F3327656FFD5505DFC26F959FBB6288A" box="[1017,1121,1716,1740]" name="Thailand" pageId="44" pageNumber="43">Thailand</collectingCountry>
|
||
</emphasis>
|
||
,
|
||
<emphasis id="B951EAD4FFD5505DFBA4F959FAB3288A" box="[1147,1380,1716,1740]" italics="true" pageId="44" pageNumber="43">
|
||
<collectingRegion id="49E1F824FFD5505DFBA4F959FAB3288A" box="[1147,1380,1716,1740]" country="Thailand" name="Chumphon" pageId="44" pageNumber="43">Chumphon Province</collectingRegion>
|
||
</emphasis>
|
||
,
|
||
<location id="8EFA601DFFD5505DFAA0F959FA69288A" LSID="urn:lsid:plazi:treatment:038C87D0FFD55040FC32FA5FFB7E2DA5:8EFA601DFFD5505DFAA0F959FA69288A" box="[1407,1470,1716,1740]" country="Thailand" latitude="9.829299" longLatPrecision="1" longitude="98.77315" name="Phato" pageId="44" pageNumber="43" stateProvince="Chumphon">Phato</location>
|
||
,
|
||
<geoCoordinate id="EE115001FFD5505DFCEFF939FC6828AA" box="[816,959,1748,1772]" degrees="9.8292990" direction="north" orientation="latitude" pageId="44" pageNumber="43" precision="1" value="9.829299">9.8292990°N</geoCoordinate>
|
||
<geoCoordinate id="EE115001FFD5505DFC18F939FBB428AA" box="[967,1123,1748,1772]" degrees="98.7731480" direction="east" orientation="longitude" pageId="44" pageNumber="43" precision="1" value="98.77315">98.7731480°E</geoCoordinate>
|
||
,
|
||
<elevation id="0008D1F5FFD5505DFBB1F939FB2D28AA" box="[1134,1274,1748,1772]" metricMagnitude="2" metricUnit="m" metricValue="2.2199999999999998" pageId="44" pageNumber="43" unit="m" value="222.0">
|
||
<quantity id="4CDD9B23FFD5505DFBB1F939FB6528AA" box="[1134,1202,1748,1772]" metricMagnitude="2" metricUnit="m" metricValue="2.2199999999999998" pageId="44" pageNumber="43" unit="m" value="222.0">222 m</quantity>
|
||
a. s. l.
|
||
</elevation>
|
||
,
|
||
<date id="FF9B1006FFD5505DFAD8F939FAAD28AA" box="[1287,1402,1748,1772]" pageId="44" pageNumber="43" value="2019-10-28">
|
||
<collectingDate id="EFDFE9EEFFD5505DFAD8F939FAAD28AA" box="[1287,1402,1748,1772]" pageId="44" pageNumber="43" value="2019-10-28">28.X.2019</collectingDate>
|
||
</date>
|
||
, 22:00 h
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="3B4D3C9BFFD5505DFC90F918FA69296A" collectionCode="UV" collectorName="M. Stockmann" country="Thailand" pageId="44" pageNumber="43" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
|
||
<collectionCode id="ED34AE03FFD5505DFC90F918FCA2294A" box="[847,885,1781,1804]" country="Colombia" name="Departamento de Biologia de la Universidad del Valle" pageId="44" pageNumber="43">UV</collectionCode>
|
||
detection, scorpion walking around on tree, primary forest, new moon,
|
||
<specimenCount id="9D23FD4FFFD5505DFC21F8F9FBF7296A" box="[1022,1056,1812,1836]" pageId="44" pageNumber="43" type="male">1♂</specimenCount>
|
||
(
|
||
<typeStatus id="549E8864FFD5505DFBEFF8F9FB45296A" box="[1072,1170,1812,1836]" pageId="44" pageNumber="43" type="holotype">holotype</typeStatus>
|
||
, 1800), leg.
|
||
<collectorName id="26D05310FFD5505DFAC3F8F8FA69296A" box="[1308,1470,1812,1836]" pageId="44" pageNumber="43">M. Stockmann</collectorName>
|
||
</materialsCitation>
|
||
,
|
||
<collectionCode id="ED34AE03FFD5505DFCEFF8D9FCA4290A" box="[816,883,1844,1868]" pageId="44" pageNumber="43">FKCP</collectionCode>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33F654DFFD5505DFCEFF899FAD229AA" pageId="44" pageNumber="43" type="etymology">
|
||
<paragraph id="8B9A36C6FFD5505DFCEFF899FAD229AA" blockId="44.[816,1474,1908,2028]" pageId="44" pageNumber="43">ETYMOLOGY. The specific epithet is a patronym honoring Alexei Andreevich Byalynitskii-Birulya (or A. A. Birula) (1864–1938), a renowned Russian zoologist and arachnologist who specialized in scorpions and solpugids.</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF5A664EFFD4505CFFACF856FAB629AA" ID-DOI="http://doi.org/10.5281/zenodo.5741895" ID-Zenodo-Dep="5741895" httpUri="https://zenodo.org/record/5741895/files/figure.png" pageId="45" pageNumber="44" startId="45.[115,192,1979,2001]" targetBox="[119,1470,262,1938]" targetPageId="45">
|
||
<paragraph id="8B9A36C6FFD4505CFFACF856FAB629AA" blockId="45.[115,1473,1979,2029]" pageId="45" pageNumber="44">
|
||
<emphasis id="B951EAD4FFD4505CFFACF856FECD2996" bold="true" box="[115,282,1979,2001]" pageId="45" pageNumber="44">Figures 331–336</emphasis>
|
||
:
|
||
<taxonomicName id="4C254D45FFD4505CFEF8F856FE192996" authority="Kovařík & Lowe & Stockmann & Šťáhlavský, 2020" authorityName="Kovařík & Lowe & Stockmann & Šťáhlavský" authorityYear="2020" box="[295,462,1979,2000]" class="Arachnida" family="Euscorpiidae" genus="Scorpiops" kingdom="Animalia" order="Scorpiones" pageId="45" pageNumber="44" phylum="Arthropoda" rank="species" species="birulai" status="sp. nov.">
|
||
<emphasis id="B951EAD4FFD4505CFEF8F856FE192996" box="[295,462,1979,2000]" italics="true" pageId="45" pageNumber="44">Scorpiops birulai</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26257AFFFD4505CFE0AF856FDDC2996" box="[469,523,1979,2000]" pageId="45" pageNumber="44" rank="species">
|
||
<emphasis id="B951EAD4FFD4505CFE0AF856FE3B2996" bold="true" box="[469,492,1979,2000]" pageId="45" pageNumber="44">sp</emphasis>
|
||
.
|
||
<emphasis id="B951EAD4FFD4505CFE27F856FDD22996" bold="true" box="[504,517,1979,2000]" pageId="45" pageNumber="44">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
, male holotype.
|
||
<emphasis id="B951EAD4FFD4505CFD71F856FC822996" bold="true" box="[686,853,1979,2001]" pageId="45" pageNumber="44">Figures 331–332</emphasis>
|
||
. dorsal (331) and ventral (332) views.
|
||
<emphasis id="B951EAD4FFD4505CFB11F856FAA22996" bold="true" box="[1230,1397,1979,2001]" pageId="45" pageNumber="44">Figures 333–336</emphasis>
|
||
. Telson lateral (333), and metasoma and telson lateral (334), ventral (335), and dorsal (336) views. Scale bars: 10 mm (331–332, 334–336).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF5A664EFFD7505FFFABF840FBC729BD" ID-DOI="http://doi.org/10.5281/zenodo.5741897" ID-Zenodo-Dep="5741897" httpUri="https://zenodo.org/record/5741897/files/figure.png" pageId="46" pageNumber="45" startId="46.[116,193,1965,1987]" targetBox="[122,1465,230,1951]" targetPageId="46">
|
||
<paragraph id="8B9A36C6FFD7505FFFABF840FBC729BD" blockId="46.[116,1475,1965,2043]" pageId="46" pageNumber="45">
|
||
<emphasis id="B951EAD4FFD7505FFFABF840FECA2984" bold="true" box="[116,285,1965,1987]" pageId="46" pageNumber="45">Figures 337–346</emphasis>
|
||
:
|
||
<taxonomicName id="4C254D45FFD7505FFEF4F843FE032984" authority="Kovařík & Lowe & Stockmann & Šťáhlavský, 2020" authorityName="Kovařík & Lowe & Stockmann & Šťáhlavský" authorityYear="2020" box="[299,468,1965,1987]" class="Arachnida" family="Euscorpiidae" genus="Scorpiops" kingdom="Animalia" order="Scorpiones" pageId="46" pageNumber="45" phylum="Arthropoda" rank="species" species="birulai" status="sp. nov.">
|
||
<emphasis id="B951EAD4FFD7505FFEF4F843FE032984" box="[299,468,1965,1987]" italics="true" pageId="46" pageNumber="45">Scorpiops birulai</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26257AFFFD7505FFE02F843FDC32985" box="[477,532,1966,1987]" pageId="46" pageNumber="45" rank="species">
|
||
<emphasis id="B951EAD4FFD7505FFE02F843FE232985" bold="true" box="[477,500,1966,1987]" pageId="46" pageNumber="45">sp</emphasis>
|
||
.
|
||
<emphasis id="B951EAD4FFD7505FFDDEF843FDD92985" bold="true" box="[513,526,1966,1987]" pageId="46" pageNumber="45">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
, male holotype, pedipalp segments. Chela dorsal (337), external (338) and ventral (339) views. Patella dorsal (340), external (341) and ventral (342) views. Femur and trochanter dorsal (343), and ventral (344) views. Fixed (345) and movable (346) finger dentition. Trichobothrial pattern is indicated by white circles (337a–342a).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF5A664EFFD6505EFFABF871FB6429AC" ID-DOI="http://doi.org/10.5281/zenodo.5741899" ID-Zenodo-Dep="5741899" httpUri="https://zenodo.org/record/5741899/files/figure.png" pageId="47" pageNumber="46" startId="47.[116,193,1948,1970]" targetBox="[113,1474,238,1923]" targetPageId="47">
|
||
<paragraph id="8B9A36C6FFD6505EFFABF871FB6429AC" blockId="47.[116,1474,1948,2026]" pageId="47" pageNumber="46">
|
||
<emphasis id="B951EAD4FFD6505EFFABF871FECC29F4" bold="true" box="[116,283,1948,1970]" pageId="47" pageNumber="46">Figures 347–358</emphasis>
|
||
:
|
||
<emphasis id="B951EAD4FFD6505EFEF7F871FE0729F4" bold="true" box="[296,464,1948,1970]" pageId="47" pageNumber="46">Figures 347–352</emphasis>
|
||
.
|
||
<taxonomicName id="4C254D45FFD6505EFE03F870FD5429F4" authority="Kovařík & Lowe & Stockmann & Šťáhlavský, 2020" authorityName="Kovařík & Lowe & Stockmann & Šťáhlavský" authorityYear="2020" box="[476,643,1949,1970]" class="Arachnida" family="Euscorpiidae" genus="Scorpiops" kingdom="Animalia" order="Scorpiones" pageId="47" pageNumber="46" phylum="Arthropoda" rank="species" species="birulai" status="sp. nov.">
|
||
<emphasis id="B951EAD4FFD6505EFE03F870FD5429F4" box="[476,643,1949,1970]" italics="true" pageId="47" pageNumber="46">Scorpiops birulai</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26257AFFFD6505EFD55F870FD6829F4" box="[650,703,1949,1970]" pageId="47" pageNumber="46" rank="species">
|
||
<emphasis id="B951EAD4FFD6505EFD55F870FD7629F4" bold="true" box="[650,673,1949,1970]" pageId="47" pageNumber="46">sp</emphasis>
|
||
.
|
||
<emphasis id="B951EAD4FFD6505EFD72F870FD6D29F4" bold="true" box="[685,698,1949,1970]" pageId="47" pageNumber="46">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
, male holotype, carapace and tergites I–IV (347), coxosternal area and sternites (348), left legs I–IV, retrolateral aspect (349–352 respectively).
|
||
<emphasis id="B951EAD4FFD6505EFD05F855FC56298B" bold="true" box="[730,897,1976,1998]" pageId="47" pageNumber="46">Figures 353–358</emphasis>
|
||
.
|
||
<taxonomicName id="4C254D45FFD6505EFC52F854FBB0298B" authority="Kovařík & Lowe & Stockmann & Šťáhlavský, 2020" authorityName="Kovařík & Lowe & Stockmann & Šťáhlavský" authorityYear="2020" box="[909,1127,1976,1998]" class="Arachnida" family="Euscorpiidae" genus="Scorpiops" kingdom="Animalia" order="Scorpiones" pageId="47" pageNumber="46" phylum="Arthropoda" rank="species" species="sherwoodae" status="sp. nov.">
|
||
<emphasis id="B951EAD4FFD6505EFC52F854FBB0298B" box="[909,1127,1976,1998]" italics="true" pageId="47" pageNumber="46">Scorpiops sherwoodae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26257AFFFD6505EFBB1F855FB742988" box="[1134,1187,1976,1998]" pageId="47" pageNumber="46" rank="species">
|
||
<emphasis id="B951EAD4FFD6505EFBB1F855FB52298B" bold="true" box="[1134,1157,1976,1997]" pageId="47" pageNumber="46">sp</emphasis>
|
||
.
|
||
<emphasis id="B951EAD4FFD6505EFB4EF855FB49298B" bold="true" box="[1169,1182,1976,1997]" pageId="47" pageNumber="46">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
, male holotype, carapace and tergites I–IV (353), coxosternal area and sternites (354), left legs I–IV, retrolateral aspect (355–358 respectively).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF5A664EFFC95041FFABF825FF3C29BC" ID-DOI="http://doi.org/10.5281/zenodo.5741901" ID-Zenodo-Dep="5741901" httpUri="https://zenodo.org/record/5741901/files/figure.png" pageId="48" pageNumber="47" startId="48.[116,193,1992,2014]" targetBox="[113,1474,232,1980]" targetPageId="48">
|
||
<paragraph id="8B9A36C6FFC95041FFABF825FF3C29BC" blockId="48.[116,1474,1992,2042]" pageId="48" pageNumber="47">
|
||
<emphasis id="B951EAD4FFC95041FFABF825FECD299B" bold="true" box="[116,282,1992,2014]" pageId="48" pageNumber="47">Figures 359–362</emphasis>
|
||
. Thailand, type localities of
|
||
<taxonomicName id="4C254D45FFC95041FDF4F825FD05299B" authority="Kovařík & Lowe & Stockmann & Šťáhlavský, 2020" authorityName="Kovařík & Lowe & Stockmann & Šťáhlavský" authorityYear="2020" box="[555,722,1992,2013]" class="Arachnida" family="Euscorpiidae" genus="Scorpiops" kingdom="Animalia" order="Scorpiones" pageId="48" pageNumber="47" phylum="Arthropoda" rank="species" species="birulai" status="sp. nov.">
|
||
<emphasis id="B951EAD4FFC95041FDF4F825FD05299B" box="[555,722,1992,2013]" italics="true" pageId="48" pageNumber="47">Scorpiops birulai</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26257AFFFC95041FD07F825FCDA2998" box="[728,781,1992,2014]" pageId="48" pageNumber="47" rank="species">
|
||
<emphasis id="B951EAD4FFC95041FD07F825FD38299B" bold="true" box="[728,751,1992,2013]" pageId="48" pageNumber="47">sp</emphasis>
|
||
.
|
||
<emphasis id="B951EAD4FFC95041FD25F825FCD0299B" bold="true" box="[762,775,1992,2013]" pageId="48" pageNumber="47">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
(359),
|
||
<taxonomicName id="4C254D45FFC95041FC8DF825FC76299B" authority="Kovařík & Lowe & Stockmann & Šťáhlavský, 2020" authorityName="Kovařík & Lowe & Stockmann & Šťáhlavský" authorityYear="2020" box="[850,929,1992,2013]" class="Arachnida" family="Euscorpiidae" genus="Scorpiops" kingdom="Animalia" order="Scorpiones" pageId="48" pageNumber="47" phylum="Arthropoda" rank="species" species="kautti" status="sp. nov.">
|
||
<emphasis id="B951EAD4FFC95041FC8DF825FC76299B" box="[850,929,1992,2013]" italics="true" pageId="48" pageNumber="47">S. kautti</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26257AFFFC95041FC79F825FC0B2998" box="[934,988,1992,2014]" pageId="48" pageNumber="47" rank="species">
|
||
<emphasis id="B951EAD4FFC95041FC79F825FC6A299B" bold="true" box="[934,957,1992,2013]" pageId="48" pageNumber="47">sp</emphasis>
|
||
.
|
||
<emphasis id="B951EAD4FFC95041FC17F825FC02299B" bold="true" box="[968,981,1992,2013]" pageId="48" pageNumber="47">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
(360),
|
||
<taxonomicName id="4C254D45FFC95041FBFEF825FB4B299B" authority="Kovařík & Lowe & Stockmann & Šťáhlavský, 2020" authorityName="Kovařík & Lowe & Stockmann & Šťáhlavský" authorityYear="2020" box="[1057,1180,1992,2013]" class="Arachnida" family="Euscorpiidae" genus="Scorpiops" kingdom="Animalia" order="Scorpiones" pageId="48" pageNumber="47" phylum="Arthropoda" rank="species" species="krabiensis" status="sp. nov.">
|
||
<emphasis id="B951EAD4FFC95041FBFEF825FB4B299B" box="[1057,1180,1992,2013]" italics="true" pageId="48" pageNumber="47">S. krabiensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26257AFFFC95041FB7EF825FB012998" box="[1185,1238,1992,2014]" pageId="48" pageNumber="47" rank="species">
|
||
<emphasis id="B951EAD4FFC95041FB7EF825FB6F299B" bold="true" box="[1185,1208,1992,2013]" pageId="48" pageNumber="47">sp</emphasis>
|
||
.
|
||
<emphasis id="B951EAD4FFC95041FB1CF825FB07299B" bold="true" box="[1219,1232,1992,2013]" pageId="48" pageNumber="47">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
(361), and
|
||
<taxonomicName id="4C254D45FFC95041FA9BF825FA15299B" authority="Kovařík & Lowe & Stockmann & Šťáhlavský, 2020" authorityName="Kovařík & Lowe & Stockmann & Šťáhlavský" authorityYear="2020" box="[1348,1474,1992,2013]" class="Arachnida" family="Euscorpiidae" genus="Scorpiops" kingdom="Animalia" order="Scorpiones" pageId="48" pageNumber="47" phylum="Arthropoda" rank="species" species="phatoensis" status="sp. nov.">
|
||
<emphasis id="B951EAD4FFC95041FA9BF825FA15299B" box="[1348,1474,1992,2013]" italics="true" pageId="48" pageNumber="47">S. phatoensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26257AFFFC95041FFABF809FF7D29BF" box="[116,170,2020,2041]" pageId="48" pageNumber="47" rank="species">
|
||
<emphasis id="B951EAD4FFC95041FFABF809FF5C29BF" bold="true" box="[116,139,2020,2041]" pageId="48" pageNumber="47">sp</emphasis>
|
||
.
|
||
<emphasis id="B951EAD4FFC95041FF48F809FF7329BF" bold="true" box="[151,164,2020,2041]" pageId="48" pageNumber="47">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
(362).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C33F654DFFC85040FFADFF06FECE2D45" pageId="49" pageNumber="48" type="diagnosis">
|
||
<paragraph id="8B9A36C6FFC85040FFADFF06FECE2D45" blockId="49.[114,772,234,771]" pageId="49" pageNumber="48">
|
||
DIAGNOSIS (
|
||
<specimenCount id="9D23FD4FFFC85040FF28FF06FEDB2F45" box="[247,268,235,259]" pageId="49" pageNumber="48" type="male">♂</specimenCount>
|
||
). Total length
|
||
<quantity id="4CDD9B23FFC85040FE69FF07FDDE2F44" box="[438,521,234,258]" metricMagnitude="-2" metricUnit="m" metricValue="6.5" pageId="49" pageNumber="48" unit="mm" value="65.0">65 mm</quantity>
|
||
. Base color uniformly reddish black to black. Telson and tarsomere II of legs reddish brown. Pectine teeth number
|
||
<quantity id="4CDD9B23FFC85040FE60FEC6FE2D2F05" box="[447,506,299,323]" metricMagnitude="-1" metricUnit="m" metricValue="2.794" pageId="49" pageNumber="48" unit="in" value="11.0">11 in</quantity>
|
||
male; fulcra present; 3 marginal and 4 middle lamellae. Patella of pedipalp with 27 (5
|
||
<emphasis id="B951EAD4FFC85040FF4DFE86FF7C2FC5" box="[146,171,363,387]" italics="true" pageId="49" pageNumber="48">eb</emphasis>
|
||
, 7
|
||
<emphasis id="B951EAD4FFC85040FF0BFE86FF2F2FC5" box="[212,248,363,387]" italics="true" pageId="49" pageNumber="48">esb</emphasis>
|
||
, 2
|
||
<emphasis id="B951EAD4FFC85040FEFFFE86FEE82FC4" box="[288,319,363,386]" italics="true" pageId="49" pageNumber="48">em</emphasis>
|
||
, 6
|
||
<emphasis id="B951EAD4FFC85040FEB8FE86FE522FC4" box="[359,389,363,386]" italics="true" pageId="49" pageNumber="48">est</emphasis>
|
||
, and 7
|
||
<emphasis id="B951EAD4FFC85040FE01FE86FE262FC4" box="[478,497,363,386]" italics="true" pageId="49" pageNumber="48">et</emphasis>
|
||
) external and 18 ventral trichobothria. Chela of pedipalp with 12
|
||
<emphasis id="B951EAD4FFC85040FDFEFE66FDE62FE4" box="[545,561,395,418]" italics="true" pageId="49" pageNumber="48">V</emphasis>
|
||
series trichobothria located on ventral surface. Chelal trichobothrium
|
||
<emphasis id="B951EAD4FFC85040FD55FE46FD7F2F85" box="[650,680,427,451]" italics="true" pageId="49" pageNumber="48">Eb</emphasis>
|
||
<subScript id="17A13483FFC85040FD78FE56FD782F8F" attach="left" box="[679,687,443,457]" fontSize="6" pageId="49" pageNumber="48">3</subScript>
|
||
located in distal half of manus between trichobothria
|
||
<emphasis id="B951EAD4FFC85040FD5EFE26FD4B2FA4" box="[641,668,459,482]" italics="true" pageId="49" pageNumber="48">Dt</emphasis>
|
||
and
|
||
<emphasis id="B951EAD4FFC85040FD04FE26FD2A2FA4" box="[731,765,459,482]" italics="true" pageId="49" pageNumber="48">Est</emphasis>
|
||
. Fingers of pedipalps undulate in male. Chela length to width ratio 5.8. Pedipalp movable finger with ca. 70 IAD, which have the same size as MD (more than
|
||
<quantity id="4CDD9B23FFC85040FDF8FDC6FDB92C05" box="[551,622,555,579]" metricMagnitude="0" metricUnit="m" metricValue="2.54" pageId="49" pageNumber="48" unit="in" value="100.0">100 in</quantity>
|
||
number) and create a second row; there are also 7–8 ID and 18 OD present. Tarsomere II of leg III with row of six stout median ventral spinules, and two pairs of flanking setae. Metasoma I with 10 and metasoma II–IV with 8 carinae. Telson elongate and sparsely granulate, length to depth ratio
|
||
<quantity id="4CDD9B23FFC85040FDF6FD26FDBF2CA5" box="[553,616,715,739]" metricMagnitude="-2" metricUnit="m" metricValue="9.652" pageId="49" pageNumber="48" unit="in" value="3.8">3.8 in</quantity>
|
||
male; annular ring developed.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33F654DFFC85040FFADFCC6FB7E2DA5" pageId="49" pageNumber="48" type="description">
|
||
<paragraph id="8B9A36C6FFC85040FFADFCC6FD6E2A45" blockId="49.[114,773,810,2018]" pageId="49" pageNumber="48">
|
||
DESCRIPTION (
|
||
<specimenCount id="9D23FD4FFFC85040FECBFCC6FEFF2D05" box="[276,296,811,835]" pageId="49" pageNumber="48" type="male">♂</specimenCount>
|
||
<typeStatus id="549E8864FFC85040FEE7FCC6FE4D2D05" box="[312,410,811,835]" pageId="49" pageNumber="48" type="holotype">holotype</typeStatus>
|
||
). Total length
|
||
<quantity id="4CDD9B23FFC85040FD93FCC7FD742D04" box="[588,675,810,834]" metricMagnitude="-2" metricUnit="m" metricValue="6.5" pageId="49" pageNumber="48" unit="mm" value="65.0">65 mm</quantity>
|
||
. Female unknown. Habitus as shown in
|
||
<figureCitation id="131E2A43FFC85040FE0DFCA6FDA62D25" box="[466,625,843,867]" captionStart="Figures 331–336" captionStartId="45.[115,192,1979,2001]" captionTargetBox="[119,1470,262,1938]" captionTargetId="figure-9@45.[113,1474,249,1943]" captionTargetPageId="45" captionText="Figures 331–336: Scorpiops birulai sp. n., male holotype. Figures 331–332. dorsal (331) and ventral (332) views. Figures 333–336. Telson lateral (333), and metasoma and telson lateral (334), ventral (335), and dorsal (336) views. Scale bars: 10 mm (331–332, 334–336)." figureDoi="http://doi.org/10.5281/zenodo.5741895" httpUri="https://zenodo.org/record/5741895/files/figure.png" pageId="49" pageNumber="48">Figs. 331–332</figureCitation>
|
||
. For position and distribution of trichobothria on pedipalps, see
|
||
<figureCitation id="131E2A43FFC85040FD10FC86FF3C2DE5" captionStart="Figures 337–346" captionStartId="46.[116,193,1965,1987]" captionTargetBox="[122,1465,230,1951]" captionTargetId="figure-14@46.[113,1474,227,1958]" captionTargetPageId="46" captionText="Figures 337–346: Scorpiops birulai sp. n., male holotype, pedipalp segments. Chela dorsal (337), external (338) and ventral (339) views. Patella dorsal (340), external (341) and ventral (342) views. Femur and trochanter dorsal (343), and ventral (344) views. Fixed (345) and movable (346) finger dentition. Trichobothrial pattern is indicated by white circles (337a–342a)." figureDoi="http://doi.org/10.5281/zenodo.5741897" httpUri="https://zenodo.org/record/5741897/files/figure.png" pageId="49" pageNumber="48">Figs. 337a–342a</figureCitation>
|
||
. Fingers of pedipalps undulate in male (
|
||
<figureCitation id="131E2A43FFC85040FD4AFC66FD242DE5" box="[661,755,907,931]" captionStart="Figures 337–346" captionStartId="46.[116,193,1965,1987]" captionTargetBox="[122,1465,230,1951]" captionTargetId="figure-14@46.[113,1474,227,1958]" captionTargetPageId="46" captionText="Figures 337–346: Scorpiops birulai sp. n., male holotype, pedipalp segments. Chela dorsal (337), external (338) and ventral (339) views. Patella dorsal (340), external (341) and ventral (342) views. Femur and trochanter dorsal (343), and ventral (344) views. Fixed (345) and movable (346) finger dentition. Trichobothrial pattern is indicated by white circles (337a–342a)." figureDoi="http://doi.org/10.5281/zenodo.5741897" httpUri="https://zenodo.org/record/5741897/files/figure.png" pageId="49" pageNumber="48">Fig. 338</figureCitation>
|
||
).
|
||
<emphasis id="B951EAD4FFC85040FFADFC47FF3A2D84" bold="true" box="[114,237,938,962]" pageId="49" pageNumber="48">Coloration</emphasis>
|
||
(
|
||
<figureCitation id="131E2A43FFC85040FEDEFC46FE722D85" box="[257,421,939,963]" captionStart="Figures 331–336" captionStartId="45.[115,192,1979,2001]" captionTargetBox="[119,1470,262,1938]" captionTargetId="figure-9@45.[113,1474,249,1943]" captionTargetPageId="45" captionText="Figures 331–336: Scorpiops birulai sp. n., male holotype. Figures 331–332. dorsal (331) and ventral (332) views. Figures 333–336. Telson lateral (333), and metasoma and telson lateral (334), ventral (335), and dorsal (336) views. Scale bars: 10 mm (331–332, 334–336)." figureDoi="http://doi.org/10.5281/zenodo.5741895" httpUri="https://zenodo.org/record/5741895/files/figure.png" pageId="49" pageNumber="48">Figs. 331–332</figureCitation>
|
||
). Base color uniformly reddish black to black, telson, legs, and sternites are reddish brown. Chelicerae reddish brown and reticulate, fingers black.
|
||
</paragraph>
|
||
<paragraph id="8B9A36C6FFC85040FFADFBE7FF1F2B04" blockId="49.[114,773,810,2018]" pageId="49" pageNumber="48">
|
||
<emphasis id="B951EAD4FFC85040FFADFBE7FE432A65" bold="true" box="[114,404,1034,1059]" pageId="49" pageNumber="48">Carapace and mesosoma</emphasis>
|
||
(
|
||
<figureCitation id="131E2A43FFC85040FE7BFBE6FD912A65" box="[420,582,1035,1059]" captionStart="Figures 347–358" captionStartId="47.[116,193,1948,1970]" captionTargetBox="[113,1474,238,1923]" captionTargetId="figure-8@47.[113,1474,238,1923]" captionTargetPageId="47" captionText="Figures 347–358: Figures 347–352. Scorpiops birulai sp. n., male holotype, carapace and tergites I–IV (347), coxosternal area and sternites (348), left legs I–IV, retrolateral aspect (349–352 respectively). Figures 353–358. Scorpiops sherwoodae sp. n., male holotype, carapace and tergites I–IV (353), coxosternal area and sternites (354), left legs I–IV, retrolateral aspect (355–358 respectively)." figureDoi="http://doi.org/10.5281/zenodo.5741899" httpUri="https://zenodo.org/record/5741899/files/figure.png" pageId="49" pageNumber="48">Figs. 347–348</figureCitation>
|
||
). Entire carapace covered with large granules; carinae absent. Anterior margin of carapace with a deep median notch. Carapace with 3 pairs of lateral eyes of which two are normal and one is reduced. Mesosoma granulated, with one median carina. Tergite VII is pentacarinate. Sternites smooth to very finely granulated with two parallel furrows except sternite VII which has four granulate carinae. Pectine teeth number
|
||
<quantity id="4CDD9B23FFC85040FDF9FB06FD8B2B45" box="[550,604,1259,1283]" metricMagnitude="-1" metricUnit="m" metricValue="2.794" pageId="49" pageNumber="48" unit="in" value="11.0">11 in</quantity>
|
||
male
|
||
<typeStatus id="549E8864FFC85040FD41FB06FCD62B45" box="[670,769,1259,1283]" pageId="49" pageNumber="48" type="holotype">holotype</typeStatus>
|
||
. Pectines with 3 marginal and 4 middle lamellae; fulcra present.
|
||
</paragraph>
|
||
<paragraph id="8B9A36C6FFC85040FFADFAA6FD4C2805" blockId="49.[114,773,810,2018]" pageId="49" pageNumber="48">
|
||
<emphasis id="B951EAD4FFC85040FFADFAA6FEBC2B25" bold="true" box="[114,363,1355,1379]" pageId="49" pageNumber="48">Metasoma and telson</emphasis>
|
||
(
|
||
<figureCitation id="131E2A43FFC85040FEA3FAA6FDC92B25" box="[380,542,1355,1379]" captionStart="Figures 331–336" captionStartId="45.[115,192,1979,2001]" captionTargetBox="[119,1470,262,1938]" captionTargetId="figure-9@45.[113,1474,249,1943]" captionTargetPageId="45" captionText="Figures 331–336: Scorpiops birulai sp. n., male holotype. Figures 331–332. dorsal (331) and ventral (332) views. Figures 333–336. Telson lateral (333), and metasoma and telson lateral (334), ventral (335), and dorsal (336) views. Scale bars: 10 mm (331–332, 334–336)." figureDoi="http://doi.org/10.5281/zenodo.5741895" httpUri="https://zenodo.org/record/5741895/files/figure.png" pageId="49" pageNumber="48">Figs. 333–336</figureCitation>
|
||
). Metasoma sparsely hirsute and granulated, with sparse, relatively large granules. Metasomal segment I with 10 carinae, II–IV with 8 carinae, and V with 5 carinae. Median lateral carinae of metasoma II and V indicated by isolated granules that may coalesce into carinae. Dorsolateral carinae of the second to fourth segments terminate posteriorly in a pronounced tooth. Telson elongate and sparsely granulate with annular ring developed.
|
||
</paragraph>
|
||
<paragraph id="8B9A36C6FFC85040FFADF9A7FF1029A4" blockId="49.[114,773,810,2018]" pageId="49" pageNumber="48">
|
||
<emphasis id="B951EAD4FFC85040FFADF9A7FF362824" bold="true" box="[114,225,1610,1634]" pageId="49" pageNumber="48">Pedipalps</emphasis>
|
||
(
|
||
<figureCitation id="131E2A43FFC85040FF2CF9A6FE432825" box="[243,404,1611,1635]" captionStart="Figures 337–346" captionStartId="46.[116,193,1965,1987]" captionTargetBox="[122,1465,230,1951]" captionTargetId="figure-14@46.[113,1474,227,1958]" captionTargetPageId="46" captionText="Figures 337–346: Scorpiops birulai sp. n., male holotype, pedipalp segments. Chela dorsal (337), external (338) and ventral (339) views. Patella dorsal (340), external (341) and ventral (342) views. Femur and trochanter dorsal (343), and ventral (344) views. Fixed (345) and movable (346) finger dentition. Trichobothrial pattern is indicated by white circles (337a–342a)." figureDoi="http://doi.org/10.5281/zenodo.5741897" httpUri="https://zenodo.org/record/5741897/files/figure.png" pageId="49" pageNumber="48">Figs. 337–346</figureCitation>
|
||
). Pedipalps very sparsely hirsute. Patella with 27 (5
|
||
<emphasis id="B951EAD4FFC85040FEEAF986FE9928C5" box="[309,334,1643,1667]" italics="true" pageId="49" pageNumber="48">eb</emphasis>
|
||
, 7
|
||
<emphasis id="B951EAD4FFC85040FEB0F986FE4428C5" box="[367,403,1643,1667]" italics="true" pageId="49" pageNumber="48">esb</emphasis>
|
||
, 2
|
||
<emphasis id="B951EAD4FFC85040FE6CF986FE0528C4" box="[435,466,1643,1666]" italics="true" pageId="49" pageNumber="48">em</emphasis>
|
||
, 6
|
||
<emphasis id="B951EAD4FFC85040FE2CF986FDC628C4" box="[499,529,1643,1666]" italics="true" pageId="49" pageNumber="48">est</emphasis>
|
||
, and 7
|
||
<emphasis id="B951EAD4FFC85040FD81F986FDA628C4" box="[606,625,1643,1666]" italics="true" pageId="49" pageNumber="48">et</emphasis>
|
||
) external and 18 ventral trichobothria. Chela with 12 ventral trichobothria located on the ventral surface. Femur and patella granulated. Femur with 5–6 granulose carinae, and patella with 5 carinae with dorsal and ventral patellar spurs pronounced. Manus dorsally with fine, rounded granules, which in the central area form an irregular dorsal secondary carina. External surface of chela densely covered by minute granules, with a regular external secondary carina. Movable fingers with ca. 70 IAD, which have the same size as MD (more than
|
||
<quantity id="4CDD9B23FFC85040FD82F866FD7529E5" box="[605,674,1931,1955]" metricMagnitude="0" metricUnit="m" metricValue="2.54" pageId="49" pageNumber="48" unit="in" value="100.0">100 in</quantity>
|
||
number) and create a second row; there are also 7–8 ID and 18 OD present.
|
||
</paragraph>
|
||
<paragraph id="8B9A36C6FFC85040FCEEFF06FC202F85" blockId="49.[817,1475,234,483]" pageId="49" pageNumber="48">
|
||
<emphasis id="B951EAD4FFC85040FCEEFF06FCB12F45" bold="true" box="[817,870,235,259]" pageId="49" pageNumber="48">Legs</emphasis>
|
||
(
|
||
<figureCitation id="131E2A43FFC85040FCADFF06FBD92F44" box="[882,1038,234,259]" captionStart="Figures 347–358" captionStartId="47.[116,193,1948,1970]" captionTargetBox="[113,1474,238,1923]" captionTargetId="figure-8@47.[113,1474,238,1923]" captionTargetPageId="47" captionText="Figures 347–358: Figures 347–352. Scorpiops birulai sp. n., male holotype, carapace and tergites I–IV (347), coxosternal area and sternites (348), left legs I–IV, retrolateral aspect (349–352 respectively). Figures 353–358. Scorpiops sherwoodae sp. n., male holotype, carapace and tergites I–IV (353), coxosternal area and sternites (354), left legs I–IV, retrolateral aspect (355–358 respectively)." figureDoi="http://doi.org/10.5281/zenodo.5741899" httpUri="https://zenodo.org/record/5741899/files/figure.png" pageId="49" pageNumber="48">Figs. 349–352</figureCitation>
|
||
). Tibia and tarsomeres of legs with several setae not arranged into bristle combs on dorsal surfaces but with rows of spinules on dorsolateral surfaces and on legs I
|
||
<emphasis id="B951EAD4FFC85040FA7BFEC6FA662F04" box="[1444,1457,299,322]" italics="true" pageId="49" pageNumber="48">–</emphasis>
|
||
II also on ventrolateral surface. Tarsomere II of legs I–IV with 5–7 stout median ventral spinules and two pairs of flanking setae. Femur with 3–4 and patella 4–5 carinae; both femur and patella granulated.
|
||
</paragraph>
|
||
<paragraph id="8B9A36C6FFC85040FCEEFE26FBB02FA5" blockId="49.[817,1475,234,483]" box="[817,1127,459,483]" pageId="49" pageNumber="48">
|
||
<emphasis id="B951EAD4FFC85040FCEEFE26FC0F2FA5" bold="true" box="[817,984,459,483]" pageId="49" pageNumber="48">Measurements</emphasis>
|
||
. See
|
||
<tableCitation id="C6A7037DFFC85040FBCDFE26FBB42FA5" box="[1042,1123,459,483]" captionStart="Table 3" captionStartId="44.[117,173,1000,1021]" captionText="Table 3. Comparative measurements of adults of Scorpiops birulai sp. n., S. ciki sp. n., and S. dii sp. n. Abbreviations: length (L), width (W, in carapace it corresponds to posterior width), depth (D)." pageId="49" pageNumber="48">Table 3</tableCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9A36C6FFC85040FCEEFDE6FA592DE4" blockId="49.[817,1475,523,931]" pageId="49" pageNumber="48">
|
||
AFFINITIES. Among species which have 11–14 ventral trichobothria on the chela manus, only four other species have 18 ventral trichobothria on the pedipalp patella. However, they differ as follows:
|
||
<taxonomicName id="4C254D45FFC85040FBF7FD86FAB02CC5" authority="(Lourenco, 2013)" authorityName="Kovařík & Lowe & Stockmann & Šťáhlavský" authorityYear="2020" baseAuthorityName="Lourenco" baseAuthorityYear="2013" box="[1064,1383,619,643]" class="Arachnida" family="Euscorpiidae" genus="Scorpiops" kingdom="Animalia" order="Scorpiones" pageId="49" pageNumber="48" phylum="Arthropoda" rank="species" species="calmonti">
|
||
<emphasis id="B951EAD4FFC85040FBF7FD86FB752CC5" box="[1064,1186,619,643]" italics="true" pageId="49" pageNumber="48">S. calmonti</emphasis>
|
||
(Lourenço, 2013)
|
||
</taxonomicName>
|
||
has one or two trichobothria of the ventral
|
||
<emphasis id="B951EAD4FFC85040FB1AFD66FB022CE4" box="[1221,1237,651,674]" italics="true" pageId="49" pageNumber="48">V</emphasis>
|
||
series traversing the ventroexternal carina, and positioned on the external surface of the manus; males of
|
||
<taxonomicName id="4C254D45FFC85040FBFEFD26FB6C2CA5" authority="Kovařík & Lowe & Stockmann & Šťáhlavský, 2020" authorityName="Kovařík & Lowe & Stockmann & Šťáhlavský" authorityYear="2020" box="[1057,1211,715,739]" class="Arachnida" family="Euscorpiidae" genus="Scorpiops" kingdom="Animalia" order="Scorpiones" pageId="49" pageNumber="48" phylum="Arthropoda" rank="species" species="sherwoodae" status="sp. nov.">
|
||
<emphasis id="B951EAD4FFC85040FBFEFD26FB6C2CA5" box="[1057,1211,715,739]" italics="true" pageId="49" pageNumber="48">S. sherwoodae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26257AFFFC85040FB1FFD26FB2C2CA4" box="[1216,1275,715,739]" pageId="49" pageNumber="48" rank="species">
|
||
<emphasis id="B951EAD4FFC85040FB1FFD26FB0E2CA5" bold="true" box="[1216,1241,715,739]" pageId="49" pageNumber="48">sp</emphasis>
|
||
.
|
||
<emphasis id="B951EAD4FFC85040FB3AFD26FB232CA5" bold="true" box="[1253,1268,715,739]" pageId="49" pageNumber="48">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="4C254D45FFC85040FAF5FD26FC172D45" authority="Simon, 1887" authorityName="Simon" authorityYear="1887" baseAuthorityName="Simon" baseAuthorityYear="1887" class="Arachnida" family="Euscorpiidae" genus="Scorpiops" kingdom="Animalia" order="Scorpiones" pageId="49" pageNumber="48" phylum="Arthropoda" rank="species" species="anthracinus">
|
||
<emphasis id="B951EAD4FFC85040FAF5FD26FA152CA5" box="[1322,1474,715,739]" italics="true" pageId="49" pageNumber="48">S. anthracinus</emphasis>
|
||
<bibRefCitation id="EFB44B37FFC85040FCEEFD06FC172D45" author="SIMON" box="[817,960,747,771]" pageId="49" pageNumber="48" pagination="101 - 117" refId="ref65986" refString="SIMON, E. 1887. Etude sur les Arachnides de l'Asie meridionale faisant partie des collections de l'Indian Museum (Calcutta). I. Arachnides recueillis a Tavoy (Tenasserim) par Moti Ram. Journal of the Asiatic Society of Bengal, 56 (2): 101 - 117." type="journal article" year="1887">Simon, 1887</bibRefCitation>
|
||
</taxonomicName>
|
||
have fingers of pedipalps strongly undulate; and
|
||
<taxonomicName id="4C254D45FFC85040FCBFFCE6FB582D65" authority="(Kovarik, 2013)" authorityName="Kovařík & Lowe & Stockmann & Šťáhlavský" authorityYear="2020" baseAuthorityName="Kovarik" baseAuthorityYear="2013" box="[864,1167,779,803]" class="Arachnida" family="Euscorpiidae" genus="Scorpiops" kingdom="Animalia" order="Scorpiones" pageId="49" pageNumber="48" phylum="Arthropoda" rank="species" species="citadelle">
|
||
<emphasis id="B951EAD4FFC85040FCBFFCE6FC0E2D65" box="[864,985,779,803]" italics="true" pageId="49" pageNumber="48">S. citadelle</emphasis>
|
||
(Kovařík, 2013)
|
||
</taxonomicName>
|
||
has total length
|
||
<quantity id="4CDD9B23FFC85040FA98FCE7FA692D64" box="[1351,1470,778,802]" metricMagnitude="-2" metricUnit="m" metricValue="4.5" metricValueMax="5.3" metricValueMin="3.7" pageId="49" pageNumber="48" unit="mm" value="45.0" valueMax="53.0" valueMin="37.0">37–53 mm</quantity>
|
||
, reduced fulcra, and almost straight pedipalp fingers in males. In contrast,
|
||
<taxonomicName id="4C254D45FFC85040FC6BFCA6FBCD2D25" authority="Kovařík & Lowe & Stockmann & Šťáhlavský, 2020" authorityName="Kovařík & Lowe & Stockmann & Šťáhlavský" authorityYear="2020" box="[948,1050,843,867]" class="Arachnida" family="Euscorpiidae" genus="Scorpiops" kingdom="Animalia" order="Scorpiones" pageId="49" pageNumber="48" phylum="Arthropoda" rank="species" species="birulai" status="sp. nov.">
|
||
<emphasis id="B951EAD4FFC85040FC6BFCA6FBCD2D25" box="[948,1050,843,867]" italics="true" pageId="49" pageNumber="48">S. birulai</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26257AFFFC85040FBFCFCA6FBB52D24" box="[1059,1122,843,867]" pageId="49" pageNumber="48" rank="species">
|
||
<emphasis id="B951EAD4FFC85040FBFCFCA6FBEB2D25" bold="true" box="[1059,1084,843,867]" pageId="49" pageNumber="48">sp</emphasis>
|
||
.
|
||
<emphasis id="B951EAD4FFC85040FB93FCA6FB8C2D25" bold="true" box="[1100,1115,843,867]" pageId="49" pageNumber="48">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
has all
|
||
<emphasis id="B951EAD4FFC85040FB64FCA6FB1C2D24" box="[1211,1227,843,866]" italics="true" pageId="49" pageNumber="48">V</emphasis>
|
||
series trichobothria of the chela manus on the ventral surface, fingers of pedipalps undulate in male, total length
|
||
<quantity id="4CDD9B23FFC85040FBAFFC67FB6A2DE4" box="[1136,1213,906,930]" metricMagnitude="-2" metricUnit="m" metricValue="6.5" pageId="49" pageNumber="48" unit="mm" value="65.0">65 mm</quantity>
|
||
, and fulcra present.
|
||
</paragraph>
|
||
<paragraph id="8B9A36C6FFC85040FCEEFC26FB7E2DA5" blockId="49.[817,1193,971,995]" box="[817,1193,971,995]" pageId="49" pageNumber="48">
|
||
DISTRIBUTION.
|
||
<collectingCountry id="F3327656FFC85040FC0EFC26FBE72DA5" box="[977,1072,971,995]" name="Thailand" pageId="49" pageNumber="48">Thailand</collectingCountry>
|
||
(
|
||
<figureCitation id="131E2A43FFC85040FBE0FC26FB4B2DA5" box="[1087,1180,971,995]" captionStart="Figure 799" captionStartId="128.[117,185,1143,1165]" captionTargetBox="[113,1474,231,1335]" captionTargetId="figure-14@128.[113,1474,231,1337]" captionTargetPageId="128" captionText="Figure 799. Geographic distribution of the family Scorpiopidae. The map plots locality" figureDoi="http://doi.org/10.5281/zenodo.5742043" httpUri="https://zenodo.org/record/5742043/files/figure.png" pageId="49" pageNumber="48">Fig. 799</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |